treatments-xml/data/03/91/EC/0391EC25FFCE0B499A9CFD34FE6EFEF9.xml

<document ID-DOI="10.11646/zootaxa.5264.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="7836130" ID-ZooBank="1F30E79F-6E77-46AE-A537-098A9549E628" approvalRequired="8" approvalRequired_for_taxonomicNames="5" approvalRequired_for_textStreams="3" checkinTime="1681725752866" checkinUser="plazi" docAuthor="Sittichaya, Wisut, Smith, Sarah M. &amp; Beaver, Roger A." docDate="2023" docId="0391EC25FFCE0B499A9CFD34FE6EFEF9" docLanguage="en" docName="zootaxa.5264.1.3.pdf" docOrigin="Zootaxa 5264 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5264.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Webbia granulosa Sittichaya, Smith &amp; Beaver 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="53" masterDocId="FFA8945DFFCA0B4F9A0BFF86FFBDFFE4" masterDocTitle="New species, taxonomic changes and newly recorded species of Webbia Hopkins, ambrosia beetles from Thailand and neighbouring countries (Coleoptera: Curculionidae: Scolytinae: Xyleborini)" masterLastPageNumber="63" masterPageNumber="47" pageNumber="51" updateTime="1681726304425" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New species, taxonomic changes and newly recorded species of Webbia Hopkins, ambrosia beetles from Thailand and neighbouring countries (Coleoptera: Curculionidae: Scolytinae: Xyleborini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sittichaya, Wisut</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-6200-1285</mods:nameIdentifier>
<mods:affiliation>Agricultural Innovation and Management Division, Faculty of Natural Resources, Prince of Songkla University, Songkhla, 90110, Thailand. wanakorn 62 @ hotmail. com; https: // orcid. org / 0000 - 0001 - 6200 - 1285</mods:affiliation>
<mods:nameIdentifier type="email">wanakorn62@hotmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Smith, Sarah M.</mods:namePart>
<mods:affiliation>Department of Entomology, Michigan State University, 288 Farm Lane, 243 Natural Science Bldg., East Lansing, MI 48824, USA.</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Beaver, Roger A.</mods:namePart>
<mods:affiliation>161 / 2 Mu 5, Soi Wat Pranon, T. Donkaew, A. Maerim, Chiangmai 50180, Thailand.</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-04-12</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5264</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>47</mods:start>
<mods:end>63</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5264.1.3</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5264.1.3</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7836130</mods:identifier>
<mods:identifier type="ZooBank">1F30E79F-6E77-46AE-A537-098A9549E628</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7836154" ID-Zenodo-Dep="7836154" LSID="urn:lsid:plazi:treatment:0391EC25FFCE0B499A9CFD34FE6EFEF9" httpUri="http://treatment.plazi.org/id/0391EC25FFCE0B499A9CFD34FE6EFEF9" lastPageId="6" lastPageNumber="53" pageId="4" pageNumber="51">
<subSubSection box="[151,820,690,717]" pageId="4" pageNumber="51" type="nomenclature">
<paragraph blockId="4.[151,820,690,753]" box="[151,820,690,717]" pageId="4" pageNumber="51">
<heading bold="true" box="[151,820,690,717]" fontSize="11" level="1" pageId="4" pageNumber="51" reason="1">
<emphasis bold="true" box="[151,820,690,717]" pageId="4" pageNumber="51">
<taxonomicName authority="Sittichaya, Smith &amp; Beaver" authorityName="Sittichaya, Smith &amp; Beaver" authorityYear="2023" box="[151,719,690,717]" class="Insecta" family="Curculionidae" genus="Webbia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="granulosa" status="sp. nov.">
