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353 lines
66 KiB
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<document id="E350B93DCD86BD2A01A0625CB0BB53AE" ID="10.11646/zootaxa.4139.1.4" ID-DOI="10.11646/zootaxa.4139.1.4" ID-GBIF-Dataset="3cf8edb7-4be5-4ec3-803e-ca1f224d703b" ID-ISSN="1175-5326" ID-Zenodo-Dep="271999" ID-ZooBank="2744EC46-70A1-4267-AAB8-2131F5DC21DC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1468909435473" checkinUser="plazi" docAuthor="Nunes, Rafael V. & Vaz-De-Mello, Fernando Z." docDate="2016" docId="03C187AE453AFF830EC8F8AEFA00F94C" docLanguage="en" docName="zootaxa.4139.1.4.pdf" docOrigin="Zootaxa 4139 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Selenocopris Burmeister" docType="key" docVersion="7" masterDocId="FFF8FFD6453BFF810E5FFF9FFF9CFFFA" masterDocTitle="New brachypterous species of Dichotomius (Selenocopris) Burmeister (Coleoptera: Scarabaeidae: Scarabaeinae) with the definition of species groups and taxonomic notes in the subgenus" updateTime="1698365276952" updateUser="plazi">
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<mods:titleInfo id="812F4ED0D7D2D78B3FDB7D176F5FA701">
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<mods:title id="2888AA7F15FF343658F4FCA6987456CD">New brachypterous species of Dichotomius (Selenocopris) Burmeister (Coleoptera: Scarabaeidae: Scarabaeinae) with the definition of species groups and taxonomic notes in the subgenus</mods:title>
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</mods:titleInfo>
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<mods:name id="9E133A82F61E93D8F88D4E6AE690A462" type="personal">
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<mods:roleTerm id="7AA9FC1EF3A6B0C8DF4AF9E403623D3F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="87AF19CA8CD0B40D648A58D0274123F2">Nunes, Rafael V.</mods:namePart>
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</mods:name>
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<mods:name id="F96E1CB1086BF541A64B2C9EFB6E347A" type="personal">
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<mods:role id="592715EDCE26F5BCB7B39894AD522482">
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<mods:roleTerm id="519F93889BB06F249E8AEC5A18C88205">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="623903D84074D9E69886E37977C9A168">Vaz-De-Mello, Fernando Z.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="9DB8A12FDE2641C5D8B0B8F3A4E41C65">text</mods:typeOfResource>
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<mods:relatedItem id="7FE3C96C8CDCC08901CF3BB5B1E777E1" type="host">
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<mods:titleInfo id="5293A97590D434FAA3DD1903976C6919">
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<mods:title id="6FD070C2471C92C0EBF36220ED211CBD">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="EC0ED5718DB175D8873DEAD8EE12D4E5">
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<mods:date id="D8AE8D90B2DF7AAF72D07EA787412CFD">2016</mods:date>
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<mods:detail id="5C4F8691E6EA3A15137B12A6D24E5504" type="volume">
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<mods:number id="539B101AC244AE9679F60A7F29069FD2">4139</mods:number>
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</mods:detail>
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<mods:detail id="0A96E3A7B1E9370128FB5F01F215D235" type="issue">
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<mods:number id="E39EEEA2029760020C30F5256231BB07">1</mods:number>
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</mods:detail>
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<mods:classification id="6EA208FDD3041D368FF2BC98B080BEB7">journal volume</mods:classification>
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<mods:identifier id="883C4E5D6C402024F3643D17A8082AC3" type="DOI">10.11646/zootaxa.4139.1.4</mods:identifier>
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<mods:identifier id="95B3A46E5EEA89DCFF511237136C5824" type="GBIF-Dataset">3cf8edb7-4be5-4ec3-803e-ca1f224d703b</mods:identifier>
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<mods:identifier id="D81D14EE4AE74A5145C7755E3B4EDDBC" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="1E14068066BB1E98EF7D5E0D930A562E" type="Zenodo-Dep">271999</mods:identifier>
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</mods:mods>
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<treatment id="03C187AE453AFF830EC8F8AEFA00F94C" ID-DOI="http://doi.org/10.5281/zenodo.6090767" ID-GBIF-Taxon="123198325" ID-Zenodo-Dep="6090767" LSID="urn:lsid:plazi:treatment:03C187AE453AFF830EC8F8AEFA00F94C" httpUri="http://treatment.plazi.org/id/03C187AE453AFF830EC8F8AEFA00F94C" lastPageId="2" pageId="1">
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<subSubSection id="C3726533453AFF830EC8F8AEFDB1FF2E" lastPageId="2" pageId="1" type="nomenclature">
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<paragraph id="8BD736B8453AFF800EC8F8AEFBC5F8B1" blockId="1.[151,1113,1841,1867]" box="[151,1113,1841,1867]" pageId="1">
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<heading id="D09F81D4453AFF800EC8F8AEFBC5F8B1" bold="true" box="[151,1113,1841,1867]" fontSize="11" level="1" pageId="1" reason="1">
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<emphasis id="B91CEAAA453AFF800EC8F8AEFBC5F8B1" bold="true" box="[151,1113,1841,1867]" pageId="1">
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Key for species groups and brachypterous species of
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<taxonomicName id="4C684D3B453AFF800D76F8AEFBC5F8B1" ID-CoL="7PZD7" authority="Burmeister" authorityName="Burmeister" box="[809,1113,1841,1867]" class="Insecta" family="Scarabaeidae" genus="Selenocopris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="genus">
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<emphasis id="B91CEAAA453AFF800D76F8AEFC5FF8B1" bold="true" box="[809,963,1841,1867]" italics="true" pageId="1">Selenocopris</emphasis>
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Burmeister
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD736B8453AFF830EC8F8E8FDB1FF2E" blockId="1.[151,1436,1911,2011]" lastBlockId="2.[151,1436,151,213]" lastPageId="2" pageId="1">
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The following key allows the identification of the
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<taxonomicName id="4C684D3B453AFF800C99F8E7FCCBF86A" box="[710,855,1912,1936]" class="Insecta" family="Scarabaeidae" genus="Selenocopris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="genus">
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<emphasis id="B91CEAAA453AFF800C99F8E7FCCBF86A" box="[710,855,1912,1936]" italics="true" pageId="1">Selenocopris</emphasis>
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</taxonomicName>
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species groups and the brachypterous species of the subgenus, now assigned to the
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<taxonomicName id="4C684D3B453AFF800FA4F803FD11F84E" box="[507,653,1948,1972]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="quadraticeps">
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<emphasis id="B91CEAAA453AFF800FA4F803FD11F84E" box="[507,653,1948,1972]" italics="true" pageId="1">quadraticeps</emphasis>
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</taxonomicName>
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species group. When not specified, the characters fit both for males and females. Females differ from males by always having the 6th abdominal sternite not shortened medially (usually three times wider than the 5th sternite) and/or having modifications on the median portion of it or at the apex of pygidium (
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<figureCitation id="13532A3D4539FF830F33FF23FE29FF2E" box="[364,437,188,213]" captionStart="FIGURE 1" captionStartId="3.[151,250,1738,1760]" captionTargetBox="[255,1318,202,1704]" captionTargetId="figure@3.[249,1332,187,1718]" captionTargetPageId="3" captionText="FIGURE 1. (A) Ventral habitus of a female of Dichotomius (Selenocopris) periotoi n. sp. The white arrow at the head indicates the position of the ventral clypeal process and the white at the abdomen indicates the modification at the medial portion of the 6 th sternite; (B) lateral view of the same specimen. The white arrow indicating the position of the mesepisternum.; (C) detail of the mesepisternum of the same specimen. The white arrow indicates the posterior portion of the upper margin of the mesepisternum ending in an acute angle, which is typical of Selenocopris; (D) elytral striae and interstriae of D. (S.) fissus (Gillet); (E) elytral striae and interstriae of D. (S.) fissiceps (Felsche) (quadraticeps group); (F) head and clypeal teeth of D. (S.) bicuspis (Germar); (G) head and clypeal teeth of D. (S.) fissiceps (quadraticeps group); (H) 6 th abdominal sternite of the female of D. (S.) spadiceus (Luederwaldt); (I) 6 th abdominal sternite of the female of D. (S.) punctulatipennis (Luederwaldt) (bicuspis group); (J) 6 th abdominal sternite of the male of D. (S.) ascanius (Harold); (K) 6 th abdominal sternite of the female of D. (S.) ascanius (Luederwaldt); (L) 6 th abdominal sternite of the female of D. (S.) quadraticeps (Felsche)." httpUri="https://zenodo.org/record/272000/files/figure.png" pageId="2">Figs. 1</figureCitation>
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A, 1H–L).
