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<document id="03E61A329D4DB52E2A9A1CC6D30AD16E" ID-DOI="10.5281/zenodo.6464279" ID-ISBN="978-1-7397467-0-4" ID-ISSN="2754-9844" ID-ZooBank="FE033A11-ACCB-4C6D-B185-63420F8CA79B" IM.tables_requiresApprovalFor="GgImagineBatch" checkinTime="1699386246229" checkinUser="felipe" docAuthor="Hansson, Christer" docDate="2022" docId="03B387856524B81FFDFC6CCC21D9C854" docLanguage="en" docName="TaxMonogNeotropHymenopt.1.1-137.pdf" docOrigin="Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 1 (1)" docSource="http://dx.doi.org/10.5281/zenodo.6464278" docStyle="DocumentStyle:DAA2ADC4E839520FEEF1E0A10B782A41.1:TaxMonogNeotropHymenopt.2022-.journal_article" docStyleId="DAA2ADC4E839520FEEF1E0A10B782A41" docStyleName="TaxMonogNeotropHymenopt.2022-.journal_article" docStyleVersion="1" docTitle="Closterocerus barbatus Hansson 2022, sp.nov." docType="treatment" docUuid="4892A7CE-DFB1-445A-A640-3FD59B798DE6" docUuidSource="ZooBank" docVersion="2" lastPageNumber="29" masterDocId="FF8AFFFD653AB801FFC86C732579CE20" masterDocTitle="Eulophidae Of Costa Rica (Hymenoptera: Chalcidoidea), 4 The Genus Closterocerus Westwood S. Str." masterLastPageNumber="137" masterPageNumber="1" pageNumber="29" updateTime="1699839903426" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="E1B6AE4E09BEC6BBF14F735ED8823B0E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="3CE56076B95146C341235A859899B3E7">Eulophidae Of Costa Rica (Hymenoptera: Chalcidoidea), 4 The Genus Closterocerus Westwood S. Str.</mods:title>
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<mods:name id="0890E48BF0E151ABFC834A4A8B95BDF3" type="personal">
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<mods:namePart id="44681A384EFB9837BDDE1186F1143A9D">Hansson, Christer</mods:namePart>
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<mods:title id="6F682D4AA402CA5DFAA71123BE299710">Taxonomic Monographs on Neotropical Hymenoptera</mods:title>
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<mods:part id="294DF7CABFB1765D6DC5756B46569218">
<mods:date id="1F33F7BF84A315DF5F463C5336324B04">2022</mods:date>
<mods:detail id="256FC88294AC3735037DDAAB67C5A12B" type="series">
<mods:title id="3B779F1126D314CAC34CCC5D91F2FDF0">Oxford, England</mods:title>
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<mods:number id="650F62393029A6A7452037CD49AA48F6">2022-04-28</mods:number>
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<mods:detail id="DF20B9639BD13D9F8F93D40881D065AF" type="volume">
<mods:number id="12F209C9DB17C53AEAE099C02197020F">1</mods:number>
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<mods:classification id="4DCA5607BAD76CC83E5CE178F727C901">journal article</mods:classification>
<mods:identifier id="7020A3F479664F5B0C5E8935CA75CA0E" type="DOI">10.5281/zenodo.6464279</mods:identifier>
<mods:identifier id="A1AD5D2B376435B6B42C949DDC7A4E32" type="ISBN">978-1-7397467-0-4</mods:identifier>
<mods:identifier id="4992DEB7E08D4B02AF5B27A5A2513345" type="ISSN">2754-9844</mods:identifier>
<mods:identifier id="199F69E7665F4E2B62A00FE6FC570108" type="ZooBank">FE033A11-ACCB-4C6D-B185-63420F8CA79B</mods:identifier>
</mods:mods>