<emphasis bold="true" box="[151,370,690,717]" italics="true" pageId="4" pageNumber="51">Webbia granulosa</emphasis>
Sittichaya, Smith &amp; Beaver
</taxonomicName>
<taxonomicNameLabel box="[727,820,691,717]" pageId="4" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,217,727,753]" pageId="4" pageNumber="51" type="description">
<paragraph blockId="4.[151,820,690,753]" box="[151,217,727,753]" pageId="4" pageNumber="51">
<figureCitation box="[151,217,727,753]" captionStart="FIGURE 2" captionStartId="5.[151,250,1912,1936]" captionTargetBox="[182,1432,563,1889]" captionTargetId="figure-188@5.[155,1433,561,1889]" captionTargetPageId="5" captionText="FIGURE 2. Webbia granulosa sp. nov. holotype A) dorsal view; B) lateral view; C) postero-lateral view; D) frons; E) head lateral and antenna; F) declivital face." figureDoi="http://doi.org/10.5281/zenodo.7836136" httpUri="https://zenodo.org/record/7836136/files/figure.png" pageId="4" pageNumber="51">Fig. 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="51" type="materials_examined">
<paragraph blockId="4.[151,1437,798,2013]" pageId="4" pageNumber="51">
<emphasis bold="true" box="[151,321,798,825]" pageId="4" pageNumber="51">Type material.</emphasis>
<materialsCitation collectingDate="2020-03-01" collectionCode="NHMW" collectorName="W. Sittichaya" country="Thailand" latitude="17.065416" location="Phu Pan National Park" longLatPrecision="1" longitude="103.969475" pageId="4" pageNumber="51" specimenCount="1" stateProvince="Sakon Nakhon" typeStatus="holotype">
<emphasis bold="true" box="[325,436,798,825]" pageId="4" pageNumber="51">
<emphasis bold="true" box="[325,429,798,825]" italics="true" pageId="4" pageNumber="51">
<typeStatus box="[325,429,798,825]" pageId="4" pageNumber="51">Holotype</typeStatus>
</emphasis>
,
</emphasis>
female, 
<collectingCountry box="[526,674,799,825]" name="Thailand" pageId="4" pageNumber="51">THAILAND</collectingCountry>
, 
<collectingRegion box="[681,846,799,825]" country="Thailand" name="Sakon Nakhon" pageId="4" pageNumber="51">Sakon Nakhon</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:0391EC25FFCE0B499A9CFD34FE6EFEF9:8EE70BE8FFCE0B4B995DFC99FBF1FCDD" box="[854,1100,799,825]" country="Thailand" latitude="17.065416" longLatPrecision="1" longitude="103.969475" name="Phu Pan National Park" pageId="4" pageNumber="51" stateProvince="Sakon Nakhon">Phu Pan National Park</location>
, 
<geoCoordinate box="[1108,1262,800,827]" degrees="17" direction="north" minutes="03" orientation="latitude" pageId="4" pageNumber="51" precision="1" seconds="55.5" value="17.065416">17°03′55.5&quot;N</geoCoordinate>
<geoCoordinate box="[1265,1431,798,825]" degrees="103" direction="east" minutes="58" orientation="longitude" pageId="4" pageNumber="51" precision="1" seconds="10.1" value="103.969475">103°58'10.1&quot;E</geoCoordinate>
, dry deciduous dipterocarp forest, 340msl, 
<collectingMethod box="[647,866,835,861]" pageId="4" pageNumber="51">ethanol baited trap</collectingMethod>
, 
<date box="[882,1006,835,861]" pageId="4" pageNumber="51" value="2020-03-01">
<collectingDate box="[882,1006,835,861]" pageId="4" pageNumber="51" value="2020-03-01">01.iii.2020</collectingDate>
</date>
, 
<collectorName box="[1021,1176,835,861]" pageId="4" pageNumber="51">W. Sittichaya</collectorName>
(
<collectionCode box="[1195,1294,835,861]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="4" pageNumber="51" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
. 
