128 lines
17 KiB
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128 lines
17 KiB
XML
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<document id="7E3A2D4A48A1774CBD0519DA945A7AC6" ID-DOI="10.11646/zootaxa.3635.5" ID-GBIF-Dataset="6c160e64-940a-41f6-89d4-e7299281bc6e" ID-ISSN="1175-5326" ID-Zenodo-Dep="217427" ID-ZooBank="5E6A3AFD-89BF-441F-A826-A075C932D42D" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460587389638" checkinUser="plazi" docAuthor="Anisyutkin, Leonid N., Anichkin, Alexandr E. & Thinh, Nguyen Van" docDate="2013" docId="8C4987926728FFFFC7B0FC58FC67D673" docLanguage="en" docName="zt03635p532.pdf" docOrigin="Zootaxa 3635 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macrostylopyga laosana Anisyutkin 2010" docType="treatment" docVersion="6" lastPageNumber="530" masterDocId="7070FFEA6722FFF5C727FF8CFF81D059" masterDocTitle="Macrostylopyga gen. nov., a new genus of cockroaches (Dictyoptera: Blattidae), with descriptions of two new species" masterLastPageNumber="532" masterPageNumber="520" pageNumber="530" updateTime="1698325378222" updateUser="plazi">
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<mods:titleInfo id="6678129D4F4119A026B03FB739B06AD4">
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<mods:title id="B80A2DEADD4F94B9B4F0E43B36746697">Macrostylopyga gen. nov., a new genus of cockroaches (Dictyoptera: Blattidae), with descriptions of two new species</mods:title>
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</mods:titleInfo>
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<mods:name id="1AED19384AA5E4D00816EFAF75DDE0D8" type="personal">
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<mods:roleTerm id="60A7D71833E9A95F4603EE8744D9392E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0FA82AC68EBEF19D1348210D37E99EE3">Anisyutkin, Leonid N.</mods:namePart>
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</mods:name>
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<mods:name id="251BE68D01986D9038436D50BB6CD306" type="personal">
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<mods:role id="39DCBBE806CCEA2A866FDABB2F358B61">
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<mods:roleTerm id="B2CF238B177BA43EC86D03129EE401BD">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9C440A0B002C1255730F7EAC0E2A312C">Anichkin, Alexandr E.</mods:namePart>
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</mods:name>
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<mods:name id="AF6119BF91E50C1E1B429F729B7F48DB" type="personal">
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<mods:role id="60A4FC0F3564F4D927A023DE6F7FADDB">
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<mods:roleTerm id="DFDF5C5F316E185B91A1BF662E90D3B8">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E347AF47E854FC904F98D2070983A9D2">Thinh, Nguyen Van</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="F06BC0CDEC1E039E2E106CE6995059BE">text</mods:typeOfResource>
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<mods:titleInfo id="BC43B4EF6915EE75ABD8D2A57168E056">
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<mods:title id="A13F5F47B35A8EFE4CD877BACD466B17">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="D30B2A53EBA1AA23D4B26CCDA42B0B76">
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<mods:date id="86D6198269D921C30DB0DF2197157798">2013</mods:date>
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<mods:detail id="387BCF1972C710CDE20E1FBC0189EDCF" type="volume">
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<mods:number id="8DC0FFE75B3BE2741ED87EFC04D3BE6A">3635</mods:number>
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</mods:detail>
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<mods:detail id="9B548D9912460C14F6E9BDCBA3B189A9" type="issue">
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<mods:number id="A99C74BE6F0A9A2CEBC2C0AE2BA3C647">5</mods:number>
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</mods:detail>
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<mods:extent id="80B48BD4CA09AB09FF3AC512C86D735B" unit="page">
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<mods:start id="1439A8E9BD34E408B636359465A0AEA3">520</mods:start>
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<mods:end id="20B423A7610BE88D81F05E36F1284665">532</mods:end>
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</mods:extent>
