<documentid="66381E29B7052CCDAF3CAA4CFCA87BA7"ID-DOI="10.5852/ejt.2023.865.2097"ID-GBIF-Dataset="3a63a183-743e-4185-8fdc-1080072ada0a"ID-ISSN="2118-9773"ID-Zenodo-Dep="7867448"ID-ZooBank="845BAE1A-0E6E-484B-BE6D-17F866777B51"IM.bibliography_approvedBy="diego"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="felipe"IM.taxonomicNames_approvedBy="felipe"IM.treatments_approvedBy="diego"checkinTime="1682512407979"checkinUser="felipe"docAuthor="Rivera-Quiroz, F. Andrés & Álvarez-Padilla, Fernando"docDate="2023"docId="03C8E927FFF7FFD5FDE5FB44FEBC6453"docLanguage="en"docName="ejt-865_1-94.pdf"docOrigin="European Journal of Taxonomy 865"docSource="http://dx.doi.org/10.5852/ejt.2023.865.2097"docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1"docStyleId="EF2B578F1D15862ADE45B0C07C620911"docStyleName="EJT.2018-.journal_article.type1"docStyleVersion="14"docTitle="Anyphaena salgueiroi Rivera-Quiroz & Álvarez-Padilla 2023, sp. nov."docType="treatment"docVersion="10"lastPageNumber="70"masterDocId="FFF1915FFFB4FF90FF81FFF8FF856160"masterDocTitle="Integration or minimalism: twenty-one new species of ghost spiders (Anyphaenidae: Anyphaena) from Mexico"masterLastPageNumber="94"masterPageNumber="1"pageNumber="68"updateTime="1732489834501"updateUser="ExternalLinkService"zenodo-license-document="CC-BY-4.0"zenodo-license-figures="CC-BY-4.0"zenodo-license-treatments="UNSPECIFIED">
<mods:titleid="32EF046942EDA9C477F90EDBF763BFFD">Integration or minimalism: twenty-one new species of ghost spiders (Anyphaenidae: Anyphaena) from Mexico</mods:title>
<mods:affiliationid="0B10A9A11AE14840B2E371340F014D35">Understanding Evolution Research Group, Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, the Netherlands.</mods:affiliation>
<mods:affiliationid="E375ACFF449C5B3C847F8897AAD6D61E">Laboratorio de Aracnología, Departamento de Biología Comparada, Facultad de Ciencias, Universidad Nacional Autónoma de México, Distrito Federal, México Ciudad Universitaria, México D. F. Del. Coyoacán, C. P. 04510, Mexico.</mods:affiliation>
<taxonomicNameid="4C6123B2FFF7FFD3FDE5FB44FCEA65B7"ID-CoL="C66LQ"authority="Rivera-Quiroz & Álvarez-Padilla, 2023"authorityName="Rivera-Quiroz & Álvarez-Padilla"authorityYear="2023"box="[612,879,1212,1239]"class="Arachnida"family="Anyphaenidae"genus="Anyphaena"kingdom="Animalia"order="Araneae"pageId="67"pageNumber="68"phylum="Arthropoda"rank="species"species="salgueiroi"status="sp. nov.">
<figureCitationid="135A44B4FFF7FFD3FD01FAFBFD7E647D"box="[640,763,1283,1309]"captionStart="Fig"captionStartId="7.[189,232,1783,1809]"captionTargetBox="[192,1396,266,1720]"captionTargetId="figure-80@7.[189,1399,265,1728]"captionTargetPageId="7"captionText="Fig. 2. Anyphaena spp., epigynum anatomies. A–B. A. adnani sp. nov., paratype, ♀ (CNAN-T01546). A. Epigynum, ventral view. B. Epigynum, dorsal view. – C–D. A. rebecae sp. nov., holotype, ♀ (CNAN-T01536). C. Epigynum, ventral view. D. Epigynum, dorsal view. – E–F. A. salgueiroi sp. nov., holotype, ♀ (CNAN-T01537). E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.7867461"httpUri="https://zenodo.org/record/7867461/files/figure.png"pageId="67"pageNumber="68">Figs 2E–F</figureCitation>
,
<figureCitationid="135A44B4FFF7FFD3FC87FAFBFCA3647D"box="[774,806,1283,1309]"captionStart="Fig"captionStartId="68.[190,233,1928,1954]"captionTargetBox="[192,1395,685,1875]"captionTargetId="figure-160@68.[189,1398,684,1885]"captionTargetPageId="68"captionText="Fig. 42. Anyphaena salgueiroi sp. nov., holotype, ♀ (CNAN-T01537). A. Dorsal habitus. B. Lateral habitus. C. Ventral habitus. D. Prosoma, anterior view. E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: A–C = 1.0 mm; D =0.5 mm; E–F= 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.7867580"httpUri="https://zenodo.org/record/7867580/files/figure.png"pageId="67"pageNumber="68">42</figureCitation>
