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116 lines
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<document id="6B4B54EA2302639577F845E9143F02BE" ID-CLB-Dataset="28466" ID-DOI="http://dx.doi.org/10.3897/dez.65.27424" ID-GBIF-Dataset="290e099d-cb77-4ad7-975e-32ce4a51926f" ID-Pensoft-Pub="1860-1324-2-177" ID-ZooBank="A047B48DD161424FB8800428DCC5888A" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1860-1324-2-177" ModsDocOrigin="Deutsche Entomologische Zeitschrift 65 (2)" ModsDocTitle="Taxonomic review of Australian Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) with special emphasis on the M.lophoides (Chaudoir) species complex" checkinTime="1552154842744" checkinUser="pensoft" docAuthor="Liebherr, James K." docDate="2018" docId="1703D5BCE180D81D9746CAD0FBA02F98" docLanguage="en" docName="DeutEntomolZeit 65(2): 177-224" docOrigin="Deutsche Entomologische Zeitschrift 65 (2)" docSource="http://dx.doi.org/10.3897/dez.65.27424" docTitle="Moriomorphina (Mecyclothorax) Sharp" docType="treatment" docVersion="7" lastPageNumber="177" masterDocId="FFB33A56FFD86772FFAAFFA9FF882245" masterDocTitle="Taxonomic review of Australian Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) with special emphasis on the M. lophoides (Chaudoir) species complex" masterLastPageNumber="224" masterPageNumber="177" pageNumber="177" updateTime="1701378747773" updateUser="ExternalLinkService">
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<mods:title id="619A7D97A2F75834324B569A24046BFA">Taxonomic review of Australian Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) with special emphasis on the M. lophoides (Chaudoir) species complex</mods:title>
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<mods:namePart id="D97F4BC058EA437CF1F5C28820055FE1">Liebherr, James K.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="CA983FC2D64374FED496AE6E92B0EB34">text</mods:typeOfResource>
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<mods:title id="EEBDA84E632E212F97836B390401E891">Deutsche Entomologische Zeitschrift</mods:title>
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<mods:part id="482D3D659F03A7AC9877840AE506CFA8">
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<mods:date id="FCB51BAC1AEA58909D58F27B6D942316">2018</mods:date>
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<mods:detail id="9DD9A72C9696C0820AF350D93B12698B" type="volume">
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<mods:number id="31EA28AE942B8669060717B9F5571EB6">65</mods:number>
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<mods:number id="C16B579327CD8E34A363048A384648D4">2</mods:number>
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<mods:url id="9D16B8BF5AF6263C56AA6AE8075CFDFE">http://dx.doi.org/10.3897/dez.65.27424</mods:url>
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</mods:location>
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<mods:classification id="95D165AC0D4BC890C1BEEF169EFF2D48">journal article</mods:classification>
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<mods:identifier id="A7936804F95033B0608ED698D7B42AEA" type="DOI">http://dx.doi.org/10.3897/dez.65.27424</mods:identifier>
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</mods:mods>
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<treatment id="1703D5BCE180D81D9746CAD0FBA02F98" ID-GBIF-Taxon="154365998" LSID="urn:lsid:plazi:treatment:1703D5BCE180D81D9746CAD0FBA02F98" httpUri="http://treatment.plazi.org/id/1703D5BCE180D81D9746CAD0FBA02F98" lastPageNumber="177" pageId="0" pageNumber="177">
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<subSubSection id="35263CF54BA303F0521AE9A5EE26F206" pageId="0" pageNumber="177" type="nomenclature">
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<paragraph id="464D124F5D7A293CF435446D75226C63" pageId="0" pageNumber="177">
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<taxonomicName id="AEF542849C91C21E728A600CCF776C8B" authority="Sharp" class="Insecta" family="Carabidae" genus="Moriomorphina" higherTaxonomySource="GBIF" infraspecific-rank="Subgenus" kingdom="Animalia" lsidName="Moriomorphina (Mecyclothorax)" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="subGenus" subGenus="Mecyclothorax">Subgenus Mecyclothorax Sharp</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="FEDB2210B4A126D0B74A74B213443867" pageId="0" pageNumber="177" type="diagnosis">
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<paragraph id="912D85268A96F3A10DD272B76AD0B07B" pageId="0" pageNumber="177">Diagnosis.</paragraph>
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<paragraph id="7D0FAC8D5022C0783612DB4A06D3E9E5" pageId="0" pageNumber="177">
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This subgenus comprises over 350 species (
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<bibRefCitation id="0EB7B9488D15C9F87D07FF9E7DBF3C7B" author="Liebherr, JK" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="0" pageNumber="177" pagination="1 - 63" title="Cladistic classification of Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) and taxonomic revision of the New Caledonian subgenus Phacothorax Jeannel." url="https://doi.org/10.3897/dez.65.21000" volume="65" year="2018">Liebherr 2018</bibRefCitation>
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), and so morphological variation among the species is the most extreme observed among the various
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<taxonomicName id="E7A8695E9C62371FAB6CD1A8107F7DB5" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
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subgenera. However, throughout taxa of this subgenus, the labrum is emarginate apically, either distinctly and angularly as in
