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<document id="93737F44056BA4190FFFFCBFF280BE63" ID-CLB-Dataset="29691" ID-DOI="10.11646/zootaxa.4444.4.7" ID-GBIF-Dataset="9629e519-7bd2-42e6-aea6-7d480ddd8392" ID-ISSN="1175-5326" ID-Zenodo-Dep="1309921" ID-ZooBank="D943FB45-6B57-430D-9B77-3B0FB0F647DE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1531314576115" checkinUser="plazi" docAuthor="Winterbottom, Richard &amp; Erdmann, Mark V." docDate="2018" docId="3E6F87A3FFC3FF8A8EF5F9DC1A7AF84F" docLanguage="en" docName="zootaxa.4444.4.7.pdf" docOrigin="Zootaxa 4444 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trimma meityae Winterbottom &amp; Erdmann 2018, new species" docType="treatment" docUuid="7A218DC5-BADD-46F5-AF4B-699D538BF4D3" docUuidSource="ZooBank" docVersion="8" lastPageNumber="480" masterDocId="C256FFDBFFC6FF838E62FFBD1E7CFFA7" masterDocTitle="Two new species of blue-eyed Trimma (Pisces; Gobiidae) from New Guinea" masterLastPageNumber="483" masterPageNumber="471" pageNumber="476" updateTime="1698553880082" updateUser="ExternalLinkService">
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<mods:title id="E9679E2028D53A788F55148394090775">Two new species of blue-eyed Trimma (Pisces; Gobiidae) from New Guinea</mods:title>
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<mods:namePart id="EA0BA7234F7A587D97F6699BDFABC51D">Winterbottom, Richard</mods:namePart>
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<mods:namePart id="5287859F41F32F22495D0849CB22E5A5">Erdmann, Mark V.</mods:namePart>
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<mods:title id="545BBD0D6F84ABD0F32A023F42EB45AD">Zootaxa</mods:title>
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<mods:date id="0553336CFE983CABC3F1850846D18666">2018</mods:date>
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<mods:number id="8401A337F73AD979951B58BF10EBF920">2018-06-11</mods:number>
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<mods:identifier id="67840F295DFD4A6FC9B89A7584A406DD" type="CLB-Dataset">29691</mods:identifier>
<mods:identifier id="56D396FE2E91950E535E99442C32203F" type="DOI">10.11646/zootaxa.4444.4.7</mods:identifier>
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<treatment id="3E6F87A3FFC3FF8A8EF5F9DC1A7AF84F" ID-DOI="http://doi.org/10.5281/zenodo.5997057" ID-GBIF-Taxon="146045464" ID-Zenodo-Dep="5997057" LSID="urn:lsid:zoobank.org:act:7A218DC5-BADD-46F5-AF4B-699D538BF4D3" httpUri="http://treatment.plazi.org/id/3E6F87A3FFC3FF8A8EF5F9DC1A7AF84F" lastPageId="9" lastPageNumber="480" pageId="5" pageNumber="476">
<subSubSection id="FEDC653EFFC3FF868EF5F9DC1F06F93A" pageId="5" pageNumber="476" type="nomenclature">
<paragraph id="B67936B5FFC3FF868EF5F9DC1F8FF9DC" blockId="5.[151,636,1633,1765]" box="[151,499,1633,1659]" pageId="5" pageNumber="476">
<heading id="ED3181D9FFC3FF868EF5F9DC1F8FF9DC" bold="true" box="[151,499,1633,1659]" fontSize="11" level="1" pageId="5" pageNumber="476" reason="1">
<emphasis id="84B2EAA7FFC3FF868EF5F9DC1F8FF9DC" bold="true" box="[151,499,1633,1659]" pageId="5" pageNumber="476">
<taxonomicName id="71C64D36FFC3FF868EF5F9DC1F20F9DC" authority="Winterbottom &amp; Erdmann, 2018" authorityName="Winterbottom &amp; Erdmann" authorityYear="2018" box="[151,348,1633,1659]" class="Actinopterygii" family="Gobiidae" genus="Trimma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="476" phylum="Chordata" rank="species" species="meityae" status="sp. nov.">
<emphasis id="84B2EAA7FFC3FF868EF5F9DC1F20F9DC" bold="true" box="[151,348,1633,1659]" italics="true" pageId="5" pageNumber="476">Trimma meityae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F8157DCFFC3FF868F01F9DC1F8FF9DC" box="[355,499,1633,1659]" pageId="5" pageNumber="476" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="B67936B5FFC3FF868EF5F9391F06F93A" blockId="5.[151,636,1633,1765]" box="[151,378,1668,1693]" pageId="5" pageNumber="476">Meity’s Pygmygoby</paragraph>
</subSubSection>
<subSubSection id="FEDC653EFFC3FF868EF5F9151C00F942" pageId="5" pageNumber="476" type="description">
<paragraph id="B67936B5FFC3FF868EF5F9151F7AF967" blockId="5.[151,636,1633,1765]" box="[151,262,1704,1729]" pageId="5" pageNumber="476">
<figureCitation id="2EFD2A30FFC3FF868EF5F9151F7EF967" box="[151,258,1704,1729]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStart-3="FIGURE 7" captionStartId-0="7.[151,250,1463,1485]" captionStartId-1="8.[151,250,1968,1990]" captionStartId-2="9.[151,250,1139,1161]" captionStartId-3="10.[151,250,1896,1918]" captionTargetBox-0="[151,1433,217,1416]" captionTargetBox-1="[164,1428,301,1949]" captionTargetBox-2="[151,1436,228,1085]" captionTargetBox-3="[151,1436,255,1819]" captionTargetId-0="figure@7.[151,1436,193,1442]" captionTargetId-1="figure@8.[164,1428,301,1949]" captionTargetId-2="figure@9.[151,1436,228,1085]" captionTargetId-3="figure@10.[151,1436,255,1819]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="10" captionText-0="FIGURE 4. Neighbour Joining network derived from a BOL analysis of the COI gene of the Trimma haplogroups associated with T. blematium and T. meityae. Image by R. Winterbottom." captionText-1="FIGURE 5. Trimma meityae, live; A &amp; B—Cendrawasih Bay; C—freshly collected, Cendrawasih Bay. Photos: A &amp; B Figure by M. V. Erdmann; C by G. R. Allen." captionText-2="FIGURE 6. Trimma meityae, 17.3 mm SL male paratype, ROM 106364, Cendrawasih Bay, lateral (and slightly dorsal) view of left nasal apparatus, specimen stained with cyanine blue. Photo by R. Winterbottom." captionText-3="FIGURE 7. Trimma meityae, 18.6 male holotype, ROM 106350. Left lateral (A) and dorsal (B) views of head papillae, specimen stained with cyanine blue. Papillae in a given row connected by thin green lines, red arrow points to supernumerary papilla. Second letters of abbreviations in ea, ep and ip stand for anterior and posterior respectively. Photos by R. Winterbottom." httpUri-0="https://zenodo.org/record/1309933/files/figure.png" httpUri-1="https://zenodo.org/record/1309935/files/figure.png" httpUri-2="https://zenodo.org/record/1309937/files/figure.png" httpUri-3="https://zenodo.org/record/1309939/files/figure.png" pageId="5" pageNumber="476">Figs. 4–7</figureCitation>
.
</paragraph>
<paragraph id="B67936B5FFC3FF868EF5F9701C00F942" blockId="5.[151,636,1633,1765]" box="[151,636,1740,1765]" pageId="5" pageNumber="476">No published names pertain to this species.</paragraph>
</subSubSection>
<subSubSection id="FEDC653EFFC3FF868EF5F8A91CF9F846" pageId="5" pageNumber="476" type="materials_examined">
<paragraph id="B67936B5FFC3FF868EF5F8A91A2AF8F6" blockId="5.[151,1436,1812,2017]" pageId="5" pageNumber="476">
<materialsCitation id="06AE3CE8FFC3FF868EF5F8A91A2AF8F6" ID-GBIF-Occurrence="1880439934" collectingDate="4 Aug." collectionCode="ROM" collectorName="M. V. Erdmann." country="Indonesia" county="Material" elevation="56" latitude="-2.0531332" location="Cendrawasih Bay" longLatPrecision="1" longitude="134.1553" municipality="Pulau Purup" pageId="5" pageNumber="476" specimenCode="ROM 106350, 18.6" specimenCount="1" stateProvince="West Papua" typeStatus="holotype">
<emphasis id="84B2EAA7FFC3FF868EF5F8A91C72F88A" bold="true" box="[151,526,1812,1837]" pageId="5" pageNumber="476">
<collectingCounty id="5F184E39FFC3FF868EF5F8A91E83F88A" box="[151,255,1812,1837]" pageId="5" pageNumber="476">Material</collectingCounty>
examined.
