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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1081.71872" ID-GBIF-Dataset="1ca2a495-97f3-4226-8c61-df278e3a66e7" ID-PMC="PMC8786815" ID-Pensoft-Pub="1313-2970-1081-137" ID-Pensoft-UUID="B8C62CFD7F495CEE8F18C5BD0A8B8281" ID-PubMed="35087300" ID-ZooBank="8ED559A3F2D444F88A13AF880D6B5A6C" ModsDocID="1313-2970-1081-137" checkinTime="1642466624738" checkinUser="pensoft" docAuthor="Rasoamanana, Nicole &amp; Fisher, Brian L." docDate="2022" docId="B07D7FC5250752D48742BC8001EFFC5C" docLanguage="en" docName="ZooKeys 1081: 137-231" docOrigin="ZooKeys 1081" docPubDate="2022-01-17" docSource="http://dx.doi.org/10.3897/zookeys.1081.71872" docTitle="Camponotus andrianjaka Rasoamanana &amp; Fisher 2022, sp. nov." docType="treatment" docUuid="85932AC5-3790-4161-B72C-134BF27EAC0A" docUuidSource="ZooBank" docVersion="4" id="B8C62CFD7F495CEE8F18C5BD0A8B8281" lastPageNumber="137" masterDocId="B8C62CFD7F495CEE8F18C5BD0A8B8281" masterDocTitle="A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses" masterLastPageNumber="231" masterPageNumber="137" pageNumber="137" updateTime="1668151311846" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rasoamanana, Nicole</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9702-7231</mods:nameIdentifier>
<mods:affiliation>Madagascar Biodiversity Center, BP 6257, Parc Botanique et Zoologique de Tsimbazaza, Antananarivo, Madagascar</mods:affiliation>
<mods:nameIdentifier type="email">rsoanarivo35@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fisher, Brian L.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4653-3270</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-01-17</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1081</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>137</mods:start>
<mods:end>231</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1081.71872</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1081.71872</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1081-137</mods:identifier>
<mods:identifier type="ZooBank">8ED559A3F2D444F88A13AF880D6B5A6C</mods:identifier>
<mods:identifier type="Pensoft-UUID">B8C62CFD7F495CEE8F18C5BD0A8B8281</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="192358979" LSID="urn:lsid:zoobank.org:act:85932AC5-3790-4161-B72C-134BF27EAC0A" httpUri="http://treatment.plazi.org/id/B07D7FC5250752D48742BC8001EFFC5C" lastPageNumber="137" pageId="0" pageNumber="137">
<subSubSection pageId="0" pageNumber="137" type="nomenclature">
<paragraph pageId="0" pageNumber="137">
<taxonomicName LSID="http://zoobank.org/85932AC5-3790-4161-B72C-134BF27EAC0A" authority="Rasoamanana &amp; Fisher, 2022" authorityName="Rasoamanana &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus andrianjaka" order="Hymenoptera" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="andrianjaka" status="sp. nov.">Camponotus andrianjaka</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="137">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="137" type="description">
<paragraph pageId="0" pageNumber="137">
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Morphological character of minor and major workers of C. ellioti species group A head of minor worker of C. ellioti (CASENT 0450893) B minor worker of C. maintikibo in lateral view (CASENT 0062631) C head of major worker of C. maintikibo (CASENT 0062173) D minor worker of C. andrianjaka in lateral view (CASENT 0243690)." figureDoi="10.3897/zookeys.1081.71872.figure8" httpUri="https://binary.pensoft.net/fig/634927" pageId="0" pageNumber="137">Figures 8D</figureCitation>
<figureCitation captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Mesosoma in lateral view A C. maintilany (CASENT 0120678) B C. andrianjaka (CASENT 0243690)." figureDoi="10.3897/zookeys.1081.71872.figure39" httpUri="https://binary.pensoft.net/fig/634958" pageId="0" pageNumber="137">, 39B</figureCitation>
<figureCitation captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Camponotus andrianjaka minor worker (CASENT 0243690) 1 body in lateral view 2 head in full-face view 3 body in dorsal view." figureDoi="10.3897/zookeys.1081.71872.figure61" httpUri="https://binary.pensoft.net/fig/634980" pageId="0" pageNumber="137">, 61A-C</figureCitation>