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</paragraph>
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</subSubSection>
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<subSubSection id="C37265334539FF830EC8FE9CFA00F94C" pageId="2" type="key">
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<keyStep id="309C2E1D4539FF830EC8FE9CFA07FDDB" pageId="2">
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<paragraph id="8BD736B84539FF830EC8FE9CFA00FE74" blockId="2.[151,1436,259,1718]" pageId="2">
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<keyLead id="3099958D4539FF830EC8FE9CFA00FE74" pageId="2">
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1 Clypeo-genal junction border rounded, lacking angulation (
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<figureCitation id="13532A3D4539FF830D40FE9CFCCAFEE3" box="[799,854,259,281]" captionStart="FIGURE 1" captionStartId="3.[151,250,1738,1760]" captionTargetBox="[255,1318,202,1704]" captionTargetId="figure@3.[249,1332,187,1718]" captionTargetPageId="3" captionText="FIGURE 1. (A) Ventral habitus of a female of Dichotomius (Selenocopris) periotoi n. sp. The white arrow at the head indicates the position of the ventral clypeal process and the white at the abdomen indicates the modification at the medial portion of the 6 th sternite; (B) lateral view of the same specimen. The white arrow indicating the position of the mesepisternum.; (C) detail of the mesepisternum of the same specimen. The white arrow indicates the posterior portion of the upper margin of the mesepisternum ending in an acute angle, which is typical of Selenocopris; (D) elytral striae and interstriae of D. (S.) fissus (Gillet); (E) elytral striae and interstriae of D. (S.) fissiceps (Felsche) (quadraticeps group); (F) head and clypeal teeth of D. (S.) bicuspis (Germar); (G) head and clypeal teeth of D. (S.) fissiceps (quadraticeps group); (H) 6 th abdominal sternite of the female of D. (S.) spadiceus (Luederwaldt); (I) 6 th abdominal sternite of the female of D. (S.) punctulatipennis (Luederwaldt) (bicuspis group); (J) 6 th abdominal sternite of the male of D. (S.) ascanius (Harold); (K) 6 th abdominal sternite of the female of D. (S.) ascanius (Luederwaldt); (L) 6 th abdominal sternite of the female of D. (S.) quadraticeps (Felsche)." httpUri="https://zenodo.org/record/272000/files/figure.png" pageId="2">Fig. 1</figureCitation>
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F) and/or clypeal teeth separated from adjacent clypeal bor- der by external emarginations (
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<figureCitation id="13532A3D4539FF830C45FEBFFDCFFECC" box="[538,595,288,310]" captionStart="FIGURE 1" captionStartId="3.[151,250,1738,1760]" captionTargetBox="[255,1318,202,1704]" captionTargetId="figure@3.[249,1332,187,1718]" captionTargetPageId="3" captionText="FIGURE 1. (A) Ventral habitus of a female of Dichotomius (Selenocopris) periotoi n. sp. The white arrow at the head indicates the position of the ventral clypeal process and the white at the abdomen indicates the modification at the medial portion of the 6 th sternite; (B) lateral view of the same specimen. The white arrow indicating the position of the mesepisternum.; (C) detail of the mesepisternum of the same specimen. The white arrow indicates the posterior portion of the upper margin of the mesepisternum ending in an acute angle, which is typical of Selenocopris; (D) elytral striae and interstriae of D. (S.) fissus (Gillet); (E) elytral striae and interstriae of D. (S.) fissiceps (Felsche) (quadraticeps group); (F) head and clypeal teeth of D. (S.) bicuspis (Germar); (G) head and clypeal teeth of D. (S.) fissiceps (quadraticeps group); (H) 6 th abdominal sternite of the female of D. (S.) spadiceus (Luederwaldt); (I) 6 th abdominal sternite of the female of D. (S.) punctulatipennis (Luederwaldt) (bicuspis group); (J) 6 th abdominal sternite of the male of D. (S.) ascanius (Harold); (K) 6 th abdominal sternite of the female of D. (S.) ascanius (Luederwaldt); (L) 6 th abdominal sternite of the female of D. (S.) quadraticeps (Felsche)." httpUri="https://zenodo.org/record/272000/files/figure.png" pageId="2">Fig. 1</figureCitation>
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F). Female pygidium swollen, apex excavated in some species (
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<figureCitation id="13532A3D4539FF830A84FEBFFABDFECC" box="[1243,1313,288,310]" captionStart="FIGURE 2" captionStartId="4.[151,250,1123,1145]" captionTargetBox="[400,1183,199,1090]" captionTargetId="figure@4.[397,1189,193,1102]" captionTargetPageId="4" captionText="FIGURE 2. (A) Pygidium of D. (S.) punctulatipennis (bicuspis group). The white arrow indicates the excavation at the pygidium apex; (B) swollen pygidium of D. (S.) bicuspis; (C) pygidium of D. (S.) periotoi; (D) pygidium of D. (S.) ingens. The white arrow indicates the ocellate punctures near the basis of the pygidium; (E) macropterous hind wing of D. (S.) fissceps and brachypterous hind wing of D. (S.) ingens. The black arrows in the macropterous wing (larger one) indicates the anal and cubital vein. This wing is damaged at two points, one along the coastal vein and other on the posterior portion. Scale bar = 5 mm." httpUri="https://zenodo.org/record/272001/files/figure.png" pageId="2">Figs. 2</figureCitation>
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B–C) and/or female 6th abdominal sternite bearing a rounded central lobe advancing above pygidium apex. (
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<figureCitation id="13532A3D4539FF830AC1FEA2FB44FEA9" box="[1182,1240,317,339]" captionStart="FIGURE 1" captionStartId="3.[151,250,1738,1760]" captionTargetBox="[255,1318,202,1704]" captionTargetId="figure@3.[249,1332,187,1718]" captionTargetPageId="3" captionText="FIGURE 1. (A) Ventral habitus of a female of Dichotomius (Selenocopris) periotoi n. sp. The white arrow at the head indicates the position of the ventral clypeal process and the white at the abdomen indicates the modification at the medial portion of the 6 th sternite; (B) lateral view of the same specimen. The white arrow indicating the position of the mesepisternum.; (C) detail of the mesepisternum of the same specimen. The white arrow indicates the posterior portion of the upper margin of the mesepisternum ending in an acute angle, which is typical of Selenocopris; (D) elytral striae and interstriae of D. (S.) fissus (Gillet); (E) elytral striae and interstriae of D. (S.) fissiceps (Felsche) (quadraticeps group); (F) head and clypeal teeth of D. (S.) bicuspis (Germar); (G) head and clypeal teeth of D. (S.) fissiceps (quadraticeps group); (H) 6 th abdominal sternite of the female of D. (S.) spadiceus (Luederwaldt); (I) 6 th abdominal sternite of the female of D. (S.) punctulatipennis (Luederwaldt) (bicuspis group); (J) 6 th abdominal sternite of the male of D. (S.) ascanius (Harold); (K) 6 th abdominal sternite of the female of D. (S.) ascanius (Luederwaldt); (L) 6 th abdominal sternite of the female of D. (S.) quadraticeps (Felsche)." httpUri="https://zenodo.org/record/272000/files/figure.png" pageId="2">
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Fig.
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<date id="FFD610784539FF830A91FEA2FB44FEA9" box="[1230,1240,317,339]" pageId="2">1</date>
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</figureCitation>
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I). South portion of Atlantic Forest (
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<collectingCountry id="F37F76284539FF830F23FEC4FE26FE8B" box="[380,442,347,369]" name="Brazil" pageId="2">Brazil</collectingCountry>
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,
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<collectingCountry id="F37F76284539FF830F9BFEC4FD82FE8A" box="[452,542,347,368]" name="Paraguay" pageId="2">
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<collectingRegion id="49ACF85A4539FF830F9BFEC4FD82FE8A" box="[452,542,347,368]" country="Paraguay" name="Asuncion" pageId="2">Paraguay</collectingRegion>
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</collectingCountry>
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,
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<collectingCountry id="F37F76284539FF830C77FEC4FD15FE8B" box="[552,649,347,369]" name="Argentina" pageId="2">Argentina</collectingCountry>
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) and gallery forests at
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<collectingRegion id="49ACF85A4539FF830D3BFEC4FC36FE8B" box="[868,938,347,369]" country="Paraguay" name="Central" pageId="2">Central</collectingRegion>
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<collectingCountry id="F37F76284539FF830DEFFEC4FC77FE8B" box="[944,1003,347,369]" name="Brazil" pageId="2">Brazil</collectingCountry>
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and South America
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<collectingRegion id="49ACF85A4539FF830AEFFEC4FB71FE8B" box="[1200,1261,347,369]" country="Argentina" name="Chaco" pageId="2">Chaco</collectingRegion>
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.................................................................................................
