<documentid="B4FD13D2E772CEA62B5FF998AE3526A2"ID-DOI="10.5852/ejt.2024.963.2687"ID-ISSN="2118-9773"ID-Zenodo-Dep="13963130"ID-ZooBank="BA331360-A678-4233-A7CC-7308EF8B6D7E"IM.bibliography_approvedBy="felipe"IM.illustrations_approvedBy="felipe"IM.materialsCitations_approvedBy="guilherme"IM.metadata_approvedBy="guilherme"IM.tables_requiresApprovalFor="GgImagineBatch"IM.taxonomicNames_approvedBy="guilherme"IM.treatmentCitations_approvedBy="guilherme"IM.treatments_approvedBy="guilherme"checkinTime="1729527971783"checkinUser="plazi"docAuthor="Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M. & Carvalho, Leonardo S."docDate="2024"docId="552AE709E805FFCCF43DFA60647B8338"docLanguage="en"docName="EJT.2024.963.1-169.pdf"docOrigin="European Journal of Taxonomy 963"docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2687/12427"docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1"docStyleId="EF2B578F1D15862ADE45B0C07C620911"docStyleName="EJT.2018-.journal_article.type1"docStyleVersion="14"docTitle="Ibotyporanga walekeru Huber & Meng & Král & Ávila Herrera & Carvalho 2024, sp. nov."docType="treatment"docUuid="295920A0-330E-42FD-B2D0-9982066BDB0B"docUuidSource="ZooBank"docVersion="4"lastPageNumber="44"masterDocId="A9139F71E823FFE7F609FFBB6751811D"masterDocTitle="Diamonds in the rough: Ibotyporanga (Araneae, Pholcidae) spiders in semi-arid Neotropical environments"masterLastPageNumber="169"masterPageNumber="1"pageNumber="39"updateTime="1730213223309"updateUser="guilherme"zenodo-license-document="CC-BY-4.0"zenodo-license-figures="CC-BY-4.0"zenodo-license-treatments="UNSPECIFIED">
<mods:titleid="868037330CEA1C8E8B14A177099B19C5">Diamonds in the rough: Ibotyporanga (Araneae, Pholcidae) spiders in semi-arid Neotropical environments</mods:title>
<mods:affiliationid="5D0F4756E07CB4718F221F89B9632E9C">Department of Genetics and Microbiology, Faculty of Science, Charles University, Prague, Czech Republic.</mods:affiliation>
<mods:affiliationid="778F2562486C6FD84815983070CFA22C">Department of Genetics and Microbiology, Faculty of Science, Charles University, Prague, Czech Republic.</mods:affiliation>
<mods:affiliationid="742D6448784D6B2220888CF8E35D5692">Campus Amílcar Ferreira Sobral, Universidade Federal do Piauí, Floriano, Piauí, Brazil.</mods:affiliation>
<taxonomicNameid="1A832D9CE805FFC1F43DFA6064CF84E8"authority="Huber"authorityName="Huber & Meng & Král & Ávila Herrera & Carvalho"authorityYear="2024"box="[564,926,1499,1526]"class="Arachnida"family="Pholcidae"genus="Ibotyporanga"kingdom="Animalia"order="Araneae"pageId="38"pageNumber="39"phylum="Arthropoda"rank="species"species="walekeru"status="sp. nov.">
<figureCitationid="45B84A9AE805FFC1F6D7F99A663F8721"box="[222,366,1569,1596]"captionStart="Fig"captionStartId="34.[189,232,1820,1846]"captionTargetBox="[189,1398,265,1792]"captionTargetId="figure-125@34.[189,1398,265,1792]"captionTargetPageId="34"captionText="Fig. 25. Ibotyporanga spp., live specimens, part one (cf. Figs 55, 73). A. I. ziruma Huber sp. nov., female with egg sac from Colombia, Magdalena, Santa Marta. B–C. I. walekeru Huber sp. nov., male from Colombia, La Guajira, S of Riohacha, and female with egg sac from Colombia, Cesar, ESE of Pueblo Bello. D–E. I. piojo Huber sp. nov., male and female with egg sac from Colombia, Atlántico, near Piojo. F–G. I. itatim Huber sp. nov., male and female with egg sac from Brazil, Bahia, W of Itatim. H–I. I. xique Huber sp. nov., male and female with egg sac from Brazil, Bahia, S of Xique-Xique."figureDoi="http://doi.org/10.5281/zenodo.13963192"httpUri="https://zenodo.org/record/13963192/files/figure.png"pageId="38"pageNumber="39">Figs 25B–C</figureCitation>
<figureCitationid="45B84A9AE805FFC1F773F99A66CB8726"box="[378,410,1569,1595]"captionStart="Fig"captionStartId="35.[189,232,1241,1267]"captionTargetBox="[189,1398,583,1206]"captionTargetId="figure-147@35.[189,1398,583,1206]"captionTargetPageId="35"captionText="Fig. 26. Known geographic distribution of Northern SouthAmerican species.Abbreviation:A=Atlántico."figureDoi="http://doi.org/10.5281/zenodo.13963194"httpUri="https://zenodo.org/record/13963194/files/figure.png"pageId="38"pageNumber="39">26</figureCitation>
<figureCitationid="45B84A9AE805FFC1F7AFF99966978721"box="[422,454,1570,1596]"captionStart="Fig"captionStartId="39.[189,232,1962,1988]"captionTargetBox="[189,1398,1313,1923]"captionTargetId="figure-271@39.[189,1398,1313,1923]"captionTargetPageId="39"captionText="Fig. 31. Ibotyporanga walekeru Huber sp. nov., male from Colombia, Cesar, ESE of Pueblo Bello, ZFMK Ar 24350. Left palp, prolateral, dorsal, and retrolateral views. Scale line: 0.3 mm."figureDoi="http://doi.org/10.5281/zenodo.13963206"httpUri="https://zenodo.org/record/13963206/files/figure.png"pageId="38"pageNumber="39">31</figureCitation>