<figureCitation captionStart="Figure 62" captionStartId="F62" captionText="Figure 62. A minor worker of C. andrianjaka (CASENT 0243690) B minor worker of C. MG 132 (CASENT 0188254)." figureDoi="10.3897/zookeys.1081.71872.figure62" httpUri="https://binary.pensoft.net/fig/634981" pageId="0" pageNumber="137">, 62</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="137" type="holotype worker">
<paragraph pageId="0" pageNumber="137">Holotype worker.</paragraph>
<paragraph pageId="0" pageNumber="137">
Madagascar, Province Fianarantsoa, Isalo III National Parc, 12.02 km SW Ranohira,
<geoCoordinate degrees="22.6158" direction="south" orientation="latitude" precision="5" value="-22.6158">-22.6158</geoCoordinate>
,
<geoCoordinate degrees="45.31084" direction="east" orientation="longitude" precision="1" value="45.31084">45.31084</geoCoordinate>
, 870 m, Bismarckia woodland, ex soil, 15-24 February 2010 (ARA), collection code: ARA0306, specimen code: CASENT0243690 (CAS).
<emphasis bold="true" pageId="0" pageNumber="137">Paratype.</emphasis>
one worker with same data as holotype but specimen coded as CASENT0243954 (CAS).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="137" type="worker diagnosis">
<paragraph pageId="0" pageNumber="137">Worker diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="137">
<taxonomicName authorityName="Rasoamanana &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus andrianjaka" order="Hymenoptera" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="andrianjaka">
<emphasis italics="true" pageId="0" pageNumber="137">Camponotus andrianjaka</emphasis>
</taxonomicName>
can be easily distinguished from other species of the
<taxonomicName authorityName="Forel" authorityYear="1891" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus ellioti" order="Hymenoptera" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="ellioti">
<emphasis italics="true" pageId="0" pageNumber="137">Camponotus ellioti</emphasis>
</taxonomicName>
group by the following character set: body bicolored, head and mesosoma reddish brown, gaster dark brown to black; anterior clypeal margin rounded, not produced; propodeum dorsum slightly concave at the level of metanotal suture; petiole nodiform and surmounted with standing, whitish hairs on its posterior margin.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="137" type="description">
<paragraph pageId="0" pageNumber="137">Description of minor worker.</paragraph>
<paragraph pageId="0" pageNumber="137">
Medium-sized species. Absolute cephalic size (CS: 0.84
<normalizedToken originalValue="±">+/-</normalizedToken>
0.07; 0.81-0.88). In full-face view, head subovate, usually somewhat longer than broad, posterior cephalic margin roundly convex; lateral margin of head straight and tapering to front (CWb/CL: 0.31
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01, 0.30-0.32). Eyes subelliptical, protruding, its border smoothly aligned to lateral margins of head (PoOC/CL: 0.08
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01, 0.08-0.09). Mandibles triangular with six teeth. Clypeus carinate, anterior clypeal margin with rounded triangular projection (ClyL/GPD: 0.24
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01, 0.23-0.25). Antennal scape long, surpassing the occiput by the length of two basal funiculi (SL/CS: 0.36
<normalizedToken originalValue="±">+/-</normalizedToken>
0.04, 0.35-0.38). In lateral view, mesosoma without anterolateral margination, promesonotum convex, mesopropodeal suture shallowly impressed, propodeum rather angulate with slightly concave basal and declivitous surface (MW/ML: 0.19
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02, 0.18-0.21, MPH/ML: 0.15
