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<document id="742D6CEE1EDAC778E90B758D1D923D9F" ID-DOI="10.1046/j.1096-3642.2002.00039.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="10114672" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630436109371" checkinUser="carolina" docAuthor="Valdés, Ángel" docDate="2002" docId="03B5879A75756C65937EFB9220AEAC96" docLanguage="en" docName="j.1096-3642.2002.00039.x.pdf" docOrigin="Zoological Journal of the Linnean Society 136 (4)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1046/j.1096-3642.2002.00039.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Discodoris ketos" docType="treatment" docVersion="4" lastPageNumber="571" masterDocId="FF8CFFE275576C419042FF872440A839" masterDocTitle="A phylogenetic analysis and systematic revision of the cryptobranch dorids (Mollusca, Nudibranchia, Anthobranchia)" masterLastPageNumber="636" masterPageNumber="535" pageNumber="569" updateTime="1700038700340" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="523792BC3F5D237D2293BC4D75183755">A phylogenetic analysis and systematic revision of the cryptobranch dorids (Mollusca, Nudibranchia, Anthobranchia)</mods:title>
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<mods:namePart id="4921AAF5245C1C4D5FBC021C40CFD853">Valdés, Ángel</mods:namePart>
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<mods:typeOfResource id="51F62ADC2551746008CC5A8254C5A34B">text</mods:typeOfResource>
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<mods:title id="230BA170BC706926E2707C7EC31C5872">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="4C052E5A4D3FD8CB85C86B0E738F47AE">2002</mods:date>
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<mods:number id="3F27614483C2EB7C86552C0951E1CE8D">2002-12-31</mods:number>
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<mods:url id="3F99AB9479C9C921046447847997B478">https://academic.oup.com/zoolinnean/article-lookup/doi/10.1046/j.1096-3642.2002.00039.x</mods:url>
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<mods:classification id="DCC0FE49C0C565D1DCB6154120D0771B">journal article</mods:classification>
<mods:identifier id="961F19E145D449B8401F55FEBF520CE9" type="DOI">10.1046/j.1096-3642.2002.00039.x</mods:identifier>
<mods:identifier id="1DA2A5FA018749C5F514399E8845964A" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="CEC7F77D673B650CE071D070F6D8AFF1" type="Zenodo-Dep">10114672</mods:identifier>
</mods:mods>
<treatment id="03B5879A75756C65937EFB9220AEAC96" LSID="urn:lsid:plazi:treatment:03B5879A75756C65937EFB9220AEAC96" httpUri="http://treatment.plazi.org/id/03B5879A75756C65937EFB9220AEAC96" lastPageId="36" lastPageNumber="571" pageId="34" pageNumber="569">
<subSubSection id="C306650775756C63937EFB9220A2AC75" pageId="34" pageNumber="569" type="nomenclature">
<paragraph id="8BA3368C75756C63937EFB922139AC15" blockId="34.[828,1401,1044,1068]" box="[828,1401,1044,1068]" pageId="34" pageNumber="569">
<heading id="D0EB81E075756C63937EFB922139AC15" box="[828,1401,1044,1068]" centered="true" fontSize="9" level="2" pageId="34" pageNumber="569" reason="2">
<taxonomicName id="4C1C4D0F75756C63937EFB922139AC15" ID-CoL="36TJF" authority="(EV. MARCUS &amp; ER. MARCUS" box="[828,1401,1044,1068]" class="Gastropoda" family="Discodorididae" genus="Discodoris" kingdom="Animalia" order="Nudibranchia" pageId="34" pageNumber="569" phylum="Mollusca" rank="species" species="ketos">
<emphasis id="B968EA9E75756C63937EFB922059AC12" box="[828,1049,1045,1068]" italics="true" pageId="34" pageNumber="569">DISCODORIS KETOS</emphasis>
(EV. MARCUS &amp; ER. MARCUS,
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BA3368C75756C639391FBB320A2AC75" blockId="34.[979,1250,1076,1100]" box="[979,1250,1076,1100]" pageId="34" pageNumber="569">
<heading id="D0EB81E075756C639391FBB320A2AC75" box="[979,1250,1076,1100]" centered="true" fontSize="9" level="2" pageId="34" pageNumber="569" reason="8">
1967) (