<figureCitationid="45B84A9AE805FFC1F7DBF999657B8721"box="[466,554,1570,1596]"captionStart="Fig"captionStartId="40.[191,234,1894,1920]"captionTargetBox="[189,1398,804,1851]"captionTargetId="figure-166@40.[189,1398,804,1851]"captionTargetPageId="40"captionText="Fig. 32. Ibotyporanga spp., tips of left procursi, prolateral (A, C, E) and retrolateral (B, D, F) views A–B. I. walekeru Huber sp. nov., from Colombia, Cesar, ESE of Pueblo Bello, ZFMK Ar 24350. C–D. I. bariro Huber, 2020, holotype from Venezuela, Falcón, SE of Bariro, ZFMK Ar 21862. E–F. I. itatim Huber sp. nov., from Brazil, Bahia, W of Itatim, ZFMK Ar 24354. Scale lines: 0.05 mm."figureDoi="http://doi.org/10.5281/zenodo.13963208"httpUri="https://zenodo.org/record/13963208/files/figure.png"pageId="38"pageNumber="39">32A–B</figureCitation>
<figureCitationid="45B84A9AE805FFC1F43FF99A65D38721"box="[566,642,1569,1596]"captionStart-0="Fig"captionStart-1="Fig"captionStart-2="Fig"captionStartId-0="41.[189,232,1893,1919]"captionStartId-1="42.[189,232,1928,1954]"captionStartId-2="43.[189,232,1859,1885]"captionTargetBox-0="[231,1365,688,1841]"captionTargetBox-1="[191,1389,1357,1886]"captionTargetBox-2="[189,1398,611,1821]"captionTargetId-0="figure-194@41.[189,1398,688,1854]"captionTargetId-1="figure-334@42.[189,1398,1352,1886]"captionTargetId-2="figure-194@43.[189,1398,611,1821]"captionTargetPageId-0="41"captionTargetPageId-1="42"captionTargetPageId-2="43"captionText-0="Fig. 33. Ibotyporanga walekeru Huber sp. nov., male from Colombia, Cesar, ESE of Pueblo Bello, ZFMK Ar 24350. A–C. Left tarsus and procursus, prolateral, dorsal, and retrolateral views. D–F. Left genital bulb, prolateral, dorsal, and retrolateral views. Abbreviations: lb=light band; ps =prolateral sclerite. Scale lines: 0.2 mm."captionText-1="Fig. 34. Ibotyporanga walekeru Huber sp. nov. A–B. Male chelicerae, frontal and lateral views, from Colombia, Cesar, ESE of Pueblo Bello, ZFMK Ar 24350. C. Cleared female genitalia, dorsal view, female from Colombia, La Guajira, S of Riohacha, ZFMK Ar 24351. Scale lines: 0.2 mm."captionText-2="Fig. 35. Ibotyporanga walekeru Huber sp. nov. A–C. Abdomen, ventral view, and cleared female genitalia, ventral and dorsal views, female from Colombia, Cesar, ESE of Pueblo Bello, ZFMK Ar 24350. D–F. Same as for preceding, female from Colombia, La Guajira, S of Riohacha, ZFMK Ar 24351. G–I. Same as for preceding, female from Colombia, La Guajira, near Papayal, ZFMK Ar 24352. Abbreviation: ep =epigynal pocket. Scale lines: 0.2 mm."figureDoi-0="http://doi.org/10.5281/zenodo.13963210"figureDoi-1="http://doi.org/10.5281/zenodo.13963212"figureDoi-2="http://doi.org/10.5281/zenodo.13963214"httpUri-0="https://zenodo.org/record/13963210/files/figure.png"httpUri-1="https://zenodo.org/record/13963212/files/figure.png"httpUri-2="https://zenodo.org/record/13963214/files/figure.png"pageId="38"pageNumber="39">33–35</figureCitation>
<figureCitationid="45B84A9AE805FFC1F487F99A65B78726"box="[654,742,1569,1595]"captionStart="Fig"captionStartId="44.[189,232,1043,1069]"captionTargetBox="[189,1398,265,997]"captionTargetId="figure-317@44.[189,1398,265,997]"captionTargetPageId="44"captionText="Fig. 36. Ibotyporanga walekeru Huber sp. nov. (A–C) and I. bariro Huber, 2020 (D), details of cleared female genitalia, dorsal views (arrows: internal tubes). A. Colombia, Cesar, ESE of Pueblo Bello, ZFMK Ar 24350. B. Colombia, La Guajira, S of Riohacha, ZFMK Ar 24351. C. Colombia, La Guajira, near Papayal, ZFMK Ar 24352. D. Venezuela, Falcón, SE of Bariro, ZFMK Ar 21863. Abbreviation: pp =pore plate. Scale lines: 0.1 mm."figureDoi="http://doi.org/10.5281/zenodo.13963216"httpUri="https://zenodo.org/record/13963216/files/figure.png"pageId="38"pageNumber="39">36A–C</figureCitation>
<figureCitationid="45B84A9AE805FFC1F5ABF99964978721"box="[930,966,1570,1596]"captionStart="Fig"captionStartId="13.[189,232,1927,1953]"captionTargetBox="[189,1398,706,1907]"captionTargetId="figure-228@13.[189,1398,706,1907]"captionTargetPageId="13"captionText="Fig. 4. Ibotyporanga spp., male gonopore and epiandrous spigots. A. I. ziruma Huber sp. nov. B. I. walekeru Huber sp. nov. C. I. itatim Huber sp. nov.D. I. naideaeMello-Leitão, 1944. E. I. guanambi Huber sp. nov. F. I. payaya Huber sp. nov. Scale lines: 10 µm."figureDoi="http://doi.org/10.5281/zenodo.13963140"httpUri="https://zenodo.org/record/13963140/files/figure.png"pageId="38"pageNumber="39">4B</figureCitation>
<figureCitationid="45B84A9AE805FFC1F5DAF99963488721"box="[979,1049,1570,1596]"captionStart="Fig"captionStartId="16.[189,232,1927,1953]"captionTargetBox="[189,1398,669,1891]"captionTargetId="figure-230@16.[189,1398,669,1891]"captionTargetPageId="16"captionText="Fig. 7. Ibotyporanga spp., spinnerets. A. I. walekeru Huber sp. nov., female AMS area. B. I. naideae Mello-Leitão, 1944, female AMS area. C–D. I. ziruma Huber sp. nov., male and female ALS and PMS. E. I. walekeru Huber sp. nov., male ALS. F. I. itatim Huber sp. nov., male ALS. Scale lines: 10 µm."figureDoi="http://doi.org/10.5281/zenodo.13963146"httpUri="https://zenodo.org/record/13963146/files/figure.png"pageId="38"pageNumber="39">7A, E</figureCitation>