,
<figureCitationid="135A44B4FFF7FFD3FCB3FAFBFC0C647D"box="[818,905,1283,1309]"captionStart="Fig"captionStartId="71.[189,232,1928,1954]"captionTargetBox="[195,1379,615,1895]"captionTargetId="figure-143@71.[189,1398,605,1903]"captionTargetPageId="71"captionText="Fig. 44. Anyphaena spp. A–B. A. salgueiroi sp. nov., holotype, ♀ (CNAN-T01537). A. Epigynum, ventral view. B. Epigynum, dorsal view. – C–D. A. ibarrai sp. nov., holotype, ♀ (CNAN-T01528). C. Epigynum, ventral view. D. Epigynum, dorsal view. Scale bars: 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.7867586"httpUri="https://zenodo.org/record/7867586/files/figure.png"pageId="67"pageNumber="68">44A–B</figureCitation>
,
<figureCitationid="135A44B4FFF7FFD3FC17FAFAFC36647D"box="[918,947,1282,1309]"captionStart="Fig"captionStartId="82.[189,232,1018,1044]"captionTargetBox="[193,1392,269,964]"captionTargetId="figure-209@82.[189,1398,265,976]"captionTargetPageId="82"captionText="Fig. 52. Localities for the species of Anyphaena. Black square=Atotonilco Plots I and II, separated by 400 m, nine species registered: A. bifurcata sp. nov., A. bromelicola Platnick, 1977 (ON619634- 38 Supp. file 1), A. catalina Platnick, 1974 (ON619673-75), A. dulceae sp. nov., A. epicardia sp. nov., A. fernandae sp. nov., A. porta sp. nov., A. salgueiroi sp. nov. and A. sofiae sp. nov. White square=Xamaticpac Plots I and II, plots contiguous, eleven species registered: A. adnani sp. nov., A. alachua Platnick, 1974 (ON619665), A. fraterna (Banks, 1896) (ON619671), A. megamedia sp. nov., A. miniducta sp. nov., A. natachae sp. nov., A. porta sp. nov., A. quadrata sp. nov., A. rebecae sp. nov., A. stigma sp. nov. and A. tonoi sp. nov. Black circle =Xilitla, four species registered: A. franciscoi sp. nov., A. ibarrai sp. nov., A. noctua sp. nov. and A. triangularis sp. nov. White circle =Los Tuxtlas, two species registered: A. jimenezi sp. nov. and A. urieli sp. nov."figureDoi="http://doi.org/10.5281/zenodo.7867604"httpUri="https://zenodo.org/record/7867604/files/figure.png"pageId="67"pageNumber="68">52</figureCitation>
<taxonomicNameid="4C6123B2FFF7FFD3FECDFA8BFE6664EC"authority="Rivera-Quiroz & Álvarez-Padilla, 2023"authorityName="Rivera-Quiroz & Álvarez-Padilla"authorityYear="2023"box="[332,483,1394,1421]"class="Arachnida"family="Anyphaenidae"genus="Anyphaena"kingdom="Animalia"order="Araneae"pageId="67"pageNumber="68"phylum="Arthropoda"rank="species"species="salgueiroi"status="sp. nov.">
<taxonomicNameid="4C6123B2FFF7FFD3FC7FFA8BFBE564EC"authority="Rivera-Quiroz & Álvarez-Padilla, 2023"authorityName="Rivera-Quiroz & Álvarez-Padilla"authorityYear="2023"box="[1022,1120,1395,1421]"class="Arachnida"family="Anyphaenidae"genus="Anyphaena"kingdom="Animalia"order="Araneae"pageId="67"pageNumber="68"phylum="Arthropoda"rank="species"species="porta"status="sp. nov.">
<taxonomicNameid="4C6123B2FFF7FFD3FA82FA8BFAFD64EC"authority="Rivera-Quiroz & Álvarez-Padilla, 2023"authorityName="Rivera-Quiroz & Álvarez-Padilla"authorityYear="2023"box="[1283,1400,1394,1421]"class="Arachnida"family="Anyphaenidae"genus="Anyphaena"kingdom="Animalia"order="Araneae"pageId="67"pageNumber="68"phylum="Arthropoda"rank="species"species="ibarrai"status="sp. nov.">
<paragraphid="8BDE5831FFF7FFD3FF3CF9B2FE2D67E7"blockId="67.[189,1399,1610,1671]"pageId="67"pageNumber="68">The species epithet is dedicated to Dr Francisco Javier Salgueiro-Sepúlveda, former member of the second author’s lab.</paragraph>
<collectingRegionid="49A596D3FFF7FFD3FEFAF8F9FE63667B"box="[379,486,1793,1819]"country="Mexico"name="Veracruz de Ignacio de la Llave"pageId="67"pageNumber="68">Veracruz</collectingRegion>