<treatment id="03B387856524B81FFDFC6CCC21D9C854" LSID="urn:lsid:zoobank.org:act:4892A7CE-DFB1-445A-A640-3FD59B798DE6" httpUri="http://treatment.plazi.org/id/03B387856524B81FFDFC6CCC21D9C854" lastPageNumber="29" pageId="30" pageNumber="29">
<subSubSection id="C30065186524B81FFDFC6CCC26F1CEF7" box="[564,904,191,215]" pageId="30" pageNumber="29" type="nomenclature">
<paragraph id="8BA536936524B81FFDFC6CCC26F1CEF7" blockId="30.[564,904,191,247]" box="[564,904,191,215]" pageId="30" pageNumber="29">
<heading id="D0ED81FF6524B81FFDFC6CCC26F1CEF7" bold="true" box="[564,904,191,215]" centered="true" fontSize="10" level="2" pageId="30" pageNumber="29" reason="2">
<emphasis id="B96EEA816524B81FFDFC6CCC26F1CEF7" bold="true" box="[564,904,191,215]" pageId="30" pageNumber="29">
<taxonomicName id="4C1A4D106524B81FFDFC6CCC264BCEF7" authority="Hansson, 2022" authorityName="Hansson" authorityYear="2022" box="[564,818,191,215]" class="Insecta" family="Eulophidae" genus="Closterocerus" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="29" phylum="Arthropoda" rank="species" species="barbatus" status="sp.nov.">
<emphasis id="B96EEA816524B81FFDFC6CCC264BCEF7" bold="true" box="[564,818,191,215]" italics="true" pageId="30" pageNumber="29">Closterocerus barbatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25D57FA6524B81FFCF16CCC26F1CEF7" box="[825,904,191,215]" pageId="30" pageNumber="29" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30065186524B81FFD526CAC265ACED7" box="[666,803,223,247]" pageId="30" pageNumber="29" type="description">
<paragraph id="8BA536936524B81FFD526CAC265ACED7" blockId="30.[564,904,191,247]" box="[666,803,223,247]" pageId="30" pageNumber="29">
(
<figureCitation id="13212A166524B81FFD696CAC278DCED7" box="[673,756,223,247]" captionStart="Figs 47–52" captionStartId="107.[181,228,1662,1686]" captionTargetBox="[183,1224,190,1657]" captionTargetId="figure-8@107.[183,1224,190,1659]" captionTargetPageId="107" captionText="Figs 47–52. Closterocerus spp. C. aureopurpureus, paratype ♂ – (47) habitus dorsal view; (48) habitus lateral view. C. azofeifai – (49) habitus dorsal view, holotype ♀; (50) habitus lateral view, holotype ♀; (51) habitus lateral view, paratype ♂. C. barbatus, holotype ♀ – (52) habitus dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10117000" httpUri="https://zenodo.org/record/10117000/files/figure.png" pageId="30" pageNumber="29">Figs 52</figureCitation>
,
<figureCitation id="13212A166524B81FFD376CAC2662CED7" box="[767,795,223,247]" captionStart="Figs 53–58" captionStartId="108.[213,260,1552,1576]" captionTargetBox="[213,1251,190,1542]" captionTargetId="figure-7@108.[213,1251,190,1542]" captionTargetPageId="108" captionText="Figs 53–58. Closterocerus spp. C. brevipes – (53) habitus dorsal view, holotype ♀; (54) habitus lateral view, holotype ♀; (55) habitus dorsal view, paratype ♂; (56) habitus lateral view, paratype ♂; (57) hind leg, holotype ♀. C. barbatus, holotype ♀ – (58) head frontal view." figureDoi="http://doi.org/10.5281/zenodo.10117002" httpUri="https://zenodo.org/record/10117002/files/figure.png" pageId="30" pageNumber="29">58</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30065186524B81FFF1D6D6C24F0CB77" pageId="30" pageNumber="29" type="diagnosis">
<paragraph id="8BA536936524B81FFF1D6D6C2191CFD7" blockId="30.[213,1256,287,504]" pageId="30" pageNumber="29">
