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169 lines
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.669.11773" ID-GBIF-Dataset="e2f4b6e8-d909-4231-867a-08808dfc66ca" ID-Pensoft-Pub="1313-2970-669-113" ID-ZBK="13D2ECC5A53C4A19A00C406E940BADD2" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-669-113" ModsDocOrigin="ZooKeys 669" ModsDocTitle="Revision of the Neotropical water scavenger beetle genus Tobochares Short & García, 2007 (Coleoptera, Hydrophilidae, Acidocerinae)" checkinTime="1492859332780" checkinUser="pensoft" docAuthor="Kohlenberg, Alex T. & Short, Andrew Edward Z." docDate="2017" docId="052959C9FB4D39D1C553F393CFF2BFD0" docLanguage="en" docName="ZooKeys 669: 113-146" docOrigin="ZooKeys 669" docSource="http://dx.doi.org/10.3897/zookeys.669.11773" docTitle="Tobochares sp. A" docType="treatment" docVersion="4" lastPageNumber="138" masterDocId="FFB58A38FFA2FFE95A3EFF9EA741B933" masterDocTitle="Revision of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae)" masterLastPageNumber="146" masterPageNumber="113" pageNumber="138" updateTime="1643557144543" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kohlenberg, Alex T.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Short, Andrew Edward Z.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>669</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>113</mods:start>
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<mods:end>146</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.669.11773</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.669.11773</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-669-113</mods:identifier>
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<mods:identifier type="ZBK">13D2ECC5A53C4A19A00C406E940BADD2</mods:identifier>
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<mods:identifier type="ZooBank">13D2ECC5A53C4A19A00C406E940BADD2</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="128696888" LSID="urn:lsid:plazi:treatment:052959C9FB4D39D1C553F393CFF2BFD0" httpUri="http://treatment.plazi.org/id/052959C9FB4D39D1C553F393CFF2BFD0" lastPageNumber="138" pageId="25" pageNumber="138">
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<subSubSection pageId="25" pageNumber="138" type="nomenclature">
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<paragraph pageId="25" pageNumber="138">
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<taxonomicName authority="sp. A" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="25" pageNumber="138" phylum="Arthropoda" rank="genus">Tobochares sp. A</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="25" pageNumber="138" type="materials_examined">
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<paragraph pageId="25" pageNumber="138">Material examined</paragraph>
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<paragraph pageId="25" pageNumber="138">(1). VENEZUELA: Amazonas: Tobogan de la Selva, leg. M. Balke (1 female, SEMC; DNA voucher SLE526).</paragraph>
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</subSubSection>
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<subSubSection pageId="25" pageNumber="138" type="differential diagnosis">
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<paragraph pageId="25" pageNumber="138">Differential diagnosis.</paragraph>
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<paragraph pageId="25" pageNumber="138">
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This species is morphologically very similar to
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<taxonomicName genus="T." lsidName="T. pallidus" pageId="25" pageNumber="138" rank="species" species="pallidus">T. pallidus</taxonomicName>
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, and shares most diagnostic features of that species (in particular its very pale coloration). This species can be separated from
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<taxonomicName genus="T." lsidName="T. pallidus" pageId="25" pageNumber="138" rank="species" species="pallidus">T. pallidus</taxonomicName>
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by the faint but distinctly impressed striae in the posterior quarter of the elytra, and the slightly less emarginated eyes.
