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<document id="479109F1B266417041EAA2D98C8DA2B6" ID-CLB-Dataset="4077" ID-DOI="10.11646/phytotaxa.522.3.5" ID-GBIF-Dataset="f73db7bf-31a4-4d2c-85ef-55c99a91286a" ID-ISSN="1179-3163" ID-Zenodo-Dep="5560427" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1633933481003" checkinUser="plazi" docAuthor="Svensson, Måns &amp; Westberg, Martin" docDate="2021" docId="522A16765A091014FF50FF6BFD02FBDC" docLanguage="en" docName="phytotaxa.522.3.5.pdf" docOrigin="Phytotaxa 522 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.522.3.5" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Carbonea tephromelae M. Svensson &amp; M. Westb. 2021, sp. nov." docType="treatment" docVersion="8" lastPageNumber="226" masterDocId="AE136E0E5A0D1011FFD8FF9FFF82FFC4" masterDocTitle="A new lichenicolous species of Carbonea (Ascomycota, Lecanoraceae) from northern Sweden" masterLastPageNumber="230" masterPageNumber="221" pageNumber="225" updateTime="1699459414505" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="ED9F502C427641AC00FB9149FBFA51E8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1DC33E3B491E1E9682DABE0AE81D18EB">
<mods:title id="411D93C316CAE0AB24E6CBF52A636D39">A new lichenicolous species of Carbonea (Ascomycota, Lecanoraceae) from northern Sweden</mods:title>
</mods:titleInfo>
<mods:name id="83C8CECC8BAE42FC21849DF51BB998A8" type="personal">
<mods:role id="15E241E9682BAD8B5992A5A930CD7C6F">
<mods:roleTerm id="985EF3455350C94E75899C4FC26A1232">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="51E018BB4272B07C3C9361B60C0CEE6F">Svensson, Måns</mods:namePart>
<mods:nameIdentifier id="4E22AE4C7D42A918AA4CBA3EBF3266CB" type="ORCID">0000-0003-1664-8226</mods:nameIdentifier>
<mods:affiliation id="98A33589D1E56ADAFE080A0E26170612">Museum of Evolution, Uppsala University, Norbyvägen 16, SE- 752 36, Uppsala, Sweden. &amp; mans. svensson @ em. uu. se; https: // orcid. org / 0000 - 0003 - 1664 - 8226</mods:affiliation>
<mods:nameIdentifier id="59AC25DDACE62CE3B8BEF906B4C2A6EB" type="email">mans.svensson@em.uu.se</mods:nameIdentifier>
</mods:name>
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<mods:role id="1ED4F5F11BA85CBAAE927E4A23409832">
<mods:roleTerm id="65A04E53B9148E6511AF77CB345AA455">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AE11BA786FAB622C63D090EAB504D80E">Westberg, Martin</mods:namePart>
<mods:nameIdentifier id="422662CF6CCE293932AAE954DE536390" type="ORCID">0000-0002-8346-0322</mods:nameIdentifier>
<mods:affiliation id="23034DAAD2372AC3FFCDBB698365300E">Museum of Evolution, Uppsala University, Norbyvägen 16, SE- 752 36, Uppsala, Sweden. &amp; martin. westberg @ em. uu. se; https: // orcid. org / 0000 - 0002 - 8346 - 0322</mods:affiliation>
<mods:nameIdentifier id="F7599A2558CC04F441DEC788FE699795" type="email">martin.westberg@em.uu.se</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="F51E77A6BEEACDE3EE8AA0F05C82F0F5">text</mods:typeOfResource>
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<mods:title id="78159F582F9645BEF91342B4BE59DA11">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="A4CFB52DB181F64EE68E2CFC2135CDA7">
<mods:date id="7E05B8FED048879E15B10DAE1E08C0AF">2021</mods:date>
<mods:detail id="647AE1D7E1D5E53938EF9A2427512430" type="pubDate">
<mods:number id="8FE0DF92924381CDCB153A0AF3709BF4">2021-10-11</mods:number>
</mods:detail>
<mods:detail id="15CFA01EACF075EBFDBA2BA30FEB0E21" type="volume">
<mods:number id="9E746A56089E3ADF75E28723A35DE86E">522</mods:number>
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<mods:detail id="F66A54B926E9208C3C68F31461847E84" type="issue">