<figureCitation id="13272A0975756C639467FBB320C4AC75" box="[1061,1156,1076,1100]" captionStart="Figure 4" captionStartId="12.[142,220,1147,1166]" captionTargetBox="[142,1422,195,1115]" captionTargetId="figure-362@12.[142,1422,195,1115]" captionTargetPageId="12" captionText="Figure 4. Living animals. A, Doris pseudoargus (CASIZ 121105). B, Doris immonda (CASIZ 089023), photo by T. M. Gosliner. C, Doris granulosa (CASIZ 073536), photo by T. M. Gosliner. D, Discodoris boholiensis (CASIZ 083654), photo by T. M. Gosliner. E, Discodoris ketos, San Pedrillo, Puntarenas, Costa Rica, photo by T. M. Gosliner. F, Thordisa rubescens (CASIZ 015860), photo by T. M. Gosliner. G, Aphelodoris antillensis (CASIZ 077289), photo by T. M. Gosliner. H, Peltodoris atromaculata (CASIZ 119474). I, Peltodoris nobilis, Monterey Bay, California, photo by A. Smith." figureDoi="http://doi.org/10.5281/zenodo.10131873" httpUri="https://zenodo.org/record/10131873/files/figure.png" pageId="34" pageNumber="569">FIGS 4E</figureCitation>
,
<figureCitation id="13272A0975756C6394D3FBB320EFAC75" box="[1169,1199,1076,1100]" captionStart="Figure 19" captionStartId="37.[155,233,1731,1750]" captionTargetBox="[155,1433,194,1696]" captionTargetId="figure-85@37.[155,1435,194,1698]" captionTargetPageId="37" captionText="Figure 19. Discodoris ketos (CASIZ 081808), SEM images of the radula, jaws and dorsal tubercles. A, inner lateral teeth; scale bar = 75 Mm. B, mid-lateral teeth; scale bar = 71 Mm. C, outer lateral teeth; scale bar = 42 Mm. D, dorsal tubercles; scale bar = 750 Mm. E, jaw elements; scale bar = 30 Mm." figureDoi="http://doi.org/10.5281/zenodo.10131907" httpUri="https://zenodo.org/record/10131907/files/figure.png" pageId="34" pageNumber="569">19</figureCitation>
,
<figureCitation id="13272A0975756C6394FFFBB3209AAC75" box="[1213,1242,1076,1100]" captionStart="Figure 20" captionStartId="38.[155,233,1789,1808]" captionTargetBox="[166,1426,204,1755]" captionTargetId="figure-83@38.[155,1435,195,1757]" captionTargetPageId="38" captionText="Figure 20. Discodoris ketos (CASIZ 081808). A, general view of the anatomy; scale bar = 1 mm. B, reproductive system; scale bar = 1 mm. C, lateral view of the buccal bulb; scale bar = 1 mm. D, central nervous system; scale bar = 0.5 mm. E, ventral view of the mouth area; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.10131909" httpUri="https://zenodo.org/record/10131909/files/figure.png" pageId="34" pageNumber="569">20</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C306650775756C639364FBD927F9ACAB" pageId="34" pageNumber="569" type="reference_group">
<paragraph id="8BA3368C75756C639364FBD927F9ACAB" blockId="34.[806,1422,1117,1170]" pageId="34" pageNumber="569">
<treatmentCitationGroup id="AB0C11A275756C639364FBD927F9ACAB" pageId="34" pageNumber="569">
<treatmentCitation id="0ABD109D75756C639364FBD921CAAC4B" author="Marcus Ev &amp; Marcus Er" box="[806,1418,1117,1139]" page="194" pageId="34" pageNumber="569" year="1967">
<taxonomicName id="4C1C4D0F75756C639364FBD921CAAC4B" ID-CoL="36TJF" authority="Marcus &amp; Marcus, 1967 b: 192 - 194" authorityName="Ev. Marcus &amp; Er. Marcus" authorityPageNumber="192 - 194" authorityYear="1967" box="[806,1418,1117,1139]" class="Gastropoda" family="Discodorididae" genus="Tayuva" kingdom="Animalia" order="Nudibranchia" pageId="34" pageNumber="569" phylum="Mollusca" rank="species" species="ketos">
<emphasis id="B968EA9E75756C639364FBD92780AC4B" box="[806,960,1117,1139]" italics="true" pageId="34" pageNumber="569">Tayuva ketos</emphasis>
<bibRefCitation id="EF8D4B7D75756C639393FBD921CAAC4B" author="Marcus Ev &amp; Marcus Er" box="[977,1418,1117,1139]" pageId="34" pageNumber="569" pagination="1 - 256" refId="ref57192" refString="Marcus Ev, Marcus Er. 1967 b. American opisthobranch molluks. Studies in Tropical Oceanography 6: 1 - 256, pl. 1." type="journal article" year="1967">Marcus &amp; Marcus, 1967b: 192–194</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 52-56.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C306650775756C659364FB5F20AEAC96" lastPageId="36" lastPageNumber="571" pageId="34" pageNumber="569" type="materials_examined">
<paragraph id="8BA3368C75756C639364FB5F2781ACD5" blockId="34.[806,1422,1239,1354]" box="[806,961,1239,1261]" pageId="34" pageNumber="569">
<emphasis id="B968EA9E75756C639364FB5F2781ACD5" box="[806,961,1239,1261]" italics="true" pageId="34" pageNumber="569">Type material</emphasis>
</paragraph>
<paragraph id="8BA3368C75756C639364FB702123AD73" blockId="34.[806,1422,1239,1354]" pageId="34" pageNumber="569">
<materialsCitation id="3B743CD175756C639364FB702123AD73" collectingDate="1963-11-02" collectionCode="USNM" collectorName="P. Pickens" country="Mexico" location="Playa Norse" municipality="Puerto Penasco" pageId="34" pageNumber="569" specimenCode="USNM 678409" specimenCount="1" stateProvince="Sonora" typeStatus="lectotype">