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<taxonomicName id="4C684D3B4539FF830A9CFEE6FA8EFE74" ID-CoL="6D436" box="[1219,1298,377,398]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="bicuspis">
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<emphasis id="B91CEAAA4539FF830A9CFEE6FA8EFE74" box="[1219,1298,377,398]" italics="true" pageId="2">bicuspis</emphasis>
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</taxonomicName>
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species group
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</keyLead>
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</paragraph>
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<paragraph id="8BD736B84539FF830EC8FE09FA07FDDB" blockId="2.[151,1436,259,1718]" pageId="2">
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<keyLead id="3099958D4539FF830EC8FE09FA07FDDB" pageId="2">
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- Clypeo-genal junction border not rounded, forming at least an obtuse angle, which may be very obtuse on smaller males and females (
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<figureCitation id="13532A3D4539FF830F64FE2CFEEFFE33" box="[315,371,435,457]" captionStart="FIGURE 1" captionStartId="3.[151,250,1738,1760]" captionTargetBox="[255,1318,202,1704]" captionTargetId="figure@3.[249,1332,187,1718]" captionTargetPageId="3" captionText="FIGURE 1. (A) Ventral habitus of a female of Dichotomius (Selenocopris) periotoi n. sp. The white arrow at the head indicates the position of the ventral clypeal process and the white at the abdomen indicates the modification at the medial portion of the 6 th sternite; (B) lateral view of the same specimen. The white arrow indicating the position of the mesepisternum.; (C) detail of the mesepisternum of the same specimen. The white arrow indicates the posterior portion of the upper margin of the mesepisternum ending in an acute angle, which is typical of Selenocopris; (D) elytral striae and interstriae of D. (S.) fissus (Gillet); (E) elytral striae and interstriae of D. (S.) fissiceps (Felsche) (quadraticeps group); (F) head and clypeal teeth of D. (S.) bicuspis (Germar); (G) head and clypeal teeth of D. (S.) fissiceps (quadraticeps group); (H) 6 th abdominal sternite of the female of D. (S.) spadiceus (Luederwaldt); (I) 6 th abdominal sternite of the female of D. (S.) punctulatipennis (Luederwaldt) (bicuspis group); (J) 6 th abdominal sternite of the male of D. (S.) ascanius (Harold); (K) 6 th abdominal sternite of the female of D. (S.) ascanius (Luederwaldt); (L) 6 th abdominal sternite of the female of D. (S.) quadraticeps (Felsche)." httpUri="https://zenodo.org/record/272000/files/figure.png" pageId="2">Fig. 1</figureCitation>
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G), clypeal teeth never separated from adjacent clypeal border by external emarginations (
|
|||
|
|
<figureCitation id="13532A3D4539FF830AADFE2CFAB6FE33" box="[1266,1322,435,457]" captionStart="FIGURE 1" captionStartId="3.[151,250,1738,1760]" captionTargetBox="[255,1318,202,1704]" captionTargetId="figure@3.[249,1332,187,1718]" captionTargetPageId="3" captionText="FIGURE 1. (A) Ventral habitus of a female of Dichotomius (Selenocopris) periotoi n. sp. The white arrow at the head indicates the position of the ventral clypeal process and the white at the abdomen indicates the modification at the medial portion of the 6 th sternite; (B) lateral view of the same specimen. The white arrow indicating the position of the mesepisternum.; (C) detail of the mesepisternum of the same specimen. The white arrow indicates the posterior portion of the upper margin of the mesepisternum ending in an acute angle, which is typical of Selenocopris; (D) elytral striae and interstriae of D. (S.) fissus (Gillet); (E) elytral striae and interstriae of D. (S.) fissiceps (Felsche) (quadraticeps group); (F) head and clypeal teeth of D. (S.) bicuspis (Germar); (G) head and clypeal teeth of D. (S.) fissiceps (quadraticeps group); (H) 6 th abdominal sternite of the female of D. (S.) spadiceus (Luederwaldt); (I) 6 th abdominal sternite of the female of D. (S.) punctulatipennis (Luederwaldt) (bicuspis group); (J) 6 th abdominal sternite of the male of D. (S.) ascanius (Harold); (K) 6 th abdominal sternite of the female of D. (S.) ascanius (Luederwaldt); (L) 6 th abdominal sternite of the female of D. (S.) quadraticeps (Felsche)." httpUri="https://zenodo.org/record/272000/files/figure.png" pageId="2">Fig. 1</figureCitation>
|
|||
|
|
G). Female pygidium normally convex, apex never excavated; female 6th abdominal sternite either
|
|||
|
|
<date id="FFD610784539FF830A78FE4FFBA3FE1C" box="[1063,1087,464,486]" pageId="2">3x</date>
|
|||
|
|
wider than other segments (
|
|||
|
|
<figureCitation id="13532A3D4539FF830B13FE4FFA1EFE1C" box="[1356,1410,464,486]" captionStart="FIGURE 1" captionStartId="3.[151,250,1738,1760]" captionTargetBox="[255,1318,202,1704]" captionTargetId="figure@3.[249,1332,187,1718]" captionTargetPageId="3" captionText="FIGURE 1. (A) Ventral habitus of a female of Dichotomius (Selenocopris) periotoi n. sp. The white arrow at the head indicates the position of the ventral clypeal process and the white at the abdomen indicates the modification at the medial portion of the 6 th sternite; (B) lateral view of the same specimen. The white arrow indicating the position of the mesepisternum.; (C) detail of the mesepisternum of the same specimen. The white arrow indicates the posterior portion of the upper margin of the mesepisternum ending in an acute angle, which is typical of Selenocopris; (D) elytral striae and interstriae of D. (S.) fissus (Gillet); (E) elytral striae and interstriae of D. (S.) fissiceps (Felsche) (quadraticeps group); (F) head and clypeal teeth of D. (S.) bicuspis (Germar); (G) head and clypeal teeth of D. (S.) fissiceps (quadraticeps group); (H) 6 th abdominal sternite of the female of D. (S.) spadiceus (Luederwaldt); (I) 6 th abdominal sternite of the female of D. (S.) punctulatipennis (Luederwaldt) (bicuspis group); (J) 6 th abdominal sternite of the male of D. (S.) ascanius (Harold); (K) 6 th abdominal sternite of the female of D. (S.) ascanius (Luederwaldt); (L) 6 th abdominal sternite of the female of D. (S.) quadraticeps (Felsche)." httpUri="https://zenodo.org/record/272000/files/figure.png" pageId="2">Fig. 1</figureCitation>
|
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|
|
L) or having other modifications at the median portion (emarginations, grooves, projecions or tubercles) (
|
|||
|
|
<figureCitation id="13532A3D4539FF830A94FE72FA97FDF9" box="[1227,1291,493,515]" captionStart="FIGURE 1" captionStartId="3.[151,250,1738,1760]" captionTargetBox="[255,1318,202,1704]" captionTargetId="figure@3.[249,1332,187,1718]" captionTargetPageId="3" captionText="FIGURE 1. (A) Ventral habitus of a female of Dichotomius (Selenocopris) periotoi n. sp. The white arrow at the head indicates the position of the ventral clypeal process and the white at the abdomen indicates the modification at the medial portion of the 6 th sternite; (B) lateral view of the same specimen. The white arrow indicating the position of the mesepisternum.; (C) detail of the mesepisternum of the same specimen. The white arrow indicates the posterior portion of the upper margin of the mesepisternum ending in an acute angle, which is typical of Selenocopris; (D) elytral striae and interstriae of D. (S.) fissus (Gillet); (E) elytral striae and interstriae of D. (S.) fissiceps (Felsche) (quadraticeps group); (F) head and clypeal teeth of D. (S.) bicuspis (Germar); (G) head and clypeal teeth of D. (S.) fissiceps (quadraticeps group); (H) 6 th abdominal sternite of the female of D. (S.) spadiceus (Luederwaldt); (I) 6 th abdominal sternite of the female of D. (S.) punctulatipennis (Luederwaldt) (bicuspis group); (J) 6 th abdominal sternite of the male of D. (S.) ascanius (Harold); (K) 6 th abdominal sternite of the female of D. (S.) ascanius (Luederwaldt); (L) 6 th abdominal sternite of the female of D. (S.) quadraticeps (Felsche)." httpUri="https://zenodo.org/record/272000/files/figure.png" pageId="2">Figs. 1</figureCitation>
|
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|
|
H, 1K). Forest habitats in South America............................................................................... 2
|
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|
</keyLead>
|
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|
|
</paragraph>
|
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</keyStep>
|
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|
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<keyStep id="309C2E1D4539FF830EC8FDB7FA00FCF1" pageId="2">
|
|||
|
|
<paragraph id="8BD736B84539FF830EC8FDB7FA0AFD6C" blockId="2.[151,1436,259,1718]" pageId="2">
|
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|
|
<keyLead id="3099958D4539FF830EC8FDB7FA0AFD6C" pageId="2">
|
|||
|
|
<specimenCount id="9D6EFD314539FF830EC8FDB7FE8EFDC4" box="[151,274,552,574]" pageId="2" type="male">2 Male</specimenCount>
|
|||
|
|
6th abdominal ventrite with strong transverse groove all along its extension (
|
|||
|
|
<figureCitation id="13532A3D4539FF830DADFDB7FBB4FDC4" box="[1010,1064,552,574]" captionStart="FIGURE 1" captionStartId="3.[151,250,1738,1760]" captionTargetBox="[255,1318,202,1704]" captionTargetId="figure@3.[249,1332,187,1718]" captionTargetPageId="3" captionText="FIGURE 1. (A) Ventral habitus of a female of Dichotomius (Selenocopris) periotoi n. sp. The white arrow at the head indicates the position of the ventral clypeal process and the white at the abdomen indicates the modification at the medial portion of the 6 th sternite; (B) lateral view of the same specimen. The white arrow indicating the position of the mesepisternum.; (C) detail of the mesepisternum of the same specimen. The white arrow indicates the posterior portion of the upper margin of the mesepisternum ending in an acute angle, which is typical of Selenocopris; (D) elytral striae and interstriae of D. (S.) fissus (Gillet); (E) elytral striae and interstriae of D. (S.) fissiceps (Felsche) (quadraticeps group); (F) head and clypeal teeth of D. (S.) bicuspis (Germar); (G) head and clypeal teeth of D. (S.) fissiceps (quadraticeps group); (H) 6 th abdominal sternite of the female of D. (S.) spadiceus (Luederwaldt); (I) 6 th abdominal sternite of the female of D. (S.) punctulatipennis (Luederwaldt) (bicuspis group); (J) 6 th abdominal sternite of the male of D. (S.) ascanius (Harold); (K) 6 th abdominal sternite of the female of D. (S.) ascanius (Luederwaldt); (L) 6 th abdominal sternite of the female of D. (S.) quadraticeps (Felsche)." httpUri="https://zenodo.org/record/272000/files/figure.png" pageId="2">Fig. 1</figureCitation>
|
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|
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J). Female 6th abdominal ventrite with a "U" shape emargination at medial portion (
|
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|
|
<figureCitation id="13532A3D4539FF830CD3FDDAFD5EFDA1" box="[652,706,581,603]" captionStart="FIGURE 1" captionStartId="3.[151,250,1738,1760]" captionTargetBox="[255,1318,202,1704]" captionTargetId="figure@3.[249,1332,187,1718]" captionTargetPageId="3" captionText="FIGURE 1. (A) Ventral habitus of a female of Dichotomius (Selenocopris) periotoi n. sp. The white arrow at the head indicates the position of the ventral clypeal process and the white at the abdomen indicates the modification at the medial portion of the 6 th sternite; (B) lateral view of the same specimen. The white arrow indicating the position of the mesepisternum.; (C) detail of the mesepisternum of the same specimen. The white arrow indicates the posterior portion of the upper margin of the mesepisternum ending in an acute angle, which is typical of Selenocopris; (D) elytral striae and interstriae of D. (S.) fissus (Gillet); (E) elytral striae and interstriae of D. (S.) fissiceps (Felsche) (quadraticeps group); (F) head and clypeal teeth of D. (S.) bicuspis (Germar); (G) head and clypeal teeth of D. (S.) fissiceps (quadraticeps group); (H) 6 th abdominal sternite of the female of D. (S.) spadiceus (Luederwaldt); (I) 6 th abdominal sternite of the female of D. (S.) punctulatipennis (Luederwaldt) (bicuspis group); (J) 6 th abdominal sternite of the male of D. (S.) ascanius (Harold); (K) 6 th abdominal sternite of the female of D. (S.) ascanius (Luederwaldt); (L) 6 th abdominal sternite of the female of D. (S.) quadraticeps (Felsche)." httpUri="https://zenodo.org/record/272000/files/figure.png" pageId="2">Fig. 1</figureCitation>
|
|||
|
|
K). Apex of male paramera curved inward (dorsal and ventral view) lacking rounded or circular excavations near basis (dorsal view). Pronotum simply convex, having at most a pair of tubercles at anterior portion.........................................................................