<mods:number id="5B8CCFA60FBAD0699A1360FFF11C8C5F">3</mods:number>
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<mods:location id="0CB9C46A25D1D885F4CD6E46CEE06F73">
<mods:url id="546308BB16F7C4C0CB5D61BF7508BCF2">http://dx.doi.org/10.11646/phytotaxa.522.3.5</mods:url>
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<mods:classification id="BB211372B8530F0982F096203F81E41F">journal article</mods:classification>
<mods:identifier id="974A2D055335BDCAB249CAD9D1FD992E" type="CLB-Dataset">4077</mods:identifier>
<mods:identifier id="B2D15495C71777232DAF81443949815A" type="DOI">10.11646/phytotaxa.522.3.5</mods:identifier>
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<treatment id="522A16765A091014FF50FF6BFD02FBDC" ID-DOI="http://doi.org/10.5281/zenodo.5562611" ID-GBIF-Taxon="188613985" ID-Zenodo-Dep="5562611" LSID="urn:lsid:plazi:treatment:522A16765A091014FF50FF6BFD02FBDC" httpUri="http://treatment.plazi.org/id/522A16765A091014FF50FF6BFD02FBDC" lastPageId="5" lastPageNumber="226" pageId="4" pageNumber="225">
<subSubSection id="9299F4EB5A091015FF50FF6BFE3EFEF7" pageId="4" pageNumber="225" type="nomenclature">
<paragraph id="DA3CA7605A091015FF50FF6BFC15FECA" blockId="4.[136,919,244,271]" box="[136,919,244,271]" pageId="4" pageNumber="225">
<heading id="8174100C5A091015FF50FF6BFC15FECA" box="[136,919,244,271]" fontSize="11" level="2" pageId="4" pageNumber="225" reason="3">
<taxonomicName id="1D83DCE35A091015FF50FF6BFD33FECA" ID-CoL="9Z2LS" authority="M.Svensson &amp; M.Westb." authorityName="M. Svensson &amp; M. Westb." authorityYear="2021" box="[136,689,244,271]" class="Lecanoromycetes" family="Lecanoraceae" genus="Carbonea" kingdom="Fungi" order="Lecanorales" pageId="4" pageNumber="225" phylum="Ascomycota" rank="species" species="tephromelae" status="sp. nov.">
<emphasis id="E8F77B725A091015FF50FF6BFE12FECB" bold="true" box="[136,400,244,271]" italics="true" pageId="4" pageNumber="225">Carbonea tephromelae</emphasis>
M.Svensson &amp; M.Westb.
</taxonomicName>
<taxonomicNameLabel id="F3C4C6095A091015FD6FFF6AFC8CFECB" box="[695,782,245,271]" pageId="4" pageNumber="225" rank="species">
<emphasis id="E8F77B725A091015FD6FFF6AFD52FECA" box="[695,720,245,270]" italics="true" pageId="4" pageNumber="225">sp</emphasis>
.
<emphasis id="E8F77B725A091015FD06FF6AFC84FECA" box="[734,774,245,270]" italics="true" pageId="4" pageNumber="225">nov</emphasis>
.
</taxonomicNameLabel>
<figureCitation id="42B8BBE55A091015FCCDFF6BFC15FECA" box="[789,919,244,270]" captionStart="FIGURE 2" captionStartId="4.[136,229,1925,1947]" captionTargetBox="[240,1348,727,1900]" captionTargetId="figure-192@4.[240,1348,727,1900]" captionTargetPageId="4" captionText="FIGURE 2A–E. Carbonea tephromelae. A. Habitus - apothecia of C. tephromelae on the thallus of Tephromela atra (UPS L-994014, holotype). B. Section of an apothecium showing the colourless hypothecium, blue-green epihymenium and the dark excipulum (UPS L- 994013). C–E. Showing the range of ascospores shapes from globose to broadly ellipsoid (UPS L-119284). Scale bars: A=1 mm; B=100 µm; C–E=5 µm." figureDoi="http://doi.org/10.5281/zenodo.5560431" httpUri="https://zenodo.org/record/5560431/files/figure.png" pageId="4" pageNumber="225">Fig. 2A–2E</figureCitation>
</heading>
</paragraph>
<paragraph id="DA3CA7605A091015FF50FE87FE3EFEF7" blockId="4.[136,444,280,307]" box="[136,444,280,307]" pageId="4" pageNumber="225">MycoBank No: MB840305</paragraph>
</subSubSection>
<subSubSection id="9299F4EB5A091015FF50FEFFFAEAFE06" pageId="4" pageNumber="225" type="materials_examined">