<typeStatus id="54A7882E75756C639364FB702783AD35" box="[806,963,1271,1292]" pageId="34" pageNumber="569" type="lectotype">LECTOTYPE</typeStatus>
(here designated):
<location id="8EC3605775756C6394E7FB712174AD35" LSID="urn:lsid:plazi:treatment:03B5879A75756C65937EFB9220AEAC96:8EC3605775756C6394E7FB712174AD35" box="[1189,1332,1270,1292]" country="Mexico" municipality="Puerto Penasco" name="Playa Norse" pageId="34" pageNumber="569" stateProvince="Sonora">Playa Norse</location>
,
<collectingMunicipality id="6BC7ACF675756C639503FB7027C3AD12" pageId="34" pageNumber="569">Puerto Peñasco</collectingMunicipality>
,
<collectingRegion id="49D8F86E75756C6393D6FA9227A6AD13" box="[916,998,1301,1322]" country="Mexico" name="Sonora" pageId="34" pageNumber="569">Sonora</collectingRegion>
,
<collectingCountry id="F30B761C75756C6393B5FA922009AD12" box="[1015,1097,1301,1323]" name="Mexico" pageId="34" pageNumber="569">Mexico</collectingCountry>
,
<date id="FFA2104C75756C639418FA92216FAD12" box="[1114,1327,1301,1323]" pageId="34" pageNumber="569" value="1963-11-02">
<collectingDate id="EFE6E9A475756C639418FA92216FAD12" box="[1114,1327,1301,1323]" pageId="34" pageNumber="569" value="1963-11-02">
2 November
<quantity id="4CE49B6975756C6394B4FA92216FAD12" box="[1270,1327,1301,1323]" metricMagnitude="0" metricUnit="m" metricValue="1.96328" pageId="34" pageNumber="569" unit="mm" value="1963.28">1963</quantity>
</collectingDate>
</date>
, 28 mm preserved length, leg.
<collectorName id="26E9535A75756C639467FAB320DAAD73" box="[1061,1178,1332,1354]" pageId="34" pageNumber="569">P. Pickens</collectorName>
(
<specimenCode id="DBBA9EF775756C6394EBFAB32118AD73" box="[1193,1368,1332,1354]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="34" pageNumber="569" type="Museum">USNM 678409</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BA3368C75756C639364FA082044AD9D" blockId="34.[806,1423,1423,1630]" box="[806,1028,1423,1444]" pageId="34" pageNumber="569">
<emphasis id="B968EA9E75756C639364FA082044AD9D" box="[806,1028,1423,1444]" italics="true" pageId="34" pageNumber="569">Additional material</emphasis>
</paragraph>
<paragraph id="8BA3368C75756C639364FA2927A5AE67" blockId="34.[806,1423,1423,1630]" pageId="34" pageNumber="569">
<materialsCitation id="3B743CD175756C639364FA2920BEAE06" collectingDate="1953-12-01" collectingDateMax="1984-01-25" collectingDateMin="1953-12-01" collectionCode="T" collectorName="L. O. Miles" country="Mexico" county="Las Cruces" location="North of Gauiola" municipality="Mazatlan" pageId="34" pageNumber="569" specimenCode="CASIZ 081808" specimenCount="2" stateProvince="Sinaloa">
<location id="8EC3605775756C639364FA292044ADFD" LSID="urn:lsid:plazi:treatment:03B5879A75756C65937EFB9220AEAC96:8EC3605775756C639364FA292044ADFD" box="[806,1028,1454,1476]" country="Mexico" county="Las Cruces" municipality="Mazatlan" name="North of Gauiola" pageId="34" pageNumber="569" stateProvince="Sinaloa">North of Gauiola</location>
,
<collectingMunicipality id="6BC7ACF675756C63945FFA2920CCADFD" box="[1053,1164,1454,1476]" pageId="34" pageNumber="569">Mazatlán</collectingMunicipality>
,
<collectingRegion id="49D8F86E75756C6394E7FA2920BDADFD" box="[1189,1277,1454,1476]" country="Mexico" name="Sinaloa" pageId="34" pageNumber="569">Sinaloa</collectingRegion>
,
<collectingCountry id="F30B761C75756C639554FA292128ADFD" box="[1302,1384,1454,1476]" name="Mexico" pageId="34" pageNumber="569">Mexico</collectingCountry>
,
<date id="FFA2104C75756C6395C3FA2827A5ADDA" pageId="34" pageNumber="569" value="1953-12-01">
<collectingDate id="EFE6E9A475756C6395C3FA2827A5ADDA" pageId="34" pageNumber="569" value="1953-12-01">1 December 1953</collectingDate>
</date>
,
<specimenCount id="9D1AFD0575756C6393BFFA4920E4ADDA" box="[1021,1188,1485,1507]" pageId="34" pageNumber="569" type="generic">one specimen</specimenCount>
,
<quantity id="4CE49B6975756C6394FEFA4A214AADDA" box="[1212,1290,1485,1507]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" pageId="34" pageNumber="569" unit="mm" value="42.0">42 mm</quantity>
preserved length, leg.
<collectorName id="26E9535A75756C6393F3FA6B2072AE3B" box="[945,1074,1516,1538]" pageId="34" pageNumber="569">L. O. Miles</collectorName>
(
<specimenCode id="DBBA9EF775756C639401FA6B20B0AE3B" box="[1091,1264,1516,1538]" collectionCode="CASIZ" pageId="34" pageNumber="569">CASIZ 081808</specimenCode>
).