<materialsCitation collectionCode="WSTC, MSUC, RABC" country="Thailand" pageId="4" pageNumber="51" specimenCount="2" stateProvince="Sakon Nakhon" typeStatus="paratype">
<emphasis bold="true" box="[1316,1430,835,861]" italics="true" pageId="4" pageNumber="51">
<typeStatus box="[1316,1430,835,861]" pageId="4" pageNumber="51">Paratypes</typeStatus>
</emphasis>
, female, as 
<typeStatus box="[272,370,871,897]" pageId="4" pageNumber="51">holotype</typeStatus>
(4) (1, 
<collectionCode box="[453,533,871,897]" pageId="4" pageNumber="51">WSTC</collectionCode>
) (1, 
<collectionCode box="[585,666,871,897]" pageId="4" pageNumber="51">MSUC</collectionCode>
) (1, 
<collectionCode box="[718,796,871,897]" pageId="4" pageNumber="51">RABC</collectionCode>
)
</materialsCitation>
<materialsCitation box="[810,963,871,897]" collectionCode="THNHM" country="Thailand" isTailEnumeration="true" pageId="4" pageNumber="51" specimenCount="1" stateProvince="Sakon Nakhon" typeStatus="paratype">
(1, 
<collectionCode box="[847,955,871,897]" pageId="4" pageNumber="51">THNHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="51" type="diagnosis">
<paragraph blockId="4.[151,1437,798,2013]" pageId="4" pageNumber="51">
<emphasis bold="true" box="[199,322,906,932]" pageId="4" pageNumber="51">Diagnosis.</emphasis>
<quantity box="[331,497,906,933]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.15" metricValueMin="1.75" pageId="4" pageNumber="51" unit="mm" value="1.95" valueMax="2.15" valueMin="1.75">1.75–2.15 mm</quantity>
long (mean = 
<quantity box="[671,776,907,933]" metricMagnitude="-3" metricUnit="m" metricValue="1.92" pageId="4" pageNumber="51" unit="mm" value="1.92">1.92 mm</quantity>
, n=5); 2.92–3.09× as long as wide (mean = 2.97×, n=5). This species is distinguished by the elytral declivity subshining, declivital face flat not impressed, circumdeclivital margin with small, acute denticles; declivital striae and interstriae granulate, granules small, stronger on interstriae, bearing short, brownish, hair‒like setae; posterolateral declivital margin with a short acute process; elytral apex with small, triangular processes.
</paragraph>
<paragraph blockId="4.[151,1437,798,2013]" pageId="4" pageNumber="51">
The species is similar to 
<taxonomicName authority="Sittichaya &amp; Smith &amp; Beaver, 2023" authorityName="Sittichaya &amp; Smith &amp; Beaver" authorityYear="2023" box="[466,650,1087,1113]" class="Insecta" family="Curculionidae" genus="Webbia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="aculeata" status="sp. nov.">
<emphasis box="[466,650,1087,1113]" italics="true" pageId="4" pageNumber="51">Webbia aculeata</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[655,740,1087,1113]" pageId="4" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
, 
<taxonomicName authorityName="Schedl Beaver &amp; Browne" authorityYear="1975" box="[751,871,1087,1112]" class="Insecta" family="Curculionidae" genus="Webbia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis box="[751,871,1087,1112]" italics="true" pageId="4" pageNumber="51">W. cornuta</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Browne" authorityYear="1986" box="[921,1066,1087,1113]" class="Insecta" family="Curculionidae" genus="Webbia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="hatanakai">
<emphasis box="[921,1066,1087,1113]" italics="true" pageId="4" pageNumber="51">W. hatanakai</emphasis>
</taxonomicName>
. 
<taxonomicName authorityName="Sittichaya &amp; Smith &amp; Beaver" authorityYear="2023" box="[1078,1206,1087,1113]" class="Insecta" family="Curculionidae" genus="Webbia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="aculeata">
<emphasis box="[1078,1206,1087,1113]" italics="true" pageId="4" pageNumber="51">W. aculeata</emphasis>
</taxonomicName>
can be distinguished by the much larger size and greater length of the processes on the posterolateral margin of the elytral declivity (
<figureCitation captionStart="FIGURE 1" captionStartId="3.[151,250,1540,1564]" captionTargetBox="[153,1435,182,1515]" captionTargetId="figure-18@3.[152,1436,181,1516]" captionTargetPageId="3" captionText="FIGURE 1. Webbia aculeata sp. nov. holotype A) dorsal view; B) lateral view; C) postero-lateral view; D) frons; E) venter and antennae; F) declivital face." figureDoi="http://doi.org/10.5281/zenodo.7836134" httpUri="https://zenodo.org/record/7836134/files/figure.png" pageId="4" pageNumber="51">Fig. 1</figureCitation>
); 
<taxonomicName authorityName="Browne" authorityYear="1986" box="[193,343,1159,1185]" class="Insecta" family="Curculionidae" genus="Webbia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="hatanakai">
<emphasis box="[193,343,1159,1185]" italics="true" pageId="4" pageNumber="51">W. hatanakai</emphasis>
</taxonomicName>
by the very coarse, closely placed strial punctures on the elytral declivity; 
<taxonomicName authorityName="Schedl Beaver &amp; Browne" authorityYear="1975" box="[1227,1353,1159,1184]" class="Insecta" family="Curculionidae" genus="Webbia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis box="[1227,1353,1159,1184]" italics="true" pageId="4" pageNumber="51">W. cornuta</emphasis>
</taxonomicName>
by the smooth, shining elytral declivity lacking granules.