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</mods:part>
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<mods:classification id="EDC5D1D90088DC199A7BADADA927447A">journal article</mods:classification>
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<mods:identifier id="F1638CBF08B803133288E5395EB5DE38" type="DOI">10.11646/zootaxa.3635.5</mods:identifier>
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<mods:identifier id="81CE05C2EC2DA838319994CC7E482B47" type="GBIF-Dataset">6c160e64-940a-41f6-89d4-e7299281bc6e</mods:identifier>
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<mods:identifier id="1D0E3156CDB886C24F33854C3DC18399" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="0E14D720090A1BB97EA73F50877F6B17" type="Zenodo-Dep">217427</mods:identifier>
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<mods:identifier id="60E0E6B21B336B001893B0A518F77FAC" type="ZooBank">5E6A3AFD-89BF-441F-A826-A075C932D42D</mods:identifier>
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</mods:mods>
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<treatment id="8C4987926728FFFFC7B0FC58FC67D673" ID-DOI="http://doi.org/10.5281/zenodo.6165484" ID-GBIF-Taxon="119565077" ID-Zenodo-Dep="6165484" LSID="urn:lsid:plazi:treatment:8C4987926728FFFFC7B0FC58FC67D673" httpUri="http://treatment.plazi.org/id/8C4987926728FFFFC7B0FC58FC67D673" lastPageNumber="530" pageId="10" pageNumber="530">
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<subSubSection id="4CFA650F6728FFFFC7B0FC58FE2DD457" pageId="10" pageNumber="530" type="nomenclature">
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<paragraph id="045F36846728FFFFC7B0FC58FD2AD3B4" blockId="10.[151,683,979,1038]" box="[151,683,979,1006]" pageId="10" pageNumber="530">
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<heading id="5F1781E86728FFFFC7B0FC58FD2AD3B4" bold="true" box="[151,683,979,1006]" fontSize="11" level="1" pageId="10" pageNumber="530" reason="1">
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<taxonomicName id="C3E04D076728FFFFC7B0FC58FD2AD3B4" ID-CoL="3XBG8" authority="Anisyutkin, 2010" authorityName="Anisyutkin" authorityYear="2010" box="[151,683,979,1006]" class="Insecta" family="Blattidae" genus="Macrostylopyga" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="530" phylum="Arthropoda" rank="species" species="laosana">
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<emphasis id="3694EA966728FFFFC7B0FC58FD2AD3B4" bold="true" box="[151,683,979,1006]" pageId="10" pageNumber="530">
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<emphasis id="3694EA966728FFFFC7B0FC58FE3CD3B7" bold="true" box="[151,445,980,1006]" italics="true" pageId="10" pageNumber="530">Macrostylopyga laosana</emphasis>
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(Anisyutkin, 2010)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="045F36846728FFFFC7B0FC7AFE2DD457" blockId="10.[151,683,979,1038]" box="[151,428,1014,1038]" pageId="10" pageNumber="530">
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(
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<figureCitation id="9CDB2A016728FFFFC7B8FC7AFEA7D457" box="[159,294,1014,1038]" captionStart="FIGURES 12 – 26" captionStartId="6.[151,264,1797,1820]" captionTargetBox="[253,1329,203,1766]" captionTargetId="figure@6.[239,1348,193,1776]" captionTargetPageId="6" captionText="FIGURES 12 – 26. Macrostylopyga spp., male, holotypes (12 – 17, 23, 24 — M. bidupi sp. nov., 18 – 22 — M. grandis sp. nov., 25, 26 — M. laosana (Anisyutkin, 2010). 12 — facial part of head; 13 — head, thorax and 1 – 7 th abdominal tergites from above; 14, 15 — abdominal apex from above (14) and below (15, hypandrium and genitalia removed); 16 — caudo-medial processes of paraprocts; 17 — hypandrium from below; 18, 22 — hind tarsus from outside (18) and below (22); 19 — arolium and claws; 20, 21, 23 – 26 — metatarsi: right fore (20, 23), right middle (21, 24), left hind (25, 26) from below (20, 21, 23, 24, 26) and outside (25). Dotted area shows membranous parts. Abbreviations: I – X — 1 st – 10 th abdominal tergites respectively, a. s.; ar.; e. r.; i. r.; pr.; par. — see text. Arrows show places of styli attachment. Scale bar (1 mm = e; 2 mm = h, i, k; 3 mm = d, j, l; 5 mm = a – c, f, g): a = 12; b = 13; c = 14; d = 15; e = 16; f = 17; g = 18; h = 19; i = 20, 21; j = 22; k = 23, 24; l = 25, 26." httpUri="https://zenodo.org/record/217429/files/figure.png" pageId="10" pageNumber="530">Figs. 25, 26</figureCitation>