<collectingCounty id="62C24E0075756C639547FA6B21CAAE38" box="[1285,1418,1516,1537]" pageId="34" pageNumber="569">Las Cruces</collectingCounty>
,
<collectingRegion id="49D8F86E75756C639364F98D2052AE19" box="[806,1042,1546,1568]" country="Mexico" name="Baja California Sur" pageId="34" pageNumber="569">Baja California Sur</collectingRegion>
,
<collectingCountry id="F30B761C75756C639466F98D2036AE19" box="[1060,1142,1546,1568]" name="Mexico" pageId="34" pageNumber="569">Mexico</collectingCountry>
,
<date id="FFA2104C75756C6394C5F98D2115AE19" box="[1159,1365,1546,1568]" pageId="34" pageNumber="569" value="1984-01-25">
<collectingDate id="EFE6E9A475756C6394C5F98D2115AE19" box="[1159,1365,1546,1568]" pageId="34" pageNumber="569" value="1984-01-25">25 January 1984</collectingDate>
</date>
,
<specimenCount id="9D1AFD0575756C639524F98C27D2AE06" pageId="34" pageNumber="569" type="generic">one specimen</specimenCount>
,
<quantity id="4CE49B6975756C6393DCF9AE27ACAE06" box="[926,1004,1577,1599]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="34" pageNumber="569" unit="mm" value="30.0">30 mm</quantity>
preserved length, leg.
<collectionCode id="ED0DAE4975756C6394ACF9AD20BEAE06" box="[1262,1278,1578,1599]" name="Tavera, Department of Geology and Geophysics" pageId="34" pageNumber="569">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B743CD175756C63954BF9AD27A1AE67" collectionCode="CASIZ" collectorName="M. Gosliner" country="Mexico" pageId="34" pageNumber="569" specimenCode="CASIZ 072843" specimenCount="1" stateProvince="Sinaloa">
<collectorName id="26E9535A75756C63954BF9AD21CEAE06" box="[1289,1422,1577,1599]" pageId="34" pageNumber="569">M. Gosliner</collectorName>
(
<specimenCode id="DBBA9EF775756C63936FF9CF2799AE67" box="[813,985,1608,1630]" collectionCode="CASIZ" pageId="34" pageNumber="569">CASIZ 072843</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA3368C75756C639364F925204FAE8E" blockId="34.[806,1422,1698,1904]" box="[806,1039,1698,1719]" pageId="34" pageNumber="569">
<emphasis id="B968EA9E75756C639364F925204FAE8E" box="[806,1039,1698,1719]" italics="true" pageId="34" pageNumber="569">External morphology</emphasis>
</paragraph>
<paragraph id="8BA3368C75756C659364F9462511AA51" blockId="34.[806,1422,1698,1904]" lastBlockId="36.[142,759,196,708]" lastPageId="36" lastPageNumber="571" pageId="34" pageNumber="569">
The background colour of the living animals is pale grey (
<figureCitation id="13272A0975756C63932BF96727FDAECF" box="[873,957,1760,1782]" captionStart="Figure 4" captionStartId="12.[142,220,1147,1166]" captionTargetBox="[142,1422,195,1115]" captionTargetId="figure-362@12.[142,1422,195,1115]" captionTargetPageId="12" captionText="Figure 4. Living animals. A, Doris pseudoargus (CASIZ 121105). B, Doris immonda (CASIZ 089023), photo by T. M. Gosliner. C, Doris granulosa (CASIZ 073536), photo by T. M. Gosliner. D, Discodoris boholiensis (CASIZ 083654), photo by T. M. Gosliner. E, Discodoris ketos, San Pedrillo, Puntarenas, Costa Rica, photo by T. M. Gosliner. F, Thordisa rubescens (CASIZ 015860), photo by T. M. Gosliner. G, Aphelodoris antillensis (CASIZ 077289), photo by T. M. Gosliner. H, Peltodoris atromaculata (CASIZ 119474). I, Peltodoris nobilis, Monterey Bay, California, photo by A. Smith." figureDoi="http://doi.org/10.5281/zenodo.10131873" httpUri="https://zenodo.org/record/10131873/files/figure.png" pageId="34" pageNumber="569">Fig. 4E</figureCitation>
). The dorsum is covered with numerous rounded or oval dark brown patches, which are larger in the centre of the dorsum. There are a few darker patches, almost black, situated in two rows on both sides of the visceral hump and several opaque white spots irregularly distributed. The rhinophoral and branchial sheaths are low and surrounded by several white spots. The rhinophores are grey, with a number of small dark brown spots. The branchial leaves are also grey, having dark grey spots and white patches. The whole dorsum is covered with small, conical tubercles (
<figureCitation id="13272A0975736C659146FEFB2528A9AB" box="[260,360,380,402]" captionStart="Figure 19" captionStartId="37.[155,233,1731,1750]" captionTargetBox="[155,1433,194,1696]" captionTargetId="figure-85@37.[155,1435,194,1698]" captionTargetPageId="37" captionText="Figure 19. Discodoris ketos (CASIZ 081808), SEM images of the radula, jaws and dorsal tubercles. A, inner lateral teeth; scale bar = 75 Mm. B, mid-lateral teeth; scale bar = 71 Mm. C, outer lateral teeth; scale bar = 42 Mm. D, dorsal tubercles; scale bar = 750 Mm. E, jaw elements; scale bar = 30 Mm." figureDoi="http://doi.org/10.5281/zenodo.10131907" httpUri="https://zenodo.org/record/10131907/files/figure.png" pageId="36" pageNumber="571">Fig. 19D</figureCitation>
). The largest tubercles are situated in the central region of the body. The rhinophoral and branchial sheaths have tubercles similar to those on the rest of the dorsum. There are six tripinnate branchial leaves. The anal papilla is situated in the centre of the branchial circle of leaves. The rhinophores are elongate, having 21 lamellae in a 30-mm preserved length specimen.