<emphasis id="B96EEA816524B81FFF1D6D6C2430CF17" bold="true" box="[213,329,287,311]" pageId="30" pageNumber="29">Diagnosis.</emphasis>
Scape and pedicel flattened, flagellum less flattened, scape subtriangular and widest just below apex, apical three flagellomeres ±fused to form a clava (
<figureCitation id="13212A166524B81FFCAF6D4C26CCCF77" box="[871,949,319,343]" captionStart="Figs 53–58" captionStartId="108.[213,260,1552,1576]" captionTargetBox="[213,1251,190,1542]" captionTargetId="figure-7@108.[213,1251,190,1542]" captionTargetPageId="108" captionText="Figs 53–58. Closterocerus spp. C. brevipes – (53) habitus dorsal view, holotype ♀; (54) habitus lateral view, holotype ♀; (55) habitus dorsal view, paratype ♂; (56) habitus lateral view, paratype ♂; (57) hind leg, holotype ♀. C. barbatus, holotype ♀ – (58) head frontal view." figureDoi="http://doi.org/10.5281/zenodo.10117002" httpUri="https://zenodo.org/record/10117002/files/figure.png" pageId="30" pageNumber="29">Fig. 58</figureCitation>
); occiput strongly hairy (
<figureCitation id="13212A166524B81FFB746D4C258ACF57" captionStart="Figs 47–52" captionStartId="107.[181,228,1662,1686]" captionTargetBox="[183,1224,190,1657]" captionTargetId="figure-8@107.[183,1224,190,1659]" captionTargetPageId="107" captionText="Figs 47–52. Closterocerus spp. C. aureopurpureus, paratype ♂ – (47) habitus dorsal view; (48) habitus lateral view. C. azofeifai – (49) habitus dorsal view, holotype ♀; (50) habitus lateral view, holotype ♀; (51) habitus lateral view, paratype ♂. C. barbatus, holotype ♀ – (52) habitus dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10117000" httpUri="https://zenodo.org/record/10117000/files/figure.png" pageId="30" pageNumber="29">Fig. 52</figureCitation>
); pronotal collar long and smooth, with a strong carina along anterior margin (
<figureCitation id="13212A166524B81FFBDF6D2C211DCF57" box="[1047,1124,351,375]" captionStart="Figs 47–52" captionStartId="107.[181,228,1662,1686]" captionTargetBox="[183,1224,190,1657]" captionTargetId="figure-8@107.[183,1224,190,1659]" captionTargetPageId="107" captionText="Figs 47–52. Closterocerus spp. C. aureopurpureus, paratype ♂ – (47) habitus dorsal view; (48) habitus lateral view. C. azofeifai – (49) habitus dorsal view, holotype ♀; (50) habitus lateral view, holotype ♀; (51) habitus lateral view, paratype ♂. C. barbatus, holotype ♀ – (52) habitus dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10117000" httpUri="https://zenodo.org/record/10117000/files/figure.png" pageId="30" pageNumber="29">Fig. 52</figureCitation>
); midlobe of mesoscutum with one pair of setae (
<figureCitation id="13212A166524B81FFD856D0C27E2CFB7" box="[589,667,383,407]" captionStart="Figs 47–52" captionStartId="107.[181,228,1662,1686]" captionTargetBox="[183,1224,190,1657]" captionTargetId="figure-8@107.[183,1224,190,1659]" captionTargetPageId="107" captionText="Figs 47–52. Closterocerus spp. C. aureopurpureus, paratype ♂ – (47) habitus dorsal view; (48) habitus lateral view. C. azofeifai – (49) habitus dorsal view, holotype ♀; (50) habitus lateral view, holotype ♀; (51) habitus lateral view, paratype ♂. C. barbatus, holotype ♀ – (52) habitus dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10117000" httpUri="https://zenodo.org/record/10117000/files/figure.png" pageId="30" pageNumber="29">Fig. 52</figureCitation>