<collectingDateid="EF9B8719FFF7FFD3FCDFF852FBB466A4"box="[862,1073,1962,1989]"pageId="67"pageNumber="68"value="2012-05-21"valueMax="2012-05-30"valueMin="2012-05-21">21–30 May 2012</collectingDate>
Total length 8.2. Carapace orange over cephalic area with two red longitudinal bands over most of thoracic area, lateral margins yellow, clypeus red. Sternum pale orange, darker at margins, intercoxal triangles present on all legs. Labium red, white at tip, longer than wide. Endites pale orange, rectangular, slightly broader at tip. Chelicerae orange, promargin with five teeth, retromargin with six to seven denticles. Leg coloration: yellow at base, turning red after femora to tarsi (
<figureCitationid="135A44B4FFF0FFD4FBB9FDB4FB496306"box="[1080,1228,588,614]"captionStart="Fig"captionStartId="68.[190,233,1928,1954]"captionTargetBox="[192,1395,685,1875]"captionTargetId="figure-160@68.[189,1398,684,1885]"captionTargetPageId="68"captionText="Fig. 42. Anyphaena salgueiroi sp. nov., holotype, ♀ (CNAN-T01537). A. Dorsal habitus. B. Lateral habitus. C. Ventral habitus. D. Prosoma, anterior view. E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: A–C = 1.0 mm; D =0.5 mm; E–F= 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.7867580"httpUri="https://zenodo.org/record/7867580/files/figure.png"pageId="68"pageNumber="69">Fig. 42A–D</figureCitation>
). Lateral sides of abdomen dorsum covered with red reticulated pattern leaving clearer central longitudinal band, lateral sides with same pattern, less dense ventrally. Tracheal spiracle at center, anterior half of abdomen surface almost without red pattern, posterior half dominated by reddish longitudinal lines (
<figureCitationid="135A44B4FFF1FFD5FB5BFEABFAEF600D"box="[1242,1386,339,365]"captionStart="Fig"captionStartId="68.[190,233,1928,1954]"captionTargetBox="[192,1395,685,1875]"captionTargetId="figure-160@68.[189,1398,684,1885]"captionTargetPageId="68"captionText="Fig. 42. Anyphaena salgueiroi sp. nov., holotype, ♀ (CNAN-T01537). A. Dorsal habitus. B. Lateral habitus. C. Ventral habitus. D. Prosoma, anterior view. E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: A–C = 1.0 mm; D =0.5 mm; E–F= 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.7867580"httpUri="https://zenodo.org/record/7867580/files/figure.png"pageId="69"pageNumber="70">Fig. 42A–C</figureCitation>
). Epigynum atrium rectangular, longer than wide, door-shaped. Hood narrow and straight. Copulatory openings inside two semicircular sclerotized plates (
<figureCitationid="135A44B4FFF1FFD5FCA9FE64FC3660D7"box="[808,947,412,439]"captionStart="Fig"captionStartId="68.[190,233,1928,1954]"captionTargetBox="[192,1395,685,1875]"captionTargetId="figure-160@68.[189,1398,684,1885]"captionTargetPageId="68"captionText="Fig. 42. Anyphaena salgueiroi sp. nov., holotype, ♀ (CNAN-T01537). A. Dorsal habitus. B. Lateral habitus. C. Ventral habitus. D. Prosoma, anterior view. E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: A–C = 1.0 mm; D =0.5 mm; E–F= 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.7867580"httpUri="https://zenodo.org/record/7867580/files/figure.png"pageId="69"pageNumber="70">Fig. 42E–F</figureCitation>
). Copulatory ducts weakly sclerotized following external edge of semicircular plates, turning 180° beyond hood, and entering spermathecae ventrally at junction between long seminal receptacles and fertilization duct origin. Fertilization ducts short and curved (