</paragraph>
<caption id="DF63660475746C6290D9F944270CAF28" ID-DOI="http://doi.org/10.5281/zenodo.10131905" ID-Zenodo-Dep="10131905" httpUri="https://zenodo.org/record/10131905/files/figure.png" pageId="35" pageNumber="570" startId="35.[155,233,1731,1750]" targetBox="[155,1433,194,1698]" targetPageId="35">
<paragraph id="8BA3368C75746C6290D9F944270CAF28" blockId="35.[155,1435,1731,1809]" pageId="35" pageNumber="570">
<emphasis id="B968EA9E75746C6290D9F9442550AEEF" bold="true" box="[155,272,1731,1750]" pageId="35" pageNumber="570">Figure 18.</emphasis>
<taxonomicName id="4C1C4D0F75746C62915EF94425B0AEEF" baseAuthorityName="Bergh" baseAuthorityYear="1878" box="[284,496,1731,1750]" class="Gastropoda" family="Discodorididae" genus="Discodoris" kingdom="Animalia" order="Nudibranchia" pageId="35" pageNumber="570" phylum="Mollusca" rank="species" species="zibethina">
<emphasis id="B968EA9E75746C62915EF94425B0AEEF" box="[284,496,1731,1750]" italics="true" pageId="35" pageNumber="570">Discodoris zibethina</emphasis>
</taxonomicName>
(ZMUC GAS-2112), SEM images of the radula, jaws and dorsal tubercles. A, inner lateral teeth; scale bar = 100 Mm. B, mid-lateral teeth; scale bar = 150 Mm. C, outer lateral teeth; scale bar = 100 Mm. D, jaw elements; scale bar = 30 Mm. E, dorsal tubercles; scale bar = 430 Mm.
</paragraph>
</caption>
<paragraph id="8BA3368C75736C6590E5FDF62558AAFD" blockId="36.[142,759,196,708]" pageId="36" pageNumber="571">
Ventrally the anterior border of the foot is grooved and notched (
<figureCitation id="13272A0975736C659173FD1725D3AA9F" box="[305,403,656,678]" captionStart="Figure 20" captionStartId="38.[155,233,1789,1808]" captionTargetBox="[166,1426,204,1755]" captionTargetId="figure-83@38.[155,1435,195,1757]" captionTargetPageId="38" captionText="Figure 20. Discodoris ketos (CASIZ 081808). A, general view of the anatomy; scale bar = 1 mm. B, reproductive system; scale bar = 1 mm. C, lateral view of the buccal bulb; scale bar = 1 mm. D, central nervous system; scale bar = 0.5 mm. E, ventral view of the mouth area; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.10131909" httpUri="https://zenodo.org/record/10131909/files/figure.png" pageId="36" pageNumber="571">Fig. 20E</figureCitation>
). The oral tentacles are short and conical.
</paragraph>
<paragraph id="8BA3368C75736C6590CCFC8D24B1AB26" blockId="36.[142,759,778,1904]" box="[142,241,778,799]" pageId="36" pageNumber="571">
<emphasis id="B968EA9E75736C6590CCFC8D24B1AB26" box="[142,241,778,799]" italics="true" pageId="36" pageNumber="571">Anatomy</emphasis>
</paragraph>
<paragraph id="8BA3368C75736C6590CCFCAE26C1AD32" blockId="36.[142,759,778,1904]" pageId="36" pageNumber="571">
The posterior end of the glandular portion of the oral tube has six strong retractor muscles (
<figureCitation id="13272A0975736C659206FCCF26E4AB67" box="[580,676,840,862]" captionStart="Figure 20" captionStartId="38.[155,233,1789,1808]" captionTargetBox="[166,1426,204,1755]" captionTargetId="figure-83@38.[155,1435,195,1757]" captionTargetPageId="38" captionText="Figure 20. Discodoris ketos (CASIZ 081808). A, general view of the anatomy; scale bar = 1 mm. B, reproductive system; scale bar = 1 mm. C, lateral view of the buccal bulb; scale bar = 1 mm. D, central nervous system; scale bar = 0.5 mm. E, ventral view of the mouth area; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.10131909" httpUri="https://zenodo.org/record/10131909/files/figure.png" pageId="36" pageNumber="571">Fig. 20C</figureCitation>
) which attach to the body wall. The oval, muscular buccal bulb has two large additional muscles attached; two long salivary glands connect with it at each side of the oesophageal junction. The buccal bulb is longer than the glandular portion of the oral tube. The labial cuticle is armed with a number of small rodlets (
<figureCitation id="13272A0975736C6592C5FB8726AAAC2F" box="[647,746,1024,1046]" captionStart="Figure 19" captionStartId="37.[155,233,1731,1750]" captionTargetBox="[155,1433,194,1696]" captionTargetId="figure-85@37.[155,1435,194,1698]" captionTargetPageId="37" captionText="Figure 19. Discodoris ketos (CASIZ 081808), SEM images of the radula, jaws and dorsal tubercles. A, inner lateral teeth; scale bar = 75 Mm. B, mid-lateral teeth; scale bar = 71 Mm. C, outer lateral teeth; scale bar = 42 Mm. D, dorsal tubercles; scale bar = 750 Mm. E, jaw elements; scale bar = 30 Mm." figureDoi="http://doi.org/10.5281/zenodo.10131907" httpUri="https://zenodo.org/record/10131907/files/figure.png" pageId="36" pageNumber="571">Fig. 19E</figureCitation>
). The radular formula is 23 ¥ 32.0.