); mesoscutum and mesoscutellum with strong engraved reticulation (
<figureCitation id="13212A166524B81FFE976DEC24C8CF97" box="[351,433,415,439]" captionStart="Figs 47–52" captionStartId="107.[181,228,1662,1686]" captionTargetBox="[183,1224,190,1657]" captionTargetId="figure-8@107.[183,1224,190,1659]" captionTargetPageId="107" captionText="Figs 47–52. Closterocerus spp. C. aureopurpureus, paratype ♂ – (47) habitus dorsal view; (48) habitus lateral view. C. azofeifai – (49) habitus dorsal view, holotype ♀; (50) habitus lateral view, holotype ♀; (51) habitus lateral view, paratype ♂. C. barbatus, holotype ♀ – (52) habitus dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10117000" httpUri="https://zenodo.org/record/10117000/files/figure.png" pageId="30" pageNumber="29">Fig. 52</figureCitation>
); petiole 0.5× as long as wide with strong sculpture on dorsal surface; fore wing hyaline with median one-third infuscate (
<figureCitation id="13212A166524B81FFD746DCC2673CFF7" box="[700,778,447,471]" captionStart="Figs 47–52" captionStartId="107.[181,228,1662,1686]" captionTargetBox="[183,1224,190,1657]" captionTargetId="figure-8@107.[183,1224,190,1659]" captionTargetPageId="107" captionText="Figs 47–52. Closterocerus spp. C. aureopurpureus, paratype ♂ – (47) habitus dorsal view; (48) habitus lateral view. C. azofeifai – (49) habitus dorsal view, holotype ♀; (50) habitus lateral view, holotype ♀; (51) habitus lateral view, paratype ♂. C. barbatus, holotype ♀ – (52) habitus dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10117000" httpUri="https://zenodo.org/record/10117000/files/figure.png" pageId="30" pageNumber="29">Fig. 52</figureCitation>
), speculum large and open below and towards base of wing; gaster with tuft of white hairs anterolaterally (
<figureCitation id="13212A166524B81FFC816DAC26EECFD7" box="[841,919,479,503]" captionStart="Figs 47–52" captionStartId="107.[181,228,1662,1686]" captionTargetBox="[183,1224,190,1657]" captionTargetId="figure-8@107.[183,1224,190,1659]" captionTargetPageId="107" captionText="Figs 47–52. Closterocerus spp. C. aureopurpureus, paratype ♂ – (47) habitus dorsal view; (48) habitus lateral view. C. azofeifai – (49) habitus dorsal view, holotype ♀; (50) habitus lateral view, holotype ♀; (51) habitus lateral view, paratype ♂. C. barbatus, holotype ♀ – (52) habitus dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10117000" httpUri="https://zenodo.org/record/10117000/files/figure.png" pageId="30" pageNumber="29">Fig. 52</figureCitation>
); length of body female
<quantity id="4CE29B766524B81FFB5E6DAC219CCFD7" box="[1174,1253,479,503]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="30" pageNumber="29" unit="mm" value="1.9">1.9mm</quantity>
.
</paragraph>
<paragraph id="8BA536936524B81FFF1D6E6C27F3CC17" blockId="30.[213,1257,543,1304]" box="[213,650,543,567]" pageId="30" pageNumber="29">
Female
<typeStatus id="54A188316524B81FFEE26E6C24F5CC17" box="[298,396,543,567]" pageId="30" pageNumber="29">holotype</typeStatus>
: length of body
<quantity id="4CE29B766524B81FFDFF6E6C27FFCC17" box="[567,646,543,567]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="30" pageNumber="29" unit="mm" value="1.9">1.9mm</quantity>
.