<figureCitationid="45B84A9AE805FFC1F22FF99963088721"box="[1062,1113,1570,1596]"captionStart="Fig"captionStartId="19.[189,232,1894,1920]"captionTargetBox="[189,1398,782,1857]"captionTargetId="figure-230@19.[189,1398,782,1857]"captionTargetPageId="19"captionText="Fig. 10. Ibotyporanga spp., stridulatory files on male chelicerae, lateral views (A, B, E are mirror images of left chelicerae). A. I. ziruma Huber sp. nov. B. I. walekeru Huber sp. nov. C. I. itatim Huber sp. nov. D. I. naideae Mello-Leitão, 1944. E. I. guanambi Huber sp. nov. F. I. sertao Huber sp. nov. Scale lines: A–D, F =10 µm; E= 20 µm."figureDoi="http://doi.org/10.5281/zenodo.13963156"httpUri="https://zenodo.org/record/13963156/files/figure.png"pageId="38"pageNumber="39">10B</figureCitation>
<figureCitationid="45B84A9AE805FFC1F26CF99A63CA8721"box="[1125,1179,1569,1596]"captionStart="Fig"captionStartId="24.[189,232,1821,1847]"captionTargetBox="[189,1398,583,1782]"captionTargetId="figure-202@24.[189,1398,583,1782]"captionTargetPageId="24"captionText="Fig. 15. Ibotyporanga spp., leg femora and trichobothria. A. I. ziruma Huber sp. nov., right male femur 1, retrolateral view; arrows point at slightly stronger ventral hairs. B. I. guanambi Huber sp. nov., right male femur 3, retrolateral view; arrow points at slightly stronger ventral hair. C. I. ziruma, retrolateral trichobothrium on right male tibia 3. D. I. walekeru Huber sp. nov., retrolateral trichobothrium on right female tibia 2. E. I. itatim Huber sp. nov., retrolateral trichobothrium on right female tibia 2. F. I. naideae Mello-Leitão, 1944, trichobothrium on right female metatarsus 1. Scale lines: 10 µm."figureDoi="http://doi.org/10.5281/zenodo.13963168"httpUri="https://zenodo.org/record/13963168/files/figure.png"pageId="38"pageNumber="39">15D</figureCitation>
<figureCitationid="45B84A9AE805FFC1F2AEF999638B8721"box="[1191,1242,1570,1596]"captionStart="Fig"captionStartId="26.[189,232,1856,1882]"captionTargetBox="[189,1398,656,1818]"captionTargetId="figure-217@26.[189,1398,656,1818]"captionTargetPageId="26"captionText="Fig. 17. Ibotyporanga spp., putative chemoreceptors on male and female legs. A–B. I. ziruma Huber sp. nov., right female tarsus 1 and right male metatarsus 1. C. I. walekeru Huber sp. nov., left female metatarsus 1, near tip, prolateral view. D–E. I. itatim Huber sp. nov., right female metatarsus 2 and right female metatarsus 3. F. I. canudos Huber sp. nov., left female tarsus 4. Scale lines: A, E–F =2 µm; B–D =10 µm."figureDoi="http://doi.org/10.5281/zenodo.13963172"httpUri="https://zenodo.org/record/13963172/files/figure.png"pageId="38"pageNumber="39">17C</figureCitation>
<figureCitationid="45B84A9AE805FFC1F2EFF99962498721"box="[1254,1304,1570,1596]"captionStart="Fig"captionStartId="29.[189,232,1892,1918]"captionTargetBox="[189,1398,718,1847]"captionTargetId="figure-230@29.[189,1398,718,1847]"captionTargetPageId="29"captionText="Fig. 20. Ibotyporanga spp., left leg tarsi, prolateral views. A. I. sertao Huber sp. nov., male tarsus 1. B. I. canudosHuber sp. nov., female tarsus 2. C. I. naideaeMello-Leitão, 1944, male tarsus 3. D. I. ziruma Huber sp. nov., female tarsus 4 (arrows: comb-hairs). E. I. walekeru Huber sp. nov., female tarsus 4 (arrows: comb-hairs). F. I. naideae, male tarsus 4 (arrows: comb-hairs). Scale lines: 10 µm."figureDoi="http://doi.org/10.5281/zenodo.13963182"httpUri="https://zenodo.org/record/13963182/files/figure.png"pageId="38"pageNumber="39">20E</figureCitation>
<figureCitationid="45B84A9AE805FFC1F32DF99962078721"box="[1316,1366,1570,1596]"captionStart="Fig"captionStartId="30.[189,232,1891,1917]"captionTargetBox="[189,1398,700,1852]"captionTargetId="figure-174@30.[189,1398,700,1852]"captionTargetPageId="30"captionText="Fig. 21. Ibotyporanga spp., right leg tarsi, retrolateral views. A. I. walekeru Huber sp. nov., female tarsus 1. B. I. itatim Huber sp. nov., female tarsus 2. C. I. naideae Mello-Leitão, 1944, female tarsus 2 (arrows: membranous areas between pseudosegments). D. I. itatim, female tarsus 3. E. I. itatim, female tarsus 4. F. I. naideae, male tarsus 4. Scale lines: 10 µm."figureDoi="http://doi.org/10.5281/zenodo.13963184"httpUri="https://zenodo.org/record/13963184/files/figure.png"pageId="38"pageNumber="39">21A</figureCitation>