<quantity id="4CE49B6975736C659252FB98260CAC0D" box="[528,588,1054,1076]" metricMagnitude="-1" metricUnit="m" metricValue="8.128" pageId="36" pageNumber="571" unit="in" value="32.0">32 in</quantity>
a 42-mm long specimen. Rachidian teeth are absent. The lateral teeth are hamate and lack denticles (
<figureCitation id="13272A0975736C659210FBDB26F5AC4B" box="[594,693,1116,1138]" captionStart="Figure 19" captionStartId="37.[155,233,1731,1750]" captionTargetBox="[155,1433,194,1696]" captionTargetId="figure-85@37.[155,1435,194,1698]" captionTargetPageId="37" captionText="Figure 19. Discodoris ketos (CASIZ 081808), SEM images of the radula, jaws and dorsal tubercles. A, inner lateral teeth; scale bar = 75 Mm. B, mid-lateral teeth; scale bar = 71 Mm. C, outer lateral teeth; scale bar = 42 Mm. D, dorsal tubercles; scale bar = 750 Mm. E, jaw elements; scale bar = 30 Mm." figureDoi="http://doi.org/10.5281/zenodo.10131907" httpUri="https://zenodo.org/record/10131907/files/figure.png" pageId="36" pageNumber="571">Fig. 19A</figureCitation>
). The teeth from the middle portion of the half-row are larger than those closer to the medial portion of the radula (
<figureCitation id="13272A0975736C6590ADFB3F2511ACF7" box="[239,337,1208,1230]" captionStart="Figure 19" captionStartId="37.[155,233,1731,1750]" captionTargetBox="[155,1433,194,1696]" captionTargetId="figure-85@37.[155,1435,194,1698]" captionTargetPageId="37" captionText="Figure 19. Discodoris ketos (CASIZ 081808), SEM images of the radula, jaws and dorsal tubercles. A, inner lateral teeth; scale bar = 75 Mm. B, mid-lateral teeth; scale bar = 71 Mm. C, outer lateral teeth; scale bar = 42 Mm. D, dorsal tubercles; scale bar = 750 Mm. E, jaw elements; scale bar = 30 Mm." figureDoi="http://doi.org/10.5281/zenodo.10131907" httpUri="https://zenodo.org/record/10131907/files/figure.png" pageId="36" pageNumber="571">Fig. 19B</figureCitation>
). The outermost teeth are smaller and also lack denticles (
<figureCitation id="13272A0975736C6591E8FB51264CACD4" box="[426,524,1238,1261]" captionStart="Figure 19" captionStartId="37.[155,233,1731,1750]" captionTargetBox="[155,1433,194,1696]" captionTargetId="figure-85@37.[155,1435,194,1698]" captionTargetPageId="37" captionText="Figure 19. Discodoris ketos (CASIZ 081808), SEM images of the radula, jaws and dorsal tubercles. A, inner lateral teeth; scale bar = 75 Mm. B, mid-lateral teeth; scale bar = 71 Mm. C, outer lateral teeth; scale bar = 42 Mm. D, dorsal tubercles; scale bar = 750 Mm. E, jaw elements; scale bar = 30 Mm." figureDoi="http://doi.org/10.5281/zenodo.10131907" httpUri="https://zenodo.org/record/10131907/files/figure.png" pageId="36" pageNumber="571">Fig. 19C</figureCitation>
). The oesophagus is short and connects directly to the stomach.
