<documentid="A460582D3C654CC8876881EC359F70F2"ID-CLB-Dataset="8437"ID-DOI="10.11646/zootaxa.4853.3.1"ID-GBIF-Dataset="9ed37c70-0d85-41bf-809f-58349df9a4a7"ID-ISSN="1175-5326"ID-Zenodo-Dep="4410727"ID-ZooBank="802D1C38-1186-4187-8289-5AC125242802"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="felipe"IM.tables_requiresApprovalFor="existingObjects,plazi"IM.taxonomicNames_approvedBy="felipe"checkinTime="1609603243845"checkinUser="plazi"docAuthor="Marques, Mariana P., Ceríaco, Luis M. P., Buehler, Matthew D., Bandei- Ra, Suzana A., Janota, Joyce M. & Bauer, Aaron M."docDate="2020"docId="1970878C5466FFDC9EB0F9521BCFFA30"docLanguage="en"docName="zootaxa.4853.3.1.pdf"docOrigin="Zootaxa 4853 (3)"docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article"docStyleId="647186512141C8FC8976D5BCC54AEB7D"docStyleName="Zootaxa.2013-.journal_article"docStyleVersion="9"docTitle="Lygodactylus angolensis Bocage 1896"docType="treatment"docVersion="9"lastPageNumber="324"masterDocId="E549FFF45473FFCB9E27FFFE1875FFE7"masterDocTitle="A revision of the Dwarf Geckos, genus Lygodactylus (Squamata: Gekkonidae) from Angola, with the description of three new species"masterLastPageNumber="352"masterPageNumber="301"pageNumber="322"updateTime="1720041580985"updateUser="ExternalLinkService"zenodo-license-document="CLOSED">
<mods:titleid="AE0632AA5A46DFB0EA09BF15D662BF7B">A revision of the Dwarf Geckos, genus Lygodactylus (Squamata: Gekkonidae) from Angola, with the description of three new species</mods:title>
<mods:affiliationid="C267830B771503C2F7EDF85A9D0644C4">Research Center in Biodiversity and Genetic Resources (CIBIO), InBIO, University of Porto, Rua Padre Armando Quintas 7, Vairão, 4485 - 661 Porto, Portugal & mpltmarques @ gmail. com; https: // orcid. org / 0000 - 0002 - 1712 - 2632 & Museu Nacional de História Natural e da Ciência, Universidade de Lisboa, Rua da Escola Politécnica, 58, 1269 - 102 Lisboa, Portugal & mpltmarques @ gmail. com; https: // orcid. org / 0000 - 0002 - 1712 - 2632 & lmceriaco @ mhnc. up. pt; https: // orcid. org / 0000 - 0002 - 0591 - 9978 & Museu de História Natural e da Ciência da Universidade do Porto, Praça Gomes Teixeira 4099 - 002 Porto, Portugal & lmceriaco @ mhnc. up. pt; https: // orcid. org / 0000 - 0002 - 0591 - 9978 & mpltmarques @ gmail. com; https: // orcid. org / 0000 - 0002 - 1712 - 2632</mods:affiliation>
<mods:affiliationid="087321678DBDB2379EC1C3D93DD904D1">Museu de História Natural e da Ciência da Universidade do Porto, Praça Gomes Teixeira 4099 - 002 Porto, Portugal & lmceriaco @ mhnc. up. pt; https: // orcid. org / 0000 - 0002 - 0591 - 9978 & Museu Nacional de História Natural e da Ciência, Universidade de Lisboa, Rua da Escola Politécnica, 58, 1269 - 102 Lisboa, Portugal & mpltmarques @ gmail. com; https: // orcid. org / 0000 - 0002 - 1712 - 2632 & lmceriaco @ mhnc. up. pt; https: // orcid. org / 0000 - 0002 - 0591 - 9978</mods:affiliation>
<mods:affiliationid="C9DA19BA3AA8FECF57224B41B516834F">Department of Biology and Center for Biodiversity and Ecosystem Stewardship, Villanova University, 800 Lancaster Avenue, Villanova, Pennsylvania 19085 - 1699, USA & mbuehle 3 @ gmail. com; https: // orcid. org / 0000 - 0001 - 5460 - 7770 & adez. bandeira @ gmail. com; https: // orcid. org / 0000 - 0002 - 7496 - 9119 & Instituto Nacional da Biodiversidade e Áreas de Conservação, Ministério do Ambiente de Angola, Centralidade do Kilamba, Rua 26 de Fevereiro, Quarteirão Nimi Ya Lukemi, edifício Q 11, 3 º andar, Luanda, Angola. & adez. bandeira @ gmail. com; https: // orcid. org / 0000 - 0002 - 7496 - 9119</mods:affiliation>
<mods:affiliationid="B214990DACE7FC6C75225A0746FB87F1">Instituto Nacional da Biodiversidade e Áreas de Conservação, Ministério do Ambiente de Angola, Centralidade do Kilamba, Rua 26 de Fevereiro, Quarteirão Nimi Ya Lukemi, edifício Q 11, 3 º andar, Luanda, Angola. & adez. bandeira @ gmail. com; https: // orcid. org / 0000 - 0002 - 7496 - 9119</mods:affiliation>