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="52" pageId="4" pageNumber="51" type="description">
<paragraph blockId="4.[151,1437,798,2013]" lastBlockId="5.[151,1437,151,537]" lastPageId="5" lastPageNumber="52" pageId="4" pageNumber="51">
<emphasis bold="true" box="[199,446,1230,1256]" pageId="4" pageNumber="51">Description (female)</emphasis>
<quantity box="[456,622,1230,1257]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.15" metricValueMin="1.75" pageId="4" pageNumber="51" unit="mm" value="1.95" valueMax="2.15" valueMin="1.75">1.75–2.15 mm</quantity>
long (mean = 
<quantity box="[798,904,1231,1257]" metricMagnitude="-3" metricUnit="m" metricValue="1.92" pageId="4" pageNumber="51" unit="mm" value="1.92">1.92 mm</quantity>
; n = 5); 2.92–3.09× as long as wide (mean = 2.97, n = 5). Frons, pronotum and elytral bases dark brown (brown in 
<specimenCount box="[959,1122,1267,1293]" pageId="4" pageNumber="51" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
), elytral disc, abdomen and appendages pale yellowish, declivital summit and declivity ferruginous. 
<emphasis bold="true" box="[970,1042,1303,1329]" pageId="4" pageNumber="51">Head:</emphasis>
epistoma entire, transverse, with a row of short, hair-like setae. Frons feebly convex to upper level of eyes, shining, shagreened, with feebly elevated, vertical median line extending nearly to vertex, sparsely, shallowly punctured below upper level of eyes, each puncture with an erect hair-like seta, setae much finer than those on epistoma. Eyes moderately deeply emarginate just above antennal insertion, upper and lower portions subequal in size. Submentum triangular, deeply impressed. Antennal scape thickened, slightly longer than club. Pedicel narrower than scape, 2× longer than the 2-segmented funicle. Club flattened, circular, as broad as tall (slightly deformed, appearing taller in 
<typeStatus box="[1118,1220,1519,1545]" pageId="4" pageNumber="51">holotype</typeStatus>
), type 2, segment 1 corneous, corneous portion small, occupying approximately ¼ of club, its margin concave on anterior face; segments 1 and 2 visible on posterior face, segment 2 narrow, corneous part an anterior side slender. 
<emphasis bold="true" box="[1171,1301,1591,1617]" pageId="4" pageNumber="51">Pronotum:</emphasis>
1.25‒1.31× as long as wide (mean = 1.28, n = 5). In dorsal view elongate, quadrate frontally, type a, sides parallel in basal 4/5; anterior margin without serrations. In lateral view elongate, flattened, bulging frontally, type b, disc flat, summit approximately at anterior 1/5. Anterior slope subshining, with transverse, fine, narrow asperities, becoming lower and more strongly transverse towards summit, apical half bearing short, fine, semi-recumbent hair-like setae; setae longest at anterior margin becoming shorter posteriorly. Disc subshining, alutaceous, moderately densely, finely punctate, punctures with a very short, pale yellowish hair-like seta. Lateral margins weakly carinate. Base feebly bisinuate, margin oblique, posterior angles angulate. 
<emphasis bold="true" box="[762,891,1842,1868]" pageId="4" pageNumber="51">Scutellum:</emphasis>
not visible from above. 