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<taxonomicName id="48BAAC7F981E2274C993F337A082DB4D" lsidName="M. goweri" pageId="0" pageNumber="177" rank="species" species="goweri">M. goweri</taxonomicName>
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Moore of Lord Howe Island (
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<bibRefCitation id="B8BFFCFFCBEF57FBA53DFDAC10903D29" author="Liebherr, JK" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="0" pageNumber="177" pagination="1 - 63" title="Cladistic classification of Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) and taxonomic revision of the New Caledonian subgenus Phacothorax Jeannel." url="https://doi.org/10.3897/dez.65.21000" volume="65" year="2018">Liebherr 2018</bibRefCitation>
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, fig. 1A), or less so as in
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<taxonomicName id="8C97785FF03AB9E09CAE3A4449471AE8" lsidName="M. aeneipennis" pageId="0" pageNumber="177" rank="species" species="aeneipennis">M. aeneipennis</taxonomicName>
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Liebherr of
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<normalizedToken id="3FC96D2CA3BCD575129FFA89C45F0834" originalValue="Haleakalā">Haleakala</normalizedToken>
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, Maui,
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<normalizedToken id="4D567AA3998BC752E1473A404D5A8153" originalValue="Hawaiʻi">Hawaiʻi</normalizedToken>
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(
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<bibRefCitation id="64B334798C0544637A7C1654213D73F9" author="Liebherr, JK" journalOrPublisher="ZooKeys" pageId="0" pageNumber="177" pagination="1 - 407" title="The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala, Maui: Keystone of a hyperdiverse Hawaiian radiation." url="https://doi.org/10.3897/zookeys.544.6074" volume="544" year="2015">Liebherr 2015</bibRefCitation>
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, fig. 7). The ligular margin is generally truncate with the ligular setae well separated (
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<bibRefCitation id="AD8D3B4EC2978D276D141C6EA6ECD7E4" author="Liebherr, JK" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="0" pageNumber="177" pagination="1 - 63" title="Cladistic classification of Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) and taxonomic revision of the New Caledonian subgenus Phacothorax Jeannel." url="https://doi.org/10.3897/dez.65.21000" volume="65" year="2018">Liebherr 2018</bibRefCitation>
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, fig. 1G), though as exceptions, the ligula is apically rounded in the Papuan taxa
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<taxonomicName id="A15CC37E5528046DBAE016350E97FFD2" lsidName="M. brispex" pageId="0" pageNumber="177" rank="species" species="brispex">M. brispex</taxonomicName>
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and
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<taxonomicName id="D9B792201C5CB45ABE8F428FE5ACA8BD" lsidName="M. andersoni" pageId="0" pageNumber="177" rank="species" species="andersoni">M. andersoni</taxonomicName>
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(
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<bibRefCitation id="D5ABF1D2605393D7CE5722AEA3E0E504" author="Liebherr, JK" journalOrPublisher="Coleopterists Bulletin" pageId="0" pageNumber="177" pagination="679 - 703" title="Review of Mecyclothorax Sharp (Coleoptera: Carabidae: Moriomorphini) from Papua New Guinea with descriptions of five new species." url="https://doi.org/10.1649/0010-065X-71.4.679" volume="71" year="2017">Liebherr 2017</bibRefCitation>
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,
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<bibRefCitation id="BB9F909A54F37272B642FF99076C8C80" author="Liebherr, JK" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="0" pageNumber="177" pagination="1 - 63" title="Cladistic classification of Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) and taxonomic revision of the New Caledonian subgenus Phacothorax Jeannel." url="https://doi.org/10.3897/dez.65.21000" volume="65" year="2018">2018</bibRefCitation>
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, fig. 7). The prosternum exhibits a smooth to distinctly punctate anteapical groove, though never any other punctures, such as in the median depression observed among species of subgenus
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<taxonomicName id="5F639EBC2745BD411E7A81417B099CCA" class="Insecta" family="Carabidae" genus="Moriomorphina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Moriomorphina (Eucyclothorax)" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="subGenus" subGenus="Eucyclothorax">Eucyclothorax</taxonomicName>
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. The parascutellar striole is present, and may be smooth or punctate, with up to 8 punctures along its length (Fig. 6). Among the mainland Australian species, the pronotal median base is depressed relative to the disc, and punctate; a distinction noted by
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<bibRefCitation id="81D104AC0598677C623B05CA500E9953" author="Blackburn, T" journalOrPublisher=")" pageId="0" pageNumber="177" pagination="1387 - 1506" title="Further notes on Australian Coleoptera, with descriptions of new genera and species. Proceedings of the Linnean Society of New South Wales (2 nd ser." volume="3" year="1889">Blackburn (1889)</bibRefCitation>
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.