<typeStatus id="697D8817FFC3FF868FF9F8A91C76F88A" box="[411,522,1812,1837]" pageId="5" pageNumber="476" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="E6609ECEFFC3FF868C7DF8A81CBBF88A" box="[543,711,1812,1837]" collectionCode="ROM" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="5" pageNumber="476">ROM 106350</specimenCode>
,
<quantity id="713E9B50FFC3FF868CB9F8A91D35F88B" box="[731,841,1812,1837]" metricMagnitude="-2" metricUnit="m" metricValue="1.86" pageId="5" pageNumber="476" unit="mm" value="18.6">
<specimenCode id="E6609ECEFFC3FF868CB9F8A91D70F88A" box="[731,780,1812,1837]" collectionCode="ROM" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="5" pageNumber="476">18.6</specimenCode>
mm
</quantity>
SL male,
<collectingCountry id="CED17625FFC3FF868DBBF8A91A35F88A" box="[985,1097,1812,1837]" name="Indonesia" pageId="5" pageNumber="476">Indonesia</collectingCountry>
,
<collectingRegion id="7402F857FFC3FF868A3CF8A81A91F88B" box="[1118,1261,1813,1836]" country="Indonesia" name="Papua Barat" pageId="5" pageNumber="476">West Papua</collectingRegion>
,
<collectingMunicipality id="561DACCFFFC3FF868B60F8A91BE4F88B" box="[1282,1432,1812,1837]" pageId="5" pageNumber="476">Pulau Purup</collectingMunicipality>
,
<location id="B319606EFFC3FF868EF5F8851F22F8F6" LSID="urn:lsid:plazi:treatment:3E6F87A3FFC3FF8A8EF5F9DC1A7AF84F:B319606EFFC3FF868EF5F8851F22F8F6" box="[151,350,1848,1873]" country="Indonesia" county="Material" latitude="-2.0531332" longLatPrecision="1" longitude="134.1553" municipality="Pulau Purup" name="Cendrawasih Bay" pageId="5" pageNumber="476" stateProvince="West Papua">Cendrawasih Bay</location>
,
<geoCoordinate id="D3F25072FFC3FF868F0BF8851C7DF8F7" box="[361,513,1848,1872]" direction="south" orientation="latitude" pageId="5" pageNumber="476" precision="1" value="-2.0531332">02° 03.188' S</geoCoordinate>
,
<geoCoordinate id="D3F25072FFC3FF868C6CF8851CCAF8F7" box="[526,694,1848,1872]" direction="east" orientation="longitude" pageId="5" pageNumber="476" precision="1" value="134.1553">134° 09.318' E</geoCoordinate>
,
<elevation id="3DEBD186FFC3FF868CA0F8851C82F8F7" box="[706,766,1848,1873]" metricMagnitude="1" metricUnit="m" metricValue="5.6" pageId="5" pageNumber="476" unit="m" value="56.0">
<quantity id="713E9B50FFC3FF868CA0F8851C82F8F7" box="[706,766,1848,1873]" metricMagnitude="1" metricUnit="m" metricValue="5.6" pageId="5" pageNumber="476" unit="m" value="56.0">56 m</quantity>
</elevation>
,
<collectingDate id="D23CE99DFFC3FF868D6BF8851D24F8F6" box="[777,856,1848,1873]" pageId="5" pageNumber="476">
<date id="C2781075FFC3FF868D6BF8851D24F8F6" box="[777,856,1848,1873]" pageId="5" pageNumber="476">4 Aug.</date>
</collectingDate>
, 2017,
<collectorName id="1B335363FFC3FF868DCBF8841A2AF8F6" box="[937,1110,1848,1873]" pageId="5" pageNumber="476">M.V. Erdmann.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="B67936B5FFC3FF868EA5F8E11CF9F846" blockId="5.[151,1436,1812,2017]" pageId="5" pageNumber="476">
<materialsCitation id="06AE3CE8FFC3FF868EA5F8E11F94F81A" ID-GBIF-Occurrence="1880439933" collectingDate="1924-10" collectionCode="MZB, ROM" collectorName="M. V. Erdmann" country="Indonesia" elevation="52" latitude="-2.0531332" location="Pulau Purup" longLatPrecision="1" longitude="134.1553" municipality="Cendrawasih Bay" pageId="5" pageNumber="476" specimenCode="2" specimenCount="2" stateProvince="West Papua" typeStatus="paratype">
<emphasis id="84B2EAA7FFC3FF868EA5F8E11F34F8D2" bold="true" box="[199,328,1884,1909]" pageId="5" pageNumber="476">
<typeStatus id="697D8817FFC3FF868EA5F8E11F3FF8D2" box="[199,323,1884,1909]" pageId="5" pageNumber="476" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectionCode id="D0D7AE70FFC3FF868F31F8E01FECF8D3" box="[339,400,1885,1908]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="5" pageNumber="476">MZB</collectionCode>
24599,
<specimenCode id="E6609ECEFFC3FF868F90F8E01C7CF8D2" box="[498,512,1885,1909]" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="5" pageNumber="476">2</specimenCode>
(17.1–17.2), collected with the
<typeStatus id="697D8817FFC3FF868D12F8E11DAAF8D2" box="[880,982,1884,1909]" pageId="5" pageNumber="476" type="holotype">holotype</typeStatus>
.
<collectionCode id="D0D7AE70FFC3FF868D81F8E01A5FF8D2" box="[995,1059,1885,1909]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="5" pageNumber="476">ROM</collectionCode>
T21296, (
<quantity id="713E9B50FFC3FF868AC2F8E11B75F8D3" box="[1184,1289,1884,1909]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="5" pageNumber="476" unit="mm" value="15.5">15.5 mm</quantity>
SL female),
<collectingCountry id="CED17625FFC3FF868EF5F83D1F7BF83E" box="[151,263,1920,1945]" name="Indonesia" pageId="5" pageNumber="476">Indonesia</collectingCountry>
,
<collectingRegion id="7402F857FFC3FF868F70F83C1FEBF83F" box="[274,407,1921,1944]" country="Indonesia" name="Papua Barat" pageId="5" pageNumber="476">West Papua</collectingRegion>
,
<collectingMunicipality id="561DACCFFFC3FF868FC1F83D1C16F83E" box="[419,618,1920,1945]" pageId="5" pageNumber="476">Cendrawasih Bay</collectingMunicipality>
,
<location id="B319606EFFC3FF868C17F83D1D7EF83F" LSID="urn:lsid:plazi:treatment:3E6F87A3FFC3FF8A8EF5F9DC1A7AF84F:B319606EFFC3FF868C17F83D1D7EF83F" box="[629,770,1920,1945]" country="Indonesia" latitude="-2.0531332" longLatPrecision="1" longitude="134.1553" municipality="Cendrawasih Bay" name="Pulau Purup" pageId="5" pageNumber="476" stateProvince="West Papua">Pulau Purup</location>
,
<geoCoordinate id="D3F25072FFC3FF868D6FF83D1DD9F83F" box="[781,933,1920,1944]" direction="south" orientation="latitude" pageId="5" pageNumber="476" precision="1" value="-2.0531332">02° 03.188' S</geoCoordinate>
,
<geoCoordinate id="D3F25072FFC3FF868DD0F83D1A26F83F" box="[946,1114,1920,1944]" direction="east" orientation="longitude" pageId="5" pageNumber="476" precision="1" value="134.1553">134° 09.318' E</geoCoordinate>
,
<elevation id="3DEBD186FFC3FF868A04F83D1ADDF83F" box="[1126,1185,1920,1945]" metricMagnitude="1" metricUnit="m" metricValue="5.2" pageId="5" pageNumber="476" unit="m" value="52.0">
<quantity id="713E9B50FFC3FF868A04F83D1ADDF83F" box="[1126,1185,1920,1945]" metricMagnitude="1" metricUnit="m" metricValue="5.2" pageId="5" pageNumber="476" unit="m" value="52.0">52 m</quantity>
</elevation>
,
<collectingDate id="D23CE99DFFC3FF868ACFF83D1B7FF83F" box="[1197,1283,1920,1944]" pageId="5" pageNumber="476" value="1924-10">
<date id="C2781075FFC3FF868ACFF83D1B7FF83F" box="[1197,1283,1920,1944]" pageId="5" pageNumber="476" value="1924-10">24 Oct.</date>
</collectingDate>
, 2016, field # MVE-16-070,
<collectorName id="1B335363FFC3FF868F5FF8181F94F81A" box="[317,488,1956,1981]" pageId="5" pageNumber="476">M.V. Erdmann</collectorName>
</materialsCitation>
. ROM 106364, 3(17.3–17.7), collected with the holotype. ROM T25134, (16.1), &amp; T25135, (16.8), collected with the holotype.
</paragraph>
</subSubSection>
<subSubSection id="FEDC653EFFC0FF858EA5FF2A1C2AFE2C" pageId="6" pageNumber="477" type="diagnosis">
<paragraph id="B67936B5FFC0FF858EA5FF2A1C2AFE2C" blockId="6.[151,1437,151,1999]" pageId="6" pageNumber="477">
<emphasis id="84B2EAA7FFC0FF858EA5FF2A1F3DFF17" bold="true" box="[199,321,151,176]" pageId="6" pageNumber="477">Diagnosis.</emphasis>
A species of
<taxonomicName id="71C64D36FFC0FF858FBCFF241C49FF17" authorityName="Jordan &amp; Seale" authorityYear="1906" box="[478,565,153,176]" class="Actinopterygii" family="Gobiidae" genus="Trimma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="477" phylum="Chordata" rank="genus">
<emphasis id="84B2EAA7FFC0FF858FBCFF241C49FF17" box="[478,565,153,176]" italics="true" pageId="6" pageNumber="477">Trimma</emphasis>
</taxonomicName>
with scales present on cheeks and opercle, 8–10 scales in predorsal midline, 17–18 unbranched pectoral fin rays, unbranched 5th pelvic fin ray 40–47% length of 4th ray, 21–22 total gill rakers, a broad interorbital (56–79% pupil width) with narrow crease-like postorbital groove, ending at posteriormost papilla in row
<emphasis id="84B2EAA7FFC0FF858F26FEB51F2EFEB8" box="[324,338,264,287]" italics="true" pageId="6" pageNumber="477">p</emphasis>
, 8 papillae in row
<emphasis id="84B2EAA7FFC0FF858C54FEB51C38FEB8" box="[566,580,264,287]" italics="true" pageId="6" pageNumber="477">p</emphasis>
(with 2 papillae ventral to papilla row
<emphasis id="84B2EAA7FFC0FF858A74FEB51A58FEB8" box="[1046,1060,264,287]" italics="true" pageId="6" pageNumber="477">n</emphasis>
), and nasal apparatus small and situated on anterior one-third of snout with posterior nares forming posterodorsal margin of nasal sac. When live or freshly collected, dorsal surface of eye light blue; preserved specimens with melanophores on dorsal surface of snout mostly adjacent to nasal capsules.