<bibRefCitationid="F5484B6B5466FFDE9FE2F9531A1DF921"author="Bocage, J. V. B."box="[453,616,1708,1735]"pageId="21"pageNumber="322"pagination="105 - 114"refId="ref25633"refString="Bocage, J. V. B. (1896) Mammiferos, aves e reptis da Hanha, no sertao de Benguella. Jornal de Sciencias Mathematicas, Physicas e Naturaes, Segunda Serie, 4, 105 - 114."type="journal article"year="1896">Bocage, 1896</bibRefCitation>
<tableCitationid="DC5B03215466FFDE9EB8F9311974F90E"box="[159,257,1743,1769]"captionStart="TABLE 4"captionStartId="18.[151,239,404,428]"captionText="TABLE 4. Comparison between the different species of Lygodactylus occurring in Angola. Measurements are presented in millimeters (mm). Abbreviations are those described in Materials and Methods section."pageId="21"pageNumber="322">Tables 4</tableCitation>
;
<figureCitationid="09E22A1F5466FFDE9F2AF931190DF90E"box="[269,376,1743,1769]"captionStart-0="FIGURE 1"captionStart-1="FIGURE 2"captionStart-10="FIGURE 11"captionStart-2="FIGURE 3"captionStart-3="FIGURE 4"captionStart-4="FIGURE 5"captionStart-5="FIGURE 6"captionStart-6="FIGURE 7"captionStart-7="FIGURE 8"captionStart-8="FIGURE 9"captionStart-9="FIGURE 10"captionStartId-0="19.[151,250,1799,1823]"captionStartId-1="20.[151,250,1754,1778]"captionStartId-10="28.[151,250,1970,1994]"captionStartId-2="21.[151,248,1463,1487]"captionStartId-3="22.[151,250,1008,1032]"captionStartId-4="22.[151,250,1956,1980]"captionStartId-5="24.[151,250,1070,1094]"captionStartId-6="24.[151,250,1967,1991]"captionStartId-7="25.[151,250,1452,1476]"captionStartId-8="26.[151,250,1943,1967]"captionStartId-9="28.[151,250,1008,1032]"captionTargetBox-0="[203,1396,960,1759]"captionTargetBox-1="[151,1436,200,1730]"captionTargetBox-10="[151,1436,1092,1946]"captionTargetBox-2="[161,1426,190,1426]"captionTargetBox-3="[151,1437,181,985]"captionTargetBox-4="[151,1437,1129,1933]"captionTargetBox-5="[201,1386,306,1046]"captionTargetBox-6="[202,1386,1154,1943]"captionTargetBox-7="[157,1428,189,1420]"captionTargetBox-8="[152,1433,184,1919]"captionTargetBox-9="[151,1436,181,984]"captionTargetId-0="figure-386@19.[180,1408,915,1775]"captionTargetId-1="figure-22@20.[151,1436,200,1730]"captionTargetId-10="figure-56@28.[151,1436,1092,1946]"captionTargetId-2="figure-19@21.[151,1436,181,1440]"captionTargetId-3="figure-22@22.[151,1437,181,985]"captionTargetId-4="figure-74@22.[151,1437,1129,1933]"captionTargetId-5="figure-102@24.[201,1386,306,1046]"captionTargetId-6="figure-131@24.[202,1386,1154,1943]"captionTargetId-7="figure-19@25.[151,1436,181,1428]"captionTargetId-8="figure-22@26.[152,1436,181,1919]"captionTargetId-9="figure-22@28.[151,1436,181,984]"captionTargetPageId-0="19"captionTargetPageId-1="20"captionTargetPageId-10="28"captionTargetPageId-2="21"captionTargetPageId-3="22"captionTargetPageId-4="22"captionTargetPageId-5="24"captionTargetPageId-6="24"captionTargetPageId-7="25"captionTargetPageId-8="26"captionTargetPageId-9="28"captionText-0="FIGURE 1. Maximum likelihood phylogeny of Lygodactylus using ND2. Nodes supported by ML and BI are indicated by black dots. nodes only supported by the ML tree are indicated with half-filled black dots. nodes only supported by BI are indicated by half-filled red dots. and nodes with low support in both the ML and BI trees have red dots. Species of Lygodactylus occurring in Angola have a star next to them in the tree. Species within the Madgascan Lygodactylus clade include L. tolampyae, L. heterurus, L. arnoulti. Species in the Greater Drakensberg group include; L. nigropunctatus, L. graniticolus, L. ocellatus, L. soutpansbergensis. L. incognitus, L. montiscaeruli, L. waterbergensis, and L. methueni. Note that this tree should be used only as a guide to shallow relationships in the genus and to indicate relative divergence."captionText-1="FIGURE 2. Comparison of the number of precloacal pores in Angolan individuals of Lygodactylus angolensis, L. baptistai sp. nov., L. nyaneka sp. nov., L. lawrencei and L. tchokwe sp. nov."captionText-10="FIGURE 11. Lygodactylus baptistai sp. nov. in life (Holotype CAS 263557). Mamué. Serra da Neve. southwestern Angola Photos by Ishan Agarwal."captionText-2="FIGURE 3. Comparison of the nostrils position inAngolan individuals of:A. L. angolensis (AMB 9216); B. Lygodactylus baptistai sp. nov. (CAS 263557); C. L. nyaneka sp. nov. (UF 187229); D. L. lawrencei (CAS 264259); E. L. tchokwe sp. nov. (MD 5312b)."captionText-3="FIGURE 4. Comparison of the tail scales arrangement in Angolan individuals of A. Lygodactylus angolensis (MHNC-UP/REP 533); B. L. nyaneka sp. nov. (CAS 263531); C. L. baptistai sp. nov. (CAS 263557); D. L. lawrencei (CAS 264259); E. L. tchokwe sp. nov. (MD 5312d)."captionText-4="FIGURE5.Dorsalandventralwhole-bodyviews(left)anddorsal.lateralandventralviewsofthehead(right)oftheholotypeofLygodactylyslaurea