<figureCitationid="45B84A9AE805FFC1F297F92A627E87B1"box="[1182,1327,1681,1708]"captionStart="Fig"captionStartId="41.[189,232,1893,1919]"captionTargetBox="[231,1365,688,1841]"captionTargetId="figure-194@41.[189,1398,688,1854]"captionTargetPageId="41"captionText="Fig. 33. Ibotyporanga walekeru Huber sp. nov., male from Colombia, Cesar, ESE of Pueblo Bello, ZFMK Ar 24350. A–C. Left tarsus and procursus, prolateral, dorsal, and retrolateral views. D–F. Left genital bulb, prolateral, dorsal, and retrolateral views. Abbreviations: lb=light band; ps =prolateral sclerite. Scale lines: 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13963210"httpUri="https://zenodo.org/record/13963210/files/figure.png"pageId="38"pageNumber="39">Fig. 33A–C</figureCitation>
<bibRefCitationid="B9122BEEE805FFC1F7E9F963652087EF"author="Huber B. A. & Villarreal O."box="[480,625,1752,1778]"pageId="38"pageNumber="39"pagination="1 - 317"refId="ref66069"refString="Huber B. A. & Villarreal O. 2020. On Venezuelan pholcid spiders (Araneae, Pholcidae). European Journal of Taxonomy 718: 1 - 317. https: // doi. org / 10.5852 / ejt. 2020.718.1101"type="journal article"year="2020">Huber, 2020</bibRefCitation>
</taxonomicName>
by presence of only a few hair-like processes distally on procursus (many large fringes in
<figureCitationid="45B84A9AE805FFC1F4AEF94064638608"box="[679,818,1787,1813]"captionStart="Fig"captionStartId="40.[191,234,1894,1920]"captionTargetBox="[189,1398,804,1851]"captionTargetId="figure-166@40.[189,1398,804,1851]"captionTargetPageId="40"captionText="Fig. 32. Ibotyporanga spp., tips of left procursi, prolateral (A, C, E) and retrolateral (B, D, F) views A–B. I. walekeru Huber sp. nov., from Colombia, Cesar, ESE of Pueblo Bello, ZFMK Ar 24350. C–D. I. bariro Huber, 2020, holotype from Venezuela, Falcón, SE of Bariro, ZFMK Ar 21862. E–F. I. itatim Huber sp. nov., from Brazil, Bahia, W of Itatim, ZFMK Ar 24354. Scale lines: 0.05 mm."figureDoi="http://doi.org/10.5281/zenodo.13963208"httpUri="https://zenodo.org/record/13963208/files/figure.png"pageId="38"pageNumber="39">Fig. 32A–B</figureCitation>
<figureCitationid="45B84A9AE805FFC1F57CF940635D8608"box="[885,1036,1787,1813]"captionStart="Fig"captionStartId="40.[191,234,1894,1920]"captionTargetBox="[189,1398,804,1851]"captionTargetId="figure-166@40.[189,1398,804,1851]"captionTargetPageId="40"captionText="Fig. 32. Ibotyporanga spp., tips of left procursi, prolateral (A, C, E) and retrolateral (B, D, F) views A–B. I. walekeru Huber sp. nov., from Colombia, Cesar, ESE of Pueblo Bello, ZFMK Ar 24350. C–D. I. bariro Huber, 2020, holotype from Venezuela, Falcón, SE of Bariro, ZFMK Ar 21862. E–F. I. itatim Huber sp. nov., from Brazil, Bahia, W of Itatim, ZFMK Ar 24354. Scale lines: 0.05 mm."figureDoi="http://doi.org/10.5281/zenodo.13963208"httpUri="https://zenodo.org/record/13963208/files/figure.png"pageId="38"pageNumber="39">Fig. 32 C–D</figureCitation>
<taxonomicNameid="1A832D9CE805FFC1F6B4F8A4664A8625"authorityName="Huber & Meng & Král & Ávila Herrera & Carvalho"authorityYear="2024"box="[189,283,1823,1848]"class="Arachnida"family="Pholcidae"genus="Ibotyporanga"kingdom="Animalia"order="Araneae"pageId="38"pageNumber="39"phylum="Arthropoda"rank="species"species="itatim"status="sp. nov.">
<taxonomicNameid="1A832D9CE805FFC1F59FF8A464A58625"authorityName="Huber & Meng & Král & Ávila Herrera & Carvalho"authorityYear="2024"box="[918,1012,1823,1848]"class="Arachnida"family="Pholcidae"genus="Ibotyporanga"kingdom="Animalia"order="Araneae"pageId="38"pageNumber="39"phylum="Arthropoda"rank="species"species="itatim">
<figureCitationid="45B84A9AE805FFC1F456F8DE65BC8662"box="[607,749,1893,1919]"captionStart="Fig"captionStartId="44.[189,232,1043,1069]"captionTargetBox="[189,1398,265,997]"captionTargetId="figure-317@44.[189,1398,265,997]"captionTargetPageId="44"captionText="Fig. 36. Ibotyporanga walekeru Huber sp. nov. (A–C) and I. bariro Huber, 2020 (D), details of cleared female genitalia, dorsal views (arrows: internal tubes). A. Colombia, Cesar, ESE of Pueblo Bello, ZFMK Ar 24350. B. Colombia, La Guajira, S of Riohacha, ZFMK Ar 24351. C. Colombia, La Guajira, near Papayal, ZFMK Ar 24352. D. Venezuela, Falcón, SE of Bariro, ZFMK Ar 21863. Abbreviation: pp =pore plate. Scale lines: 0.1 mm."figureDoi="http://doi.org/10.5281/zenodo.13963216"httpUri="https://zenodo.org/record/13963216/files/figure.png"pageId="38"pageNumber="39">Fig. 36A–C</figureCitation>
<figureCitationid="45B84A9AE805FFC1F2AAF8DE625E8662"box="[1187,1295,1893,1919]"captionStart="Fig"captionStartId="44.[189,232,1043,1069]"captionTargetBox="[189,1398,265,997]"captionTargetId="figure-317@44.[189,1398,265,997]"captionTargetPageId="44"captionText="Fig. 36. Ibotyporanga walekeru Huber sp. nov. (A–C) and I. bariro Huber, 2020 (D), details of cleared female genitalia, dorsal views (arrows: internal tubes). A. Colombia, Cesar, ESE of Pueblo Bello, ZFMK Ar 24350. B. Colombia, La Guajira, S of Riohacha, ZFMK Ar 24351. C. Colombia, La Guajira, near Papayal, ZFMK Ar 24352. D. Venezuela, Falcón, SE of Bariro, ZFMK Ar 21863. Abbreviation: pp =pore plate. Scale lines: 0.1 mm."figureDoi="http://doi.org/10.5281/zenodo.13963216"httpUri="https://zenodo.org/record/13963216/files/figure.png"pageId="38"pageNumber="39">Fig. 36D</figureCitation>