</paragraph>
</subSubSection>
<subSubSection id="FEDC653EFFC0FF8B8EA5FE2B1ADEFF5F" lastPageId="8" lastPageNumber="479" pageId="6" pageNumber="477" type="description">
<paragraph id="B67936B5FFC0FF858EA5FE2B1A18F868" blockId="6.[151,1437,151,1999]" pageId="6" pageNumber="477">
<emphasis id="84B2EAA7FFC0FF858EA5FE2B1F25FE08" bold="true" box="[199,345,406,431]" pageId="6" pageNumber="477">Description.</emphasis>
The description is based on the
<typeStatus id="697D8817FFC0FF858CBEFE2B1D42FE08" box="[732,830,406,431]" pageId="6" pageNumber="477" type="holotype">holotype</typeStatus>
and
<specimenCount id="A0C0FD3CFFC0FF858D1EFE2B1A78FE17" box="[892,1028,406,432]" pageId="6" pageNumber="477" type="generic" typeStatus="paratypes">5 paratypes</specimenCount>
. Dorsal fin VI +
<date id="C2781075FFC0FF858AB6FE251A84FE08" box="[1236,1272,407,431]" pageId="6" pageNumber="477">I 8</date>
, second spine elongated, reaching posteriorly when adpressed to between base of 5th ray to 1
<taxonomicName id="71C64D36FFC0FF858D9EFE071A38FE71" box="[1020,1092,442,470]" pageId="6" pageNumber="477" rank="subSpecies" subSpecies="scale">st scale</taxonomicName>
behind last dorsal fin ray (
<figureCitation id="2EFD2A30FFC0FF858B0CFE001EDBFE5A" captionStart="FIGURE 5" captionStartId="8.[151,250,1968,1990]" captionTargetBox="[164,1428,301,1949]" captionTargetId="figure@8.[164,1428,301,1949]" captionTargetPageId="8" captionText="FIGURE 5. Trimma meityae, live; A &amp; B—Cendrawasih Bay; C—freshly collected, Cendrawasih Bay. Photos: A &amp; B Figure by M. V. Erdmann; C by G. R. Allen." httpUri="https://zenodo.org/record/1309935/files/figure.png" pageId="6" pageNumber="477">Fig. 5</figureCitation>
, mean = base of 7th ray,
<typeStatus id="697D8817FFC0FF858FA1FE591C59FE5A" box="[451,549,484,509]" pageId="6" pageNumber="477" type="holotype">holotype</typeStatus>
to base
<emphasis id="84B2EAA7FFC0FF858CEAFE591CBBFE5A" bold="true" box="[648,711,484,509]" pageId="6" pageNumber="477">ray 8</emphasis>
), first ray of second dorsal fin branched (
<emphasis id="84B2EAA7FFC0FF858ACBFE591B46FE5A" bold="true" box="[1193,1338,484,509]" pageId="6" pageNumber="477">unbranched</emphasis>
in two), remaining fin rays branched except for posterior element of last ray, fin reaches posteriorly 46–
<emphasis id="84B2EAA7FFC0FF858A8CFDB51B76FD86" bold="true" box="[1262,1290,520,545]" pageId="6" pageNumber="477">57</emphasis>
–61% (54%) distance between base of last ray and first exposed dorsal procurrent caudal fin ray; anal fin
<date id="C2781075FFC0FF858AB6FD901B6DFDE2" box="[1236,1297,556,581]" pageId="6" pageNumber="477">
I 6–
<emphasis id="84B2EAA7FFC0FF858B61FD911B6DFDE2" bold="true" box="[1283,1297,556,581]" pageId="6" pageNumber="477">8</emphasis>
</date>
(7.7, single specimen with 6 rays, clearly deformed or due to injury), first ray unbranched, fin reaches posteriorly 31–
<emphasis id="84B2EAA7FFC0FF858B5CFDED1B26FDCE" bold="true" box="[1342,1370,592,617]" pageId="6" pageNumber="477">33</emphasis>
–42% (34%) distance between base of last ray and first exposed ventral procurrent caudal fin ray; pectoral fin
<emphasis id="84B2EAA7FFC0FF858B34FDC91B0EFD2A" bold="true" box="[1366,1394,628,653]" pageId="6" pageNumber="477">17</emphasis>
–18 (17.7), all rays unbranched, fin reaching posteriorly to region above urogenital papilla to anal spine; pelvic fin
<date id="C2781075FFC0FF858B14FD241BEBFD16" box="[1398,1431,664,689]" pageId="6" pageNumber="477">I 5</date>
, fifth ray unbranched and 40–
<emphasis id="84B2EAA7FFC0FF858F87FD011C7DFD72" bold="true" box="[485,513,700,725]" pageId="6" pageNumber="477">46</emphasis>
–47% (44%) length of fourth ray, which reaches posteriorly to between bases of anal spine to 2nd anal ray, pelvic rays 1–4 with single sequential branch point; basal membrane forming fold across midline above last pre-pelvic scale; no fraenum. Lateral scales 23; anterior transverse scales 8; posterior transverse scales 7; cheek with two rows cycloid scales, upper row of
<emphasis id="84B2EAA7FFC0FF858D2DFC961D21FCE3" bold="true" box="[847,861,811,836]" pageId="6" pageNumber="477">1</emphasis>
–2 and lower row with 6–
<emphasis id="84B2EAA7FFC0FF858AEFFC961AE7FCE3" bold="true" box="[1165,1179,811,836]" pageId="6" pageNumber="477">7</emphasis>
(means = 1.5 and 6.8 respectively, midline of predorsal with
<date id="C2781075FFC0FF858C31FCF21CDBFCC0" box="[595,679,847,872]" pageId="6" pageNumber="477" value="2010-09-08">
8–
<emphasis id="84B2EAA7FFC0FF858C0DFCF21C01FCCF" bold="true" box="[623,637,847,872]" pageId="6" pageNumber="477">9</emphasis>
–10
</date>
(9.0) scales, anterior 1–4 rows may be cycloid, otherwise ctenoid; anteriormost scales on sides (may be ctenoid or cycloid) and top of nape reaching anteriorly almost to posterior margin of eye; opercle fully scaled above papilla row
<emphasis id="84B2EAA7FFC0FF858C9EFC251D6EFC08" box="[764,786,920,943]" italics="true" pageId="6" pageNumber="477">oi</emphasis>
in 4 horizontal rows, dorsalmost row of
<emphasis id="84B2EAA7FFC0FF858A81FC2A1A8DFC17" bold="true" box="[1251,1265,919,944]" pageId="6" pageNumber="477">4</emphasis>
–5, then 2–
<emphasis id="84B2EAA7FFC0FF858B13FC2A1B03FC17" bold="true" box="[1393,1407,919,944]" pageId="6" pageNumber="477">3</emphasis>
,
<emphasis id="84B2EAA7FFC0FF858BECFC2A1BE0FC17" bold="true" box="[1422,1436,919,944]" pageId="6" pageNumber="477">3</emphasis>
and
<emphasis id="84B2EAA7FFC0FF858EABFC061EABFC73" bold="true" box="[201,215,955,980]" pageId="6" pageNumber="477">1</emphasis>
–2 mostly cycloid scales, although scales in mid-upper region may be ctenoid; 3 vertical rows of cycloid scales on pectoral fin base with
<quantity id="713E9B50FFC0FF858C60FC621C36FC5F" box="[514,586,991,1016]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" metricValueMax="5.08" metricValueMin="2.54" pageId="6" pageNumber="477" unit="in" value="1.5" valueMax="2.0" valueMin="1.0">
<emphasis id="84B2EAA7FFC0FF858C60FC621C6CFC5F" bold="true" box="[514,528,991,1016]" pageId="6" pageNumber="477">1</emphasis>
–2 in
</quantity>
anteriormost row,
<quantity id="713E9B50FFC0FF858D42FC621D14FC5F" box="[800,872,991,1016]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="7.62" metricValueMin="5.08" pageId="6" pageNumber="477" unit="in" value="2.5" valueMax="3.0" valueMin="2.0">
2–
<emphasis id="84B2EAA7FFC0FF858D5EFC621D36FC5F" bold="true" box="[828,842,991,1016]" pageId="6" pageNumber="477">3</emphasis>
in
</quantity>
second row and
<quantity id="713E9B50FFC0FF858A4BFC621A29FC5F" box="[1065,1109,991,1016]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="6" pageNumber="477" unit="in" value="5.0">5 in</quantity>
outer row;
<emphasis id="84B2EAA7FFC0FF858AB8FC621A94FC5F" bold="true" box="[1242,1256,991,1016]" pageId="6" pageNumber="477">7</emphasis>
–8 (7.2) cycloid scales in midline anterior to pelvic fin base; area between pelvic spine and ventral margin of pectoral fin base with cycloid (smaller specimens) or ctenoid (larger specimens) scales; anterior few rows of scales in midline of belly cycloid; circumpeduncular scales 12, scales rows in midline between base of last anal ray and first ventral procurrent caudal fin ray
<date id="C2781075FFC0FF858FA5FBD21C60FB20" box="[455,540,1135,1160]" pageId="6" pageNumber="477" value="2010-09-08">
8–
<emphasis id="84B2EAA7FFC0FF858F81FBD21F8DFB2F" bold="true" box="[483,497,1135,1160]" pageId="6" pageNumber="477">9</emphasis>
–10
</date>
(9.2). Upper jaw with outer row of spaced, enlarged curved canines which decrease slightly in height and reach posteriorly 4/5ths of length of premaxilla, two irregular inner rows of small conical teeth (about half height of outer teeth) becoming reduced to single row at bend of premaxilla and continuing posteriorly to 3/5ths length of premaxilla. Lower jaw with short row of about 5 enlarged, spaced, curved canines from symphysis almost to bend of dentary, about 2 irregular rows of slightly curved smaller (2/3rds height of outer teeth) teeth at symphysis, grading to single row, decreasing slightly in size posteriorly, and reaching almost to tip of coronoid process of dentary. Tongue broadly rounded to spatulate with small central tip. Gill opening extending anteroventrally to below mid to posterior pupil; gill rakers
<emphasis id="84B2EAA7FFC0FF858DD7FACE1DBFFA2B" bold="true" box="[949,963,1395,1420]" pageId="6" pageNumber="477">4</emphasis>
–5 + 16–
<emphasis id="84B2EAA7FFC0FF858A49FACE1A3BFA2B" bold="true" box="[1067,1095,1395,1420]" pageId="6" pageNumber="477">18</emphasis>
= 21–
<emphasis id="84B2EAA7FFC0FF858AF6FACE1ACCFA2B" bold="true" box="[1172,1200,1395,1420]" pageId="6" pageNumber="477">22</emphasis>