<treatmentCitationid="1078108B5466FFDE9FBCF8E81A21F8C9"author="Bocage, J. V. B."box="[411,596,1814,1838]"page="110"pageId="21"pageNumber="322"year="1896">
<bibRefCitationid="F5484B6B5466FFDE9FBCF8E81A21F8C9"author="Bocage, J. V. B."box="[411,596,1814,1838]"pageId="21"pageNumber="322"pagination="105 - 114"refId="ref25633"refString="Bocage, J. V. B. (1896) Mammiferos, aves e reptis da Hanha, no sertao de Benguella. Jornal de Sciencias Mathematicas, Physicas e Naturaes, Segunda Serie, 4, 105 - 114."type="journal article"year="1896">Bocage 1896: 110</bibRefCitation>
</treatmentCitation>
,
<treatmentCitationid="1078108B5466FFDE9C79F8E81AB3F8C9"author="Bocage, J. V. B."box="[606,710,1814,1838]"page="193"pageId="21"pageNumber="322"year="1897">
<bibRefCitationid="F5484B6B5466FFDE9C79F8E81AB3F8C9"author="Bocage, J. V. B."box="[606,710,1814,1838]"pageId="21"pageNumber="322"pagination="187 - 206"refId="ref25675"refString="Bocage, J. V. B. (1897) Mammiferos, reptis e batrachios d'Africa de que exitem exemplares typicos no Museu de Lisboa. Jornal de Sciencias Mathematicas, Physicas e Naturaes, Segunda Serie, 4, 187 - 206."type="journal article"year="1897">1897: 193</bibRefCitation>
<bibRefCitationid="F5484B6B5466FFDE9DAEF8E81C2AF8C9"author="Loveridge, A."box="[905,1119,1814,1838]"pageId="21"pageNumber="322"pagination="1 - 469"refId="ref27936"refString="Loveridge, A. (1947) Revision of the African lizards of the family Gekkonidae. Bulletin of the Museum of Comparative Zoology, 98, 1 - 469, pls. 1 - 7."type="journal article"year="1947">Loveridge 1947: 207</bibRefCitation>
<bibRefCitationid="F5484B6B5466FFDE9A4FF8E81CA5F8C9"author="Loveridge, A."box="[1128,1232,1814,1838]"pageId="21"pageNumber="322"pagination="151 - 362"refId="ref28000"refString="Loveridge, A. (1957) Checklist of the reptiles and amphibians of East Africa (Uganda, Kenya, Tanganyika, Zanzibar). Bulletin of the Museum of Comparative Zoology, 117, 151 - 362 + i-xxxvi."type="journal article"year="1957">1957: 187</bibRefCitation>
<bibRefCitationid="F5484B6B5466FFDE9AFEF8E81DEDF8C9"author="Hellmich, W."box="[1241,1432,1814,1838]"pageId="21"pageNumber="322"pagination="1 - 92"refId="ref27360"refString="Hellmich, W. (1957) Herpetologische Ergebnisse einer Forschungsreise in Angola. Veroffentlichungen der Zoologischen Staatssammlung Munchen, 5, 1 - 92."type="journal article"year="1957">Hellmich 1957: 35</bibRefCitation>
</treatmentCitation>
;
<bibRefCitationid="F5484B6B5466FFDE9EE0F8C7193FF8B6"author="Pasteur, G."box="[199,330,1849,1873]"pageId="21"pageNumber="322"pagination="1 - 132"refId="ref29152"refString="Pasteur, G. (1965 [1964]) Recherches sur l'evolution des lygodactyles, lezards Afro-Malgaches actuels. Travaux de l'Institut Scientific Cherifien, Serie Zoologie, 9, 1 - 132, pls. 1 - 12."type="journal article"year="1965">Pasteur 1965</bibRefCitation>
[1964]: 56;
<treatmentCitationid="1078108B5466FFDE9FE1F8C71A08F8B6"author="Branch, W. R."box="[454,637,1849,1873]"page="245"pageId="21"pageNumber="322"year="1998">
<bibRefCitationid="F5484B6B5466FFDE9FE1F8C71A08F8B6"author="Branch, W. R."box="[454,637,1849,1873]"pageId="21"pageNumber="322"refId="ref25855"refString="Branch, W. R. (1998) Field Guide to Snakes and Other Reptiles of Southern Africa. 3 rd Edition. Ralph Curtis Books, Sanibel Island, Florida, 399 pp. https: // doi. org / 10.1111 / evo. 13062"type="book"year="1998">Branch 1998: 245</bibRefCitation>
</treatmentCitation>
;
<treatmentCitationid="1078108B5466FFDE9CA0F8C71BCCF8B6"author="Broadley, D. G. & Cotterill, F. P. D."box="[647,953,1849,1873]"page="41"pageId="21"pageNumber="322"year="2004">