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</paragraph>
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<caption pageId="25" pageNumber="138">
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<paragraph pageId="25" pageNumber="138">
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Figure 14. Aedeagi (ventral view) of
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<taxonomicName class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="25" pageNumber="138" phylum="Arthropoda" rank="genus">Tobochares</taxonomicName>
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spp. A
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<taxonomicName genus="T." lsidName="T. kusad" pageId="25" pageNumber="138" rank="species" species="kusad">T. kusad</taxonomicName>
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sp. n. B
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<taxonomicName genus="T." lsidName="T. sulcatus" pageId="25" pageNumber="138" rank="species" species="sulcatus">T. sulcatus</taxonomicName>
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C
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<taxonomicName genus="T." lsidName="T. sipaliwini" pageId="25" pageNumber="138" rank="species" species="sipaliwini">T. sipaliwini</taxonomicName>
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D
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<taxonomicName genus="T." lsidName="T. striatus" pageId="25" pageNumber="138" rank="species" species="striatus">T. striatus</taxonomicName>
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E
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<taxonomicName genus="T." lsidName="T. kasikasima" pageId="25" pageNumber="138" rank="species" species="kasikasima">T. kasikasima</taxonomicName>
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F
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<taxonomicName genus="T." lsidName="T. pallidus" pageId="25" pageNumber="138" rank="species" species="pallidus">T. pallidus</taxonomicName>
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sp. n. G
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<taxonomicName genus="T." lsidName="T. canaliculatus" pageId="25" pageNumber="138" rank="species" species="canaliculatus">T. canaliculatus</taxonomicName>
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sp. n. H
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<taxonomicName genus="T." lsidName="T. canthus" pageId="25" pageNumber="138" rank="species" species="canthus">T. canthus</taxonomicName>
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sp. n. I
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<taxonomicName genus="T." lsidName="T. emarginatus" pageId="25" pageNumber="138" rank="species" species="emarginatus">T. emarginatus</taxonomicName>
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sp. n.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="25" pageNumber="138" type="remarks">
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<paragraph pageId="25" pageNumber="138">Remarks.</paragraph>
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<paragraph pageId="25" pageNumber="138">
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This species is only known from a single specimen, which we also extracted for DNA. Molecular data also supports this taxon as sister to
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<taxonomicName genus="T." lsidName="T. pallidus" pageId="25" pageNumber="138" rank="species" species="pallidus">T. pallidus</taxonomicName>
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, to which it is also similar morphologically. We refrain from describing the species until additional specimens, including ideally a male, can be found.
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</paragraph>
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<caption pageId="25" pageNumber="138">
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<paragraph pageId="25" pageNumber="138">
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Figure 15. Distribution of
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<taxonomicName class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="25" pageNumber="138" phylum="Arthropoda" rank="genus">Tobochares</taxonomicName>
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spp.
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</paragraph>
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</caption>
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<caption pageId="25" pageNumber="138">
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<paragraph pageId="25" pageNumber="138">
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Figure 16. Habitat of
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<taxonomicName class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="25" pageNumber="138" phylum="Arthropoda" rank="genus">Tobochares</taxonomicName>
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spp. in Suriname.
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<normalizedToken originalValue="A–B">A-B</normalizedToken>
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Base of Voltzberg, collecting event SR16-0316-01C, habitat of
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<taxonomicName genus="T." lsidName="T. sipaliwini" pageId="25" pageNumber="138" rank="species" species="sipaliwini">T. sipaliwini</taxonomicName>
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<normalizedToken originalValue="C–D">C-D</normalizedToken>
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Waterfall in the upper Palumeu watershed, collecting event SR12-0311-03A, habitat of
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<taxonomicName genus="T." lsidName="T. emarginatus" pageId="25" pageNumber="138" rank="species" species="emarginatus">T. emarginatus</taxonomicName>
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sp. n. E Base of Voltzberg, collecting event SR16-0316-01A, habitat of
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<taxonomicName genus="T." lsidName="T. sipaliwini" pageId="25" pageNumber="138" rank="species" species="sipaliwini">T. sipaliwini</taxonomicName>
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F Kasikasima, collecting event SR16-0324-01C, habitat of
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<taxonomicName genus="T." lsidName="T. sipaliwini" pageId="25" pageNumber="138" rank="species" species="sipaliwini">T. sipaliwini</taxonomicName>
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,
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<taxonomicName genus="T." lsidName="T. kasikasima" pageId="25" pageNumber="138" rank="species" species="kasikasima">T. kasikasima</taxonomicName>
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,
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<taxonomicName genus="T." lsidName="T. striatus" pageId="25" pageNumber="138" rank="species" species="striatus">T. striatus</taxonomicName>
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, and
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<taxonomicName genus="T." lsidName="T. emarginatus" pageId="25" pageNumber="138" rank="species" species="emarginatus">T. emarginatus</taxonomicName>
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sp. n. Red arrows point to example microhabitats where specimens were collected
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</paragraph>
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</caption>
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<caption pageId="25" pageNumber="138">
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<paragraph pageId="25" pageNumber="138">
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Figure 17. Habitat of
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<taxonomicName class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="25" pageNumber="138" phylum="Arthropoda" rank="genus">Tobochares</taxonomicName>
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spp. in Guyana and Suriname.