<emphasis bold="true" box="[1180,1265,1843,1869]" pageId="4" pageNumber="51">Elytra:</emphasis>
1.53‒1.73× as long as wide (mean = 1.60, n = 5), 1.18‒1.37× as long as pronotum (mean = 1.27×, n = 5). Elytral base V‒shaped, concave, with tuft of mycangial setae along the margin, edge costate, humeral angles rounded; sides subparallel in basal 2/3, then angulate to apex. Elytra smooth, alutaceous, except near scutellum and declivital summit. Disc convex, shining, subtransparent, striae punctate, punctures minute, inconspicuous, separated by 1–1.5× diameters of a puncture, each puncture with a very short, fine hair‒like seta, setae longer toward lateral margin; interstriae flat, 2× as wide as striae. Declivital summit dull, weakly, irregularly rugulose; declivity abruptly truncate, declivital margins armed with short, pointed denticles, each denticle with several moderately long brownish hair‒like setae. Declivital face flat not impressed, granulate, subshining; striae weakly granulate-punctate, interstriae granulate, granules larger than on striae, all granules with a short, strong, semi-recumbent seta; strial and interstrial setae subequal in size, but many interstrial setae slightly flattened. Posterolateral margin of declivity armed with a short, acutely pointed process on each side, processes weakly curved towards midline. Apical margin with a short, triangular process on each side next to suture. 
<emphasis bold="true" box="[432,498,403,429]" pageId="5" pageNumber="52">Legs:</emphasis>
procoxae contiguous. Protibiae slender, broadest at middle; posterior face inflated, granulate; outer margin with ten short and small socketed denticles, their length shorter than basal width. Meso- and metatibiae with outer margin rounded, broadest at middle, each with nine small, socketed denticles, posterior faces flat, unarmed; anterior faces very finely granulate.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7836136" ID-Zenodo-Dep="7836136" httpUri="https://zenodo.org/record/7836136/files/figure.png" pageId="5" pageNumber="52" startId="5.[151,250,1912,1936]" targetBox="[182,1432,563,1889]" targetPageId="5">
<paragraph blockId="5.[151,1436,1912,1973]" pageId="5" pageNumber="52">
<emphasis bold="true" box="[151,277,1912,1936]" pageId="5" pageNumber="52">FIGURE 2.</emphasis>
<taxonomicName authority="Sittichaya &amp; Smith &amp; Beaver, 2023" authorityName="Sittichaya &amp; Smith &amp; Beaver" authorityYear="2023" box="[285,471,1912,1936]" class="Insecta" family="Curculionidae" genus="Webbia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="granulosa" status="sp. nov.">
<emphasis box="[285,471,1912,1936]" italics="true" pageId="5" pageNumber="52">Webbia granulosa</emphasis>
</taxonomicName>
<emphasis bold="true" box="[479,561,1913,1937]" pageId="5" pageNumber="52">
<taxonomicNameLabel box="[479,561,1913,1937]" pageId="5" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
holotype A) dorsal view; B) lateral view; C) postero-lateral view; D) frons; E) head lateral and antenna; F) declivital face.
</paragraph>
</caption>
<subSubSection pageId="6" pageNumber="53" type="etymology">
<paragraph blockId="6.[151,1436,151,285]" pageId="6" pageNumber="53">
<emphasis bold="true" box="[199,334,151,177]" pageId="6" pageNumber="53">Etymology.</emphasis>
L. 
<emphasis box="[374,498,151,177]" italics="true" pageId="6" pageNumber="53">granulosus</emphasis>
= with small granules. Adjective. The name refers to the granules on the declivital striae and interstriae.
</paragraph>
</subSubSection>
<subSubSection box="[199,657,222,249]" pageId="6" pageNumber="53" type="distribution">
<paragraph blockId="6.[151,1436,151,285]" box="[199,657,222,249]" pageId="6" pageNumber="53">
<emphasis bold="true" box="[199,353,222,248]" pageId="6" pageNumber="53">Distribution.</emphasis>
<collectingCountry box="[359,459,223,249]" name="Thailand" pageId="6" pageNumber="53">Thailand</collectingCountry>
(
<collectingRegion box="[473,646,223,249]" country="Thailand" name="Sakon Nakhon" pageId="6" pageNumber="53">Sakon Nakhon</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection box="[199,467,259,285]" pageId="6" pageNumber="53" type="biology_ecology">
<paragraph blockId="6.[151,1436,151,285]" box="[199,467,259,285]" pageId="6" pageNumber="53">
<emphasis bold="true" box="[199,342,259,285]" pageId="6" pageNumber="53">Host plants.</emphasis>
Unknown.
</paragraph>
</subSubSection>
</treatment>
</document>