<taxonomicNameid="1A832D9CE805FFC1F32BF8DD62248662"authorityName="Huber & Meng & Král & Ávila Herrera & Carvalho"authorityYear="2024"box="[1314,1397,1894,1919]"class="Arachnida"family="Pholcidae"genus="Ibotyporanga"kingdom="Animalia"order="Araneae"pageId="38"pageNumber="39"phylum="Arthropoda"rank="species"species="piojo"status="sp. nov.">
also with distinct internal tubes but with more strongly curved epigynal pocket and shorter legs (tibia 1 <0.75);
<taxonomicNameid="1A832D9CE805FFC1F777F817668A86D8"authorityName="Huber & Meng & Král & Ávila Herrera & Carvalho"authorityYear="2024"box="[382,475,1964,1989]"class="Arachnida"family="Pholcidae"genus="Ibotyporanga"kingdom="Animalia"order="Araneae"pageId="38"pageNumber="39"phylum="Arthropoda"rank="species"species="itatim">
<materialsCitationid="6DEB5C42E804FFC0F6B4FDAA65838353"collectingDate="2022-09-21"collectorName="B. A. Huber"country="Colombia"elevation="240"latitude="10.3449"location="Pueblo Bello"longLatPrecision="7"longitude="-73.4349"pageId="39"pageNumber="40"specimenCode="MUSENUV-Ar 2736"specimenCount="1"specimenCount-male="1"stateProvince="Cesar"typeStatus="holotype">
<materialsCitationid="6DEB5C42E804FFC0F6B4FD22663B83CB"collectingDate="2022-09-21"collectorName="B. A. Huber"country="Colombia"elevation="240"latitude="10.3449"location="Cesar"longLatPrecision="7"longitude="-73.4349"pageId="39"pageNumber="40"specimenCode="MUSENUV-Ar 2737"specimenCount="9"specimenCount-female="4"specimenCount-male="5"stateProvince="Cesar"typeStatus="paratype">
<materialsCitationid="6DEB5C42E804FFC0F78AFD0763F483CB"box="[387,1189,699,726]"collectingDate="2022-09-21"collectionCode="ZFMK"collectorName="B. A. Huber"country="Colombia"elevation="240"latitude="10.3449"location="Pueblo Bello"longLatPrecision="7"longitude="-73.4349"pageId="39"pageNumber="40"specimenCode="ZFMK Ar 24350"specimenCount="4"specimenCount-female="2"specimenCount-male="2"stateProvince="Cesar"typeStatus="paratype">
<materialsCitationid="6DEB5C42E804FFC0F432FCF464968291"collectingDate="2022-09-19"collectorName="B. A. Huber"country="Colombia"elevation="30"latitude="11.4848"location="5 km S of Riohacha"longLatPrecision="7"longitude="-72.9051"pageId="39"pageNumber="40"specimenCode="MUSENUV-Ar 2738"specimenCount="2"specimenCount-male="2"stateProvince="La Guajira">
<locationid="D85C00C4E804FFC0F549FCF463668274"LSID="urn:lsid:plazi:treatment:552AE709E805FFCCF43DFA60647B8338:D85C00C4E804FFC0F549FCF463668274"box="[832,1079,847,874]"country="Colombia"latitude="11.4848"longLatPrecision="7"longitude="-72.9051"name="5 km S of Riohacha"pageId="39"pageNumber="40"stateProvince="La Guajira">
<materialsCitationid="6DEB5C42E804FFC0F5EAFCC9656382AD"collectingDate="2022-09-19"collectionCode="ZFMK"collectorName="Ar"country="Colombia"elevation="30"latitude="11.4848"location="5 km S of Riohacha"longLatPrecision="7"longitude="-72.9051"pageId="39"pageNumber="40"specimenCode="ZFMK Ar 24351"specimenCount="2"specimenCount-female="1"specimenCount-male="1"stateProvince="La Guajira">
<materialsCitationid="6DEB5C42E804FFC0F445FC2E664682CF"collectingDate="2022-09-19"collectionCode="ZFMK"collectorName="Col"country="Colombia"elevation="30"latitude="11.4848"location="5 km S of Riohacha"longLatPrecision="7"longitude="-72.9051"pageId="39"pageNumber="40"specimenCode="ZFMK Col287"specimenCount="3"specimenCount-female="3"stateProvince="La Guajira">
<materialsCitationid="6DEB5C42E804FFC0F6B4FB93651E857B"collectingDate="2022-09-19"collectionCode="ZFMK"collectorName="B. A. Huber"country="Colombia"elevation="150"latitude="11.0029"location="Papayal"longLatPrecision="7"longitude="-72.7708"pageId="39"pageNumber="40"specimenCode="ZFMK Ar 24352"specimenCount="1"specimenCount-female="1"stateProvince="La Guajira">
<materialsCitationid="6DEB5C42E804FFC0F461FBF766218594"collectingDate="2022-09-19"collectionCode="ZFMK"collectorName="B. A. Huber & Ar"country="Colombia"elevation="150"latitude="11.0029"location="Papayal"longLatPrecision="7"longitude="-72.7708"pageId="39"pageNumber="40"specimenCode="ZFMK Col292"specimenCount="1"specimenCount-female="1"stateProvince="La Guajira">
<taxonomicNameid="1A832D9CE804FFC0F721F81165C386D9"authority="Huber"authorityName="Huber & Meng & Král & Ávila Herrera & Carvalho"authorityYear="2024"box="[296,658,1962,1988]"class="Arachnida"family="Pholcidae"genus="Ibotyporanga"kingdom="Animalia"order="Araneae"pageId="39"pageNumber="40"phylum="Arthropoda"rank="species"species="walekeru"status="sp. nov.">
(in ethanol). Prosoma and legs ochre-yellow, carapace medially with narrow brown mark widening posteriorly; legs without dark rings; abdomen gray with many dark internal marks dorsally and laterally; ventrally with indistinct light ochre plates in front of gonopore and in front of spinnerets.