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<normalizedToken originalValue="A–B">A-B</normalizedToken>
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Guyana, Kusad Mt., collecting event GY13-1027-03B, type locality for
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<taxonomicName genus="T." lsidName="T. kusad" pageId="25" pageNumber="138" rank="species" species="kusad">T. kusad</taxonomicName>
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sp. n.
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<normalizedToken originalValue="C–D">C-D</normalizedToken>
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Suriname, side of Kappel Airstrip, collecting event SR13-0824-02B, habitat of
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<taxonomicName genus="T." lsidName="T. kasikasima" pageId="25" pageNumber="138" rank="species" species="kasikasima">T. kasikasima</taxonomicName>
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<normalizedToken originalValue="E–F">E-F</normalizedToken>
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Suriname, summit of Tafelberg, Caiman Creek seepage, collecting event SR13-0819-01A, habitat of
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<taxonomicName genus="T." lsidName="T. kasikasima" pageId="25" pageNumber="138" rank="species" species="kasikasima">T. kasikasima</taxonomicName>
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. Red arrows point to example microhabitats where specimens were collected.
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</paragraph>
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</caption>
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<caption pageId="25" pageNumber="138">
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<paragraph pageId="25" pageNumber="138">
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Figure 18. Habitat of
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<taxonomicName class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="25" pageNumber="138" phylum="Arthropoda" rank="genus">Tobochares</taxonomicName>
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spp. in Venezuela.
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<normalizedToken originalValue="A–B">A-B</normalizedToken>
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Tobogan de la Selva, upstream, collecting event AS-08-080b, type locality for
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<taxonomicName genus="T." lsidName="T. canaliculatus" pageId="25" pageNumber="138" rank="species" species="canaliculatus">T. canaliculatus</taxonomicName>
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sp. n.,
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<taxonomicName genus="T." lsidName="T. sulcatus" pageId="25" pageNumber="138" rank="species" species="sulcatus">T. sulcatus</taxonomicName>
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,
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<taxonomicName genus="T." lsidName="T. canthus" pageId="25" pageNumber="138" rank="species" species="canthus">T. canthus</taxonomicName>
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sp. n., and
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<taxonomicName genus="T." lsidName="T. pallidus" pageId="25" pageNumber="138" rank="species" species="pallidus">T. pallidus</taxonomicName>
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sp. n. C Tobogan de la Selva, collecting event VZ09-0114-01D D Outcrop near Pijiguaos, collecting event VZ10-0708-01B, habitat of
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<taxonomicName genus="T." lsidName="T. pallidus" pageId="25" pageNumber="138" rank="species" species="pallidus">T. pallidus</taxonomicName>
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sp. n.
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<normalizedToken originalValue="E–F">E-F</normalizedToken>
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Rock outcrop near Pijiguaos, collecting event VZ10-0709-01B, habitat of
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<taxonomicName genus="T." lsidName="T. pallidus" pageId="25" pageNumber="138" rank="species" species="pallidus">T. pallidus</taxonomicName>
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sp. n. Red arrows point to example microhabitats where specimens were collected.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document>
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