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,
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<figureCitation id="9CDB2A016728FFFFC615FC7AFEFAD457" box="[306,379,1014,1038]" captionStart="FIGURES 27 – 38" captionStartId="7.[151,264,1891,1914]" captionTargetBox="[203,1384,204,1860]" captionTargetId="figure@7.[200,1387,193,1870]" captionTargetPageId="7" captionText="FIGURES 27 – 38. Macrostylopyga spp., male genitalia, holotypes (27 – 30 — M. grandis sp. nov., 31 – 34 — M. bidupi sp. nov., 35 – 38 — M. laosana (Anisyutkin, 2010). Left phallomera from above (27, 31, 35), outside (28, 32, 36), inside (29, 33, partly, 37) and below (30, 34, partly, 38). Dotted area shows membranous parts. Figs. 35 – 38 after Anisyutkin, 2010. Abbreviations: a. c. s.; a. p.; c. pr.; d. c. p.; e. c. s.; L 1; L 2; L 3; L 4 C; L 4 C, c. p.; L 4 D; L 4 F; m. l.; r. m. l.; r. o.; s. l.; sla; t. p. — see text. Scale bar 2 mm." httpUri="https://zenodo.org/record/217430/files/figure.png" pageId="10" pageNumber="530">35–38</figureCitation>
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,
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<figureCitation id="9CDB2A016728FFFFC6A1FC7AFE22D457" box="[390,419,1014,1038]" captionStart="FIGURES 50 – 56" captionStartId="9.[151,264,1862,1885]" captionTargetBox="[218,1362,203,1830]" captionTargetId="figure@9.[196,1392,193,1841]" captionTargetPageId="9" captionText="FIGURES 50 – 56. Macrostylopyga spp., male genitalia, holotypes (50 – 52 — M. grandis sp. nov., 53 – 55 — M. bidupi sp. nov., 56 — M. laosana (Anisyutkin, 2010). 50, 51, 53, 54 — right phallomere from above (50, 53) and below (51, 54); 52 — sclerite R 1 H; 55 — sclerites R 1 H and R 1 G from dorso-caudal; 56 — outline of sclerite R 1 H processes. Dotted area shows membranous parts. Fig. 56 after Anisyutkin, 2010. Abbreviations: p. s.; R 1 G; R 1 H; R 2; R 3; t. s. — see text. Scale bar (2 mm = a – c, g; 1 mm = d – f): a = 50; b = 51; c = 52; d = 53; e = 54; f = 55; g = 56." httpUri="https://zenodo.org/record/217432/files/figure.png" pageId="10" pageNumber="530">56</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="4CFA650F6728FFFFC7B0FBB2FB69D553" pageId="10" pageNumber="530" type="materials_examined">
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<paragraph id="045F36846728FFFFC7B0FBB2FE11D4C7" blockId="10.[151,1437,1086,1579]" pageId="10" pageNumber="530">
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<emphasis id="3694EA966728FFFFC7B0FBB2FE87D40E" bold="true" box="[151,262,1086,1111]" pageId="10" pageNumber="530">Material.</emphasis>
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<typeStatus id="DB5B88266728FFFFC628FBB2FEF6D40F" box="[271,375,1086,1110]" pageId="10" pageNumber="530" type="holotype">Holotype</typeStatus>
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male,
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<collectingCountry id="7CF776146728FFFFC6EFFBB3FD82D40F" box="[456,515,1087,1110]" name="Laos" pageId="10" pageNumber="530">Laos</collectingCountry>
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: Vientiane (Wieng Jan) Prov., National Park Phu Khao Khouay, Ban Vangheua, elevation =
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<quantity id="C3189B616728FFFFC63DFBEEFEE3D423" box="[282,354,1122,1146]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="530" unit="m" value="800.0">800 m</quantity>
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,
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<geoCoordinate id="61D450436728FFFFC64AFBEEFE7FD423" box="[365,510,1122,1146]" direction="north" orientation="latitude" pageId="10" pageNumber="530" precision="1" value="18.335033">18°20.102'N</geoCoordinate>
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,
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<geoCoordinate id="61D450436728FFFFC52FFBEEFD22D423" box="[520,675,1122,1146]" direction="east" orientation="longitude" pageId="10" pageNumber="530" precision="1" value="102.81638">102°48.983'E</geoCoordinate>
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, pitfall & carrion trap, mixed bamboo forest,
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<date id="705E10446728FFFFC3BCFBEEFF53D4C7" pageId="10" pageNumber="530" value="2008-02-27" valueMax="2008-03-28" valueMin="2008-02-27">27 February–28 March 2008</date>
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, coll. S. Tarasov.