</paragraph>
<paragraph id="8BA3368C75736C6590E5FA932625AEEE" blockId="36.[142,759,778,1904]" pageId="36" pageNumber="571">
The ampulla is very long and convoluted (
<figureCitation id="13272A0975736C6592C5FA9326AAAD13" box="[647,746,1300,1322]" captionStart="Figure 20" captionStartId="38.[155,233,1789,1808]" captionTargetBox="[166,1426,204,1755]" captionTargetId="figure-83@38.[155,1435,195,1757]" captionTargetPageId="38" captionText="Figure 20. Discodoris ketos (CASIZ 081808). A, general view of the anatomy; scale bar = 1 mm. B, reproductive system; scale bar = 1 mm. C, lateral view of the buccal bulb; scale bar = 1 mm. D, central nervous system; scale bar = 0.5 mm. E, ventral view of the mouth area; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.10131909" httpUri="https://zenodo.org/record/10131909/files/figure.png" pageId="36" pageNumber="571">Fig. 20B</figureCitation>
). It branches into a short oviduct and the prostate. The oviduct enters the female gland mass near to its centre. The prostate is large and flattened. It has two different portions that are clearly distinguishable in colour and texture. The prostate connects with a long duct that narrows and expands again into the large ejaculatory portion of the deferent duct. The penis is unarmed. The muscular deferent duct opens into a large, common atrium with the vagina. The vagina is short. At its proximal end it joins the bursa copulatrix. From the bursa copulatrix leads another duct connecting to the uterine duct and the seminal receptacle. The bursa copulatrix is oval in shape, about three times as large as the elongate seminal receptacle.
</paragraph>
<paragraph id="8BA3368C75736C6590E5F96720CEA9F6" blockId="36.[142,759,778,1904]" lastBlockId="36.[806,1423,196,555]" pageId="36" pageNumber="571">
In the central nervous system (
<figureCitation id="13272A0975736C65925EF967263EAECF" box="[540,638,1760,1782]" captionStart="Figure 20" captionStartId="38.[155,233,1789,1808]" captionTargetBox="[166,1426,204,1755]" captionTargetId="figure-83@38.[155,1435,195,1757]" captionTargetPageId="38" captionText="Figure 20. Discodoris ketos (CASIZ 081808). A, general view of the anatomy; scale bar = 1 mm. B, reproductive system; scale bar = 1 mm. C, lateral view of the buccal bulb; scale bar = 1 mm. D, central nervous system; scale bar = 0.5 mm. E, ventral view of the mouth area; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.10131909" httpUri="https://zenodo.org/record/10131909/files/figure.png" pageId="36" pageNumber="571">Fig. 20D</figureCitation>
) the cerebral and pleural ganglia are fused and distinct from the pedal ganglia. There are four cerebral nerves leading from each cerebral ganglion and three pleural nerves leading from each pleural ganglion. There is no separate abdominal ganglion on the right side of the visceral loop. The buccal ganglia are near to the rest of the central nervous system, joined to the cerebral ganglia by two relatively long nerves. Gastro-oesophageal, rhinophoral and optical ganglia are present. The pedal ganglia are clearly separated, having two nerves leading from the left ganglion and three from the right one. The pedal and parapedal commissures are enveloped together with the visceral loop.
</paragraph>
<paragraph id="8BA3368C75736C65937DFE5F20FDAA12" blockId="36.[806,1423,196,555]" pageId="36" pageNumber="571">
The circulatory system (
<figureCitation id="13272A0975736C659411FE5F20F5A9D7" box="[1107,1205,472,494]" captionStart="Figure 20" captionStartId="38.[155,233,1789,1808]" captionTargetBox="[166,1426,204,1755]" captionTargetId="figure-83@38.[155,1435,195,1757]" captionTargetPageId="38" captionText="Figure 20. Discodoris ketos (CASIZ 081808). A, general view of the anatomy; scale bar = 1 mm. B, reproductive system; scale bar = 1 mm. C, lateral view of the buccal bulb; scale bar = 1 mm. D, central nervous system; scale bar = 0.5 mm. E, ventral view of the mouth area; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.10131909" httpUri="https://zenodo.org/record/10131909/files/figure.png" pageId="36" pageNumber="571">Fig. 20A</figureCitation>
) consists of a large heart and two blood glands situated in front of and behind the central nervous system.
</paragraph>
<paragraph id="8BA3368C75736C659364FDF627C8AABF" blockId="36.[806,1423,625,1199]" box="[806,904,625,646]" pageId="36" pageNumber="571">
<emphasis id="B968EA9E75736C659364FDF627C8AABF" box="[806,904,625,646]" italics="true" pageId="36" pageNumber="571">Remarks</emphasis>
</paragraph>
<paragraph id="8BA3368C75736C659364FD17209DABA2" blockId="36.[806,1423,625,1199]" pageId="36" pageNumber="571">
<bibRefCitation id="EF8D4B7D75736C659364FD172015AA9F" author="Marcus Er &amp; Marcus Ev" box="[806,1109,656,678]" pageId="36" pageNumber="571" pagination="155 - 179" refId="ref57219" refString="Marcus Er, Marcus Ev. 1970 a. Opisthobranch mollusks from the Southern tropical Pacific. Pacific Science 24: 155 - 179." type="journal article" year="1970">Marcus &amp; Marcus (1970a)</bibRefCitation>