<paragraph id="DA3CA7605A091015FF50FEFFFAEAFE06" blockId="4.[136,1452,352,703]" pageId="4" pageNumber="225">
<materialsCitation id="6AEBAD3D5A091015FF50FEFFFAEAFE06" ID-GBIF-Occurrence="3382672301" collectedFrom="on Tephromela atra" collectingDate="2020-08-05" collectionCode="UPS" country="Sweden" county="Unna Duvgge" elevation="703" location="Jokkmokk" municipality="Padjelanta National Park" pageId="4" pageNumber="225" specimenCode="PAD384" specimenCount="1" typeStatus="holotype">
<typeStatus id="053819C25A091015FF50FEFFFF71FEBE" box="[136,243,352,378]" pageId="4" pageNumber="225" type="holotype">Holotype</typeStatus>
:—
<collectingCountry id="A294E7F05A091015FECCFEFFFE0FFEBE" box="[276,397,352,378]" name="Sweden" pageId="4" pageNumber="225">SWEDEN</collectingCountry>
. Lule lappmark:
<location id="DF5CF1BB5A091015FD9EFEFFFD39FEBE" LSID="urn:lsid:plazi:treatment:522A16765A091014FF50FF6BFD02FBDC:DF5CF1BB5A091015FD9EFEFFFD39FEBE" box="[582,699,352,378]" country="Sweden" county="Unna Duvgge" municipality="Padjelanta National Park" name="Jokkmokk" pageId="4" pageNumber="225">Jokkmokk</location>
par.,
<collectingMunicipality id="3A583D1A5A091015FD2FFEFFFB8CFEBE" box="[759,1038,352,378]" pageId="4" pageNumber="225">Padjelanta National Park</collectingMunicipality>
,
<locationDeviation id="CB8EBE425A091015FBC0FEFFFAD2FEBE" box="[1048,1360,352,379]" location="Staloluokta" pageId="4" pageNumber="225">11 km WNW of Staloluokta</locationDeviation>
, S slope of
<collectingCounty id="335DDFEC5A091015FF7EFE1AFEC1FE5B" box="[166,323,389,415]" pageId="4" pageNumber="225">Unna Duvgge</collectingCounty>
(Unna Toki),
<locationDeviation id="CB8EBE425A091015FE39FE1AFC94FE5A" box="[481,790,388,415]" pageId="4" pageNumber="225">c. 460 m NE of small cabin</locationDeviation>
,
<geoCoordinate id="BFB7C1A75A091015FCFAFE1BFC29FE5B" box="[802,939,388,415]" pageId="4" pageNumber="225">67.34787˚N</geoCoordinate>
,
<geoCoordinate id="BFB7C1A75A091015FC6EFE1BFBBFFE5B" box="[950,1085,388,415]" pageId="4" pageNumber="225">16.45453˚E</geoCoordinate>
, elev.
<quantity id="1D7B0A855A091015FB5BFE1BFB4EFE5B" box="[1155,1228,388,415]" metricMagnitude="2" metricUnit="m" metricValue="7.029999999999999" pageId="4" pageNumber="225" unit="m" value="703.0">
<elevation id="51AE40535A091015FB5BFE1BFB4EFE5B" box="[1155,1228,388,415]" metricMagnitude="2" metricUnit="m" metricValue="7.029999999999999" pageId="4" pageNumber="225" unit="m" value="703.0">703 m</elevation>
</quantity>
, S-exposed slope in bare mountain region,
<collectedFrom id="699FE49D5A091015FE5EFE36FDE0FE06" box="[390,610,424,451]" pageId="4" pageNumber="225">
on
<taxonomicName id="1D83DCE35A091015FE71FE37FDE0FE06" box="[425,610,424,450]" class="Lecanoromycetes" family="Tephromelataceae" genus="Tephromela" kingdom="Fungi" order="Lecanorales" pageId="4" pageNumber="225" phylum="Ascomycota" rank="species" species="atra">
<emphasis id="E8F77B725A091015FE71FE37FDE0FE06" box="[425,610,424,450]" italics="true" pageId="4" pageNumber="225">Tephromela atra</emphasis>
</taxonomicName>
</collectedFrom>
on rock,
<date id="AE3D81A05A091015FD14FE37FCF0FE06" box="[716,882,424,451]" pageId="4" pageNumber="225" value="2020-08-05">
<collectingDate id="BE7978485A091015FD14FE37FCF0FE06" box="[716,882,424,451]" pageId="4" pageNumber="225" value="2020-08-05">5 August 2020</collectingDate>
</date>
,
<emphasis id="E8F77B725A091015FCA5FE36FBBAFE06" box="[893,1080,424,450]" italics="true" pageId="4" pageNumber="225">Martin Westberg</emphasis>