|
|||
|
|
<taxonomicName id="4C684D3B4539FF830AE6FD1EFA91FD6C" ID-CoL="35N9K" box="[1209,1293,641,662]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="ascanius">
|
|||
|
|
<emphasis id="B91CEAAA4539FF830AE6FD1EFA91FD6C" box="[1209,1293,641,662]" italics="true" pageId="2">ascanius</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
species group
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BD736B84539FF830EC8FD01FA00FCF1" blockId="2.[151,1436,259,1718]" pageId="2">
|
|||
|
|
<keyLead id="3099958D4539FF830EC8FD01FA00FCF1" pageId="2">
|
|||
|
|
- Male 6th abdominal segment lacking strong transverse groove all over its extension. Female 6th abdominal sternite with tubercles or other modifications at medial portion (
|
|||
|
|
<figureCitation id="13532A3D4539FF830CF7FD24FD76FD2B" box="[680,746,699,721]" captionStart="FIGURE 1" captionStartId="3.[151,250,1738,1760]" captionTargetBox="[255,1318,202,1704]" captionTargetId="figure@3.[249,1332,187,1718]" captionTargetPageId="3" captionText="FIGURE 1. (A) Ventral habitus of a female of Dichotomius (Selenocopris) periotoi n. sp. The white arrow at the head indicates the position of the ventral clypeal process and the white at the abdomen indicates the modification at the medial portion of the 6 th sternite; (B) lateral view of the same specimen. The white arrow indicating the position of the mesepisternum.; (C) detail of the mesepisternum of the same specimen. The white arrow indicates the posterior portion of the upper margin of the mesepisternum ending in an acute angle, which is typical of Selenocopris; (D) elytral striae and interstriae of D. (S.) fissus (Gillet); (E) elytral striae and interstriae of D. (S.) fissiceps (Felsche) (quadraticeps group); (F) head and clypeal teeth of D. (S.) bicuspis (Germar); (G) head and clypeal teeth of D. (S.) fissiceps (quadraticeps group); (H) 6 th abdominal sternite of the female of D. (S.) spadiceus (Luederwaldt); (I) 6 th abdominal sternite of the female of D. (S.) punctulatipennis (Luederwaldt) (bicuspis group); (J) 6 th abdominal sternite of the male of D. (S.) ascanius (Harold); (K) 6 th abdominal sternite of the female of D. (S.) ascanius (Luederwaldt); (L) 6 th abdominal sternite of the female of D. (S.) quadraticeps (Felsche)." httpUri="https://zenodo.org/record/272000/files/figure.png" pageId="2">Figs. 1</figureCitation>
|
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|
|
A,H,L). Male paramera curved inward or outward (dorsal and ventral view) and/or truncated at apex and having rounded or circular excavations near basis (dorsal view) (
|
|||
|
|
<figureCitation id="13532A3D4539FF830AE0FD47FB65FD14" box="[1215,1273,728,750]" captionStart="FIGURE 3" captionStartId="5.[151,250,1788,1810]" captionTargetBox="[286,1317,833,1766]" captionTargetId="figure@5.[264,1323,805,1767]" captionTargetPageId="5" captionText="FIGURE 3. Dichotomius (Selenocopris) ingens (Luederwaldt): (A) male dorsal habitus; (B) female dorsal habitus; (C) dorsal view of male paramera; (D) lateral view of male paramera; (E) holotype labels (NHML). Scale bar = 5 mm." httpUri="https://zenodo.org/record/272002/files/figure.png" pageId="2">Fig. 3</figureCitation>
|
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|
|
C–D). Pronotum simply convex or having excavations, tubercles, lobes or strong declivity at anterior portion......................... 3
|
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|
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</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
</keyStep>
|
|||
|
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<keyStep id="309C2E1D4539FF830EC8FC8CFA00FC64" pageId="2">
|
|||
|
|
<paragraph id="8BD736B84539FF830EC8FC8CFA00FC99" blockId="2.[151,1436,259,1718]" pageId="2">
|
|||
|
|
<keyLead id="3099958D4539FF830EC8FC8CFA00FC99" pageId="2">
|
|||
|
|
<specimenCount id="9D6EFD314539FF830EC8FC8CFE8EFCD3" box="[151,274,787,809]" pageId="2" type="male">3 Male</specimenCount>
|
|||
|
|
clypeo-genal junction border either in straight sub-straight or even acute angle, in some cases expanded (
|
|||
|
|
<figureCitation id="13532A3D4539FF830B5BFC8CFAA6FCD3" box="[1284,1338,787,809]" captionStart="FIGURE 1" captionStartId="3.[151,250,1738,1760]" captionTargetBox="[255,1318,202,1704]" captionTargetId="figure@3.[249,1332,187,1718]" captionTargetPageId="3" captionText="FIGURE 1. (A) Ventral habitus of a female of Dichotomius (Selenocopris) periotoi n. sp. The white arrow at the head indicates the position of the ventral clypeal process and the white at the abdomen indicates the modification at the medial portion of the 6 th sternite; (B) lateral view of the same specimen. The white arrow indicating the position of the mesepisternum.; (C) detail of the mesepisternum of the same specimen. The white arrow indicates the posterior portion of the upper margin of the mesepisternum ending in an acute angle, which is typical of Selenocopris; (D) elytral striae and interstriae of D. (S.) fissus (Gillet); (E) elytral striae and interstriae of D. (S.) fissiceps (Felsche) (quadraticeps group); (F) head and clypeal teeth of D. (S.) bicuspis (Germar); (G) head and clypeal teeth of D. (S.) fissiceps (quadraticeps group); (H) 6 th abdominal sternite of the female of D. (S.) spadiceus (Luederwaldt); (I) 6 th abdominal sternite of the female of D. (S.) punctulatipennis (Luederwaldt) (bicuspis group); (J) 6 th abdominal sternite of the male of D. (S.) ascanius (Harold); (K) 6 th abdominal sternite of the female of D. (S.) ascanius (Luederwaldt); (L) 6 th abdominal sternite of the female of D. (S.) quadraticeps (Felsche)." httpUri="https://zenodo.org/record/272000/files/figure.png" pageId="2">Fig. 1</figureCitation>
|
|||
|
|
G). Genal border usually curved inward and clypeal teeth always curved upward. Female 6th abdominal sternite or
|
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|
|
<date id="FFD610784539FF830A99FCAFFB42FCBC" box="[1222,1246,816,838]" pageId="2">3x</date>
|
|||
|
|
larger then 5th segment medially or having a central pair of tubercles (
|
|||
|
|
<figureCitation id="13532A3D4539FF830C8DFCD2FC8DFC99" box="[722,785,845,867]" captionStart="FIGURE 1" captionStartId="3.[151,250,1738,1760]" captionTargetBox="[255,1318,202,1704]" captionTargetId="figure@3.[249,1332,187,1718]" captionTargetPageId="3" captionText="FIGURE 1. (A) Ventral habitus of a female of Dichotomius (Selenocopris) periotoi n. sp. The white arrow at the head indicates the position of the ventral clypeal process and the white at the abdomen indicates the modification at the medial portion of the 6 th sternite; (B) lateral view of the same specimen. The white arrow indicating the position of the mesepisternum.; (C) detail of the mesepisternum of the same specimen. The white arrow indicates the posterior portion of the upper margin of the mesepisternum ending in an acute angle, which is typical of Selenocopris; (D) elytral striae and interstriae of D. (S.) fissus (Gillet); (E) elytral striae and interstriae of D. (S.) fissiceps (Felsche) (quadraticeps group); (F) head and clypeal teeth of D. (S.) bicuspis (Germar); (G) head and clypeal teeth of D. (S.) fissiceps (quadraticeps group); (H) 6 th abdominal sternite of the female of D. (S.) spadiceus (Luederwaldt); (I) 6 th abdominal sternite of the female of D. (S.) punctulatipennis (Luederwaldt) (bicuspis group); (J) 6 th abdominal sternite of the male of D. (S.) ascanius (Harold); (K) 6 th abdominal sternite of the female of D. (S.) ascanius (Luederwaldt); (L) 6 th abdominal sternite of the female of D. (S.) quadraticeps (Felsche)." httpUri="https://zenodo.org/record/272000/files/figure.png" pageId="2">Figs. 1</figureCitation>
|
|||
|
|
A, 1L)......