</paragraph>
<paragraph id="8BA536936524B81FFF326E4C2743CCD7" blockId="30.[213,1257,543,1304]" pageId="30" pageNumber="29">Colour: Scape and pedicel dark brown to black, flagellum pale brown. Frons below frontofacial suture golden-green, above suture golden, close to vertex metallic purple. Vertex golden. Mesoscutum, mesoscutellum, dorsellum and propodeum black. Coxae black; femora, mid and hind tibiae dark brown, fore tibia brown with apex yellowish-brown; fore tarsus dark brown; mid and hind tarsi with T1–3 white, T4 black. Fore wing hyaline with median one-third infuscate, hind wing hyaline. Petiole black. Gaster dark brown.</paragraph>
<paragraph id="8BA536936524B81FFF326E8C24D2CD57" blockId="30.[213,1257,543,1304]" pageId="30" pageNumber="29">Head: Scape and pedicel flattened, flagellum less flattened, scape ±triangular and widest just below apex. Frons below frontofacial suture with weak reticulation; above suture and vertex smooth. Frontofacial suture V-shaped. Subtorular sutures missing. Occipital margin rounded. Occiput strongly hairy.</paragraph>
<paragraph id="8BA536936524B81FFF326F0C210CCA17" blockId="30.[213,1257,543,1304]" pageId="30" pageNumber="29">Pronotal collar long and smooth, with a strong carina along anterior margin. Mesoscutum with strong and engraved reticulation; notauli distinct in anterior one-half; midlobe with one pair of setae, attached in posterior part. Mesoscutellum convex, with strong and engraved reticulation. Dorsellum convex and predominantly smooth. Propodeum with a ±complete median carina; medially smooth and laterally with rather strong reticulation; callus with two setae. Fore wing with speculum large, open below and towards base of wing; with a stigmal hairline and with radial cell bare.</paragraph>
<paragraph id="8BA536936524B81FFF32684C2100CA57" blockId="30.[213,1257,543,1304]" pageId="30" pageNumber="29">Petiole 0.5× as long as wide, with strong sculpture on dorsal surface. Gaster oval-shaped and non-collapsed; with a tuft of white hair anterolaterally; tergites with strong reticulation.</paragraph>
<paragraph id="8BA536936524B81FFF32680C213ACB37" blockId="30.[213,1257,543,1304]" pageId="30" pageNumber="29">Relative measurements: head length dorsal 17; head length frontal 27.5; head width 42; POL 9; OOL 4; lateral ocellus maximum width 3; eye length 18.5; malar space 9; mouth width 12; mesosoma length 65; mesosoma width 43; mesoscutellum length 26; mesoscutellum width 28; fore wing length 111; fore wing width 59; marginal vein length 60; postmarginal vein length 1.5; stigmal vein length 6.5; fore wing marginal fringe length 7.5; gaster length 64; gaster width 53.</paragraph>
<paragraph id="8BA536936524B81FFF1D694C24F0CB77" blockId="30.[213,393,1343,1367]" box="[213,393,1343,1367]" pageId="30" pageNumber="29">Male. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C30065186524B81FFF1D690C24EBCBB7" box="[213,402,1407,1431]" pageId="30" pageNumber="29" type="biology_ecology">
<paragraph id="8BA536936524B81FFF1D690C24EBCBB7" blockId="30.[213,402,1407,1431]" box="[213,402,1407,1431]" pageId="30" pageNumber="29">
<emphasis id="B96EEA816524B81FFF1D690C246CCBB7" bold="true" box="[213,277,1407,1431]" pageId="30" pageNumber="29">Hosts</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30065186524B81FFF1D69CC24B5CBF7" box="[213,460,1471,1495]" pageId="30" pageNumber="29" type="distribution">
<paragraph id="8BA536936524B81FFF1D69CC24B5CBF7" blockId="30.[213,460,1471,1495]" box="[213,460,1471,1495]" pageId="30" pageNumber="29">
<emphasis id="B96EEA816524B81FFF1D69CC241ECBF7" bold="true" box="[213,359,1471,1495]" pageId="30" pageNumber="29">Distribution.</emphasis>
<collectingCountry id="F30D76036524B81FFEA569CC24B1CBF7" box="[365,456,1471,1495]" name="Ecuador" pageId="30" pageNumber="29">Ecuador</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C30065186524B81FFF1D698C21BCC814" pageId="30" pageNumber="29" type="materials_examined">
<paragraph id="8BA536936524B81FFF1D698C21BCC814" blockId="30.[213,1221,1535,1588]" pageId="30" pageNumber="29">
<emphasis id="B96EEA816524B81FFF1D698C24CAC837" bold="true" box="[213,435,1535,1559]" pageId="30" pageNumber="29">Material examined.</emphasis>
<materialsCitation id="3B723CCE6524B81FFF326A6D21BCC814" box="[250,1221,1566,1588]" collectingDate="1983-02-25" collectionCode="MZLU" collectorName="L. Huggert" country="Ecuador" elevation="3700" location="Papallacta" pageId="30" pageNumber="29" specimenCount="1" specimenCount-female="1" stateProvince="Napo" typeStatus="holotype">
Type material.