<bibRefCitationid="F5484B6B5466FFDE9CA0F8C71BCCF8B6"author="Broadley, D. G. & Cotterill, F. P. D."box="[647,953,1849,1873]"pageId="21"pageNumber="322"pagination="35 - 61"refId="ref26054"refString="Broadley, D. G. & Cotterill, F. P. D. (2004) The reptiles of southeast Katanga, an overlooked " hot spot. " African Journal of Herpetolgy, 53 (1), 35 - 61. https: // doi. org / 10.1080 / 21564574.2004.9635497"type="journal article"year="2004">Broadley & Cotterill 2004: 41</bibRefCitation>
</treatmentCitation>
;
<treatmentCitationid="1078108B5466FFDE9DE4F8C71CC0F8B6"author="Ceriaco, L. M. P. & de Sa, S. A. C. & Bandeira, S. & Valerio, H. & Stanley, E. L. & Khun, A. L. & Marques, M. P. & Vindum, J. V. & Blackburn, D. C. & Bauer, A. M."box="[963,1205,1849,1873]"page="57"pageId="21"pageNumber="322"year="2016">
<bibRefCitationid="F5484B6B5466FFDE9DE4F8C71CC0F8B6"author="Ceriaco, L. M. P. & de Sa, S. A. C. & Bandeira, S. & Valerio, H. & Stanley, E. L. & Khun, A. L. & Marques, M. P. & Vindum, J. V. & Blackburn, D. C. & Bauer, A. M."box="[963,1205,1849,1873]"pageId="21"pageNumber="322"pagination="15 - 61"refId="ref26185"refString="Ceriaco, L. M. P., de Sa, S. A. C., Bandeira, S., Valerio, H., Stanley, E. L., Khun, A. L., Marques, M. P., Vindum, J. V., Blackburn, D. C. & Bauer, A. M. (2016 a) Herpetological survey of Iona National Park and Namibe Regional Natural Park, with a synoptic list of the amphibians and reptiles of Namibe Province, southwestern Angola. Proceedings of the California Academy of Sciences, Series 4, 63, 15 - 61."type="journal article"year="2016">
<treatmentCitationid="1078108B5466FFDE9A98F8C71D52F8B6"author="Ceriaco, L. M. & Marques, M. P. & Bandeira, S."box="[1215,1319,1849,1873]"page="56"pageId="21"pageNumber="322"year="2016">
<bibRefCitationid="F5484B6B5466FFDE9A98F8C71D52F8B6"author="Ceriaco, L. M. & Marques, M. P. & Bandeira, S."box="[1215,1319,1849,1873]"pageId="21"pageNumber="322"refId="ref26304"refString="Ceriaco, L. M. P, Marques, M. P. & Bandeira, S. (2016 b) Anfibios e Repteis do Parque Nacional da Cangandala. Instituto Nacional da Biodiversidade e Areas de Conservacao, Angola & Museu Nacional de Historia Natural e da Ciencia, Lisboa, 91 pp."type="book"year="2016">2016b: 56</bibRefCitation>
</treatmentCitation>
, 57, 2018b: 413, 421, 422;
<bibRefCitationid="F5484B6B5466FFDE9F47F8A21A5AF893"author="Marques, M. P. & Ceriaco, L. M. P. & Blackburn, D. C. & Bauer, A. M."box="[352,559,1884,1908]"pageId="21"pageNumber="322"pagination="1 - 501"refId="ref28241"refString="Marques, M. P., Ceriaco, L. M. P., Blackburn, D. C. & Bauer, A. M. (2018) Diversity and distribution of the amphibians and terrestrial reptiles of Angola-Atlas of historical and bibliographic records (1840 - 2017). Proceedings of the California Academy of Sciences, Series 4, 65 (Supplement 2), 1 - 501."type="journal article"year="2018">
<treatmentCitationid="1078108B5466FFDE9CBFF8A21BF2F894"author="Branch, W. R. & Baptista, N. & Conradie, W."box="[664,903,1883,1908]"page="315"pageId="21"pageNumber="322"year="2019">
<bibRefCitationid="F5484B6B5466FFDE9CBFF8A21BF2F894"author="Branch, W. R. & Baptista, N. & Conradie, W."box="[664,903,1883,1908]"pageId="21"pageNumber="322"pagination="283 - 334"refId="ref25903"refString="Branch, W. R., Pinto. P. V., Baptista, N. & Conradie, W. (2019) The Reptiles of Angola: History, Diversity, Endemism and Hotspots. In: Huntley, B., Russo, V., Lages & Ferrand, N. (Eds.), Biodiversity of Angola, Science & Conservation: A Modern Synthesis. Springer, Cham, pp. 283 - 334. https: // doi. org / 10.1007 / 978 - 3 - 030 - 03083 - 4 _ 13"type="book chapter"year="2019">
<treatmentCitationid="1078108B5466FFDE9F52F8801A56F871"author="Schmidt, K. P."box="[373,547,1918,1942]"page="4"pageId="21"pageNumber="322"year="1933">
<bibRefCitationid="F5484B6B5466FFDE9F52F8801A56F871"author="Schmidt, K. P."box="[373,547,1918,1942]"pageId="21"pageNumber="322"pagination="1 - 15"refId="ref30093"refString="Schmidt, K. P. (1933) The reptiles of the Pulitzer-Angola Expedition. Annals of the Carnegie Museum, 22, 1 - 15, pls. 1 - 2."type="journal article"year="1933">Schmidt 1933: 4</bibRefCitation>