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01, 0.14-0.15). In lateral view, petiole nodiform, higher than long, its short anterior face rounding gradually to the convex posterior faces. Head, especially anterior portion, finely reticulate-areolate, gaster and mesosoma finely imbricate. Mandibles finely reticulate-punctate. Hairs whitish, one pair present on vertex, middle of mesonotum, and propodeum dorsum; three to four pairs on each side of lateral corner of propodeum; four pairs transversely arranged on petiolar posteromargin; ground pubescence delicate. Head and mesosoma reddish, legs light brown, basal face of first gastral segment light brown and the remainder of the segment dark brown to black.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="137" type="description">
<paragraph pageId="0" pageNumber="137">Description of major worker.</paragraph>
<paragraph pageId="0" pageNumber="137">
Characteristics of minor workers, except: head as long as broad, with rather straight posterior and lateral borders (CS: 1.15
<normalizedToken originalValue="±">+/-</normalizedToken>
0.05, 1.12-1.20, CWb/CL: 0.84
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01, 0.84-0.85). Eyes circular, placed dorsally next to lateral borders (PoOC/CL: 0.26
<normalizedToken originalValue="±">+/-</normalizedToken>
0.03, 0.23-0.27). Anterior clypeal margin forms a rounded rectangular lobe (ClyL/GPD: 0.89
<normalizedToken originalValue="±">+/-</normalizedToken>
0.05, 0.83-0.92). Antennal scape short, just reaching the occipital border (SL/CS: 0.70
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02, 0.69-0.72). Mesosoma the same as minor worker.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="137" type="distribution">
<paragraph pageId="0" pageNumber="137">Distribution and biology.</paragraph>
<paragraph pageId="0" pageNumber="137">
<taxonomicName authorityName="Rasoamanana &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus andrianjaka" order="Hymenoptera" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="andrianjaka">
<emphasis italics="true" pageId="0" pageNumber="137">Camponotus andrianjaka</emphasis>
</taxonomicName>
is a very distinctive species that appears endemic to
<emphasis italics="true" pageId="0" pageNumber="137">Bismarckia</emphasis>
woodland, shrubland, Uapaca woodland, and savannah woodland in the southern part of Madagascar (Fig.
<figureCitation captionStart="Figure 70" captionStartId="F70" captionText="Figure 70. Distribution maps of C. antsaraingy species group A C. antsaraingy; C. efitra species group B C. chrislaini, C. ellioti species group C C. ellioti. D C. andrianjaka. E C. maintilany and C. madagascarensis species group F C. ivadia" figureDoi="10.3897/zookeys.1081.71872.figure70" httpUri="https://binary.pensoft.net/fig/634989" pageId="0" pageNumber="137">70D</figureCitation>
). It can be found also on eucalyptus plantations and grassland. Altitude: 1300-1987 m.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="137" type="discussion">
<paragraph pageId="0" pageNumber="137">Discussion.</paragraph>
<paragraph pageId="0" pageNumber="137">
At first sight,
<taxonomicName authorityName="Rasoamanana &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus andrianjaka" order="Hymenoptera" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="andrianjaka">
<emphasis italics="true" pageId="0" pageNumber="137">Camponotus andrianjaka</emphasis>
</taxonomicName>
may be confused with another new species,
<taxonomicName authorityName="Mayr" authorityYear="1861" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus" order="Hymenoptera" pageId="0" pageNumber="137" phylum="Arthropoda" rank="genus">Camponotus</taxonomicName>
MG132, subgenus
<taxonomicName class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus (Myrmonesites)" order="Hymenoptera" pageId="0" pageNumber="137" phylum="Arthropoda" rank="subGenus" subGenus="Myrmonesites">Camponotus Myrmonesites</taxonomicName>
, but can be separated easily with mesonotum that is gradually rounded in dorsal view; petiole higher than long, its dorsolateral margin surmounted with white hairs; presence of ground pubescence; and gaster all black. In addition,
<taxonomicName genus="C." lsidName="C." pageId="0" pageNumber="137" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="137">C.</emphasis>
</taxonomicName>
MG132 has been collected from rainforest and montane rainforest and absent from woodland.