|
|||
|
|
<taxonomicName id="4C684D3B4539FF830DF8FCD1FBB8FC99" ID-CoL="6CRRC" box="[935,1060,846,867]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="quadraticeps">
|
|||
|
|
<emphasis id="B91CEAAA4539FF830DF8FCD1FBB8FC99" box="[935,1060,846,867]" italics="true" pageId="2">quadraticeps</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
species group (continues on couplet 5)
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BD736B84539FF830EC8FCF3FA00FC64" blockId="2.[151,1436,259,1718]" pageId="2">
|
|||
|
|
<keyLead id="3099958D4539FF830EC8FCF3FA00FC64" pageId="2">
|
|||
|
|
- Male clypeo-genal junction border always obtuse and never expanded. Genal border straight and clypeal teeth curved upward or not. Female 6th abdominal sternite with a pair of triangular projections advancing beneath pygidial apex (
|
|||
|
|
<figureCitation id="13532A3D4539FF830AAFFC17FABAFC64" box="[1264,1318,904,926]" captionStart="FIGURE 1" captionStartId="3.[151,250,1738,1760]" captionTargetBox="[255,1318,202,1704]" captionTargetId="figure@3.[249,1332,187,1718]" captionTargetPageId="3" captionText="FIGURE 1. (A) Ventral habitus of a female of Dichotomius (Selenocopris) periotoi n. sp. The white arrow at the head indicates the position of the ventral clypeal process and the white at the abdomen indicates the modification at the medial portion of the 6 th sternite; (B) lateral view of the same specimen. The white arrow indicating the position of the mesepisternum.; (C) detail of the mesepisternum of the same specimen. The white arrow indicates the posterior portion of the upper margin of the mesepisternum ending in an acute angle, which is typical of Selenocopris; (D) elytral striae and interstriae of D. (S.) fissus (Gillet); (E) elytral striae and interstriae of D. (S.) fissiceps (Felsche) (quadraticeps group); (F) head and clypeal teeth of D. (S.) bicuspis (Germar); (G) head and clypeal teeth of D. (S.) fissiceps (quadraticeps group); (H) 6 th abdominal sternite of the female of D. (S.) spadiceus (Luederwaldt); (I) 6 th abdominal sternite of the female of D. (S.) punctulatipennis (Luederwaldt) (bicuspis group); (J) 6 th abdominal sternite of the male of D. (S.) ascanius (Harold); (K) 6 th abdominal sternite of the female of D. (S.) ascanius (Luederwaldt); (L) 6 th abdominal sternite of the female of D. (S.) quadraticeps (Felsche)." httpUri="https://zenodo.org/record/272000/files/figure.png" pageId="2">Fig. 1</figureCitation>
|
|||
|
|
H)..... 4
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
</keyStep>
|
|||
|
|
<keyStep id="309C2E1D4539FF830EC8FC3AFA00FBB4" pageId="2">
|
|||
|
|
<paragraph id="8BD736B84539FF830EC8FC3AFA00FC0C" blockId="2.[151,1436,259,1718]" pageId="2">
|
|||
|
|
<keyLead id="3099958D4539FF830EC8FC3AFA00FC0C" pageId="2">
|
|||
|
|
4 Body almost always over
|
|||
|
|
<quantity id="4C909B5D4539FF830F83FC3AFD83FC41" box="[476,543,933,955]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="2" unit="mm" value="15.0">15 mm</quantity>
|
|||
|
|
in length. Elytral striae and its punctures shallowly and weakly impressed (
|
|||
|
|
<figureCitation id="13532A3D4539FF830AA4FC3AFAAEFC41" box="[1275,1330,933,955]" captionStart="FIGURE 1" captionStartId="3.[151,250,1738,1760]" captionTargetBox="[255,1318,202,1704]" captionTargetId="figure@3.[249,1332,187,1718]" captionTargetPageId="3" captionText="FIGURE 1. (A) Ventral habitus of a female of Dichotomius (Selenocopris) periotoi n. sp. The white arrow at the head indicates the position of the ventral clypeal process and the white at the abdomen indicates the modification at the medial portion of the 6 th sternite; (B) lateral view of the same specimen. The white arrow indicating the position of the mesepisternum.; (C) detail of the mesepisternum of the same specimen. The white arrow indicates the posterior portion of the upper margin of the mesepisternum ending in an acute angle, which is typical of Selenocopris; (D) elytral striae and interstriae of D. (S.) fissus (Gillet); (E) elytral striae and interstriae of D. (S.) fissiceps (Felsche) (quadraticeps group); (F) head and clypeal teeth of D. (S.) bicuspis (Germar); (G) head and clypeal teeth of D. (S.) fissiceps (quadraticeps group); (H) 6 th abdominal sternite of the female of D. (S.) spadiceus (Luederwaldt); (I) 6 th abdominal sternite of the female of D. (S.) punctulatipennis (Luederwaldt) (bicuspis group); (J) 6 th abdominal sternite of the male of D. (S.) ascanius (Harold); (K) 6 th abdominal sternite of the female of D. (S.) ascanius (Luederwaldt); (L) 6 th abdominal sternite of the female of D. (S.) quadraticeps (Felsche)." httpUri="https://zenodo.org/record/272000/files/figure.png" pageId="2">Fig. 1</figureCitation>
|
|||
|
|
D). Elytral interstriae flat, with either silky aspect given by its microsculpture (chagrination) and/or bearing brown or blue reflections. Male paramera bearing longitudinal excavations (dorsal and ventral view) (as those on
|
|||
|
|
<figureCitation id="13532A3D4539FF830A7BFC7FFBC6FC0C" box="[1060,1114,992,1014]" captionStart="FIGURE 3" captionStartId="5.[151,250,1788,1810]" captionTargetBox="[286,1317,833,1766]" captionTargetId="figure@5.[264,1323,805,1767]" captionTargetPageId="5" captionText="FIGURE 3. Dichotomius (Selenocopris) ingens (Luederwaldt): (A) male dorsal habitus; (B) female dorsal habitus; (C) dorsal view of male paramera; (D) lateral view of male paramera; (E) holotype labels (NHML). Scale bar = 5 mm." httpUri="https://zenodo.org/record/272002/files/figure.png" pageId="2">Fig. 3</figureCitation>
|
|||
|
|
C–D)......
|
|||
|
|
<taxonomicName id="4C684D3B4539FF830A82FC7EFA8EFC0C" ID-CoL="35N9Z" box="[1245,1298,993,1014]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="fissus">
|
|||
|
|
<emphasis id="B91CEAAA4539FF830A82FC7EFA8EFC0C" box="[1245,1298,993,1014]" italics="true" pageId="2">fissus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
species group
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BD736B84539FF830EC8FC61FA00FBB4" blockId="2.[151,1436,259,1718]" pageId="2">
|
|||
|
|
<keyLead id="3099958D4539FF830EC8FC61FA00FBB4" pageId="2">
|
|||
|
|
- Body almost always under
|
|||
|
|
<quantity id="4C909B5D4539FF830FAFFC62FDA9FBE9" box="[496,565,1021,1043]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="2" unit="mm" value="15.0">15 mm</quantity>
|
|||
|
|
in length. Elytral striae and its punctures deeply impressed (
|
|||
|
|
<figureCitation id="13532A3D4539FF830ACDFC62FB56FBE9" box="[1170,1226,1021,1043]" captionStart="FIGURE 1" captionStartId="3.[151,250,1738,1760]" captionTargetBox="[255,1318,202,1704]" captionTargetId="figure@3.[249,1332,187,1718]" captionTargetPageId="3" captionText="FIGURE 1. (A) Ventral habitus of a female of Dichotomius (Selenocopris) periotoi n. sp. The white arrow at the head indicates the position of the ventral clypeal process and the white at the abdomen indicates the modification at the medial portion of the 6 th sternite; (B) lateral view of the same specimen. The white arrow indicating the position of the mesepisternum.; (C) detail of the mesepisternum of the same specimen. The white arrow indicates the posterior portion of the upper margin of the mesepisternum ending in an acute angle, which is typical of Selenocopris; (D) elytral striae and interstriae of D. (S.) fissus (Gillet); (E) elytral striae and interstriae of D. (S.) fissiceps (Felsche) (quadraticeps group); (F) head and clypeal teeth of D. (S.) bicuspis (Germar); (G) head and clypeal teeth of D. (S.) fissiceps (quadraticeps group); (H) 6 th abdominal sternite of the female of D. (S.) spadiceus (Luederwaldt); (I) 6 th abdominal sternite of the female of D. (S.) punctulatipennis (Luederwaldt) (bicuspis group); (J) 6 th abdominal sternite of the male of D. (S.) ascanius (Harold); (K) 6 th abdominal sternite of the female of D. (S.) ascanius (Luederwaldt); (L) 6 th abdominal sternite of the female of D. (S.) quadraticeps (Felsche)." httpUri="https://zenodo.org/record/272000/files/figure.png" pageId="2">Fig. 1</figureCitation>
|
|||
|
|
E). Elytral interstriae shinny, not bearing microsculpture (chagrination), sometimes with color reflections. Male paramera lacking longitudinal excavations..........................................................................