<bibRefCitationid="F5484B6B5466FFDE9C08F8811AAFF871"author="Mertens, R."box="[559,730,1918,1942]"pageId="21"pageNumber="322"pagination="1 - 23"refId="ref28633"refString="Mertens, R. (1937) Reptilien und Amphibien aus dem sudlichen Inner-Afrika. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 435, 1 - 23."type="journal article"year="1937">Mertens 1937: 6</bibRefCitation>
</treatmentCitation>
;
<treatmentCitationid="1078108B5466FFDE9CC1F8801C43F871"author="Barbour, T. & Loveridge, A."box="[742,1078,1918,1942]"page="147"pageId="21"pageNumber="322"year="1946">
<bibRefCitationid="F5484B6B5466FFDE9CC1F8801C43F871"author="Barbour, T. & Loveridge, A."box="[742,1078,1918,1942]"pageId="21"pageNumber="322"pagination="59 - 214"refId="ref25276"refString="Barbour, T. & Loveridge, A. (1946) First Supplement to typical reptiles and amphibians, Bulletin of the Museum of Comparative Zoology, 96 (2), 59 - 214."type="journal article"year="1946">Barbour & Loveridge 1946: 147</bibRefCitation>
<bibRefCitationid="F5484B6B5466FFDE9A66F8811C9FF871"author="Marx, H."box="[1089,1258,1918,1942]"pageId="21"pageNumber="322"pagination="409 - 496"refId="ref28492"refString="Marx, H. (1959) Catalogue of type specimens of reptiles and amphibians in Chicago Natural History Museum. Fieldiana Zoology, 36 (4), 409 - 496. https: // doi. org / 10.5962 / bhl. title. 3034"type="journal article"year="1959">Marx 1959: 464</bibRefCitation>
</treatmentCitation>
;
<treatmentCitationid="1078108B5466FFDE9AD2F880191BF85E"author="McCoy, C. J. & Richmond, N. D."page="154"pageId="21"pageNumber="322"year="1966">
<bibRefCitationid="F5484B6B5466FFDE9AD2F880191BF85E"author="McCoy, C. J. & Richmond, N. D."pageId="21"pageNumber="322"pagination="233 - 264"refId="ref28597"refString="McCoy, C. J. & Richmond, N. D. (1966) Herpetological type-specimens in Carnegie Museum. Annals of Carnegie Museum, 38 (10), 233 - 264."type="journal article"year="1966">McCoy & Richmond 1966: 154</bibRefCitation>
<treatmentCitationid="1078108B5466FFDE9FE0F83A1A00F83C"author="Bocage, J. V. B."box="[455,629,1987,2012]"page="15"pageId="21"pageNumber="322"year="1895">
<bibRefCitationid="F5484B6B5466FFDE9FE0F83A1A00F83C"author="Bocage, J. V. B."box="[455,629,1987,2012]"pageId="21"pageNumber="322"refId="ref25600"refString="Bocage, J. V. B. (1895) Herpetologie d'Angola et du Congo. Ministerio da Marinha e das Colonias, Lisbonne, 203 pp., 20 pls."type="book"year="1895">Bocage 1895: 15</bibRefCitation>
<taxonomicNameid="56D94D195465FFDD9ED2FBEB1912FBCA"authority="Marques & Ceríaco & Buehler & Bandei- Ra & Janota & Bauer, 2020"authorityName="Marques & Ceríaco & Buehler & Bandei- Ra & Janota & Bauer"authorityYear="2020"box="[245,359,1045,1069]"class="Reptilia"family="Gekkonidae"genus="Lygodactylus"kingdom="Animalia"order="Squamata"pageId="22"pageNumber="323"phylum="Chordata"rank="species"species="nyaneka"status="sp. nov.">
<taxonomicNameid="56D94D195465FFDD9CB3FBEB1B7FFBCA"authority="Marques & Ceríaco & Buehler & Bandei- Ra & Janota & Bauer, 2020"authorityName="Marques & Ceríaco & Buehler & Bandei- Ra & Janota & Bauer"authorityYear="2020"box="[660,778,1045,1069]"class="Reptilia"family="Gekkonidae"genus="Lygodactylus"kingdom="Animalia"order="Squamata"pageId="22"pageNumber="323"phylum="Chordata"rank="species"species="baptistai"status="sp. nov.">
<taxonomicNameid="56D94D195465FFDD9BAFFBEB189FFBB6"authority="Marques & Ceríaco & Buehler & Bandei- Ra & Janota & Bauer, 2020"authorityName="Marques & Ceríaco & Buehler & Bandei- Ra & Janota & Bauer"authorityYear="2020"class="Reptilia"family="Gekkonidae"genus="Lygodactylus"kingdom="Animalia"order="Squamata"pageId="22"pageNumber="323"phylum="Chordata"rank="species"species="tchokwe"status="sp. nov.">
and surrounding countries, the species was only described at the end of the nineteenth century based on two specimens collected by the Portuguese explorer José d’Anchieta “on the walls of a corral” in Hanha,
<bibRefCitationid="F5484B6B5464FFDC9C96FF221B3DFF10"author="Bocage, J. V. B."box="[689,840,220,247]"pageId="23"pageNumber="324"pagination="105 - 114"refId="ref25633"refString="Bocage, J. V. B. (1896) Mammiferos, aves e reptis da Hanha, no sertao de Benguella. Jornal de Sciencias Mathematicas, Physicas e Naturaes, Segunda Serie, 4, 105 - 114."type="journal article"year="1896">Bocage 1896</bibRefCitation>
).