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</paragraph>
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<paragraph id="045F36846728FFFFC7E0FB26FB69D553" blockId="10.[151,1437,1086,1579]" pageId="10" pageNumber="530">
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Originally
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<taxonomicName id="C3E04D076728FFFFC663FB20FE47D49A" box="[324,454,1195,1219]" class="Insecta" family="Blattidae" genus="Macrostylopyga" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="530" phylum="Arthropoda" rank="species" species="laosana">
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<emphasis id="3694EA966728FFFFC663FB20FE47D49A" box="[324,454,1195,1219]" italics="true" pageId="10" pageNumber="530">M. laosana</emphasis>
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</taxonomicName>
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was assigned to the genus
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<taxonomicName id="C3E04D076728FFFFC42FFB27FC21D49A" box="[776,928,1195,1219]" class="Insecta" family="Blattidae" genus="Neostylopyga" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="530" phylum="Arthropoda" rank="genus">
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<emphasis id="3694EA966728FFFFC42FFB27FC21D49A" box="[776,928,1195,1219]" italics="true" pageId="10" pageNumber="530">Neostylopyga</emphasis>
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</taxonomicName>
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, with the comment that this species and
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<taxonomicName id="C3E04D076728FFFFC2A5FB20FEEAD4BF" authority="Stoll" authorityName="Stoll" class="Insecta" family="Blattidae" genus="Neostylopyga" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="530" phylum="Arthropoda" rank="species" species="rhombifolia">
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<emphasis id="3694EA966728FFFFC2A5FB20FE9AD4BF" italics="true" pageId="10" pageNumber="530">N. rhombifolia</emphasis>
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(Stoll)
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</taxonomicName>
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, the
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<typeStatus id="DB5B88266728FFFFC686FB43FE50D4BE" box="[417,465,1231,1255]" pageId="10" pageNumber="530">type</typeStatus>
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species of the genus
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<taxonomicName id="C3E04D076728FFFFC5E6FB42FCD8D4BF" box="[705,857,1230,1254]" class="Insecta" family="Blattidae" genus="Neostylopyga" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="530" phylum="Arthropoda" rank="genus">
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<emphasis id="3694EA966728FFFFC5E6FB42FCD8D4BF" box="[705,857,1230,1254]" italics="true" pageId="10" pageNumber="530">Neostylopyga</emphasis>
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</taxonomicName>
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, probably belonged to different genera (Anisyutkin 2010). The original description (Anisyutkin 2010) can be supplemented with the following details.
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</paragraph>
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</subSubSection>
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<subSubSection id="4CFA650F6728FFFFC7E0FA9AFC67D673" pageId="10" pageNumber="530" type="description">
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<paragraph id="045F36846728FFFFC7E0FA9AFC67D673" blockId="10.[151,1437,1086,1579]" pageId="10" pageNumber="530">
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<emphasis id="3694EA966728FFFFC7E0FA9AFED0D576" bold="true" box="[199,337,1302,1327]" pageId="10" pageNumber="530">Description</emphasis>
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. Male (
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<typeStatus id="DB5B88266728FFFFC68BFA9AFD94D577" box="[428,533,1302,1326]" pageId="10" pageNumber="530" type="holotype">holotype</typeStatus>
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). Tibiae not thickened distally, apical spines not reinforced. Metatarsus of hind tarsi (