<specimenCode id="8A250F1B5A091015FBE7FE37FB1CFE06" box="[1087,1182,424,450]" collectionCode="PAD" country="Italy" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14385" lsid="urn:lsid:biocol.org:col:14385" name="Università degli Studi di Padova" pageId="4" pageNumber="225" type="Herbarium">PAD384</specimenCode>
(
<collectionCode id="BC923FA55A091015FB75FE37FB60FE06" box="[1197,1250,424,450]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15542" lsid="urn:lsid:biocol.org:col:15542" name="Uppsala University, Museum of Evolution, Botany Section (Fytoteket)" pageId="4" pageNumber="225" type="Herbarium">UPS</collectionCode>
L-994014).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="9299F4EB5A091015FF65FE53FCB7FDCF" pageId="4" pageNumber="225" type="diagnosis">
<paragraph id="DA3CA7605A091015FF65FE53FCB7FDCF" blockId="4.[136,1452,352,703]" pageId="4" pageNumber="225">
<emphasis id="E8F77B725A091015FF65FE53FEB2FE22" bold="true" box="[189,304,460,486]" pageId="4" pageNumber="225">Diagnosis</emphasis>
:—Distinguished from other
<taxonomicName id="1D83DCE35A091015FDA5FE53FCC5FE22" box="[637,839,460,486]" class="Lecanoromycetes" family="Lecanoraceae" genus="Carbonea" kingdom="Fungi" order="Lecanorales" pageId="4" pageNumber="225" phylum="Ascomycota" rank="species" species="undetermined">
<emphasis id="E8F77B725A091015FDA5FE53FD6FFE22" box="[637,749,460,486]" italics="true" pageId="4" pageNumber="225">Carbonea</emphasis>
species
</taxonomicName>
with a colourless hypothecium by having globose to broadly ellipsoid spores and by growing on
<taxonomicName id="1D83DCE35A091015FDACFE6FFCACFDCE" box="[628,814,496,522]" class="Lecanoromycetes" family="Tephromelataceae" genus="Tephromela" kingdom="Fungi" order="Lecanorales" pageId="4" pageNumber="225" phylum="Ascomycota" rank="species" species="atra">
<emphasis id="E8F77B725A091015FDACFE6FFCACFDCE" box="[628,814,496,522]" italics="true" pageId="4" pageNumber="225">Tephromela atra</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="9299F4EB5A091015FF65FD8BFBB2FDEA" box="[189,1072,532,559]" pageId="4" pageNumber="225" type="etymology">
<paragraph id="DA3CA7605A091015FF65FD8BFBB2FDEA" blockId="4.[136,1452,352,703]" box="[189,1072,532,559]" pageId="4" pageNumber="225">
<emphasis id="E8F77B725A091015FF65FD8BFEBCFDEA" bold="true" box="[189,318,532,558]" pageId="4" pageNumber="225">Etymology</emphasis>
:—Named after its host species
<taxonomicName id="1D83DCE35A091015FD7BFD8BFBB2FDEA" authority="(Huds.) Hafellner." authorityName="Hafellner." baseAuthorityName="Huds." box="[675,1072,532,559]" class="Lecanoromycetes" family="Tephromelataceae" genus="Tephromela" kingdom="Fungi" order="Lecanorales" pageId="4" pageNumber="225" phylum="Ascomycota" rank="species" species="atra">
<emphasis id="E8F77B725A091015FD7BFD8BFCDFFDEA" box="[675,861,532,558]" italics="true" pageId="4" pageNumber="225">Tephromela atra</emphasis>
(Huds.) Hafellner.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="9299F4EB5A091014FF65FDA7FE9FFE1D" lastPageId="5" lastPageNumber="226" pageId="4" pageNumber="225" type="description">
<paragraph id="DA3CA7605A091015FF65FDA7FABDFD7A" blockId="4.[136,1452,352,703]" pageId="4" pageNumber="225">
<emphasis id="E8F77B725A091015FF65FDA7FEC5FD96" bold="true" box="[189,327,568,594]" pageId="4" pageNumber="225">Description</emphasis>
:—Lichenicolous fungus. Thallus inapparent, inhabiting the thallus of
<taxonomicName id="1D83DCE35A091015FB4AFDA7FAD0FD96" box="[1170,1362,568,594]" class="Lecanoromycetes" family="Tephromelataceae" genus="Tephromela" kingdom="Fungi" order="Lecanorales" pageId="4" pageNumber="225" phylum="Ascomycota" rank="species" species="atra">