</paragraph>
<caption doi="10.3897/zookeys.1081.71872.figure61" httpUri="https://binary.pensoft.net/fig/634980" pageId="0" pageNumber="137" start="Figure 61" startId="F61">
<paragraph pageId="0" pageNumber="137">
<emphasis bold="true" pageId="0" pageNumber="137">Figure 61.</emphasis>
<taxonomicName authorityName="Rasoamanana &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus andrianjaka" order="Hymenoptera" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="andrianjaka">
<emphasis italics="true" pageId="0" pageNumber="137">Camponotus andrianjaka</emphasis>
</taxonomicName>
minor worker (CASENT0243690)
<emphasis bold="true" pageId="0" pageNumber="137">1</emphasis>
body in lateral view
<emphasis bold="true" pageId="0" pageNumber="137">2</emphasis>
head in full-face view
<emphasis bold="true" pageId="0" pageNumber="137">3</emphasis>
body in dorsal view.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="137" type="etymology">
<paragraph pageId="0" pageNumber="137">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="137">This new species is named after the collector Andrianjaka Ravelomanana.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="137" type="materials_examined">
<paragraph pageId="0" pageNumber="137">Additional material examined.</paragraph>
<paragraph pageId="0" pageNumber="137">
<emphasis bold="true" pageId="0" pageNumber="137">Province Fianarantsoa</emphasis>
:
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Ampangabe VI Non Protected Area, Ampotoampoto II National Parc, Antohatsahomby V Non Protected Area, Isalo IV National Parc, Mampiarika IV Non Protected Area" county="Non Protected Area" elevation="869" latitude="-20.73528" location="Ampandravelo" longLatPrecision="1" longitude="47.08382" municipality="Non Protected Area" specimenCount="1">
<location LSID="urn:lsid:plazi:treatment:B07D7FC5250752D48742BC8001EFFC5C:442384C955CCD0717B7BB269D6F788E8" county="Non Protected Area" latitude="-20.73528" longLatPrecision="1" longitude="47.08382" municipality="Non Protected Area" name="Ampandravelo">Ampandravelo</location>
III
<location LSID="urn:lsid:plazi:treatment:B07D7FC5250752D48742BC8001EFFC5C:9375C2E8C319019DB2A50565F35478B2" county="Non Protected Area" latitude="-20.73528" longLatPrecision="1" longitude="47.08382" municipality="Non Protected Area" name="Non Protected Area">Non Protected Area</location>
,
<locationDeviation location="Ranohira">
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.072" unit="km" value="10.72">10.72 km</quantity>
NE Ranohira
</locationDeviation>
,
<geoCoordinate degrees="22.53944" direction="south" orientation="latitude" precision="1" value="-22.53944">-22.53944</geoCoordinate>
,
<geoCoordinate degrees="45.51497" direction="east" orientation="longitude" precision="1" value="45.51497">45.51497</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.69" unit="m" value="869.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.69" unit="m" value="869.0">869 m</elevation>
</quantity>
, shrubland (ARA) (CAS); Ampangabe III
<location LSID="urn:lsid:plazi:treatment:B07D7FC5250752D48742BC8001EFFC5C:9C15FA0ADEBB5B5FD66EF7CE02320FC9" county="Non Protected Area" latitude="-20.73528" longLatPrecision="1" longitude="47.08382" municipality="Non Protected Area" name="Non Protected Area">Non Protected Area</location>
,
<locationDeviation location="Itremo">
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.126" unit="km" value="21.26">21.26 km</quantity>
W Itremo
</locationDeviation>
,
<geoCoordinate degrees="20.6125" direction="south" orientation="latitude" precision="5" value="-20.6125">-20.6125</geoCoordinate>
,
<geoCoordinate degrees="46.60883" direction="east" orientation="longitude" precision="1" value="46.60883">46.60883</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.412" unit="m" value="1412.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.412" unit="m" value="1412.0">1412 m</elevation>
</quantity>
, savannah woodland (ARA) (CAS);
<collectorName>
Ampangabe VI
<location LSID="urn:lsid:plazi:treatment:B07D7FC5250752D48742BC8001EFFC5C:C3D10457ABAABDE5735DC4EE06FFDAF3" county="Non Protected Area" latitude="-20.73528" longLatPrecision="1" longitude="47.08382" municipality="Non Protected Area" name="Non Protected Area">Non Protected Area</location>