<typeStatus id="54A188316524B81FFE436A6D249DC814" box="[395,484,1566,1588]" pageId="30" pageNumber="29">Holotype</typeStatus>
<specimenCount id="9D1CFD1A6524B81FFE226A6D2485C813" box="[490,508,1566,1587]" pageId="30" pageNumber="29" type="female">♀</specimenCount>
“
<collectingCountry id="F30D76036524B81FFDC36A6D27FDC813" box="[523,644,1566,1587]" name="Ecuador" pageId="30" pageNumber="29">ECUADOR</collectingCountry>
:
<collectingRegion id="49DEF8716524B81FFD456A6D27BAC813" box="[653,707,1566,1587]" country="Ecuador" name="Napo" pageId="30" pageNumber="29">Napo</collectingRegion>
,
<location id="8EC560486524B81FFD056A6D264BC814" LSID="urn:lsid:plazi:treatment:03B387856524B81FFDFC6CCC21D9C854:8EC560486524B81FFD056A6D264BC814" box="[717,818,1566,1588]" country="Ecuador" name="Papallacta" pageId="30" pageNumber="29" stateProvince="Napo">Papallacta</location>
,
<quantity id="4CE29B766524B81FFCF46A6D26F8C813" box="[828,897,1566,1587]" metricMagnitude="3" metricUnit="m" metricValue="3.7" pageId="30" pageNumber="29" unit="m" value="3700.0">
<elevation id="0037D1A06524B81FFCF46A6D26F8C813" box="[828,897,1566,1587]" metricMagnitude="3" metricUnit="m" metricValue="3.7" pageId="30" pageNumber="29" unit="m" value="3700.0">3700m</elevation>
</quantity>
,
<date id="FFA410536524B81FFC436A6D2697C814" box="[907,1006,1566,1588]" pageId="30" pageNumber="29" value="1983-02-25">
<collectingDate id="EFE0E9BB6524B81FFC436A6D2697C814" box="[907,1006,1566,1588]" pageId="30" pageNumber="29" value="1983-02-25">25.ii.1983</collectingDate>
</date>
,
<collectorName id="26EF53456524B81FFC306A6D211AC813" box="[1016,1123,1566,1587]" pageId="30" pageNumber="29">L. Huggert</collectorName>
” (
<collectionCode id="ED0BAE566524B81FFBB06A6D21C6C814" box="[1144,1215,1566,1588]" country="Sweden" lsid="urn:lsid:biocol.org:col:34994" name="Lund University" pageId="30" pageNumber="29">MZLU</collectionCode>
)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30065186524B81FFF1D6A2F21D9C854" box="[213,1184,1628,1652]" pageId="30" pageNumber="29" type="etymology">
<paragraph id="8BA536936524B81FFF1D6A2F21D9C854" blockId="30.[213,1184,1628,1652]" box="[213,1184,1628,1652]" pageId="30" pageNumber="29">
<emphasis id="B96EEA816524B81FFF1D6A2F242CC854" bold="true" box="[213,341,1628,1652]" pageId="30" pageNumber="29">Etymology.</emphasis>
From the Latin
<taxonomicName id="4C1A4D106524B81FFDCA6A2F2718C854" authorityName="Hansson" authorityYear="2022" box="[514,609,1628,1652]" class="Insecta" family="Eulophidae" genus="Closterocerus" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="29" phylum="Arthropoda" rank="species" species="barbatus">
<emphasis id="B96EEA816524B81FFDCA6A2F2718C854" box="[514,609,1628,1652]" italics="true" pageId="30" pageNumber="29">barbatus</emphasis>
</taxonomicName>
= bearded, referring to the hairy temples and occiput.
</paragraph>
</subSubSection>
</treatment>
</document>
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