<emphasis id="E8F77B725A091015FB4AFDA7FAD0FD96" box="[1170,1362,568,594]" italics="true" pageId="4" pageNumber="225">Tephromela atra</emphasis>
</taxonomicName>
, which darkens and is apparently damaged due to the infection (
<figureCitation id="42B8BBE55A091015FD2FFDC3FCC8FDB2" box="[759,842,604,630]" captionStart="FIGURE 2" captionStartId="4.[136,229,1925,1947]" captionTargetBox="[240,1348,727,1900]" captionTargetId="figure-192@4.[240,1348,727,1900]" captionTargetPageId="4" captionText="FIGURE 2A–E. Carbonea tephromelae. A. Habitus - apothecia of C. tephromelae on the thallus of Tephromela atra (UPS L-994014, holotype). B. Section of an apothecium showing the colourless hypothecium, blue-green epihymenium and the dark excipulum (UPS L- 994013). C–E. Showing the range of ascospores shapes from globose to broadly ellipsoid (UPS L-119284). Scale bars: A=1 mm; B=100 µm; C–E=5 µm." figureDoi="http://doi.org/10.5281/zenodo.5560431" httpUri="https://zenodo.org/record/5560431/files/figure.png" pageId="4" pageNumber="225">Fig 2A</figureCitation>
). Apothecia black, shining, c. 0.2–0.4(–0.5) mm diam., rounded to flexuose, sessile on the thallus of the host. Disc flat or becoming somewhat convex in larger apothecia. Proper exciple concolorous with the disc, persistent, narrow, raised or level with the disc in older apothecia.
</paragraph>
<caption id="8EFCF7E85A091015FF50F81AFEA5F7C3" ID-DOI="http://doi.org/10.5281/zenodo.5560431" ID-Zenodo-Dep="5560431" httpUri="https://zenodo.org/record/5560431/files/figure.png" pageId="4" pageNumber="225" startId="4.[136,229,1925,1947]" targetBox="[240,1348,727,1900]" targetPageId="4">
<paragraph id="DA3CA7605A091015FF50F81AFEA5F7C3" blockId="4.[136,1452,1925,2055]" pageId="4" pageNumber="225">
<emphasis id="E8F77B725A091015FF50F81AFEAEF85F" bold="true" box="[136,300,1925,1947]" pageId="4" pageNumber="225">FIGURE 2A–E.</emphasis>
<taxonomicName id="1D83DCE35A091015FEECF81AFD90F85E" box="[308,530,1925,1946]" class="Lecanoromycetes" family="Lecanoraceae" genus="Carbonea" kingdom="Fungi" order="Lecanorales" pageId="4" pageNumber="225" phylum="Ascomycota" rank="species" species="tephromelae">
<emphasis id="E8F77B725A091015FEECF81AFD90F85E" box="[308,530,1925,1946]" italics="true" pageId="4" pageNumber="225">Carbonea tephromelae</emphasis>
</taxonomicName>
.
<emphasis id="E8F77B725A091015FDF8F81AFDB3F85E" bold="true" box="[544,561,1925,1946]" pageId="4" pageNumber="225">A</emphasis>
. Habitus - apothecia of
<taxonomicName id="1D83DCE35A091015FCF8F819FC36F85E" box="[800,948,1925,1947]" class="Lecanoromycetes" family="Lecanoraceae" genus="Carbonea" kingdom="Fungi" order="Lecanorales" pageId="4" pageNumber="225" phylum="Ascomycota" rank="species" species="tephromelae">
<emphasis id="E8F77B725A091015FCF8F819FC36F85E" box="[800,948,1925,1947]" italics="true" pageId="4" pageNumber="225">C. tephromelae</emphasis>
</taxonomicName>
on the thallus of
<taxonomicName id="1D83DCE35A091015FBBBF81AFA86F85F" box="[1123,1284,1925,1947]" class="Lecanoromycetes" family="Tephromelataceae" genus="Tephromela" kingdom="Fungi" order="Lecanorales" pageId="4" pageNumber="225" phylum="Ascomycota" rank="species" species="atra">
<emphasis id="E8F77B725A091015FBBBF81AFA86F85F" box="[1123,1284,1925,1947]" italics="true" pageId="4" pageNumber="225">Tephromela atra</emphasis>
</taxonomicName>
(UPS L-994014, holotype).
<emphasis id="E8F77B725A091015FF29F835FE83F87B" bold="true" box="[241,257,1962,1983]" pageId="4" pageNumber="225">B</emphasis>
. Section of an apothecium showing the colourless hypothecium, blue-green epihymenium and the dark excipulum (UPS L- 994013).