</collectorName>
,
<locationDeviation location="Itremo">
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.116" unit="km" value="21.16">21.16 km</quantity>
W Itremo
</locationDeviation>
,
<geoCoordinate degrees="20.61444" direction="south" orientation="latitude" precision="1" value="-20.61444">-20.61444</geoCoordinate>
,
<geoCoordinate degrees="46.6104" direction="east" orientation="longitude" precision="5" value="46.6104">46.6104</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.379" unit="m" value="1379.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.379" unit="m" value="1379.0">1379 m</elevation>
</quantity>
, shrubland (ARA) (CAS); Ampangabe VII
<location LSID="urn:lsid:plazi:treatment:B07D7FC5250752D48742BC8001EFFC5C:CA96A81793A7A27F55BFE39798153587" county="Non Protected Area" latitude="-20.73528" longLatPrecision="1" longitude="47.08382" municipality="Non Protected Area" name="Non Protected Area">Non Protected Area</location>
,
<locationDeviation location="Itremo">
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.12" unit="km" value="21.2">21.2 km</quantity>
W Itremo
</locationDeviation>
,
<geoCoordinate degrees="20.61417" direction="south" orientation="latitude" precision="1" value="-20.61417">-20.61417</geoCoordinate>
,
<geoCoordinate degrees="46.60989" direction="east" orientation="longitude" precision="1" value="46.60989">46.60989</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.42" unit="m" value="1420.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.42" unit="m" value="1420.0">1420 m</elevation>
</quantity>
, shrubland (ARA) (CAS);
<collectorName>Ampotoampoto II National Parc</collectorName>
,
<locationDeviation location="Ilakaka">
<quantity metricMagnitude="3" metricUnit="m" metricValue="8.07" unit="km" value="8.07">8.07 km</quantity>
NW Ilakaka
</locationDeviation>
,
<geoCoordinate degrees="22.62806" direction="south" orientation="latitude" precision="1" value="-22.62806">-22.62806</geoCoordinate>
,
<geoCoordinate degrees="45.18843" direction="east" orientation="longitude" precision="1" value="45.18843">45.18843</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="9.19" unit="m" value="919.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="9.19" unit="m" value="919.0">919 m</elevation>
</quantity>
, savannah woodland (ARA) (ATOL UCDC); Ampotoampoto III National Parc,
<locationDeviation location="Ilakaka">
<quantity metricMagnitude="3" metricUnit="m" metricValue="7.91" unit="km" value="7.91">7.91 km</quantity>
NW Ilakaka
</locationDeviation>
,
<geoCoordinate degrees="22.62944" direction="south" orientation="latitude" precision="1" value="-22.62944">-22.62944</geoCoordinate>
,
<geoCoordinate degrees="45.189" direction="east" orientation="longitude" precision="55" value="45.189">45.189</geoCoordinate>
, 919 m, savannah woodland (ARA) (ATOL voucher CAS); Antohatsahomby III
<location LSID="urn:lsid:plazi:treatment:B07D7FC5250752D48742BC8001EFFC5C:21FDB8F8B3806C9D364DDCC0A5C6E8C1" county="Non Protected Area" latitude="-20.73528" longLatPrecision="1" longitude="47.08382" municipality="Non Protected Area" name="Non Protected Area">Non Protected Area</location>
,
<locationDeviation location="Itremo">
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.279" unit="km" value="22.79">22.79 km</quantity>
NW Itremo
</locationDeviation>
,
<geoCoordinate degrees="20.54806" direction="south" orientation="latitude" precision="1" value="-20.54806">-20.54806</geoCoordinate>
,
<geoCoordinate degrees="46.58599" direction="east" orientation="longitude" precision="1" value="46.58599">46.58599</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.499" unit="m" value="1499.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.499" unit="m" value="1499.0">1499 m</elevation>
</quantity>
, Uapaca woodland (ARA) (CAS);
<collectorName>
Antohatsahomby V
<collectingMunicipality>Non Protected Area</collectingMunicipality>
</collectorName>
,
<locationDeviation location="Itremo">
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.263" unit="km" value="22.63">22.63 km</quantity>
NW Itremo
</locationDeviation>
,