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</paragraph>
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<paragraph id="6D7F06B8F54C8E8F35E9FDDA73012C07" pageId="0" pageNumber="177">
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The male aedeagus has an internal sac with an apical flagellar plate surrounding the gonopore (Fig. 15E). The female reproductive tract most often has the spermathecal duct entering the bursa copulatrix mediodorsally (Fig. 17
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<normalizedToken id="341A10B6E67379B2CEC9DC78E042A4B8" originalValue="C–D">C-D</normalizedToken>
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), however the sister-species pair
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<taxonomicName id="0C41377D3B4D638EE6FC2D9F31210BD7" lsidName="M. lateralis" pageId="0" pageNumber="177" rank="species" species="lateralis">M. lateralis</taxonomicName>
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+
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<taxonomicName id="AC8705E5DF1EED36371A2C3FC36A5B3D" lsidName="M. minutus" pageId="0" pageNumber="177" rank="species" species="minutus">M. minutus</taxonomicName>
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(Fig. 1A) revert to the plesiomorphic configuration wherein the spermathecal duct basally joins the juncture of the common oviduct and the bursa copulatrix (Fig. 17
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<normalizedToken id="CC60CAFA450061CC6A0C7803EADE7910" originalValue="A–B">A-B</normalizedToken>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="835EFB571D4F86683905DD3A3287086B" pageId="0" pageNumber="177" type="member species">
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<paragraph id="4C13A644B42469986684EB8BD717DA5C" pageId="0" pageNumber="177">Member species.</paragraph>
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<paragraph id="FE0E4D10323E37C9312F1A7989F67236" pageId="0" pageNumber="177">
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This subgenus is represented by four species in mainland Australia. Numbers of taxa in the substantial radiations from Hawaii, the Society Islands, New Guinea, New Zealand, the Sundas, and Lord Howe, Norfolk, and St. Paul and Amsterdam Islands are summarized in
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<bibRefCitation id="31B6D07F837FD20512E243BA1E094FD1" author="Liebherr, JK" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="0" pageNumber="177" pagination="1 - 63" title="Cladistic classification of Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) and taxonomic revision of the New Caledonian subgenus Phacothorax Jeannel." url="https://doi.org/10.3897/dez.65.21000" volume="65" year="2018">Liebherr (2018</bibRefCitation>
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: 15).
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</paragraph>
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<caption id="BA6E78B60C4ED493852A7C7AAEFA7D85" pageId="0" pageNumber="177">
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<paragraph id="E8F30BE67D5DEEC8FF2E7B9F1F21572D" pageId="0" pageNumber="177">
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Figure 6.
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<taxonomicName id="6C161D6356F90683887852FC23875F66" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
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(s. s.) spp., dorsal view: A,
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<taxonomicName id="C26DD821DC1A25B6631D9C45AD051285" lsidName="M. lateralis" pageId="0" pageNumber="177" rank="species" species="lateralis">M. lateralis</taxonomicName>
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; B,
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<taxonomicName id="EDB008B3B31914C20FEEA6ABF9BD622B" lsidName="M. minutus" pageId="0" pageNumber="177" rank="species" species="minutus">M. minutus</taxonomicName>
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; C,
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<taxonomicName id="AA95DBD9DA1F0C4CC29AD8E80D3382FB" lsidName="M. ambiguus" pageId="0" pageNumber="177" rank="species" species="ambiguus">M. ambiguus</taxonomicName>
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; D,
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<taxonomicName id="518A66ACB18FE2D9E281982BB4F18F46" lsidName="M. punctipennis" pageId="0" pageNumber="177" rank="species" species="punctipennis">M. punctipennis</taxonomicName>
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.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document>
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