<emphasis id="E8F77B725A091015FF3CF852FE90F827" bold="true" box="[228,274,1997,2019]" pageId="4" pageNumber="225">C–E</emphasis>
. Showing the range of ascospores shapes from globose to broadly ellipsoid (UPS L-119284). Scale bars: A=1 mm; B=100 µm; C–E=5 µm.
</paragraph>
</caption>
<paragraph id="DA3CA7605A081014FF65FF01FE9FFE1D" blockId="5.[136,1452,158,1049]" pageId="5" pageNumber="226">
Apothecial anatomy: Proper exciple opaque, appearing black even in thin sections, densely pigmented blue-green (Cinereorufa-green
<emphasis id="E8F77B725A081014FEB0FF5CFE24FF18" box="[360,422,195,220]" italics="true" pageId="5" pageNumber="226">sensu</emphasis>
<bibRefCitation id="BE12DA915A081014FE68FF5CFD41FF18" author="Meyer, B. &amp; Printzen, C." box="[432,707,194,221]" pageId="5" pageNumber="226" pagination="571 - 583" refId="ref6518" refString="Meyer, B. &amp; Printzen, C. (2000) Proposal for a standardized nomenclature and characterization of insoluble lichen pigments. Lichenologist 32: 571 - 583. https: // doi. org / 10.1006 / lich. 2000.0294" type="journal article" year="2000">Meyer &amp; Printzen 2000</bibRefCitation>
) in the upper part, in the lower part beneath the hypothecium in mixture with a dark brown pigment, c. 20–45(–85) µm thick, composed of radiating, thick-walled hyphae with c. 2 µm wide lumina. Hymenium colourless or faintly blue-green, c. 50–75 µm tall. Epihymenium strongly blue-green (Cinereorufa-green), N+ red, K−. Hypothecium colourless, without visible crystals in polarized light. Paraphyses simple or forked near the tips, septate and often slightly constricted at the septa, c. 1.5 µm in midhymenium, tips colourless, cylindrical to clavate, 1.5–2.5 µm wide. Asci of
<taxonomicName id="1D83DCE35A081014FCE5FEE8FC2AFE54" box="[829,936,375,400]" class="Lecanoromycetes" family="Lecanoraceae" genus="Lecanora" kingdom="Fungi" order="Lecanorales" pageId="5" pageNumber="226" phylum="Ascomycota" rank="genus">
<emphasis id="E8F77B725A081014FCE5FEE8FC2AFE54" box="[829,936,375,400]" italics="true" pageId="5" pageNumber="226">Lecanora</emphasis>
</taxonomicName>
-
<typeStatus id="053819C25A081014FC68FEE8FC66FE55" box="[944,996,375,401]" pageId="5" pageNumber="226">type</typeStatus>
, 8-spored, clavate, 41–49 × 12–19 µm. Ascospores broadly ellipsoid to globose, (7.5–)8.66 ±0.97(–11) × (5.5–)6.81 ±0.71(–8.5) µm; l/b ratio (1–)1.28 ±0.2(– 1.67) (n=57).
</paragraph>
</subSubSection>
<subSubSection id="9299F4EB5A081014FF65FE7DFD02FBDC" pageId="5" pageNumber="226" type="distribution">
<paragraph id="DA3CA7605A081014FF65FE7DFBE0FD74" blockId="5.[136,1452,158,1049]" pageId="5" pageNumber="226">
<emphasis id="E8F77B725A081014FF65FE7DFE6EFE38" bold="true" box="[189,492,482,508]" pageId="5" pageNumber="226">Distribution and habitat</emphasis>
:—The species is so far known from two localities in northern
<collectingCountry id="A294E7F05A081014FB30FE7DFAC6FE38" box="[1256,1348,482,508]" name="Sweden" pageId="5" pageNumber="226">Sweden</collectingCountry>
. Its host
<taxonomicName id="1D83DCE35A081014FF50FD99FEC7FDE4" box="[136,325,518,544]" class="Lecanoromycetes" family="Tephromelataceae" genus="Tephromela" kingdom="Fungi" order="Lecanorales" pageId="5" pageNumber="226" phylum="Ascomycota" rank="species" species="atra">
<emphasis id="E8F77B725A081014FF50FD99FEC7FDE4" box="[136,325,518,544]" italics="true" pageId="5" pageNumber="226">Tephromela atra</emphasis>
</taxonomicName>
, however, is a common and widespread lichen on both calcareous and non-calcareous rocks. We searched for
<taxonomicName id="1D83DCE35A081014FEC5FDB5FDA3FD80" box="[285,545,554,580]" class="Lecanoromycetes" family="Lecanoraceae" genus="Carbonea" kingdom="Fungi" order="Lecanorales" pageId="5" pageNumber="226" phylum="Ascomycota" rank="species" species="tephromelae">
<emphasis id="E8F77B725A081014FEC5FDB5FDA3FD80" box="[285,545,554,580]" italics="true" pageId="5" pageNumber="226">Carbonea tephromelae</emphasis>
</taxonomicName>
in 264 collections of
<taxonomicName id="1D83DCE35A081014FCC6FDB4FCEEFD80" box="[798,876,555,581]" class="Lecanoromycetes" family="Tephromelataceae" genus="Tephromela" kingdom="Fungi" order="Lecanorales" pageId="5" pageNumber="226" phylum="Ascomycota" rank="species" sensu="lato" species="atra">
<emphasis id="E8F77B725A081014FCC6FDB4FCACFD80" box="[798,814,555,580]" italics="true" pageId="5" pageNumber="226">T</emphasis>
.