<bibRefCitationid="F5484B6B5464FFDC9D7CFF231C75FF10"author="Bocage, J. V. B."box="[859,1024,220,247]"pageId="23"pageNumber="324"pagination="105 - 114"refId="ref25633"refString="Bocage, J. V. B. (1896) Mammiferos, aves e reptis da Hanha, no sertao de Benguella. Jornal de Sciencias Mathematicas, Physicas e Naturaes, Segunda Serie, 4, 105 - 114."type="journal article"year="1896">Bocage (1896)</bibRefCitation>
noted that some specimens from Cahata and Galanga, both in
<bibRefCitationid="F5484B6B5464FFDC9D8EFF011C4AFEFE"author="Bocage, J. V. B."box="[937,1087,254,281]"pageId="23"pageNumber="324"refId="ref25600"refString="Bocage, J. V. B. (1895) Herpetologie d'Angola et du Congo. Ministerio da Marinha e das Colonias, Lisbonne, 203 pp., 20 pls."type="book"year="1895">Bocage 1895</bibRefCitation>
<bibRefCitationid="F5484B6B5464FFDC9D96FE531C2CFE21"author="Bocage, J. V. B."box="[945,1113,428,455]"pageId="23"pageNumber="324"pagination="187 - 206"refId="ref25675"refString="Bocage, J. V. B. (1897) Mammiferos, reptis e batrachios d'Africa de que exitem exemplares typicos no Museu de Lisboa. Jornal de Sciencias Mathematicas, Physicas e Naturaes, Segunda Serie, 4, 187 - 206."type="journal article"year="1897">Bocage (1897)</bibRefCitation>
mistakenly considered both the specimens from Hanha and those from Cahata and Galanga as
<bibRefCitationid="F5484B6B5464FFDC9C9BFDEA1B00FDC9"author="Sternfeld, R."box="[700,885,532,558]"pageId="23"pageNumber="324"pagination="412 - 420"refId="ref30236"refString="Sternfeld, R. (1911) Zur Herpetologie Sudostafrikas. Mitteilungen aus dem Zoologischen Museum in Berlin, 5, 412 - 420."type="journal article"year="1911">Sternfeld (1911)</bibRefCitation>
<bibRefCitationid="F5484B6B5464FFDC9B0FFDEA18A5FDB6"author="Boulenger, G. A."pageId="23"pageNumber="324"pagination="77 - 86"refId="ref25815"refString="Boulenger, G. A. (1890) First report on additions to the lizard collection in the British Museum (Natural History). Proceedings of the Zoological Society of London, 1890, 77 - 86."type="journal article"year="1890">Boulenger 1890</bibRefCitation>
<bibRefCitationid="F5484B6B5464FFDC9A77FDC91D6DFDB6"author="Loveridge, A."box="[1104,1304,567,593]"pageId="23"pageNumber="324"pagination="1 - 469"refId="ref27936"refString="Loveridge, A. (1947) Revision of the African lizards of the family Gekkonidae. Bulletin of the Museum of Comparative Zoology, 98, 1 - 469, pls. 1 - 7."type="journal article"year="1947">Loveridge (1947)</bibRefCitation>
, which was the first record of the species out of
(in error) were later provided by Loveridge (1933) and
<bibRefCitationid="F5484B6B5464FFDC9C40FD821B79FD71"author="Pitman, C. R. S."box="[615,780,636,662]"pageId="23"pageNumber="324"pagination="292 - 312"refId="ref29385"refString="Pitman, C. R. S. (1934) A check list of Reptilia and Amphibia occurring and believed to occur in Northern Rhodesia. In: Pitman, C. R. S., Report on a Faunal Survey of Northern Rhodesia, Livingstone, 1934, pp. xii + 500 + Index of i-xxxii, maps A-K (herpetology on pp. 292 - 312)."type="book chapter"year="1934">Pitman (1934)</bibRefCitation>
, respectively.