(4.7 + 17.0 = 21.7). Nasal apparatus small, situated on anterior one-third of snout, anterior naris short tapering tube reaching anteriorly to above anterior margin of upper lip, posterior opening a large, pore-like opening with slightly raised rim covering posterodorsal width of nasal sac, transverse width of pore about 65% length of nasal capsule, posterior margin of posterior naris separated from bony front of orbit by
<emphasis id="84B2EAA7FFC0FF858C86F9BE1D7BF9BB" bold="true" box="[740,775,1539,1564]" pageId="6" pageNumber="477">3.5</emphasis>
–4.0 times its transverse width (mean = 3.9), nasal sac only very slightly raised above surrounding area of snout (
<figureCitation id="2EFD2A30FFC0FF858C9CF99A1D38F9E7" box="[766,836,1575,1600]" captionStart="FIGURE 6" captionStartId="9.[151,250,1139,1161]" captionTargetBox="[151,1436,228,1085]" captionTargetId="figure@9.[151,1436,228,1085]" captionTargetPageId="9" captionText="FIGURE 6. Trimma meityae, 17.3 mm SL male paratype, ROM 106364, Cendrawasih Bay, lateral (and slightly dorsal) view of left nasal apparatus, specimen stained with cyanine blue. Photo by R. Winterbottom." httpUri="https://zenodo.org/record/1309937/files/figure.png" pageId="6" pageNumber="477">Fig. 6</figureCitation>
). Bony interorbital width 56–
<emphasis id="84B2EAA7FFC0FF858AF8F99A1ACAF9E7" bold="true" box="[1178,1206,1575,1600]" pageId="6" pageNumber="477">70</emphasis>
–79.0% (65.4) pupil diameter; profile of snout gently convex, with shallow concave depression between eyes (between 4th and 5th papillae of row
<emphasis id="84B2EAA7FFC0FF858F29F9CE1F25F92D" box="[331,345,1651,1674]" italics="true" pageId="6" pageNumber="477">p</emphasis>
); epaxialis reaching anteriorly in midline to vertical above posterior margin of pupil; no narrow ridge of skin in midline of nape extending anteriorly from origin of first dorsal fin. Caudal peduncle depth as percentage caudal peduncle length 34.4–
<emphasis id="84B2EAA7FFC0FF858C16F9071CD9F974" bold="true" box="[628,677,1722,1747]" pageId="6" pageNumber="477">42.2</emphasis>
(38.4); head length as percentage SL 32.9–
<emphasis id="84B2EAA7FFC0FF858AD3F9071A9EF974" bold="true" box="[1201,1250,1722,1747]" pageId="6" pageNumber="477">33.2</emphasis>
–35.2 (33.7); as percentage head length: horizontal eye diameter
<emphasis id="84B2EAA7FFC0FF858CD7F9631C9AF950" bold="true" box="[693,742,1758,1783]" pageId="6" pageNumber="477">34.6</emphasis>
–38.9 (36.9); snout length 21.2–
<emphasis id="84B2EAA7FFC0FF858A2FF9631A02F950" bold="true" box="[1101,1150,1758,1783]" pageId="6" pageNumber="477">22.1</emphasis>
–22.8 (22.2); cheek depth 16.8–
<emphasis id="84B2EAA7FFC0FF858EB4F8BF1F7BF8BC" bold="true" box="[214,263,1794,1819]" pageId="6" pageNumber="477">20.9</emphasis>
–21.2 (19.1). Cephalic sensory papillae as in
<figureCitation id="2EFD2A30FFC0FF858D69F8BF1D2DF8BD" box="[779,849,1794,1819]" captionStart="FIGURE 7" captionStartId="10.[151,250,1896,1918]" captionTargetBox="[151,1436,255,1819]" captionTargetId="figure@10.[151,1436,255,1819]" captionTargetPageId="10" captionText="FIGURE 7. Trimma meityae, 18.6 male holotype, ROM 106350. Left lateral (A) and dorsal (B) views of head papillae, specimen stained with cyanine blue. Papillae in a given row connected by thin green lines, red arrow points to supernumerary papilla. Second letters of abbreviations in ea, ep and ip stand for anterior and posterior respectively. Photos by R. Winterbottom." httpUri="https://zenodo.org/record/1309939/files/figure.png" pageId="6" pageNumber="477">Fig. 7</figureCitation>
. Number of papillae in each row:
<emphasis id="84B2EAA7FFC0FF858AB8F8BE1A94F8BD" box="[1242,1256,1795,1818]" italics="true" pageId="6" pageNumber="477">a</emphasis>
= 6;
<emphasis id="84B2EAA7FFC0FF858B45F8BF1B49F8BC" box="[1319,1333,1794,1819]" italics="true" pageId="6" pageNumber="477">b</emphasis>
= 4–
<emphasis id="84B2EAA7FFC0FF858B10F8BF1BFCF8BC" bold="true" box="[1394,1408,1794,1819]" pageId="6" pageNumber="477">5</emphasis>
–6 (5.0);
<emphasis id="84B2EAA7FFC0FF858EBEF89A1E94F899" box="[220,232,1831,1854]" italics="true" pageId="6" pageNumber="477">c</emphasis>
= 6;
<emphasis id="84B2EAA7FFC0FF858F46F89A1F43F899" box="[292,319,1831,1854]" italics="true" pageId="6" pageNumber="477">cp</emphasis>
= 1;
<emphasis id="84B2EAA7FFC0FF858F19F89B1FF5F898" box="[379,393,1830,1855]" italics="true" pageId="6" pageNumber="477">d</emphasis>
= 6–
<emphasis id="84B2EAA7FFC0FF858FA6F89B1FAEF898" bold="true" box="[452,466,1830,1855]" pageId="6" pageNumber="477">7</emphasis>
(6.5);
<emphasis id="84B2EAA7FFC0FF858C7CF89B1C4EF898" box="[542,562,1830,1855]" italics="true" pageId="6" pageNumber="477">dʹ</emphasis>
= 6–
<emphasis id="84B2EAA7FFC0FF858C0EF89B1C06F898" bold="true" box="[620,634,1830,1855]" pageId="6" pageNumber="477">7</emphasis>
–8 (7.3);
<emphasis id="84B2EAA7FFC0FF858C81F89A1D28F899" box="[739,852,1831,1854]" italics="true" pageId="6" pageNumber="477">e-anterior</emphasis>
=
<date id="C2781075FFC0FF858D16F89A1D98F898" box="[884,996,1830,1855]" pageId="6" pageNumber="477" value="2013-12-14">
12–
<emphasis id="84B2EAA7FFC0FF858DFFF89B1DC5F898" bold="true" box="[925,953,1830,1855]" pageId="6" pageNumber="477">13</emphasis>
–14
</date>
(12.8);
<emphasis id="84B2EAA7FFC0FF858A5CF89A1AC6F899" box="[1086,1210,1831,1854]" italics="true" pageId="6" pageNumber="477">e-posterior</emphasis>
=
<date id="C2781075FFC0FF858ABBF89A1B35F898" box="[1241,1353,1830,1855]" pageId="6" pageNumber="477" value="2015-12-17">
12–
<emphasis id="84B2EAA7FFC0FF858B61F89B1B63F898" bold="true" box="[1283,1311,1830,1855]" pageId="6" pageNumber="477">15</emphasis>
–17
</date>
(14.5);
<emphasis id="84B2EAA7FFC0FF858EF5F8F61F7FF8C5" box="[151,259,1867,1890]" italics="true" pageId="6" pageNumber="477">i-anterior</emphasis>
=
<emphasis id="84B2EAA7FFC0FF858F40F8F71F4CF8C4" bold="true" box="[290,304,1866,1891]" pageId="6" pageNumber="477">6</emphasis>
–7 (6.7);
<emphasis id="84B2EAA7FFC0FF858FFAF8F61C6CF8C5" box="[408,528,1867,1890]" italics="true" pageId="6" pageNumber="477">i-posterior</emphasis>
= 7;
<emphasis id="84B2EAA7FFC0FF858C2EF8F61C26F8C5" box="[588,602,1867,1890]" italics="true" pageId="6" pageNumber="477">p</emphasis>
= 8, with 2 papillae just medial to posterior naris and 2 papillae ventral to row
<emphasis id="84B2EAA7FFC0FF858EA8F8D21EA4F821" box="[202,216,1903,1926]" italics="true" pageId="6" pageNumber="477">n</emphasis>
;
<emphasis id="84B2EAA7FFC0FF858E8AF8D21E8FF821" box="[232,243,1903,1926]" italics="true" pageId="6" pageNumber="477">r</emphasis>
= 2;
<emphasis id="84B2EAA7FFC0FF858F52F8D31F44F821" box="[304,312,1902,1926]" italics="true" pageId="6" pageNumber="477">f</emphasis>
=
<emphasis id="84B2EAA7FFC0FF858F35F8D31F19F820" bold="true" box="[343,357,1902,1927]" pageId="6" pageNumber="477">3</emphasis>
–4 (3.2);
<emphasis id="84B2EAA7FFC0FF858FADF8D21F8EF821" box="[463,498,1903,1926]" italics="true" pageId="6" pageNumber="477">cs&quot;</emphasis>
= 3;
<emphasis id="84B2EAA7FFC0FF858C4DF8D21C41F821" box="[559,573,1903,1926]" italics="true" pageId="6" pageNumber="477">g</emphasis>
= apparently absent, but may be represented by zig-zag line of 7 papillae in one specimen;
<emphasis id="84B2EAA7FFC0FF858F22F82E1F32F80D" box="[320,334,1939,1962]" italics="true" pageId="6" pageNumber="477">n</emphasis>
= 1;
<emphasis id="84B2EAA7FFC0FF858FE8F82E1FEAF80D" box="[394,406,1939,1962]" italics="true" pageId="6" pageNumber="477">x</emphasis>
= 6–
<emphasis id="84B2EAA7FFC0FF858FB3F82F1FA3F80C" bold="true" box="[465,479,1938,1963]" pageId="6" pageNumber="477">7</emphasis>
–8 (7.2, n = 5);
<emphasis id="84B2EAA7FFC0FF858CF3F82E1CE3F80D" box="[657,671,1939,1962]" italics="true" pageId="6" pageNumber="477">u</emphasis>
= 4–
<emphasis id="84B2EAA7FFC0FF858CBBF82F1C9BF80C" bold="true" box="[729,743,1938,1963]" pageId="6" pageNumber="477">5</emphasis>
(4.3);
<emphasis id="84B2EAA7FFC0FF858D56F82E1D43F80D" box="[820,831,1939,1962]" italics="true" pageId="6" pageNumber="477">z</emphasis>
= 5–
<emphasis id="84B2EAA7FFC0FF858D1BF82F1DFBF80C" bold="true" box="[889,903,1938,1963]" pageId="6" pageNumber="477">7</emphasis>
(6.3, n = 4);
<emphasis id="84B2EAA7FFC0FF858A7FF82E1A4FF80D" box="[1053,1075,1939,1962]" italics="true" pageId="6" pageNumber="477">ot</emphasis>
= 11–
<emphasis id="84B2EAA7FFC0FF858A19F82F1AEBF80C" bold="true" box="[1147,1175,1938,1963]" pageId="6" pageNumber="477">13</emphasis>
(12.3, n = 3);
<emphasis id="84B2EAA7FFC0FF858B58F82E1B2FF80D" box="[1338,1363,1939,1962]" italics="true" pageId="6" pageNumber="477">os</emphasis>
= 7–
<emphasis id="84B2EAA7FFC0FF858BECF82F1BE0F80C" bold="true" box="[1422,1436,1938,1963]" pageId="6" pageNumber="477">8</emphasis>
(7.7, n = 3);
<emphasis id="84B2EAA7FFC0FF858F41F80A1F45F869" box="[291,313,1975,1998]" italics="true" pageId="6" pageNumber="477">oi</emphasis>
= 4–
<emphasis id="84B2EAA7FFC0FF858F11F80B1FFDF868" bold="true" box="[371,385,1974,1999]" pageId="6" pageNumber="477">7</emphasis>
(5.5, n = 4). Abdominal/caudal vertebral transition not examined.