<emphasis id="E8F77B725A081014FCE5FDB4FCEEFD80" box="[829,876,555,580]" italics="true" pageId="5" pageNumber="226">atra</emphasis>
</taxonomicName>
<emphasis id="E8F77B725A081014FCADFDB4FC6AFD80" box="[885,1000,554,580]" italics="true" pageId="5" pageNumber="226">
<taxonomicNameLabel id="F3C4C6095A081014FCADFDB4FC6AFD80" box="[885,1000,554,580]" pageId="5" pageNumber="226" sensu="lato">sensu lato</taxonomicNameLabel>
</emphasis>
in herb. UPS, of which 183 were from Europe, 29 from North America, 22 from Oceania, 14 from Asia, 12 from South America and 4 from Africa, without finding any additional specimens. Most of the examined collections were from lowland localities, and it is possible that
<taxonomicName id="1D83DCE35A081014FF50FD08FEB6FD74" box="[136,308,662,689]" class="Lecanoromycetes" family="Lecanoraceae" genus="Carbonea" kingdom="Fungi" order="Lecanorales" pageId="5" pageNumber="226" phylum="Ascomycota" rank="species" species="tephromelae">
<emphasis id="E8F77B725A081014FF50FD08FF19FD75" box="[136,155,663,689]" italics="true" pageId="5" pageNumber="226">C</emphasis>
.
<emphasis id="E8F77B725A081014FF71FD09FEB6FD74" box="[169,308,662,688]" italics="true" pageId="5" pageNumber="226">tephromelae</emphasis>
</taxonomicName>
is restricted to or at least more common in alpine to subalpine localities.
</paragraph>
<paragraph id="DA3CA7605A081014FF65FD25FD02FBDC" blockId="5.[136,1452,158,1049]" pageId="5" pageNumber="226">
<emphasis id="E8F77B725A081014FF65FD25FDC2FD10" bold="true" box="[189,576,698,724]" pageId="5" pageNumber="226">Additional specimens examined:</emphasis>
—
<collectingCountry id="A294E7F05A081014FD85FD25FD56FD10" box="[605,724,698,724]" name="Sweden" pageId="5" pageNumber="226">SWEDEN</collectingCountry>
. Lule lappmark: Jokkmokk par., Padjelanta National Park,
<quantity id="1D7B0A855A081014FABEFD25FA2EFD10" box="[1382,1452,698,724]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="226" unit="km" value="11.0">11 km</quantity>
WNW of Staloluokta, S slope of Unna Duvgge (Unna Toki), 67.35390˚N, 16.45239˚E, elev.
<quantity id="1D7B0A855A081014FB47FD41FB6BFD3D" box="[1183,1257,734,761]" metricMagnitude="2" metricUnit="m" metricValue="8.18" pageId="5" pageNumber="226" unit="m" value="818.0">818 m</quantity>
, steep S-exposed slope in bare mountain region with partly calcareous rocks, on
<taxonomicName id="1D83DCE35A081014FC92FC9DFB86FCD8" box="[842,1028,770,796]" class="Lecanoromycetes" family="Tephromelataceae" genus="Tephromela" kingdom="Fungi" order="Lecanorales" pageId="5" pageNumber="226" phylum="Ascomycota" rank="species" species="atra">
<emphasis id="E8F77B725A081014FC92FC9DFB86FCD8" box="[842,1028,770,796]" italics="true" pageId="5" pageNumber="226">Tephromela atra</emphasis>
</taxonomicName>
on rock,
<date id="AE3D81A05A081014FBB6FC9DFA97FCD8" box="[1134,1301,770,796]" pageId="5" pageNumber="226" value="2020-08-02">2 August 2020</date>
,
<emphasis id="E8F77B725A081014FAF8FC9CFAB5FCD8" box="[1312,1335,771,796]" italics="true" pageId="5" pageNumber="226">M</emphasis>
.