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<figureCitation id="9CDB2A016728FFFFC7FCFAB6FEEAD50B" box="[219,363,1338,1362]" captionStart="FIGURES 12 – 26" captionStartId="6.[151,264,1797,1820]" captionTargetBox="[253,1329,203,1766]" captionTargetId="figure@6.[239,1348,193,1776]" captionTargetPageId="6" captionText="FIGURES 12 – 26. Macrostylopyga spp., male, holotypes (12 – 17, 23, 24 — M. bidupi sp. nov., 18 – 22 — M. grandis sp. nov., 25, 26 — M. laosana (Anisyutkin, 2010). 12 — facial part of head; 13 — head, thorax and 1 – 7 th abdominal tergites from above; 14, 15 — abdominal apex from above (14) and below (15, hypandrium and genitalia removed); 16 — caudo-medial processes of paraprocts; 17 — hypandrium from below; 18, 22 — hind tarsus from outside (18) and below (22); 19 — arolium and claws; 20, 21, 23 – 26 — metatarsi: right fore (20, 23), right middle (21, 24), left hind (25, 26) from below (20, 21, 23, 24, 26) and outside (25). Dotted area shows membranous parts. Abbreviations: I – X — 1 st – 10 th abdominal tergites respectively, a. s.; ar.; e. r.; i. r.; pr.; par. — see text. Arrows show places of styli attachment. Scale bar (1 mm = e; 2 mm = h, i, k; 3 mm = d, j, l; 5 mm = a – c, f, g): a = 12; b = 13; c = 14; d = 15; e = 16; f = 17; g = 18; h = 19; i = 20, 21; j = 22; k = 23, 24; l = 25, 26." httpUri="https://zenodo.org/record/217429/files/figure.png" pageId="10" pageNumber="530">Figs. 25, 26</figureCitation>
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) slightly longer than other segments combined, with 2 rows of spines along lower margin (exterior row with 16–19 spines, interior row with 11–12 spines); second segment with 4 spines in exterior row and single spine corresponding to interior row on left tarsus; all segments with small apical euplantulae bordered with 2 additional spines (
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<figureCitation id="9CDB2A016728FFFFC64FFA2AFE3CD5E7" box="[360,445,1446,1470]" captionStart="FIGURES 12 – 26" captionStartId="6.[151,264,1797,1820]" captionTargetBox="[253,1329,203,1766]" captionTargetId="figure@6.[239,1348,193,1776]" captionTargetPageId="6" captionText="FIGURES 12 – 26. Macrostylopyga spp., male, holotypes (12 – 17, 23, 24 — M. bidupi sp. nov., 18 – 22 — M. grandis sp. nov., 25, 26 — M. laosana (Anisyutkin, 2010). 12 — facial part of head; 13 — head, thorax and 1 – 7 th abdominal tergites from above; 14, 15 — abdominal apex from above (14) and below (15, hypandrium and genitalia removed); 16 — caudo-medial processes of paraprocts; 17 — hypandrium from below; 18, 22 — hind tarsus from outside (18) and below (22); 19 — arolium and claws; 20, 21, 23 – 26 — metatarsi: right fore (20, 23), right middle (21, 24), left hind (25, 26) from below (20, 21, 23, 24, 26) and outside (25). Dotted area shows membranous parts. Abbreviations: I – X — 1 st – 10 th abdominal tergites respectively, a. s.; ar.; e. r.; i. r.; pr.; par. — see text. Arrows show places of styli attachment. Scale bar (1 mm = e; 2 mm = h, i, k; 3 mm = d, j, l; 5 mm = a – c, f, g): a = 12; b = 13; c = 14; d = 15; e = 16; f = 17; g = 18; h = 19; i = 20, 21; j = 22; k = 23, 24; l = 25, 26." httpUri="https://zenodo.org/record/217429/files/figure.png" pageId="10" pageNumber="530">Fig. 25</figureCitation>
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,
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<emphasis id="3694EA966728FFFFC6ECFA24FE70D5E6" box="[459,497,1448,1471]" italics="true" pageId="10" pageNumber="530">a.s.</emphasis>
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); claws symmetrical, simple; arolium vestigial. Left hind tarsus abnormal, with 4 segments. Fore and middle tarsi similar to hind tarsi, but comparatively shorter; fore and middle metatarsi with
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<date id="705E10446728FFFFC25FFA46FEADD65F" pageId="10" pageNumber="530" value="1914-08-07" valueMax="1914-12-08" valueMin="1914-08-07">14/ 8 and 12/7–8</date>
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spines in exterior/interior rows respectively, second segment of left middle tarsus with single spine along lower margin; other segments without armament along lower margin.
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|
</paragraph>
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</subSubSection>
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|||
|
</treatment>
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|||
|
</document>
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