<bibRefCitationid="F5484B6B5464FFDC9D97FD821C16FD71"author="Schmidt, K. P."box="[944,1123,636,662]"pageId="23"pageNumber="324"pagination="1 - 15"refId="ref30093"refString="Schmidt, K. P. (1933) The reptiles of the Pulitzer-Angola Expedition. Annals of the Carnegie Museum, 22, 1 - 15, pls. 1 - 2."type="journal article"year="1933">Schmidt (1933)</bibRefCitation>
, based on the material collected by the Pulitzer Angolan Expedition in Chitau,
<figureCitationid="09E22A1F5464FFDC9E87FD3F1891FD3B"box="[160,228,705,732]"captionStart="FIGURE 5"captionStartId="22.[151,250,1956,1980]"captionTargetBox="[151,1437,1129,1933]"captionTargetId="figure-74@22.[151,1437,1129,1933]"captionTargetPageId="22"captionText="FIGURE 5. Dorsal and ventral whole-body views (left) and dorsal. lateral and ventral views of the head (right) of the holotype of Lygodactylys laureae (CM 5860)."figureDoi="http://doi.org/10.5281/zenodo.4410737"httpUri="https://zenodo.org/record/4410737/files/figure.png"pageId="23"pageNumber="324">Fig. 5</figureCitation>
<bibRefCitationid="F5484B6B5464FFDC9DADFD3F1C24FD3C"author="Loveridge, A."box="[906,1105,705,731]"pageId="23"pageNumber="324"pagination="1 - 469"refId="ref27936"refString="Loveridge, A. (1947) Revision of the African lizards of the family Gekkonidae. Bulletin of the Museum of Comparative Zoology, 98, 1 - 469, pls. 1 - 7."type="journal article"year="1947">Loveridge (1947)</bibRefCitation>
, and recent collection of new specimens very close to Chitau (
<collectionCodeid="F7C8AE5F5464FFDC9D63FD1A1B0FFD19"box="[836,890,740,766]"country="USA"httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690"lsid="urn:lsid:biocol.org:col:34700"name="California Academy of Sciences"pageId="23"pageNumber="324"type="Museum">CAS</collectionCode>
266003–266009) revealed that the only species present in the region corresponds to
<bibRefCitationid="F5484B6B5464FFDC9CEBFCF91BE5FCC6"author="Loveridge, A."box="[716,912,775,801]"pageId="23"pageNumber="324"pagination="1 - 469"refId="ref27936"refString="Loveridge, A. (1947) Revision of the African lizards of the family Gekkonidae. Bulletin of the Museum of Comparative Zoology, 98, 1 - 469, pls. 1 - 7."type="journal article"year="1947">Loveridge (1947)</bibRefCitation>
<bibRefCitationid="F5484B6B5464FFDC9D3DFCB31BDEFC80"author="Pasteur, G."box="[794,939,844,871]"pageId="23"pageNumber="324"pagination="1 - 132"refId="ref29152"refString="Pasteur, G. (1965 [1964]) Recherches sur l'evolution des lygodactyles, lezards Afro-Malgaches actuels. Travaux de l'Institut Scientific Cherifien, Serie Zoologie, 9, 1 - 132, pls. 1 - 12."type="journal article"year="1965">Pasteur 1965</bibRefCitation>
[1964] and
<bibRefCitationid="F5484B6B5464FFDC9A16FCB21CB5FC81"author="Branch, W. R."box="[1073,1216,844,870]"pageId="23"pageNumber="324"refId="ref25855"refString="Branch, W. R. (1998) Field Guide to Snakes and Other Reptiles of Southern Africa. 3 rd Edition. Ralph Curtis Books, Sanibel Island, Florida, 399 pp. https: // doi. org / 10.1111 / evo. 13062"type="book"year="1998">Branch 1998</bibRefCitation>
for estimates of the distribution of the species), but no precise locality records have been published from
<bibRefCitationid="F5484B6B5464FFDC9ED2FC4A1994FC29"author="Ceriaco, L. M. & Marques, M. P. & Bandeira, S."box="[245,481,948,974]"pageId="23"pageNumber="324"refId="ref26304"refString="Ceriaco, L. M. P, Marques, M. P. & Bandeira, S. (2016 b) Anfibios e Repteis do Parque Nacional da Cangandala. Instituto Nacional da Biodiversidade e Areas de Conservacao, Angola & Museu Nacional de Historia Natural e da Ciencia, Lisboa, 91 pp."type="book"year="2016">
<bibRefCitationid="F5484B6B5464FFDC9FCBFC4A1A42FC28"author="Ceriaco, L. M. P. & Marques, M. P. & Bandeira, S. & Blackburn, D. C. & Bauer, A. M."box="[492,567,948,975]"pageId="23"pageNumber="324"pagination="408 - 431"refId="ref26460"refString="Ceriaco, L. M. P., Marques, M. P., Bandeira, S., Blackburn, D. C. & Bauer, A. M. (2018 b) Herpetological survey of Cangandala National Park, with a synoptic list of the amphibians and reptiles of Malanje Province, central Angola. Herpetological Review, 49 (3), 408 - 431."type="journal article"year="2018">2018b</bibRefCitation>
by dirt road from Cassumbi village) provinces (see Specimens examined below).