<emphasis id="E8F77B725A081014FA9DFC9DFA29FCD8" box="[1349,1451,770,796]" italics="true" pageId="5" pageNumber="226">Westberg</emphasis>
PAD133 (UPS L-994013). Lycksele lappmark: Tärna par., Ume älv, Över-Umans sydvästligaste vik, ”Kobresia-ön” [= the river Ume älv, the southwesternmost inlet of Lake Över-Uman, ”the Kobresia-island”], 66˚06–07’N, 14˚32–33’E, alt.
<quantity id="1D7B0A855A081014FF6AFCF1FF79FC4D" box="[178,251,878,905]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="5" pageNumber="226" unit="m" value="530.0">530 m</quantity>
, kalkfri glimmerskiffer på holmens toppdel [= Calcium-free mica schist on the highest part of the islet],
<date id="AE3D81A05A081014FA57FCF1FE9EFC68" pageId="5" pageNumber="226" value="1960-08-19">19 August 1960</date>
,
<emphasis id="E8F77B725A081014FEF0FC0CFEBEFC69" box="[296,316,915,941]" italics="true" pageId="5" pageNumber="226">G</emphasis>
.
<emphasis id="E8F77B725A081014FE93FC0CFEDEFC68" box="[331,348,915,940]" italics="true" pageId="5" pageNumber="226">E</emphasis>
.
<emphasis id="E8F77B725A081014FEB3FC0CFE4FFC68" box="[363,461,915,940]" italics="true" pageId="5" pageNumber="226">Du Rietz</emphasis>
951d (UPS L-115703); Ume älv, Över-Umans sydvästligaste vik, ”ön 138” [= the river Ume älv, the southwesternmost inlet of Lake Över-Uman, ”the island 138”], 66˚06–07’N, 14˚32–33’E. Alt.
<specimenCount id="CC856CE95A081014FA84FC29FA2EFC15" box="[1372,1452,950,977]" pageId="5" pageNumber="226" type="generic">
<quantity id="1D7B0A855A081014FA84FC29FA2AFC15" box="[1372,1448,950,977]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="5" pageNumber="226" unit="m" value="530.0">530 m</quantity>
.
</specimenCount>
Lösa stenar i andra
<taxonomicName id="1D83DCE35A081014FEB9FC44FE27FC30" box="[353,421,987,1012]" class="Magnoliopsida" family="Rosaceae" genus="Dryas" kingdom="Plantae" order="Rosales" pageId="5" pageNumber="226" phylum="Tracheophyta" rank="genus">
<emphasis id="E8F77B725A081014FEB9FC44FE27FC30" box="[353,421,987,1012]" italics="true" pageId="5" pageNumber="226">Dryas</emphasis>
</taxonomicName>
-backen ovan stranden [= loose rocks in the second
<taxonomicName id="1D83DCE35A081014FC3DFC44FBABFC30" box="[997,1065,987,1012]" class="Magnoliopsida" family="Rosaceae" genus="Dryas" kingdom="Plantae" order="Rosales" pageId="5" pageNumber="226" phylum="Tracheophyta" rank="genus">
<emphasis id="E8F77B725A081014FC3DFC44FBABFC30" box="[997,1065,987,1012]" italics="true" pageId="5" pageNumber="226">Dryas</emphasis>
</taxonomicName>
-slope above the shore],
<date id="AE3D81A05A081014FAE1FC45FF41FBDC" pageId="5" pageNumber="226" value="1960-08-29">29 August 1960</date>
,
<emphasis id="E8F77B725A081014FF16FC60FF60FBDD" box="[206,226,1023,1049]" italics="true" pageId="5" pageNumber="226">G</emphasis>
.
<emphasis id="E8F77B725A081014FF28FC60FE83FBDC" box="[240,257,1023,1048]" italics="true" pageId="5" pageNumber="226">E</emphasis>
.
<emphasis id="E8F77B725A081014FED7FC60FEF3FBDC" box="[271,369,1023,1048]" italics="true" pageId="5" pageNumber="226">Du Rietz</emphasis>
<date id="AE3D81A05A081014FEA0FC61FE35FBDC" box="[376,439,1022,1048]" pageId="5" pageNumber="226" value="2197-01">2197i</date>
(UPS L-119284).
</paragraph>
</subSubSection>
</treatment>
</document>
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