described the new subspecies
<taxonomicName id="4C1C4D0F75736C659319FD282067AAFD" authorityName="Er. Marcus &amp; Ev. Marcus" authorityYear="1970" box="[859,1063,686,708]" class="Gastropoda" family="Discodorididae" genus="Tayuva" kingdom="Animalia" order="Nudibranchia" pageId="36" pageNumber="571" phylum="Mollusca" rank="subSpecies" species="ketos" subSpecies="juva">
<emphasis id="B968EA9E75736C659319FD282067AAFD" box="[859,1063,686,708]" italics="true" pageId="36" pageNumber="571">Tayuva ketos juva</emphasis>
</taxonomicName>
from the tropical Indo-Pacific. The description, based on a single preserved specimen, is not complete and does not include detailed anatomical information. It is very likely that this description corresponds to a tropical species of
<taxonomicName id="4C1C4D0F75736C659523FCAD2735AB64" authorityName="Bergh" authorityYear="1877" class="Gastropoda" family="Discodorididae" genus="Discodoris" kingdom="Animalia" order="Nudibranchia" pageId="36" pageNumber="571" phylum="Mollusca" rank="genus">
<emphasis id="B968EA9E75736C659523FCAD2735AB64" italics="true" pageId="36" pageNumber="571">Discodoris</emphasis>
</taxonomicName>
, rather than a subspecies of
<taxonomicName id="4C1C4D0F75736C659490FCCF21C7AB64" box="[1234,1415,840,861]" class="Gastropoda" family="Discodorididae" genus="Discodoris" kingdom="Animalia" order="Nudibranchia" pageId="36" pageNumber="571" phylum="Mollusca" rank="species" species="ketos">
<emphasis id="B968EA9E75736C659490FCCF21C7AB64" box="[1234,1415,840,861]" italics="true" pageId="36" pageNumber="571">Discodoris ketos</emphasis>
</taxonomicName>
, which is probably restricted to the Panamic biogeographical region of the eastern Pacific
</paragraph>
<paragraph id="8BA3368C75736C65937DFC2320AEAC96" blockId="36.[806,1423,625,1199]" pageId="36" pageNumber="571">
Another subspecies of
<taxonomicName id="4C1C4D0F75736C659402FC2320B2AB80" box="[1088,1266,932,953]" class="Gastropoda" family="Discodorididae" genus="Discodoris" kingdom="Animalia" order="Nudibranchia" pageId="36" pageNumber="571" phylum="Mollusca" rank="species" species="ketos">
<emphasis id="B968EA9E75736C659402FC2320B2AB80" box="[1088,1266,932,953]" italics="true" pageId="36" pageNumber="571">Discodoris ketos</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1C4D0F75736C6594BCFC222711ABEE" authorityName="Ev.Marcus &amp; Er.Marcus" authorityYear="1970" class="Gastropoda" family="Discodorididae" genus="Tayuva" kingdom="Animalia" order="Nudibranchia" pageId="36" pageNumber="571" phylum="Mollusca" rank="subSpecies" species="ketos" subSpecies="gila">
<emphasis id="B968EA9E75736C6594BCFC222711ABEE" italics="true" pageId="36" pageNumber="571">Tayuva ketos gila</emphasis>
</taxonomicName>
, was described by
<bibRefCitation id="EF8D4B7D75736C659463FC442107ABE1" author="Marcus Er &amp; Marcus Ev" box="[1057,1351,962,984]" pageId="36" pageNumber="571" pagination="1 - 129" refId="ref57244" refString="Marcus Er, Marcus Ev. 1970 b. Opisthobranchs from Curacao and faunistically related regions. Studies on the Fauna of Curacao and Other Caribbean Islands 33: 1 - 129." type="journal article" year="1970">Marcus &amp; Marcus (1970b)</bibRefCitation>
based on material from
<collectingCountry id="F30B761C75736C6593ADFC662010ABCF" box="[1007,1104,993,1014]" name="Curaçao" pageId="36" pageNumber="571">Curaçao</collectingCountry>
, Caribbean Sea. Again, it is difficult to determine the identity of the preserved animals they saw, but it is unlikely that they belong to the same species.
<bibRefCitation id="EF8D4B7D75736C659392FBB92149AC6A" author="Marcus Er &amp; Marcus Ev" box="[976,1289,1085,1107]" pageId="36" pageNumber="571" pagination="1 - 129" refId="ref57244" refString="Marcus Er, Marcus Ev. 1970 b. Opisthobranchs from Curacao and faunistically related regions. Studies on the Fauna of Curacao and Other Caribbean Islands 33: 1 - 129." type="journal article" year="1970">Marcus &amp; Marcus (1970b)</bibRefCitation>
mentioned the presence of denticles on the innermost teeth of the
<specimenCount id="9D1AFD0575736C659364FBFC278AACA9" box="[806,970,1146,1168]" pageId="36" pageNumber="571" type="generic">two specimens</specimenCount>
of
<taxonomicName id="4C1C4D0F75736C6593A9FBFC20EBACB6" authorityName="Ev.Marcus &amp; Er.Marcus" authorityYear="1970" box="[1003,1195,1146,1168]" class="Gastropoda" family="Discodorididae" genus="Tayuva" kingdom="Animalia" order="Nudibranchia" pageId="36" pageNumber="571" phylum="Mollusca" rank="subSpecies" species="ketos" subSpecies="gila">
<emphasis id="B968EA9E75736C6593A9FBFC20EBACB6" box="[1003,1195,1146,1168]" italics="true" pageId="36" pageNumber="571">Tayuva ketos gila</emphasis>
</taxonomicName>
; these are absent in the Panamic specimens examined here.
</paragraph>
</subSubSection>
</treatment>
</document>
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