<bibRefCitationid="F5484B6B5464FFDC9A0EFC291D69FC16"author="Marques, M. P. & Ceriaco, L. M. P. & Blackburn, D. C. & Bauer, A. M."box="[1065,1308,983,1009]"pageId="23"pageNumber="324"pagination="1 - 501"refId="ref28241"refString="Marques, M. P., Ceriaco, L. M. P., Blackburn, D. C. & Bauer, A. M. (2018) Diversity and distribution of the amphibians and terrestrial reptiles of Angola-Atlas of historical and bibliographic records (1840 - 2017). Proceedings of the California Academy of Sciences, Series 4, 65 (Supplement 2), 1 - 501."type="journal article"year="2018">
<bibRefCitationid="F5484B6B5464FFDC9EB0FC071920FBF3"author="Laurent, R. F."box="[151,341,1017,1044]"pageId="23"pageNumber="324"pagination="1 - 165"refId="ref27865"refString="Laurent, R. F. (1964) Reptiles et amphibiens de l'Angola (Troiseme contribution). Publicacoes Culturais. Companhia de Diamantes de Angola, 67, 1 - 165."type="journal article"year="1964">Laurent’s (1964)</bibRefCitation>
, but subsequent examination of these specimens, which are currently housed in the Museu do Dundo, do not support this synonymization (see account for
<taxonomicNameid="56D94D195464FFDC9B2FFBE21886FBBE"authority="Marques & Ceríaco & Buehler & Bandei- Ra & Janota & Bauer, 2020"authorityName="Marques & Ceríaco & Buehler & Bandei- Ra & Janota & Bauer"authorityYear="2020"class="Reptilia"family="Gekkonidae"genus="Lygodactylus"kingdom="Animalia"order="Squamata"pageId="23"pageNumber="324"phylum="Chordata"rank="species"species="tchokwe"status="sp. nov.">
supralabial, one postnasal); 2) nostrils separated from the rostral and pierced above the suture between rostral and first supralabial; 3) mental followed by three postmentals; 4) first infralabial <25% overlap with postpostmental; 5) 19 to 21 rows of ventral scales; 6) between 7 and 10 precloacal pores in males; 7) median subcaudals arranged in an irregular pattern of one row of scales about ⅓ width of the tail alternating with two rows (sometimes just one) of scales of about ½ the width of the tail (
<figureCitationid="09E22A1F5464FFDC9A5FFACF1CA7FAAC"box="[1144,1234,1329,1355]"captionStart="FIGURE 4"captionStartId="22.[151,250,1008,1032]"captionTargetBox="[151,1437,181,985]"captionTargetId="figure-22@22.[151,1437,181,985]"captionTargetPageId="22"captionText="FIGURE 4. Comparison of the tail scales arrangement in Angolan individuals of A. Lygodactylus angolensis (MHNC-UP/REP 533); B. L. nyaneka sp. nov. (CAS 263531); C. L. baptistai sp. nov. (CAS 263557); D. L. lawrencei (CAS 264259); E. L. tchokwe sp. nov. (MD 5312d)."figureDoi="http://doi.org/10.5281/zenodo.4410735"httpUri="https://zenodo.org/record/4410735/files/figure.png"pageId="23"pageNumber="324">Fig. 4A</figureCitation>
); 8) venter cream; 9) without markings on the gular region. The background coloration is usually light grey or light olive-brown. Some individuals have a dark stripe from the snout, through the eye, ending near the forearm insertion. The upper flank is marked with a series of pale spots forming a pale light line. These spots are usually edged with a dark mark above and below sometimes yielding the appearance of an ocellus (
<figureCitationid="09E22A1F5464FFDC9D67FA421BD8FA30"box="[832,941,1468,1495]"captionStart-0="FIGURE 6"captionStart-1="FIGURE 7"captionStartId-0="24.[151,250,1070,1094]"captionStartId-1="24.[151,250,1967,1991]"captionTargetBox-0="[201,1386,306,1046]"captionTargetBox-1="[202,1386,1154,1943]"captionTargetId-0="figure-102@24.[201,1386,306,1046]"captionTargetId-1="figure-131@24.[202,1386,1154,1943]"captionTargetPageId-0="24"captionTargetPageId-1="24"captionText-0="FIGURE 6. Dorsal and ventral whole-body views (left) and dorsal. lateral and ventral views of the head (right) of Lygodactylys angolensis (MHNC-UP/REP 495)."captionText-1="FIGURE 7. Lygodactylus angolensis in life (MHNC-UP/REP 495). Base camp surroundings in a banana plantation [-11.08391º, 16.66367º, 1240 m a.s.l.]. 14 km by dirt road from Cassumbi. Bié Province. Angola Plateau. Photo by Luis M. P. Ceríaco."figureDoi-0="http://doi.org/10.5281/zenodo.4410739"figureDoi-1="http://doi.org/10.5281/zenodo.4410741"httpUri-0="https://zenodo.org/record/4410739/files/figure.png"httpUri-1="https://zenodo.org/record/4410741/files/figure.png"pageId="23"pageNumber="324">Figs. 6–7</figureCitation>