</paragraph>
<caption id="E2B9663DFFC1FF848EF5FA0A1D76FA4B" httpUri="https://zenodo.org/record/1309933/files/figure.png" pageId="7" pageNumber="478" startId="7.[151,250,1463,1485]" targetBox="[151,1433,217,1416]" targetPageId="7">
<paragraph id="B67936B5FFC1FF848EF5FA0A1D76FA4B" blockId="7.[151,1436,1463,1516]" pageId="7" pageNumber="478">
<emphasis id="84B2EAA7FFC1FF848EF5FA0A1F6FFA6B" bold="true" box="[151,275,1463,1485]" pageId="7" pageNumber="478">FIGURE 4.</emphasis>
Neighbour Joining network derived from a BOL analysis of the COI gene of the
<taxonomicName id="71C64D36FFC1FF848A38FA051AD5FA6A" authorityName="Jordan &amp; Seale" authorityYear="1906" box="[1114,1193,1464,1485]" class="Actinopterygii" family="Gobiidae" genus="Trimma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="478" phylum="Chordata" rank="genus">
<emphasis id="84B2EAA7FFC1FF848A38FA051AD5FA6A" box="[1114,1193,1464,1485]" italics="true" pageId="7" pageNumber="478">Trimma</emphasis>
</taxonomicName>
haplogroups associated with
<taxonomicName id="71C64D36FFC1FF848EA8FA6A1F32FA4B" authorityName="Winterbottom &amp; Erdmann" authorityYear="2018" box="[202,334,1494,1516]" class="Actinopterygii" family="Gobiidae" genus="Trimma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="478" phylum="Chordata" rank="species" species="blematium">
<emphasis id="84B2EAA7FFC1FF848EA8FA6A1F32FA4B" box="[202,334,1494,1516]" italics="true" pageId="7" pageNumber="478">T. blematium</emphasis>
</taxonomicName>
and
<taxonomicName id="71C64D36FFC1FF848F1DFA6A1F9AFA4B" authorityName="Winterbottom &amp; Erdmann" authorityYear="2018" box="[383,486,1495,1516]" class="Actinopterygii" family="Gobiidae" genus="Trimma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="478" phylum="Chordata" rank="species" species="meityae">
<emphasis id="84B2EAA7FFC1FF848F1DFA6A1F9AFA4B" box="[383,486,1495,1516]" italics="true" pageId="7" pageNumber="478">T. meityae</emphasis>
</taxonomicName>
. Image by R. Winterbottom.
</paragraph>
</caption>
<paragraph id="B67936B5FFC1FF8B8EA5F9A51ADEFF5F" blockId="7.[151,1437,1560,2017]" lastBlockId="8.[151,1436,151,248]" lastPageId="8" lastPageNumber="479" pageId="7" pageNumber="478">
<emphasis id="84B2EAA7FFC1FF848EA5F9A51FD2F997" box="[199,430,1560,1585]" italics="true" pageId="7" pageNumber="478">Colour pattern. Live</emphasis>
, based on 12 images from Cendrawasih Bay,
<collectingRegion id="7402F857FFC1FF848DDBF9A41A43F997" box="[953,1087,1561,1584]" country="Indonesia" name="Papua Barat" pageId="7" pageNumber="478">West Papua</collectingRegion>
,
<collectingCountry id="CED17625FFC1FF848A2EF9A51AC1F996" box="[1100,1213,1560,1585]" name="Indonesia" pageId="7" pageNumber="478">Indonesia</collectingCountry>
. Specimens (
<figureCitation id="2EFD2A30FFC1FF848B30F9A51BEBF996" box="[1362,1431,1560,1585]" captionStart="FIGURE 5" captionStartId="8.[151,250,1968,1990]" captionTargetBox="[164,1428,301,1949]" captionTargetId="figure@8.[164,1428,301,1949]" captionTargetPageId="8" captionText="FIGURE 5. Trimma meityae, live; A &amp; B—Cendrawasih Bay; C—freshly collected, Cendrawasih Bay. Photos: A &amp; B Figure by M. V. Erdmann; C by G. R. Allen." httpUri="https://zenodo.org/record/1309935/files/figure.png" pageId="7" pageNumber="478">Fig. 5</figureCitation>
, A, B) almost entirely translucent light pink or off-white with yellow spinal cord, much lighter and brighter blue over dorsomedial surface of eye than
<taxonomicName id="71C64D36FFC1FF848C59F9DC1CB0F9DE" authorityName="Winterbottom &amp; Erdmann" authorityYear="2018" box="[571,716,1632,1657]" class="Actinopterygii" family="Gobiidae" genus="Trimma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="478" phylum="Chordata" rank="species" species="blematium">
<emphasis id="84B2EAA7FFC1FF848C59F9DC1CB0F9DE" box="[571,716,1632,1657]" italics="true" pageId="7" pageNumber="478">T. blematium</emphasis>
</taxonomicName>
, brown iris ring relatively indistinct and white inner ring around pupil wider and about half iris width. Fin rays pinkish, no apparent pigmentation in fin membranes. Other photographed specimens from this locality with darker pink background and blue over eyes less well developed.
<emphasis id="84B2EAA7FFC1FF848EF5F9711F2BF942" box="[151,343,1740,1765]" italics="true" pageId="7" pageNumber="478">Freshly collected</emphasis>
, (
<figureCitation id="2EFD2A30FFC1FF848F0FF9711FB7F942" box="[365,459,1740,1765]" captionStart="FIGURE 5" captionStartId="8.[151,250,1968,1990]" captionTargetBox="[164,1428,301,1949]" captionTargetId="figure@8.[164,1428,301,1949]" captionTargetPageId="8" captionText="FIGURE 5. Trimma meityae, live; A &amp; B—Cendrawasih Bay; C—freshly collected, Cendrawasih Bay. Photos: A &amp; B Figure by M. V. Erdmann; C by G. R. Allen." httpUri="https://zenodo.org/record/1309935/files/figure.png" pageId="7" pageNumber="478">Fig. 5 C</figureCitation>
). Upper and lower midlines reddish, body semi-translucent greenish with scale margins diffusely outlined with dark red, posterior region of cranium dull yellow, snout reddish, distal half of upper lip and region of snout below anterior margin of eye orange-red. Dorsal fins with basal stripe of whitish chromatophores, stripe half pupil-width anteriorly to full pupil-width posteriorly, fin elements pink above this; caudal with bluepurple margins and 3 equidistant similar streaks in body of fin, the middle medial in position, rest of fin greenishyellow; membranes of pelvic fin with some iridocytes, rays off-white, membranes of pectoral fin hyaline with rays light pink.
<emphasis id="84B2EAA7FFC1FF848F77F8191FF9F81A" box="[277,389,1956,1981]" italics="true" pageId="7" pageNumber="478">Preserved</emphasis>
.