|
|||
|
|
<taxonomicName id="4C684D3B4539FF830AEBFBA6FA8EFBB4" ID-CoL="6D43D" box="[1204,1298,1081,1102]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="spadiceus">
|
|||
|
|
<emphasis id="B91CEAAA4539FF830AEBFBA6FA8EFBB4" box="[1204,1298,1081,1102]" italics="true" pageId="2">spadiceus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
species group
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
</keyStep>
|
|||
|
|
<keyStep id="309C2E1D4539FF830EC8FBCAFA07FB39" pageId="2">
|
|||
|
|
<paragraph id="8BD736B84539FF830EC8FBCAFA07FB73" blockId="2.[151,1436,259,1718]" pageId="2">
|
|||
|
|
<keyLead id="3099958D4539FF830EC8FBCAFA07FB73" pageId="2">
|
|||
|
|
5 Brachypterous species (atrophied hind wings) (
|
|||
|
|
<figureCitation id="13532A3D4539FF830CEEFBCAFD71FB91" box="[689,749,1109,1131]" captionStart="FIGURE 2" captionStartId="4.[151,250,1123,1145]" captionTargetBox="[400,1183,199,1090]" captionTargetId="figure@4.[397,1189,193,1102]" captionTargetPageId="4" captionText="FIGURE 2. (A) Pygidium of D. (S.) punctulatipennis (bicuspis group). The white arrow indicates the excavation at the pygidium apex; (B) swollen pygidium of D. (S.) bicuspis; (C) pygidium of D. (S.) periotoi; (D) pygidium of D. (S.) ingens. The white arrow indicates the ocellate punctures near the basis of the pygidium; (E) macropterous hind wing of D. (S.) fissceps and brachypterous hind wing of D. (S.) ingens. The black arrows in the macropterous wing (larger one) indicates the anal and cubital vein. This wing is damaged at two points, one along the coastal vein and other on the posterior portion. Scale bar = 5 mm." httpUri="https://zenodo.org/record/272001/files/figure.png" pageId="2">Fig. 2</figureCitation>
|
|||
|
|
E): lacking humeral callus; pronotum wider than elytra (dorsal view); elytra strongly convex (lateral view) (
|
|||
|
|
<figureCitation id="13532A3D4539FF830C15FBECFD1CFB73" box="[586,640,1139,1161]" captionStart="FIGURE 1" captionStartId="3.[151,250,1738,1760]" captionTargetBox="[255,1318,202,1704]" captionTargetId="figure@3.[249,1332,187,1718]" captionTargetPageId="3" captionText="FIGURE 1. (A) Ventral habitus of a female of Dichotomius (Selenocopris) periotoi n. sp. The white arrow at the head indicates the position of the ventral clypeal process and the white at the abdomen indicates the modification at the medial portion of the 6 th sternite; (B) lateral view of the same specimen. The white arrow indicating the position of the mesepisternum.; (C) detail of the mesepisternum of the same specimen. The white arrow indicates the posterior portion of the upper margin of the mesepisternum ending in an acute angle, which is typical of Selenocopris; (D) elytral striae and interstriae of D. (S.) fissus (Gillet); (E) elytral striae and interstriae of D. (S.) fissiceps (Felsche) (quadraticeps group); (F) head and clypeal teeth of D. (S.) bicuspis (Germar); (G) head and clypeal teeth of D. (S.) fissiceps (quadraticeps group); (H) 6 th abdominal sternite of the female of D. (S.) spadiceus (Luederwaldt); (I) 6 th abdominal sternite of the female of D. (S.) punctulatipennis (Luederwaldt) (bicuspis group); (J) 6 th abdominal sternite of the male of D. (S.) ascanius (Harold); (K) 6 th abdominal sternite of the female of D. (S.) ascanius (Luederwaldt); (L) 6 th abdominal sternite of the female of D. (S.) quadraticeps (Felsche)." httpUri="https://zenodo.org/record/272000/files/figure.png" pageId="2">Fig. 1</figureCitation>
|
|||
|
|
B).............................................................. 6
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BD736B84539FF830EC8FB0EFA07FB39" blockId="2.[151,1436,259,1718]" pageId="2">
|
|||
|
|
<keyLead id="3099958D4539FF830EC8FB0EFA07FB39" pageId="2">
|
|||
|
|
- Macropterous species (fully developed hind wings) (
|
|||
|
|
<figureCitation id="13532A3D4539FF830C82FB0FFC84FB5C" box="[733,792,1168,1190]" captionStart="FIGURE 2" captionStartId="4.[151,250,1123,1145]" captionTargetBox="[400,1183,199,1090]" captionTargetId="figure@4.[397,1189,193,1102]" captionTargetPageId="4" captionText="FIGURE 2. (A) Pygidium of D. (S.) punctulatipennis (bicuspis group). The white arrow indicates the excavation at the pygidium apex; (B) swollen pygidium of D. (S.) bicuspis; (C) pygidium of D. (S.) periotoi; (D) pygidium of D. (S.) ingens. The white arrow indicates the ocellate punctures near the basis of the pygidium; (E) macropterous hind wing of D. (S.) fissceps and brachypterous hind wing of D. (S.) ingens. The black arrows in the macropterous wing (larger one) indicates the anal and cubital vein. This wing is damaged at two points, one along the coastal vein and other on the posterior portion. Scale bar = 5 mm." httpUri="https://zenodo.org/record/272001/files/figure.png" pageId="2">Fig. 2</figureCitation>
|
|||
|
|
E): having humeral callus; pronotum with the same width as elytra (dorsal view); elytra normally convex (lateral view).............................. macropterous species not dealt here
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
</keyStep>
|
|||
|
|
<keyStep id="309C2E1D4539FF830EC8FB54FA07FAE1" pageId="2">
|
|||
|
|
<paragraph id="8BD736B84539FF830EC8FB54FA00FB04" blockId="2.[151,1436,259,1718]" pageId="2">
|
|||
|
|
<keyLead id="3099958D4539FF830EC8FB54FA00FB04" pageId="2">
|
|||
|
|
6 Pygidium bearing ocellate punctures near anterior margin (basis) (
|
|||
|
|
<figureCitation id="13532A3D4539FF830D0AFB54FC12FB1B" box="[853,910,1227,1249]" captionStart="FIGURE 2" captionStartId="4.[151,250,1123,1145]" captionTargetBox="[400,1183,199,1090]" captionTargetId="figure@4.[397,1189,193,1102]" captionTargetPageId="4" captionText="FIGURE 2. (A) Pygidium of D. (S.) punctulatipennis (bicuspis group). The white arrow indicates the excavation at the pygidium apex; (B) swollen pygidium of D. (S.) bicuspis; (C) pygidium of D. (S.) periotoi; (D) pygidium of D. (S.) ingens. The white arrow indicates the ocellate punctures near the basis of the pygidium; (E) macropterous hind wing of D. (S.) fissceps and brachypterous hind wing of D. (S.) ingens. The black arrows in the macropterous wing (larger one) indicates the anal and cubital vein. This wing is damaged at two points, one along the coastal vein and other on the posterior portion. Scale bar = 5 mm." httpUri="https://zenodo.org/record/272001/files/figure.png" pageId="2">Fig. 2</figureCitation>
|
|||
|
|
D). Chapada dos Guimarães,
|
|||
|
|
<collectingRegion id="49ACF85A4539FF830AFBFB53FA83FB1B" box="[1188,1311,1227,1249]" country="Brazil" name="Mato Grosso" pageId="2">Mato Grosso</collectingRegion>
|
|||
|
|
state,
|
|||
|
|
<collectingCountry id="F37F76284539FF830B03FB54FA0BFB1B" box="[1372,1431,1227,1249]" name="Brazil" pageId="2">Brazil</collectingCountry>
|
|||
|
|
...........................................................