<figureCitationid="135A44B4FFF1FFD5FE13FDF3FD986345"box="[402,541,523,549]"captionStart="Fig"captionStartId="68.[190,233,1928,1954]"captionTargetBox="[192,1395,685,1875]"captionTargetId="figure-160@68.[189,1398,684,1885]"captionTargetPageId="68"captionText="Fig. 42. Anyphaena salgueiroi sp. nov., holotype, ♀ (CNAN-T01537). A. Dorsal habitus. B. Lateral habitus. C. Ventral habitus. D. Prosoma, anterior view. E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: A–C = 1.0 mm; D =0.5 mm; E–F= 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.7867580"httpUri="https://zenodo.org/record/7867580/files/figure.png"pageId="69"pageNumber="70">Figs 42E–F</figureCitation>
,
<figureCitationid="135A44B4FFF1FFD5FDA8FDF3FDCD6345"box="[553,584,523,549]"captionStart="Fig"captionStartId="71.[189,232,1928,1954]"captionTargetBox="[195,1379,615,1895]"captionTargetId="figure-143@71.[189,1398,605,1903]"captionTargetPageId="71"captionText="Fig. 44. Anyphaena spp. A–B. A. salgueiroi sp. nov., holotype, ♀ (CNAN-T01537). A. Epigynum, ventral view. B. Epigynum, dorsal view. – C–D. A. ibarrai sp. nov., holotype, ♀ (CNAN-T01528). C. Epigynum, ventral view. D. Epigynum, dorsal view. Scale bars: 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.7867586"httpUri="https://zenodo.org/record/7867586/files/figure.png"pageId="69"pageNumber="70">44</figureCitation>
A-B). Cephalothorax length 2.98, thoracic width 2.24, cephalic width 1.35. Clypeus height 0.11. Eye diameters: AME 0.1, ALE 0.13, PME 0.13, PLE 0.12. Eye interdistances: AME–AME 0.06, AME–ALE 0.04, ALE–PLE 0.18, PME–PME 0.17, PME–PLE 0.11. Femur lengths: I 2.52, II 2.42, III 1.97, IV 2.58. Leg spination: femur I d1-1-1, p0-1-1, r0-1-1. Tibia I v2-2-0, p1-1-1, r1- 1-1. Metatarsus I v2-2-0, p1-1-1, r1-1-1. Femur II d1-1-1, p0-1-1, r0-1-1. Tibia II v2-2-0, p1-1-1, r1-1-1. Metatarsus II v2-2-0, p1-1-1, r1-1-1. Femur III d1-1-1, p0-1-1, r0-1-1. Tibia III v2-2-2, p1-1-1, r1-1-1. Metatarsus III v2-2-2, p1-1-2, r1-1-2. Femur IV d1-1-1, p0-0-1, r0-0-1. Tibia IV v1-2-2, p1-1-1, r1-1-1. Metatarsus IV v2-2-2, p1-1-2, r1-1-2.
<taxonomicNameid="4C6123B2FFF0FFD4FEA7F870FDAF66C2"authority="Rivera-Quiroz & Álvarez-Padilla, 2023"authorityName="Rivera-Quiroz & Álvarez-Padilla"authorityYear="2023"box="[294,554,1928,1954]"class="Arachnida"family="Anyphaenidae"genus="Anyphaena"kingdom="Animalia"order="Araneae"pageId="68"pageNumber="69"phylum="Arthropoda"rank="species"species="salgueiroi"status="sp. nov.">
This species is found in oak and oak with pine forest fragments around Pico de Orizaba Volcano National Park (
<figureCitationid="135A44B4FFF1FFD5FE80FB5DFEDC65A0"box="[257,345,1189,1216]"captionStart="Fig"captionStartId="82.[189,232,1018,1044]"captionTargetBox="[193,1392,269,964]"captionTargetId="figure-209@82.[189,1398,265,976]"captionTargetPageId="82"captionText="Fig. 52. Localities for the species of Anyphaena. Black square=Atotonilco Plots I and II, separated by 400 m, nine species registered: A. bifurcata sp. nov., A. bromelicola Platnick, 1977 (ON619634- 38 Supp. file 1), A. catalina Platnick, 1974 (ON619673-75), A. dulceae sp. nov., A. epicardia sp. nov., A. fernandae sp. nov., A. porta sp. nov., A. salgueiroi sp. nov. and A. sofiae sp. nov. White square=Xamaticpac Plots I and II, plots contiguous, eleven species registered: A. adnani sp. nov., A. alachua Platnick, 1974 (ON619665), A. fraterna (Banks, 1896) (ON619671), A. megamedia sp. nov., A. miniducta sp. nov., A. natachae sp. nov., A. porta sp. nov., A. quadrata sp. nov., A. rebecae sp. nov., A. stigma sp. nov. and A. tonoi sp. nov. Black circle =Xilitla, four species registered: A. franciscoi sp. nov., A. ibarrai sp. nov., A. noctua sp. nov. and A. triangularis sp. nov. White circle =Los Tuxtlas, two species registered: A. jimenezi sp. nov. and A. urieli sp. nov."figureDoi="http://doi.org/10.5281/zenodo.7867604"httpUri="https://zenodo.org/record/7867604/files/figure.png"pageId="69"pageNumber="70">Fig. 52</figureCitation>