<typeStatus id="697D8817FFC1FF848FF4F8191F82F81A" box="[406,510,1956,1981]" pageId="7" pageNumber="478" type="holotype">Holotype</typeStatus>
pale off-white with scale margins in predorsal region and dorsal-most scale row along entire dorsum outlined with brown melanophores; a thin line of melanophores in midline from end of anal fin base almost to first ventral procurrent ray, a few melanophores medial to posterior nares (
<figureCitation id="2EFD2A30FFCEFF8B8A1CFF2A1AA0FF17" box="[1150,1244,151,176]" captionStart="FIGURE 7" captionStartId="10.[151,250,1896,1918]" captionTargetBox="[151,1436,255,1819]" captionTargetId="figure@10.[151,1436,255,1819]" captionTargetPageId="10" captionText="FIGURE 7. Trimma meityae, 18.6 male holotype, ROM 106350. Left lateral (A) and dorsal (B) views of head papillae, specimen stained with cyanine blue. Papillae in a given row connected by thin green lines, red arrow points to supernumerary papilla. Second letters of abbreviations in ea, ep and ip stand for anterior and posterior respectively. Photos by R. Winterbottom." httpUri="https://zenodo.org/record/1309939/files/figure.png" pageId="8" pageNumber="479">Fig. 7 B</figureCitation>
), on snout below anteroventral margin of eye, and from upper operculum anterior to posterior margin of eye (
<figureCitation id="2EFD2A30FFCEFF8B8AD7FF011B66FF73" box="[1205,1306,188,213]" captionStart="FIGURE 7" captionStartId="10.[151,250,1896,1918]" captionTargetBox="[151,1436,255,1819]" captionTargetId="figure@10.[151,1436,255,1819]" captionTargetPageId="10" captionText="FIGURE 7. Trimma meityae, 18.6 male holotype, ROM 106350. Left lateral (A) and dorsal (B) views of head papillae, specimen stained with cyanine blue. Papillae in a given row connected by thin green lines, red arrow points to supernumerary papilla. Second letters of abbreviations in ea, ep and ip stand for anterior and posterior respectively. Photos by R. Winterbottom." httpUri="https://zenodo.org/record/1309939/files/figure.png" pageId="8" pageNumber="479">Fig. 7 A</figureCitation>
).
<typeStatus id="697D8817FFCEFF8B8B4DFF001BE0FF72" box="[1327,1436,189,213]" pageId="8" pageNumber="479" type="paratype">Paratypes</typeStatus>
similar, but lack melanophores anteroventral to the eye, and may lack those over operculum.
</paragraph>
</subSubSection>
<caption id="E2B9663DFFCEFF8B8EF5F80D1C73F843" httpUri="https://zenodo.org/record/1309935/files/figure.png" pageId="8" pageNumber="479" startId="8.[151,250,1968,1990]" targetBox="[164,1428,301,1949]" targetPageId="8">
<paragraph id="B67936B5FFCEFF8B8EF5F80D1C73F843" blockId="8.[151,1436,1968,2020]" pageId="8" pageNumber="479">
<emphasis id="84B2EAA7FFCEFF8B8EF5F80D1F6FF862" bold="true" box="[151,275,1968,1990]" pageId="8" pageNumber="479">FIGURE 5.</emphasis>
<taxonomicName id="71C64D36FFCEFF8B8F78F80C1FC2F861" authorityName="Winterbottom &amp; Erdmann" authorityYear="2018" box="[282,446,1969,1990]" class="Actinopterygii" family="Gobiidae" genus="Trimma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="479" phylum="Chordata" rank="species" species="meityae">
<emphasis id="84B2EAA7FFCEFF8B8F78F80C1FC2F861" box="[282,446,1969,1990]" italics="true" pageId="8" pageNumber="479">Trimma meityae</emphasis>
</taxonomicName>
, live; A &amp; B—Cendrawasih Bay; C—freshly collected, Cendrawasih Bay. Photos: A &amp; B Figure by M. V. Erdmann; C by G. R. Allen.
</paragraph>
</caption>
<caption id="E2B9663DFFCFFF8A8EF5FBCE1D92FB00" httpUri="https://zenodo.org/record/1309937/files/figure.png" pageId="9" pageNumber="480" startId="9.[151,250,1139,1161]" targetBox="[151,1436,228,1085]" targetPageId="9">
<paragraph id="B67936B5FFCFFF8A8EF5FBCE1D92FB00" blockId="9.[151,1435,1139,1191]" pageId="9" pageNumber="480">
<emphasis id="84B2EAA7FFCFFF8A8EF5FBCE1F6EFB2F" bold="true" box="[151,274,1139,1161]" pageId="9" pageNumber="480">FIGURE 6.</emphasis>
<taxonomicName id="71C64D36FFCFFF8A8F7AFBC91FC7FB2E" authorityName="Winterbottom &amp; Erdmann" authorityYear="2018" box="[280,443,1140,1161]" class="Actinopterygii" family="Gobiidae" genus="Trimma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="480" phylum="Chordata" rank="species" species="meityae">
<emphasis id="84B2EAA7FFCFFF8A8F7AFBC91FC7FB2E" box="[280,443,1140,1161]" italics="true" pageId="9" pageNumber="480">Trimma meityae</emphasis>
</taxonomicName>
, 17.3 mm SL male paratype, ROM 106364, Cendrawasih Bay, lateral (and slightly dorsal) view of left nasal apparatus, specimen stained with cyanine blue. Photo by R. Winterbottom.
</paragraph>
</caption>
<subSubSection id="FEDC653EFFCFFF8A8EA5FB691F2DFAFE" pageId="9" pageNumber="480" type="etymology">
<paragraph id="B67936B5FFCFFF8A8EA5FB691F2DFAFE" blockId="9.[151,1437,1236,2025]" pageId="9" pageNumber="480">
<emphasis id="84B2EAA7FFCFFF8A8EA5FB691F32FB4A" bold="true" box="[199,334,1236,1261]" pageId="9" pageNumber="480">Etymology.</emphasis>
The species is name “
<taxonomicName id="71C64D36FFCFFF8A8C21FB691CDCFB4A" authorityName="Winterbottom &amp; Erdmann" authorityYear="2018" box="[579,672,1236,1261]" class="Actinopterygii" family="Gobiidae" genus="Trimma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="480" phylum="Chordata" rank="species" species="meityae">meityae</taxonomicName>
” in honour of Meity Mongdong, one of Indonesia’s foremost marine conservationists, who has dedicated the past several decades of her career towards expanding and improving the management of marine protected areas in
<collectingRegion id="7402F857FFCFFF8A8C18FAA01D7EFA93" box="[634,770,1309,1332]" country="Indonesia" name="Papua Barat" pageId="9" pageNumber="480">West Papua</collectingRegion>
, including the Cendrawasih Bay National Park where this species is found.
</paragraph>
</subSubSection>
<subSubSection id="FEDC653EFFCFFF8A8EA5FAD91D1FF862" pageId="9" pageNumber="480" type="distribution">
<paragraph id="B67936B5FFCFFF8A8EA5FAD91DAEFA62" blockId="9.[151,1437,1236,2025]" pageId="9" pageNumber="480">
<emphasis id="84B2EAA7FFCFFF8A8EA5FAD91F88FADA" bold="true" box="[199,500,1380,1405]" pageId="9" pageNumber="480">Distribution and habitat.</emphasis>
This species is currently known only from Cendrawasih Bay (Pulau Purup),
<collectingRegion id="7402F857FFCFFF8A8B07FAD81EA2FA07" country="Indonesia" name="Papua Barat" pageId="9" pageNumber="480">West Papua</collectingRegion>
,
<collectingCountry id="CED17625FFCFFF8A8E8CFA351F23FA06" box="[238,351,1416,1441]" name="Indonesia" pageId="9" pageNumber="480">Indonesia</collectingCountry>
. It was observed and collected from a nearshore reef with almost no exposure to waves or currents, in the
<quantity id="713E9B50FFCFFF8A8F27FA111FDFFA63" box="[325,419,1452,1477]" metricMagnitude="1" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="9" pageNumber="480" unit="m" value="55.0" valueMax="60.0" valueMin="50.0">50-60 m</quantity>
depth zone only, on a silty sand and rubble slope.
</paragraph>
<paragraph id="B67936B5FFCFFF8A8EA5FA6D1D1FF862" blockId="9.[151,1437,1236,2025]" pageId="9" pageNumber="480">
<emphasis id="84B2EAA7FFCFFF8A8EA5FA6D1F11FA4E" bold="true" box="[199,365,1488,1513]" pageId="9" pageNumber="480">Comparisons.</emphasis>
There are 9 other described species of
<taxonomicName id="71C64D36FFCFFF8A8D52FA6C1DFBFA4F" authorityName="Jordan &amp; Seale" authorityYear="1906" box="[816,903,1489,1512]" class="Actinopterygii" family="Gobiidae" genus="Trimma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="480" phylum="Chordata" rank="genus">
<emphasis id="84B2EAA7FFCFFF8A8D52FA6C1DFBFA4F" box="[816,903,1489,1512]" italics="true" pageId="9" pageNumber="480">Trimma</emphasis>
</taxonomicName>
which possess predorsal scales in the midline, scales on the cheek, unbranched pectoral fin rays, an unbranched 5th pelvic fin ray, and a bony interorbital width that is ~50% or more of the pupil width. These are:
<taxonomicName id="71C64D36FFCFFF8A8C82F9A11D96F993" authority="Allen, 2015" authorityName="Allen" authorityYear="2015" box="[736,1002,1563,1588]" class="Actinopterygii" family="Gobiidae" genus="Trimma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="480" phylum="Chordata" rank="species" species="abyssum">
<emphasis id="84B2EAA7FFCFFF8A8C82F9A11C91F994" box="[736,749,1564,1587]" italics="true" pageId="9" pageNumber="480">T</emphasis>
.
<emphasis id="84B2EAA7FFCFFF8A8C9EF9A61D20F993" box="[764,860,1563,1588]" italics="true" pageId="9" pageNumber="480">abyssum</emphasis>
<bibRefCitation id="D2574B44FFCFFF8A8D01F9A61D96F993" box="[867,1002,1563,1588]" pageId="9" pageNumber="480" refId="ref6320" refString="Allen, G. R. (2015) Four new species of Trimma (Pisces: Gobiidae) from Papua New Guinea and Indonesia. Journal of the Ocean Science Foundation, 14, 28 - 48. https: // doi. org / 10.5281 / zenodo. 1037603" type="journal article">Allen, 2015</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="71C64D36FFCFFF8A8D95F9A11B00F993" authority="Winterbottom, 2016" authorityName="Winterbottom" authorityYear="2016" box="[1015,1404,1563,1588]" class="Actinopterygii" family="Gobiidae" genus="Trimma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="480" phylum="Chordata" rank="species" species="burridgeae">
<emphasis id="84B2EAA7FFCFFF8A8D95F9A11A78F994" box="[1015,1028,1564,1587]" italics="true" pageId="9" pageNumber="480">T</emphasis>
.