|
|||
|
|
<taxonomicName id="4C684D3B4539FF830DC3FB76FAD2FB04" ID-CoL="6D43K" authority="Luederwaldt 1935" authorityName="Luederwaldt" authorityYear="1935" box="[924,1358,1256,1278]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="ingens" subGenus="Selenocopris">
|
|||
|
|
<emphasis id="B91CEAAA4539FF830DC3FB76FB14FB04" box="[924,1160,1257,1278]" italics="true" pageId="2">Dichotomius (S.) ingens</emphasis>
|
|||
|
|
(Luederwaldt 1935)
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="13532A3D4539FF830B04FB77FA08FB04" box="[1371,1428,1256,1278]" captionStart="FIGURE 3" captionStartId="5.[151,250,1788,1810]" captionTargetBox="[286,1317,833,1766]" captionTargetId="figure@5.[264,1323,805,1767]" captionTargetPageId="5" captionText="FIGURE 3. Dichotomius (Selenocopris) ingens (Luederwaldt): (A) male dorsal habitus; (B) female dorsal habitus; (C) dorsal view of male paramera; (D) lateral view of male paramera; (E) holotype labels (NHML). Scale bar = 5 mm." httpUri="https://zenodo.org/record/272002/files/figure.png" pageId="2">Fig. 3</figureCitation>
|
|||
|
|
)
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BD736B84539FF830EC8FA99FA07FAE1" blockId="2.[151,1436,259,1718]" box="[151,1435,1285,1307]" pageId="2">
|
|||
|
|
<keyLead id="3099958D4539FF830EC8FA99FA07FAE1" box="[151,1435,1285,1307]" pageId="2">
|
|||
|
|
- Pygidium lacking ocellate punctures (
|
|||
|
|
<figureCitation id="13532A3D4539FF830C14FA9AFD19FAE1" box="[587,645,1285,1307]" captionStart="FIGURE 2" captionStartId="4.[151,250,1123,1145]" captionTargetBox="[400,1183,199,1090]" captionTargetId="figure@4.[397,1189,193,1102]" captionTargetPageId="4" captionText="FIGURE 2. (A) Pygidium of D. (S.) punctulatipennis (bicuspis group). The white arrow indicates the excavation at the pygidium apex; (B) swollen pygidium of D. (S.) bicuspis; (C) pygidium of D. (S.) periotoi; (D) pygidium of D. (S.) ingens. The white arrow indicates the ocellate punctures near the basis of the pygidium; (E) macropterous hind wing of D. (S.) fissceps and brachypterous hind wing of D. (S.) ingens. The black arrows in the macropterous wing (larger one) indicates the anal and cubital vein. This wing is damaged at two points, one along the coastal vein and other on the posterior portion. Scale bar = 5 mm." httpUri="https://zenodo.org/record/272001/files/figure.png" pageId="2">Fig. 2</figureCitation>
|
|||
|
|
C), at most having very fine punctures (
|
|||
|
|
<date id="FFD610784539FF830A5EFA9AFBB5FAE1" box="[1025,1065,1285,1307]" pageId="2" value="1920-10">20x</date>
|
|||
|
|
)............................. 7
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
</keyStep>
|
|||
|
|
<keyStep id="309C2E1D4539FF830EC8FABBFA00FA6B" pageId="2">
|
|||
|
|
<paragraph id="8BD736B84539FF830EC8FABBFA09FAAC" blockId="2.[151,1436,259,1718]" pageId="2">
|
|||
|
|
<keyLead id="3099958D4539FF830EC8FABBFA09FAAC" pageId="2">
|
|||
|
|
7 Posterior portion of pronotal disc bearing ocellate punctures (
|
|||
|
|
<figureCitation id="13532A3D4539FF830D66FABCFCE1FAC3" box="[825,893,1315,1337]" captionStart="FIGURE 6" captionStartId="10.[151,250,1362,1384]" captionTargetBox="[283,1306,458,1329]" captionTargetId="figure@10.[267,1320,445,1340]" captionTargetPageId="10" captionText="FIGURE 6. Paratypes of Dichotomius (Selenocopris) paresi n. sp.: (A) male dorsal habitus; (B) female dorsal habitus (black arrows indicate the ocellate punctures at the posterior portion of the pronotal disc); (C) dorsal view of male paramera; (D) lateral view of male paramera; (E) detail of elytra, white arrow indicating the elytral striae widened near the basis. Scale bar = 5 mm." httpUri="https://zenodo.org/record/272005/files/figure.png" pageId="2">Figs. 6</figureCitation>
|
|||
|
|
A–B). Elytral striae widened, lacking distinct punctures (
|
|||
|
|
<figureCitation id="13532A3D4539FF830EB7FADFFEB6FAAC" box="[232,298,1344,1366]" captionStart="FIGURE 6" captionStartId="10.[151,250,1362,1384]" captionTargetBox="[283,1306,458,1329]" captionTargetId="figure@10.[267,1320,445,1340]" captionTargetPageId="10" captionText="FIGURE 6. Paratypes of Dichotomius (Selenocopris) paresi n. sp.: (A) male dorsal habitus; (B) female dorsal habitus (black arrows indicate the ocellate punctures at the posterior portion of the pronotal disc); (C) dorsal view of male paramera; (D) lateral view of male paramera; (E) detail of elytra, white arrow indicating the elytral striae widened near the basis. Scale bar = 5 mm." httpUri="https://zenodo.org/record/272005/files/figure.png" pageId="2">Figs. 6</figureCitation>
|
|||
|
|
A–B, E). Chapada dos Parecis at
|
|||
|
|
<collectingRegion id="49ACF85A4539FF830C36FADEFD7AFAAC" box="[617,742,1344,1366]" country="Brazil" name="Mato Grosso" pageId="2">Mato Grosso</collectingRegion>
|
|||
|
|
state,
|
|||
|
|
<collectingCountry id="F37F76284539FF830D7BFADFFCC3FAAC" box="[804,863,1344,1366]" name="Brazil" pageId="2">Brazil</collectingCountry>
|
|||
|
|
................
|
|||
|
|
<taxonomicName id="4C684D3B4539FF830A6EFADEFA89FAAC" box="[1073,1301,1345,1366]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="paresi" status="sp. nov." subGenus="Selenocopris">
|
|||
|
|
<emphasis id="B91CEAAA4539FF830A6EFADEFA89FAAC" box="[1073,1301,1345,1366]" italics="true" pageId="2">Dichotomius (S.) paresi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A22F57D14539FF830B44FADEFAD4FAAC" box="[1307,1352,1345,1366]" pageId="2" rank="species">n.sp.</taxonomicNameLabel>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="13532A3D4539FF830B0AFADFFA13FAAC" box="[1365,1423,1344,1366]" captionStart="FIGURE 6" captionStartId="10.[151,250,1362,1384]" captionTargetBox="[283,1306,458,1329]" captionTargetId="figure@10.[267,1320,445,1340]" captionTargetPageId="10" captionText="FIGURE 6. Paratypes of Dichotomius (Selenocopris) paresi n. sp.: (A) male dorsal habitus; (B) female dorsal habitus (black arrows indicate the ocellate punctures at the posterior portion of the pronotal disc); (C) dorsal view of male paramera; (D) lateral view of male paramera; (E) detail of elytra, white arrow indicating the elytral striae widened near the basis. Scale bar = 5 mm." httpUri="https://zenodo.org/record/272005/files/figure.png" pageId="2">Fig. 6</figureCitation>
|
|||
|
|
)
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BD736B84539FF830EC8FAC1FA00FA6B" blockId="2.[151,1436,259,1718]" pageId="2">
|
|||
|
|
<keyLead id="3099958D4539FF830EC8FAC1FA00FA6B" pageId="2">
|
|||
|
|
- Pronotal disc lacking ocellate punctures, those being sometimes restricted to posterior margin, anterior angles and/or to the anterior excavation. Elytral striae narrow and always bearing distinct punctures all over its extension (
|
|||
|
|
<figureCitation id="13532A3D4539FF830AECFAE4FB75FA6B" box="[1203,1257,1403,1425]" captionStart="FIGURE 1" captionStartId="3.[151,250,1738,1760]" captionTargetBox="[255,1318,202,1704]" captionTargetId="figure@3.[249,1332,187,1718]" captionTargetPageId="3" captionText="FIGURE 1. (A) Ventral habitus of a female of Dichotomius (Selenocopris) periotoi n. sp. The white arrow at the head indicates the position of the ventral clypeal process and the white at the abdomen indicates the modification at the medial portion of the 6 th sternite; (B) lateral view of the same specimen. The white arrow indicating the position of the mesepisternum.; (C) detail of the mesepisternum of the same specimen. The white arrow indicates the posterior portion of the upper margin of the mesepisternum ending in an acute angle, which is typical of Selenocopris; (D) elytral striae and interstriae of D. (S.) fissus (Gillet); (E) elytral striae and interstriae of D. (S.) fissiceps (Felsche) (quadraticeps group); (F) head and clypeal teeth of D. (S.) bicuspis (Germar); (G) head and clypeal teeth of D. (S.) fissiceps (quadraticeps group); (H) 6 th abdominal sternite of the female of D. (S.) spadiceus (Luederwaldt); (I) 6 th abdominal sternite of the female of D. (S.) punctulatipennis (Luederwaldt) (bicuspis group); (J) 6 th abdominal sternite of the male of D. (S.) ascanius (Harold); (K) 6 th abdominal sternite of the female of D. (S.) ascanius (Luederwaldt); (L) 6 th abdominal sternite of the female of D. (S.) quadraticeps (Felsche)." httpUri="https://zenodo.org/record/272000/files/figure.png" pageId="2">Fig. 1</figureCitation>
|
|||
|
|
E).......... 8
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
</keyStep>
|
|||
|
|
<keyStep id="309C2E1D4539FF830EC8FA07FA00FA13" pageId="2">
|
|||
|
|
<paragraph id="8BD736B84539FF830EC8FA07FA00FA31" blockId="2.[151,1436,259,1718]" pageId="2">
|
|||
|
|
<keyLead id="3099958D4539FF830EC8FA07FA00FA31" pageId="2">
|
|||
|
|
8 Cephalic carina surface wrinkled (striated) (
|
|||
|
|
<figureCitation id="13532A3D4539FF830CD2FA07FD4CFA54" box="[653,720,1432,1454]" captionStart="FIGURE 4" captionStartId="7.[151,250,1405,1427]" captionTargetBox="[273,1321,386,1375]" captionTargetId="figure@7.[264,1323,373,1384]" captionTargetPageId="7" captionText="FIGURE 4. Paratypes of Dichotomius (Selenocopris) darwini n. sp.: (A) male dorsal habitus; (B) female dorsal habitus; (C) dorsal view of male paramera; (D) lateral view of male paramera; (E) detail of male cephalic carina with striation; (F) detail of the anterior pronotal excavation with ocellate punctures. Scale bar = 5 mm." httpUri="https://zenodo.org/record/272003/files/figure.png" pageId="2">Figs. 4</figureCitation>
|
|||
|
|
A–B, E). Anterior excavation of pronotum bearing coarse punctures (
|
|||
|
|
<figureCitation id="13532A3D4539FF830B2AFA07FF72FA31" captionStart="FIGURE 4" captionStartId="7.[151,250,1405,1427]" captionTargetBox="[273,1321,386,1375]" captionTargetId="figure@7.[264,1323,373,1384]" captionTargetPageId="7" captionText="FIGURE 4. Paratypes of Dichotomius (Selenocopris) darwini n. sp.: (A) male dorsal habitus; (B) female dorsal habitus; (C) dorsal view of male paramera; (D) lateral view of male paramera; (E) detail of male cephalic carina with striation; (F) detail of the anterior pronotal excavation with ocellate punctures. Scale bar = 5 mm." httpUri="https://zenodo.org/record/272003/files/figure.png" pageId="2">Fig. 4</figureCitation>
|
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|
|
F). Chapadão do Céu,
|
|||
|
|
<collectingRegion id="49ACF85A4539FF830F9BFA2AFE60FA31" box="[452,508,1461,1483]" country="Brazil" name="Goias" pageId="2">Goiás</collectingRegion>
|
|||
|
|
state,
|
|||
|
|
<collectingCountry id="F37F76284539FF830C65FA2AFDE9FA31" box="[570,629,1461,1483]" name="Brazil" pageId="2">Brazil</collectingCountry>
|
|||
|
|
..............................................