<geoCoordinate degrees="20.56722" direction="south" orientation="latitude" precision="1" value="-20.56722">-20.56722</geoCoordinate>
,
<geoCoordinate degrees="46.57923" direction="east" orientation="longitude" precision="1" value="46.57923">46.57923</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.7259999999999998" unit="m" value="1726.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.7259999999999998" unit="m" value="1726.0">1726 m</elevation>
</quantity>
, Uapaca woodland (ARA) (CAS); Isalo III National Parc,
<locationDeviation location="Ranohira">
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.202" unit="km" value="12.02">12.02 km</quantity>
SW Ranohira
</locationDeviation>
,
<geoCoordinate degrees="22.61583" direction="south" orientation="latitude" precision="1" value="-22.61583">-22.61583</geoCoordinate>
,
<geoCoordinate degrees="45.31084" direction="east" orientation="longitude" precision="1" value="45.31084">45.31084</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.7" unit="m" value="870.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.7" unit="m" value="870.0">870 m</elevation>
</quantity>
, Bismarckia woodland (ARA) (CAS);
<collectorName>Isalo IV National Parc</collectorName>
,
<locationDeviation location="Ranohira">
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.2" unit="km" value="12.0">12 km</quantity>
SW Ranohira
</locationDeviation>
,
<geoCoordinate degrees="22.61472" direction="south" orientation="latitude" precision="1" value="-22.61472">-22.61472</geoCoordinate>
,
<geoCoordinate degrees="45.31304" direction="east" orientation="longitude" precision="1" value="45.31304">45.31304</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.67" unit="m" value="867.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.67" unit="m" value="867.0">867 m</elevation>
</quantity>
, Bismarckia woodland (ARA) (ATOL USNM);
<collectorName>Isalo IV National Parc</collectorName>
,
<locationDeviation location="Ranohira">
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.2" unit="km" value="12.0">12 km</quantity>
SW Ranohira
</locationDeviation>
,
<geoCoordinate degrees="22.61472" direction="south" orientation="latitude" precision="1" value="-22.61472">-22.61472</geoCoordinate>
,
<geoCoordinate degrees="45.31304" direction="east" orientation="longitude" precision="1" value="45.31304">45.31304</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.67" unit="m" value="867.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.67" unit="m" value="867.0">867 m</elevation>
</quantity>
, Bismarckia woodland (ARA) (CAS);
<collectorName>
Mampiarika IV
<collectingCounty>Non Protected Area</collectingCounty>
</collectorName>
,
<locationDeviation location="Ambositra">
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.798" unit="km" value="27.98">27.98 km</quantity>
SW Ambositra
</locationDeviation>
,
<geoCoordinate degrees="20.73528" direction="south" orientation="latitude" precision="1" value="-20.73528">-20.73528</geoCoordinate>
,
<geoCoordinate degrees="47.08382" direction="east" orientation="longitude" precision="1" value="47.08382">47.08382</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.486" unit="m" value="1486.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.486" unit="m" value="1486.0">1486 m</elevation>
</quantity>
, Uapaca woodland (ARA) (ATOL CAS)
</materialsCitation>
.
</paragraph>
<caption doi="10.3897/zookeys.1081.71872.figure62" httpUri="https://binary.pensoft.net/fig/634981" pageId="0" pageNumber="137" start="Figure 62" startId="F62">
<paragraph pageId="0" pageNumber="137">
<emphasis bold="true" pageId="0" pageNumber="137">Figure 62.</emphasis>
A minor worker of
<taxonomicName lsidName="C. andrianjaka" pageId="0" pageNumber="137" rank="species" species="andrianjaka">
<emphasis italics="true" pageId="0" pageNumber="137">C. andrianjaka</emphasis>
</taxonomicName>
(CASENT0243690)
<emphasis bold="true" pageId="0" pageNumber="137">B</emphasis>
minor worker of
<taxonomicName genus="C." lsidName="C." pageId="0" pageNumber="137" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="137">C.</emphasis>
</taxonomicName>
MG132 (CASENT0188254).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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