<emphasis id="84B2EAA7FFCFFF8A8A71F9A61AECF993" box="[1043,1168,1563,1588]" italics="true" pageId="9" pageNumber="480">burridgeae</emphasis>
<bibRefCitation id="D2574B44FFCFFF8A8AF4F9A61B00F993" box="[1174,1404,1563,1588]" pageId="9" pageNumber="480" refId="ref6699" refString="Winterbottom, R. (2016) Trimma tevegae and T. caudomaculatum revisited and redescribed (Acanthopterygii, Gobiidae), with descriptions of three new similar species from the western Pacific. Zootaxa, 4144 (1), 1 - 53. https: // doi. org / 10.11646 / zootaxa. 4144.1.1" type="journal article">Winterbottom, 2016</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="71C64D36FFCFFF8A8BE5F9A11C0BF9FF" authority="Yoshino &amp; Araga, 1975" authorityName="Yoshino &amp; Araga" authorityYear="1975" class="Actinopterygii" family="Gobiidae" genus="Trimma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="480" phylum="Chordata" rank="species" species="caudomaculatum">
<emphasis id="84B2EAA7FFCFFF8A8BE5F9A11BE8F994" box="[1415,1428,1564,1587]" italics="true" pageId="9" pageNumber="480">T</emphasis>
.
<emphasis id="84B2EAA7FFCFFF8A8EF5F9821F24F9FF" box="[151,344,1599,1624]" italics="true" pageId="9" pageNumber="480">caudomaculatum</emphasis>
<bibRefCitation id="D2574B44FFCFFF8A8F03F9821C0BF9FF" box="[353,631,1599,1624]" pageId="9" pageNumber="480" refId="ref7105" refString="Yoshino, T. &amp; Araga, C. (1975) Trimma caudomaculata Yoshino and Araga. In: Masuda, H., Araga, C. &amp; Yoshino, T. (Eds.), Coastal Fishes of Southern Japan. Tokai University Press, Tokyo, pp. 272." type="book chapter">Yoshino &amp; Araga, 1975</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="71C64D36FFCFFF8A8CE7F9FD1A7AF9FF" authority="Winterbottom, 2016" authorityName="Winterbottom" authorityYear="2016" box="[645,1030,1599,1624]" class="Actinopterygii" family="Gobiidae" genus="Trimma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="480" phylum="Chordata" rank="species" species="corerefum">
<emphasis id="84B2EAA7FFCFFF8A8CE7F9FD1CEEF9F0" box="[645,658,1600,1623]" italics="true" pageId="9" pageNumber="480">T</emphasis>
.
<emphasis id="84B2EAA7FFCFFF8A8CC1F9821D6FF9F0" box="[675,787,1599,1623]" italics="true" pageId="9" pageNumber="480">corerefum</emphasis>
<bibRefCitation id="D2574B44FFCFFF8A8D7CF9821A7AF9FF" box="[798,1030,1599,1624]" pageId="9" pageNumber="480" refId="ref6699" refString="Winterbottom, R. (2016) Trimma tevegae and T. caudomaculatum revisited and redescribed (Acanthopterygii, Gobiidae), with descriptions of three new similar species from the western Pacific. Zootaxa, 4144 (1), 1 - 53. https: // doi. org / 10.11646 / zootaxa. 4144.1.1" type="journal article">Winterbottom, 2016</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="71C64D36FFCFFF8A8A76F9FD1B05F9F0" authority="Winterbottom, 2011" authorityName="Winterbottom" authorityYear="2011" box="[1044,1401,1599,1624]" class="Actinopterygii" family="Gobiidae" genus="Trimma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="480" phylum="Chordata" rank="species" species="habrum">
<emphasis id="84B2EAA7FFCFFF8A8A76F9FD1A5DF9F0" box="[1044,1057,1600,1623]" italics="true" pageId="9" pageNumber="480">T</emphasis>
.
<emphasis id="84B2EAA7FFCFFF8A8A50F9821AF5F9FF" box="[1074,1161,1599,1624]" italics="true" pageId="9" pageNumber="480">habrum</emphasis>
<bibRefCitation id="D2574B44FFCFFF8A8AF1F9821B05F9F0" box="[1171,1401,1599,1624]" pageId="9" pageNumber="480" refId="ref6656" refString="Winterbottom, R. (2011) Six new species of the genus Trimma (Percomorpha; Gobiidae) from the Raja Ampat Islands, Indonesia. aqua, International Journal of Ichthyology, 17 (3), 127 - 162." type="journal article">Winterbottom, 2011</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="71C64D36FFCFFF8A8BE5F9FD1C51F9DB" authority="Winterbottom et al., 2014 a" authorityName="Winterbottom et al." authorityYear="2014" class="Actinopterygii" family="Gobiidae" genus="Trimma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="480" phylum="Chordata" rank="species" species="helenae">
<emphasis id="84B2EAA7FFCFFF8A8BE5F9FD1BE8F9F0" box="[1415,1428,1600,1623]" italics="true" pageId="9" pageNumber="480">T</emphasis>
.
<emphasis id="84B2EAA7FFCFFF8A8EF5F9DE1E92F9DB" box="[151,238,1635,1660]" italics="true" pageId="9" pageNumber="480">helenae</emphasis>
<bibRefCitation id="D2574B44FFCFFF8A8E95F9DE1C51F9DB" box="[247,557,1635,1660]" pageId="9" pageNumber="480" refId="ref6756" refString="Winterbottom, R., Erdmann, M. V. &amp; Cahyani, N. K. D. (2014 a). Trimma helenae (Pisces; Gobioidei), a new species of gobiid fish from Indonesia. Zootaxa, 3760 (3), 420 - 428. https: // doi. org / 10.11646 / zootaxa. 3760.3.8" type="journal article">
Winterbottom
<emphasis id="84B2EAA7FFCFFF8A8FFEF9D91FB3F9DB" box="[412,463,1635,1660]" italics="true" pageId="9" pageNumber="480">et al</emphasis>
., 2014a
</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="71C64D36FFCFFF8A8C58F9D91DC4F9DB" authority="Winterbottom, 2016" authorityName="Winterbottom" authorityYear="2016" box="[570,952,1635,1660]" class="Actinopterygii" family="Gobiidae" genus="Trimma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="480" phylum="Chordata" rank="species" species="hollemani">
<emphasis id="84B2EAA7FFCFFF8A8C58F9D91C3BF9DC" box="[570,583,1636,1659]" italics="true" pageId="9" pageNumber="480">T</emphasis>
.
<emphasis id="84B2EAA7FFCFFF8A8C3AF9DE1CB4F9DB" box="[600,712,1635,1660]" italics="true" pageId="9" pageNumber="480">hollemani</emphasis>
<bibRefCitation id="D2574B44FFCFFF8A8CB3F9DE1DC4F9DB" box="[721,952,1635,1660]" pageId="9" pageNumber="480" refId="ref6699" refString="Winterbottom, R. (2016) Trimma tevegae and T. caudomaculatum revisited and redescribed (Acanthopterygii, Gobiidae), with descriptions of three new similar species from the western Pacific. Zootaxa, 4144 (1), 1 - 53. https: // doi. org / 10.11646 / zootaxa. 4144.1.1" type="journal article">Winterbottom, 2016</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="71C64D36FFCFFF8A8DA7F9D91BE0F9DB" authority="Winterbottom &amp; Hoese, 2015" authorityName="Winterbottom &amp; Hoese" authorityYear="2015" box="[965,1436,1635,1660]" class="Actinopterygii" family="Gobiidae" genus="Trimma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="480" phylum="Chordata" rank="species" species="kitrinum">
<emphasis id="84B2EAA7FFCFFF8A8DA7F9D91DAEF9DC" box="[965,978,1636,1659]" italics="true" pageId="9" pageNumber="480">T</emphasis>
.
<emphasis id="84B2EAA7FFCFFF8A8D80F9DE1A3DF9DB" box="[994,1089,1635,1660]" italics="true" pageId="9" pageNumber="480">kitrinum</emphasis>
<bibRefCitation id="D2574B44FFCFFF8A8A28F9DE1BE0F9DB" box="[1098,1436,1635,1660]" pageId="9" pageNumber="480" refId="ref6951" refString="Winterbottom, R. &amp; Hoese, D. F. (2015) A revision of the Australian species of Trimma (Actinopterygii, Gobiidae), with descriptions of six new species and redescriptions of twenty - three valid species. Zootaxa, 3934 (1), 1 - 102. https: // doi. org / 10.11646 / zootaxa. 3934.1.1" type="journal article">Winterbottom &amp; Hoese, 2015</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="71C64D36FFCFFF8A8EA8F9351C25F907" authority="Cohen &amp; Davies, 1969" authorityName="Cohen &amp; Davies" authorityYear="1969" box="[202,601,1671,1696]" class="Actinopterygii" family="Gobiidae" genus="Trimma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="480" phylum="Chordata" rank="species" species="tevegae">
<emphasis id="84B2EAA7FFCFFF8A8EA8F9351EA4F938" box="[202,216,1672,1695]" italics="true" pageId="9" pageNumber="480">T</emphasis>
.
<emphasis id="84B2EAA7FFCFFF8A8E8BF9351F42F938" box="[233,318,1672,1695]" italics="true" pageId="9" pageNumber="480">tevegae</emphasis>
<bibRefCitation id="D2574B44FFCFFF8A8F28F93A1C25F907" box="[330,601,1671,1696]" pageId="9" pageNumber="480" refId="ref6371" refString="Cohen, D. M. &amp; Davis, W. P. (1969) Vertical orientation in a new gobioid fish from New Britain. Pacific Science, 23 (3), 317 - 324." type="journal article">Cohen &amp; Davies, 1969</bibRefCitation>
</taxonomicName>
. None of these species has 17 or more pectoral fin rays as found in
<taxonomicName id="71C64D36FFCFFF8A8BE5F9351E92F964" authorityName="Winterbottom &amp; Erdmann" authorityYear="2018" class="Actinopterygii" family="Gobiidae" genus="Trimma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="480" phylum="Chordata" rank="species" species="meityae">
<emphasis id="84B2EAA7FFCFFF8A8BE5F9351E92F964" italics="true" pageId="9" pageNumber="480">T. meityae</emphasis>
</taxonomicName>
.