|
|||
|
|
<taxonomicName id="4C684D3B4539FF830AF6FA29FA00FA31" ID-CoL="35N9S" box="[1193,1436,1462,1483]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="darwini" subGenus="Selenocopris">
|
|||
|
|
<emphasis id="B91CEAAA4539FF830AF6FA29FA00FA31" box="[1193,1436,1462,1483]" italics="true" pageId="2">Dichotomius (S.) darwini</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BD736B84539FF830EC8FA4BFA00FA13" blockId="2.[151,1436,259,1718]" box="[151,1436,1491,1513]" pageId="2">
|
|||
|
|
<keyLead id="3099958D4539FF830EC8FA4BFA00FA13" box="[151,1436,1491,1513]" pageId="2">
|
|||
|
|
- Cephalic carina lacking striation. Anterior excavation of pronotum bearing at most fine punctures (
|
|||
|
|
<date id="FFD610784539FF830ACBFA4CFB5CFA13" box="[1172,1216,1491,1513]" pageId="2" value="1920-10">20 x</date>
|
|||
|
|
) (as on
|
|||
|
|
<figureCitation id="13532A3D4539FF830B52FA4CFAD9FA13" box="[1293,1349,1491,1513]" captionStart="FIGURE 7" captionStartId="12.[151,250,1231,1253]" captionTargetBox="[268,1307,198,1194]" captionTargetId="figure@12.[265,1321,193,1210]" captionTargetPageId="12" captionText="FIGURE 7. Dichotomius (Selenocopris) periotoi n. sp.: (A) male dorsal habitus; (B) female dorsal habitus; (C) dorsal view of male paramera; (D) lateral view of male paramera; (E) detail of the fine punctures of the anterior pronotal excavation. Scale bar = 5 mm." httpUri="https://zenodo.org/record/272006/files/figure.png" pageId="2">Fig. 7</figureCitation>
|
|||
|
|
E)... 9
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
</keyStep>
|
|||
|
|
<keyStep id="309C2E1D4539FF830EC8FA6FFA00F94C" pageId="2">
|
|||
|
|
<paragraph id="8BD736B84539FF830EC8FA6FFA00F9A4" blockId="2.[151,1436,259,1718]" pageId="2">
|
|||
|
|
<keyLead id="3099958D4539FF830EC8FA6FFA00F9A4" pageId="2">
|
|||
|
|
9 Clypeo-frontal horn carinate, strongly transverse, widely emarginated at the apex and producing two tubercles at each apex (
|
|||
|
|
<figureCitation id="13532A3D4539FF830EB7F992FEBEF9D9" box="[232,290,1549,1571]" captionStart="FIGURE 7" captionStartId="12.[151,250,1231,1253]" captionTargetBox="[268,1307,198,1194]" captionTargetId="figure@12.[265,1321,193,1210]" captionTargetPageId="12" captionText="FIGURE 7. Dichotomius (Selenocopris) periotoi n. sp.: (A) male dorsal habitus; (B) female dorsal habitus; (C) dorsal view of male paramera; (D) lateral view of male paramera; (E) detail of the fine punctures of the anterior pronotal excavation. Scale bar = 5 mm." httpUri="https://zenodo.org/record/272006/files/figure.png" pageId="2">Fig. 7</figureCitation>
|
|||
|
|
A). On smaller individuals, cephalic horn not producing lateral tubercles but still with a conspicuous emargination at apex. Smaller species (maximum body length:
|
|||
|
|
<quantity id="4C909B5D4539FF830CF5F9B4FC9FF9BB" box="[682,771,1579,1601]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="2" unit="mm" value="19.5">19.5 mm</quantity>
|
|||
|
|
; maximum pronotum width:
|
|||
|
|
<quantity id="4C909B5D4539FF830A7FF9B4FBF4F9BB" box="[1056,1128,1579,1601]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="2" unit="mm" value="13.0">13 mm</quantity>
|
|||
|
|
). Descalvado Municipality,
|
|||
|
|
<collectingRegion id="49ACF85A4539FF830B27F9B4FE84F9A4" country="Brazil" name="Sao Paulo" pageId="2">São Paulo</collectingRegion>
|
|||
|
|
state,
|
|||
|
|
<collectingCountry id="F37F76284539FF830F0AF9D7FE0CF9A4" box="[341,400,1608,1630]" name="Brazil" pageId="2">Brazil</collectingCountry>
|
|||
|
|
.....................................................
|
|||
|
|
<taxonomicName id="4C684D3B4539FF830A42F9D6FA8EF9A4" ID-CoL="6D43B" box="[1053,1298,1609,1630]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="periotoi" status="sp. nov." subGenus="Selenocopris">
|
|||
|
|
<emphasis id="B91CEAAA4539FF830A42F9D6FA8EF9A4" box="[1053,1298,1609,1630]" italics="true" pageId="2">Dichotomius (S.) periotoi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B91CEAAA4539FF830B47F9D7FAD2F9A4" bold="true" box="[1304,1358,1608,1630]" pageId="2">
|
|||
|
|
<taxonomicNameLabel id="A22F57D14539FF830B47F9D7FAD2F9A4" box="[1304,1358,1608,1630]" pageId="2" rank="species">n. sp.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="13532A3D4539FF830B04F9D7FA09F9A4" box="[1371,1429,1608,1630]" captionStart="FIGURE 7" captionStartId="12.[151,250,1231,1253]" captionTargetBox="[268,1307,198,1194]" captionTargetId="figure@12.[265,1321,193,1210]" captionTargetPageId="12" captionText="FIGURE 7. Dichotomius (Selenocopris) periotoi n. sp.: (A) male dorsal habitus; (B) female dorsal habitus; (C) dorsal view of male paramera; (D) lateral view of male paramera; (E) detail of the fine punctures of the anterior pronotal excavation. Scale bar = 5 mm." httpUri="https://zenodo.org/record/272006/files/figure.png" pageId="2">Fig. 7</figureCitation>
|
|||
|
|
)
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BD736B84539FF830EC8F9F9FA00F94C" blockId="2.[151,1436,259,1718]" pageId="2">
|
|||
|
|
<keyLead id="3099958D4539FF830EC8F9F9FA00F94C" pageId="2">
|
|||
|
|
- Clypeo-frontal horn carinate but not strongly transverse, lacking emargination and tubercles at the apex (
|
|||
|
|
<figureCitation id="13532A3D4539FF830A99F9FAFB63F981" box="[1222,1279,1637,1659]" captionStart="FIGURE 5" captionStartId="9.[151,250,1708,1730]" captionTargetBox="[341,1244,193,1685]" captionTargetId="figure@9.[341,1245,193,1686]" captionTargetPageId="9" captionText="FIGURE 5. Holotype of Dichotomius (Selenocopris) manni n. sp.: (A) male dorsal habitus; (B) male paramera." httpUri="https://zenodo.org/record/272004/files/figure.png" pageId="2">Fig. 5</figureCitation>
|
|||
|
|
A). Females and smaller individuals unknown. Larger species (maximum body length:
|
|||
|
|
<quantity id="4C909B5D4539FF830DD2F91CFC48F963" box="[909,980,1667,1689]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="2" unit="mm" value="25.0">25 mm</quantity>
|
|||
|
|
; maximum pronotum width:
|
|||
|
|
<quantity id="4C909B5D4539FF830AAEF91CFAA5F963" box="[1265,1337,1667,1689]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="2" unit="mm" value="14.0">14 mm</quantity>
|
|||
|
|
). Serra da Canastra, Southwest of
|
|||
|
|
<collectingRegion id="49ACF85A4539FF830F9BF93FFDD8F94C" box="[452,580,1696,1718]" country="Brazil" name="Minas Gerais" pageId="2">Minas Gerais</collectingRegion>
|
|||
|
|
state,
|
|||
|
|
<collectingCountry id="F37F76284539FF830CDCF93FFD22F94C" box="[643,702,1696,1718]" name="Brazil" pageId="2">Brazil</collectingCountry>
|
|||
|
|
..............................
|
|||
|
|
<taxonomicName id="4C684D3B4539FF830A72F93EFA8EF94C" ID-CoL="6D3RL" box="[1069,1298,1697,1718]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="manni" status="sp. nov." subGenus="Selenocopris">
|
|||
|
|
<emphasis id="B91CEAAA4539FF830A72F93EFA8EF94C" box="[1069,1298,1697,1718]" italics="true" pageId="2">Dichotomius (S.) manni</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A22F57D14539FF830B47F93FFAD2F94C" box="[1304,1358,1696,1718]" pageId="2" rank="species">
|
|||
|
|
<emphasis id="B91CEAAA4539FF830B47F93FFAD2F94C" bold="true" box="[1304,1358,1696,1718]" pageId="2">n. sp.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="13532A3D4539FF830B04F93FFA08F94C" box="[1371,1428,1696,1718]" captionStart="FIGURE 5" captionStartId="9.[151,250,1708,1730]" captionTargetBox="[341,1244,193,1685]" captionTargetId="figure@9.[341,1245,193,1686]" captionTargetPageId="9" captionText="FIGURE 5. Holotype of Dichotomius (Selenocopris) manni n. sp.: (A) male dorsal habitus; (B) male paramera." httpUri="https://zenodo.org/record/272004/files/figure.png" pageId="2">Fig. 5</figureCitation>
|
|||
|
|
)
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
</keyStep>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|