<figureCitationid="45B84A9AE80BFFCFF7BBFD36654C83B5"box="[434,541,653,680]"captionStart="Fig"captionStartId="34.[189,232,1820,1846]"captionTargetBox="[189,1398,265,1792]"captionTargetId="figure-125@34.[189,1398,265,1792]"captionTargetPageId="34"captionText="Fig. 25. Ibotyporanga spp., live specimens, part one (cf. Figs 55, 73). A. I. ziruma Huber sp. nov., female with egg sac from Colombia, Magdalena, Santa Marta. B–C. I. walekeru Huber sp. nov., male from Colombia, La Guajira, S of Riohacha, and female with egg sac from Colombia, Cesar, ESE of Pueblo Bello. D–E. I. piojo Huber sp. nov., male and female with egg sac from Colombia, Atlántico, near Piojo. F–G. I. itatim Huber sp. nov., male and female with egg sac from Brazil, Bahia, W of Itatim. H–I. I. xique Huber sp. nov., male and female with egg sac from Brazil, Bahia, S of Xique-Xique."figureDoi="http://doi.org/10.5281/zenodo.13963192"httpUri="https://zenodo.org/record/13963192/files/figure.png"pageId="40"pageNumber="41">Fig. 25B</figureCitation>
<taxonomicNameid="1A832D9CE80BFFCFF701F83066B786B9"authority="Huber"authorityName="Huber & Meng & Král & Ávila Herrera & Carvalho"authorityYear="2024"box="[264,486,1930,1956]"class="Arachnida"family="Pholcidae"genus="Ibotyporanga"kingdom="Animalia"order="Araneae"pageId="40"pageNumber="41"phylum="Arthropoda"rank="species"species="walekeru"status="sp. nov.">
<bibRefCitationid="B9122BEEE80BFFCFF78DF816657186DA"author="Huber B. A. & Villarreal O."box="[388,544,1965,1991]"pageId="40"pageNumber="41"pagination="1 - 317"refId="ref66069"refString="Huber B. A. & Villarreal O. 2020. On Venezuelan pholcid spiders (Araneae, Pholcidae). European Journal of Taxonomy 718: 1 - 317. https: // doi. org / 10.5852 / ejt. 2020.718.1101"type="journal article"year="2020">Huber, 2020</bibRefCitation>
<taxonomicNameid="1A832D9CE80BFFCFF709F86A66FC86F7"authority="Huber"authorityName="Huber & Meng & Král & Ávila Herrera & Carvalho"authorityYear="2024"box="[256,429,2000,2026]"class="Arachnida"family="Pholcidae"genus="Ibotyporanga"kingdom="Animalia"order="Araneae"pageId="40"pageNumber="41"phylum="Arthropoda"rank="species"species="itatim"status="sp. nov.">
<figureCitationid="45B84A9AE80AFFCEF4CCFE976470805B"box="[709,801,300,326]"captionStart="Fig"captionStartId="13.[189,232,1927,1953]"captionTargetBox="[189,1398,706,1907]"captionTargetId="figure-228@13.[189,1398,706,1907]"captionTargetPageId="13"captionText="Fig. 4. Ibotyporanga spp., male gonopore and epiandrous spigots. A. I. ziruma Huber sp. nov. B. I. walekeru Huber sp. nov. C. I. itatim Huber sp. nov.D. I. naideaeMello-Leitão, 1944. E. I. guanambi Huber sp. nov. F. I. payaya Huber sp. nov. Scale lines: 10 µm."figureDoi="http://doi.org/10.5281/zenodo.13963140"httpUri="https://zenodo.org/record/13963140/files/figure.png"pageId="41"pageNumber="42">Fig. 4B</figureCitation>
<figureCitationid="45B84A9AE80AFFCEF273FE976384805B"box="[1146,1237,300,326]"captionStart="Fig"captionStartId="16.[189,232,1927,1953]"captionTargetBox="[189,1398,669,1891]"captionTargetId="figure-230@16.[189,1398,669,1891]"captionTargetPageId="16"captionText="Fig. 7. Ibotyporanga spp., spinnerets. A. I. walekeru Huber sp. nov., female AMS area. B. I. naideae Mello-Leitão, 1944, female AMS area. C–D. I. ziruma Huber sp. nov., male and female ALS and PMS. E. I. walekeru Huber sp. nov., male ALS. F. I. itatim Huber sp. nov., male ALS. Scale lines: 10 µm."figureDoi="http://doi.org/10.5281/zenodo.13963146"httpUri="https://zenodo.org/record/13963146/files/figure.png"pageId="41"pageNumber="42">Fig. 7E</figureCitation>
<figureCitationid="45B84A9AE80AFFCEF7ABFED565648095"box="[418,565,366,392]"captionStart="Fig"captionStartId="42.[189,232,1928,1954]"captionTargetBox="[191,1389,1357,1886]"captionTargetId="figure-334@42.[189,1398,1352,1886]"captionTargetPageId="42"captionText="Fig. 34. Ibotyporanga walekeru Huber sp. nov. A–B. Male chelicerae, frontal and lateral views, from Colombia, Cesar, ESE of Pueblo Bello, ZFMK Ar 24350. C. Cleared female genitalia, dorsal view, female from Colombia, La Guajira, S of Riohacha, ZFMK Ar 24351. Scale lines: 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13963212"httpUri="https://zenodo.org/record/13963212/files/figure.png"pageId="41"pageNumber="42">Fig. 34A–B</figureCitation>
<figureCitationid="45B84A9AE80AFFCEF6CFFE2A667F80B6"box="[198,302,401,427]"captionStart="Fig"captionStartId="19.[189,232,1894,1920]"captionTargetBox="[189,1398,782,1857]"captionTargetId="figure-230@19.[189,1398,782,1857]"captionTargetPageId="19"captionText="Fig. 10. Ibotyporanga spp., stridulatory files on male chelicerae, lateral views (A, B, E are mirror images of left chelicerae). A. I. ziruma Huber sp. nov. B. I. walekeru Huber sp. nov. C. I. itatim Huber sp. nov. D. I. naideae Mello-Leitão, 1944. E. I. guanambi Huber sp. nov. F. I. sertao Huber sp. nov. Scale lines: A–D, F =10 µm; E= 20 µm."figureDoi="http://doi.org/10.5281/zenodo.13963156"httpUri="https://zenodo.org/record/13963156/files/figure.png"pageId="41"pageNumber="42">Fig. 10B</figureCitation>
<figureCitationid="45B84A9AE80AFFCEF763FE69669680F1"box="[362,455,466,492]"captionStart="Fig"captionStartId="39.[189,232,1962,1988]"captionTargetBox="[189,1398,1313,1923]"captionTargetId="figure-271@39.[189,1398,1313,1923]"captionTargetPageId="39"captionText="Fig. 31. Ibotyporanga walekeru Huber sp. nov., male from Colombia, Cesar, ESE of Pueblo Bello, ZFMK Ar 24350. Left palp, prolateral, dorsal, and retrolateral views. Scale line: 0.3 mm."figureDoi="http://doi.org/10.5281/zenodo.13963206"httpUri="https://zenodo.org/record/13963206/files/figure.png"pageId="41"pageNumber="42">Fig. 31</figureCitation>