<taxonomicName id="71C64D36FFCFFF8A8E9FF9111FC0F963" authorityName="Allen" authorityYear="2015" box="[253,444,1707,1732]" class="Actinopterygii" family="Gobiidae" genus="Trimma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="480" phylum="Chordata" rank="species" species="abyssum">
<emphasis id="84B2EAA7FFCFFF8A8E9FF9111FC0F963" box="[253,444,1707,1732]" italics="true" pageId="9" pageNumber="480">Trimma abyssum</emphasis>
</taxonomicName>
has 16 such rays (the others have 15 or fewer), but has a second dorsal spine reaching posteriorly only as far as the bases of the first or second ray of the second dorsal fin (vs. to between base of 5th ray to 1
<taxonomicName id="71C64D36FFCFFF8A8EA3F9481F76F8B6" box="[193,266,1781,1809]" pageId="9" pageNumber="480" rank="subSpecies" subSpecies="scale">st scale</taxonomicName>
behind last dorsal fin ray in
<taxonomicName id="71C64D36FFCFFF8A8C29F9471CC1F8B6" authorityName="Winterbottom &amp; Erdmann" authorityYear="2018" box="[587,701,1786,1809]" class="Actinopterygii" family="Gobiidae" genus="Trimma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="480" phylum="Chordata" rank="species" species="meityae">
<emphasis id="84B2EAA7FFCFFF8A8C29F9471CC1F8B6" box="[587,701,1786,1809]" italics="true" pageId="9" pageNumber="480">T. meityae</emphasis>
</taxonomicName>
). Only three of these species have been recorded with as many as 21 total gill rakers (
<taxonomicName id="71C64D36FFCFFF8A8F11F8A31C70F892" authorityName="Winterbottom" authorityYear="2016" box="[371,524,1820,1845]" class="Actinopterygii" family="Gobiidae" genus="Trimma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="480" phylum="Chordata" rank="species" species="burridgeae">
<emphasis id="84B2EAA7FFCFFF8A8F11F8A31FFCF892" box="[371,384,1822,1845]" italics="true" pageId="9" pageNumber="480">T</emphasis>
.
<emphasis id="84B2EAA7FFCFFF8A8FEDF8A11C70F892" box="[399,524,1820,1845]" italics="true" pageId="9" pageNumber="480">burridgeae</emphasis>
</taxonomicName>
with 19-21 (mean = 19.6),
<taxonomicName id="71C64D36FFCFFF8A8D5DF8A31A67F892" authorityName="Yoshino &amp; Araga" authorityYear="1975" box="[831,1051,1820,1845]" class="Actinopterygii" family="Gobiidae" genus="Trimma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="480" phylum="Chordata" rank="species" species="caudomaculatum">
<emphasis id="84B2EAA7FFCFFF8A8D5DF8A31D30F892" box="[831,844,1822,1845]" italics="true" pageId="9" pageNumber="480">T</emphasis>
.
<emphasis id="84B2EAA7FFCFFF8A8D38F8A11A67F892" box="[858,1051,1820,1845]" italics="true" pageId="9" pageNumber="480">caudomaculatum</emphasis>
</taxonomicName>
with 17–21 (18.9) and
<taxonomicName id="71C64D36FFCFFF8A8B40F8A31BE0F892" authorityName="Winterbottom &amp; Hoese" authorityYear="2015" box="[1314,1436,1820,1845]" class="Actinopterygii" family="Gobiidae" genus="Trimma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="480" phylum="Chordata" rank="species" species="kitrinum">
<emphasis id="84B2EAA7FFCFFF8A8B40F8A31B53F892" box="[1314,1327,1822,1845]" italics="true" pageId="9" pageNumber="480">T</emphasis>
.
<emphasis id="84B2EAA7FFCFFF8A8B5FF8A11BE0F892" box="[1341,1436,1820,1845]" italics="true" pageId="9" pageNumber="480">kitrinum</emphasis>
</taxonomicName>
20–22 (21.3), versus 21–22 (21.7) in
<taxonomicName id="71C64D36FFCFFF8A8C5FF8FF1CCDF8FE" authorityName="Winterbottom &amp; Erdmann" authorityYear="2018" box="[573,689,1858,1881]" class="Actinopterygii" family="Gobiidae" genus="Trimma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="480" phylum="Chordata" rank="species" species="meityae">
<emphasis id="84B2EAA7FFCFFF8A8C5FF8FF1CCDF8FE" box="[573,689,1858,1881]" italics="true" pageId="9" pageNumber="480">T. meityae</emphasis>
</taxonomicName>
. In these species, the bony interorbital is 70–100% of pupil width (vs. 56–79.0%, mean = 65.4). A dark caudal spot is present in all the above listed species except
<taxonomicName id="71C64D36FFCFFF8A8B60F8DB1B0BF8DA" authorityName="Winterbottom &amp; Erdmann" authorityYear="2018" box="[1282,1399,1894,1917]" class="Actinopterygii" family="Gobiidae" genus="Trimma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="480" phylum="Chordata" rank="species" species="meityae">
<emphasis id="84B2EAA7FFCFFF8A8B60F8DB1B0BF8DA" box="[1282,1399,1894,1917]" italics="true" pageId="9" pageNumber="480">T. meityae</emphasis>
</taxonomicName>
,
<taxonomicName id="71C64D36FFCFFF8A8BE5F8DB1E92F806" authorityName="Winterbottom" authorityYear="2011" class="Actinopterygii" family="Gobiidae" genus="Trimma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="480" phylum="Chordata" rank="species" species="habrum">
<emphasis id="84B2EAA7FFCFFF8A8BE5F8DB1BE8F8DA" box="[1415,1428,1894,1917]" italics="true" pageId="9" pageNumber="480">T</emphasis>
.
<emphasis id="84B2EAA7FFCFFF8A8EF5F8351E92F806" box="[151,238,1928,1953]" italics="true" pageId="9" pageNumber="480">habrum</emphasis>
</taxonomicName>
,
<taxonomicName id="71C64D36FFCFFF8A8E9CF8371F0FF806" box="[254,371,1928,1953]" class="Actinopterygii" family="Gobiidae" genus="Trimma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="480" phylum="Chordata" rank="species" species="helenae">
<emphasis id="84B2EAA7FFCFFF8A8E9CF8371F70F806" box="[254,268,1930,1953]" italics="true" pageId="9" pageNumber="480">T</emphasis>
.
<emphasis id="84B2EAA7FFCFFF8A8F7EF8351F0FF806" box="[284,371,1928,1953]" italics="true" pageId="9" pageNumber="480">helenae</emphasis>
</taxonomicName>
and
<taxonomicName id="71C64D36FFCFFF8A8FCCF8371C57F806" authorityName="Winterbottom &amp; Hoese" authorityYear="2015" box="[430,555,1928,1953]" class="Actinopterygii" family="Gobiidae" genus="Trimma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="480" phylum="Chordata" rank="species" species="kitrinum">
<emphasis id="84B2EAA7FFCFFF8A8FCCF8371FC7F806" box="[430,443,1930,1953]" italics="true" pageId="9" pageNumber="480">T</emphasis>
.
<emphasis id="84B2EAA7FFCFFF8A8FAEF8351C57F806" box="[460,555,1928,1953]" italics="true" pageId="9" pageNumber="480">kitrinum</emphasis>
</taxonomicName>
, where it is absent. See
<taxonomicName id="71C64D36FFCFFF8A8D2AF8371DA0F806" authorityName="Winterbottom &amp; Erdmann" authorityYear="2018" box="[840,988,1928,1953]" class="Actinopterygii" family="Gobiidae" genus="Trimma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="480" phylum="Chordata" rank="species" species="blematium">
<emphasis id="84B2EAA7FFCFFF8A8D2AF8371DA0F806" box="[840,988,1928,1953]" italics="true" pageId="9" pageNumber="480">T. blematium</emphasis>
</taxonomicName>
under the Comparisons section for the similarities and differences between that species and
<taxonomicName id="71C64D36FFCFFF8A8C8BF8131D27F862" authorityName="Winterbottom &amp; Erdmann" authorityYear="2018" box="[745,859,1966,1989]" class="Actinopterygii" family="Gobiidae" genus="Trimma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="480" phylum="Chordata" rank="species" species="meityae">
<emphasis id="84B2EAA7FFCFFF8A8C8BF8131D27F862" box="[745,859,1966,1989]" italics="true" pageId="9" pageNumber="480">T. meityae</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="FEDC653EFFCFFF8A8EA5F86D1A7AF84F" box="[199,1030,2000,2025]" pageId="9" pageNumber="480" type="discussion">
<paragraph id="B67936B5FFCFFF8A8EA5F86D1A7AF84F" blockId="9.[151,1437,1236,2025]" box="[199,1030,2000,2025]" pageId="9" pageNumber="480">
<emphasis id="84B2EAA7FFCFFF8A8EA5F86D1F30F84E" bold="true" box="[199,332,2000,2025]" pageId="9" pageNumber="480">Discussion.</emphasis>
See
<taxonomicName id="71C64D36FFCFFF8A8FE0F86F1C68F84E" authorityName="Winterbottom &amp; Erdmann" authorityYear="2018" box="[386,532,2000,2025]" class="Actinopterygii" family="Gobiidae" genus="Trimma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="480" phylum="Chordata" rank="species" species="blematium">
<emphasis id="84B2EAA7FFCFFF8A8FE0F86F1C68F84E" box="[386,532,2000,2025]" italics="true" pageId="9" pageNumber="480">T. blematium</emphasis>
</taxonomicName>
under the Discussion section for comments.
</paragraph>
</subSubSection>
</treatment>
</document>
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