; coxa unmodified; trochanter with short rounded ventral protrusion; femur proximally with short retrolateral process not directed toward distal, with prolateral stridulatory pick, distally widened but unmodified; femur-patella joints not shifted toward one side; patella dorsally only slightly longer than medially wide; tibia-tarsus joints shifted toward retrolateral side; tarsus without dorsal process; procursus (
<figureCitationid="45B84A9AE80AFFCEF7FEFDE465D58364"box="[503,644,607,633]"captionStart="Fig"captionStartId="41.[189,232,1893,1919]"captionTargetBox="[231,1365,688,1841]"captionTargetId="figure-194@41.[189,1398,688,1854]"captionTargetPageId="41"captionText="Fig. 33. Ibotyporanga walekeru Huber sp. nov., male from Colombia, Cesar, ESE of Pueblo Bello, ZFMK Ar 24350. A–C. Left tarsus and procursus, prolateral, dorsal, and retrolateral views. D–F. Left genital bulb, prolateral, dorsal, and retrolateral views. Abbreviations: lb=light band; ps =prolateral sclerite. Scale lines: 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13963210"httpUri="https://zenodo.org/record/13963210/files/figure.png"pageId="41"pageNumber="42">Fig. 33A–C</figureCitation>
<figureCitationid="45B84A9AE809FFCDF4E4FEB2642E803E"box="[749,895,265,291]"captionStart="Fig"captionStartId="40.[191,234,1894,1920]"captionTargetBox="[189,1398,804,1851]"captionTargetId="figure-166@40.[189,1398,804,1851]"captionTargetPageId="40"captionText="Fig. 32. Ibotyporanga spp., tips of left procursi, prolateral (A, C, E) and retrolateral (B, D, F) views A–B. I. walekeru Huber sp. nov., from Colombia, Cesar, ESE of Pueblo Bello, ZFMK Ar 24350. C–D. I. bariro Huber, 2020, holotype from Venezuela, Falcón, SE of Bariro, ZFMK Ar 21862. E–F. I. itatim Huber sp. nov., from Brazil, Bahia, W of Itatim, ZFMK Ar 24354. Scale lines: 0.05 mm."figureDoi="http://doi.org/10.5281/zenodo.13963208"httpUri="https://zenodo.org/record/13963208/files/figure.png"pageId="42"pageNumber="43">Fig. 32A–B</figureCitation>
<figureCitationid="45B84A9AE809FFCDF754FE8A66BB8056"box="[349,490,305,331]"captionStart="Fig"captionStartId="41.[189,232,1893,1919]"captionTargetBox="[231,1365,688,1841]"captionTargetId="figure-194@41.[189,1398,688,1854]"captionTargetPageId="41"captionText="Fig. 33. Ibotyporanga walekeru Huber sp. nov., male from Colombia, Cesar, ESE of Pueblo Bello, ZFMK Ar 24350. A–C. Left tarsus and procursus, prolateral, dorsal, and retrolateral views. D–F. Left genital bulb, prolateral, dorsal, and retrolateral views. Abbreviations: lb=light band; ps =prolateral sclerite. Scale lines: 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13963210"httpUri="https://zenodo.org/record/13963210/files/figure.png"pageId="42"pageNumber="43">Fig. 33D–F</figureCitation>
<taxonomicNameid="1A832D9CE80AFFCEF721F8DE65C38662"authority="Huber"authorityName="Huber & Meng & Král & Ávila Herrera & Carvalho"authorityYear="2024"box="[296,658,1893,1919]"class="Arachnida"family="Pholcidae"genus="Ibotyporanga"kingdom="Animalia"order="Araneae"pageId="41"pageNumber="42"phylum="Arthropoda"rank="species"species="walekeru"status="sp. nov.">
. Without spines but with longer hairs ventrally on femora; without curved hairs; with many short vertical hairs on tibia 1; retrolateral trichobothrium of tibia 1 at 54%; prolateral trichobothrium absent on tibia 1; tarsus 1 with ~4 pseudosegments, distally fairly distinct.
<figureCitationid="45B84A9AE809FFCDF293FD3D63A383BD"box="[1178,1266,646,672]"captionStart="Fig"captionStartId="16.[189,232,1927,1953]"captionTargetBox="[189,1398,669,1891]"captionTargetId="figure-230@16.[189,1398,669,1891]"captionTargetPageId="16"captionText="Fig. 7. Ibotyporanga spp., spinnerets. A. I. walekeru Huber sp. nov., female AMS area. B. I. naideae Mello-Leitão, 1944, female AMS area. C–D. I. ziruma Huber sp. nov., male and female ALS and PMS. E. I. walekeru Huber sp. nov., male ALS. F. I. itatim Huber sp. nov., male ALS. Scale lines: 10 µm."figureDoi="http://doi.org/10.5281/zenodo.13963146"httpUri="https://zenodo.org/record/13963146/files/figure.png"pageId="42"pageNumber="43">Fig. S7</figureCitation>
) suggested that specimens from the three localities may in fact represent three distinct species. The K2P distances among them ranged from 11.0% to 13.9% (
<tableCitationid="900163A4E809FFCDF4B0FD6D647383ED"box="[697,802,726,752]"captionStart="Table 1"captionStartId="7.[183,252,568,596]"captionText="Table 1 (continued on next page). Geographic origins and GenBank accession numbers of sequenced specimens. Specimens are sorted by Code. Two previously published sequences are included for the sake of completeness: JA123 (DQ667852) from Astrin et al. (2007), and S443 (MG268742) from Eberle et al. (2018)."pageId="42"pageNumber="43">Table S1</tableCitation>
<figureCitationid="45B84A9AE809FFCDF413FCFF65D68242"box="[538,647,836,863]"captionStart="Fig"captionStartId="34.[189,232,1820,1846]"captionTargetBox="[189,1398,265,1792]"captionTargetId="figure-125@34.[189,1398,265,1792]"captionTargetPageId="34"captionText="Fig. 25. Ibotyporanga spp., live specimens, part one (cf. Figs 55, 73). A. I. ziruma Huber sp. nov., female with egg sac from Colombia, Magdalena, Santa Marta. B–C. I. walekeru Huber sp. nov., male from Colombia, La Guajira, S of Riohacha, and female with egg sac from Colombia, Cesar, ESE of Pueblo Bello. D–E. I. piojo Huber sp. nov., male and female with egg sac from Colombia, Atlántico, near Piojo. F–G. I. itatim Huber sp. nov., male and female with egg sac from Brazil, Bahia, W of Itatim. H–I. I. xique Huber sp. nov., male and female with egg sac from Brazil, Bahia, S of Xique-Xique."figureDoi="http://doi.org/10.5281/zenodo.13963192"httpUri="https://zenodo.org/record/13963192/files/figure.png"pageId="42"pageNumber="43">Fig. 25C</figureCitation>
) but with darker brown legs, dark median band on carapace extending to ocular area and clypeus, sometimes carapace also laterally with light brown bands; clypeus unmodified; tibiae with few short vertical hairs. Tibia
<figureCitationid="45B84A9AE809FFCDF6CFFC07664F82CA"box="[198,286,956,983]"captionStart="Fig"captionStartId="43.[189,232,1859,1885]"captionTargetBox="[189,1398,611,1821]"captionTargetId="figure-194@43.[189,1398,611,1821]"captionTargetPageId="43"captionText="Fig. 35. Ibotyporanga walekeru Huber sp. nov. A–C. Abdomen, ventral view, and cleared female genitalia, ventral and dorsal views, female from Colombia, Cesar, ESE of Pueblo Bello, ZFMK Ar 24350. D–F. Same as for preceding, female from Colombia, La Guajira, S of Riohacha, ZFMK Ar 24351. G–I. Same as for preceding, female from Colombia, La Guajira, near Papayal, ZFMK Ar 24352. Abbreviation: ep =epigynal pocket. Scale lines: 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13963214"httpUri="https://zenodo.org/record/13963214/files/figure.png"pageId="42"pageNumber="43">Fig. 35</figureCitation>
<figureCitationid="45B84A9AE809FFCDF224FC5F63CF82E3"box="[1069,1182,996,1022]"captionStart="Fig"captionStartId="42.[189,232,1928,1954]"captionTargetBox="[191,1389,1357,1886]"captionTargetId="figure-334@42.[189,1398,1352,1886]"captionTargetPageId="42"captionText="Fig. 34. Ibotyporanga walekeru Huber sp. nov. A–B. Male chelicerae, frontal and lateral views, from Colombia, Cesar, ESE of Pueblo Bello, ZFMK Ar 24350. C. Cleared female genitalia, dorsal view, female from Colombia, La Guajira, S of Riohacha, ZFMK Ar 24351. Scale lines: 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13963212"httpUri="https://zenodo.org/record/13963212/files/figure.png"pageId="42"pageNumber="43">Figs 34C</figureCitation>
<figureCitationid="45B84A9AE809FFCDF2A5FC5F639D82E2"box="[1196,1228,996,1023]"captionStart="Fig"captionStartId="43.[189,232,1859,1885]"captionTargetBox="[189,1398,611,1821]"captionTargetId="figure-194@43.[189,1398,611,1821]"captionTargetPageId="43"captionText="Fig. 35. Ibotyporanga walekeru Huber sp. nov. A–C. Abdomen, ventral view, and cleared female genitalia, ventral and dorsal views, female from Colombia, Cesar, ESE of Pueblo Bello, ZFMK Ar 24350. D–F. Same as for preceding, female from Colombia, La Guajira, S of Riohacha, ZFMK Ar 24351. G–I. Same as for preceding, female from Colombia, La Guajira, near Papayal, ZFMK Ar 24352. Abbreviation: ep =epigynal pocket. Scale lines: 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13963214"httpUri="https://zenodo.org/record/13963214/files/figure.png"pageId="42"pageNumber="43">35</figureCitation>
<figureCitationid="45B84A9AE809FFCDF2D3FC5F626182E3"box="[1242,1328,996,1022]"captionStart="Fig"captionStartId="44.[189,232,1043,1069]"captionTargetBox="[189,1398,265,997]"captionTargetId="figure-317@44.[189,1398,265,997]"captionTargetPageId="44"captionText="Fig. 36. Ibotyporanga walekeru Huber sp. nov. (A–C) and I. bariro Huber, 2020 (D), details of cleared female genitalia, dorsal views (arrows: internal tubes). A. Colombia, Cesar, ESE of Pueblo Bello, ZFMK Ar 24350. B. Colombia, La Guajira, S of Riohacha, ZFMK Ar 24351. C. Colombia, La Guajira, near Papayal, ZFMK Ar 24352. D. Venezuela, Falcón, SE of Bariro, ZFMK Ar 21863. Abbreviation: pp =pore plate. Scale lines: 0.1 mm."figureDoi="http://doi.org/10.5281/zenodo.13963216"httpUri="https://zenodo.org/record/13963216/files/figure.png"pageId="42"pageNumber="43">36A–C</figureCitation>
) with pair of elongate pore plates, with dome-shaped membranous structure from which pair of membranous tubes originate, apparently leading into very thin-walled globular ‘receptacles’. Cleared female genitalia of female from near Papayal very similar to cleared females from other localities.
<figureCitationid="45B84A9AE809FFCDF2AEFB6D63AE85ED"box="[1191,1279,1238,1264]"captionStart="Fig"captionStartId="35.[189,232,1241,1267]"captionTargetBox="[189,1398,583,1206]"captionTargetId="figure-147@35.[189,1398,583,1206]"captionTargetPageId="35"captionText="Fig. 26. Known geographic distribution of Northern SouthAmerican species.Abbreviation:A=Atlántico."figureDoi="http://doi.org/10.5281/zenodo.13963194"httpUri="https://zenodo.org/record/13963194/files/figure.png"pageId="42"pageNumber="43">Fig. 26</figureCitation>
<taxonomicNameid="1A832D9CE809FFCDF72DF83265DB86BE"authority="Huber"authorityName="Huber & Meng & Král & Ávila Herrera & Carvalho"authorityYear="2024"box="[292,650,1929,1955]"class="Arachnida"family="Pholcidae"genus="Ibotyporanga"kingdom="Animalia"order="Araneae"pageId="42"pageNumber="43"phylum="Arthropoda"rank="species"species="walekeru"status="sp. nov.">
locality, the spiders were found in a low, dry forest on a roadside hill. They were beaten from old dry branches lying on the ground and hollowed by termites. They shared this microhabitat with another species of
<bibRefCitationid="B9122BEEE808FFCCF53CFEC6649B8085"author="Huber B. A. & Villarreal O."box="[821,970,381,408]"pageId="43"pageNumber="44"pagination="1 - 317"refId="ref66069"refString="Huber B. A. & Villarreal O. 2020. On Venezuelan pholcid spiders (Araneae, Pholcidae). European Journal of Taxonomy 718: 1 - 317. https: // doi. org / 10.5852 / ejt. 2020.718.1101"type="journal article"year="2020">Huber, 2020</bibRefCitation>
</taxonomicName>
. The specimens from near Riohacha were beaten out of dead cactus branches lying on the ground on a degraded roadside with bushes and a few small trees. They shared this microhabitat with