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143 lines
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<document id="87CEF33C0B6EFC4511186A639CD70605" ID-DOI="10.1206/3916.1" ID-ISSN="0003-0082" ID-Zenodo-Dep="5369804" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630265248043" checkinUser="carolina" docAuthor="Morgan, Gary S., Albury, Nancy A., Rímoli, Renato, Lehman, Phillip, Rosenberger, Alfred L. & Cooke, Siobhán B." docDate="2018" docId="B871FF4B7E33D34EFE26FD24F95E91B5" docLanguage="en" docName="N3916 highres.pdf" docOrigin="American Museum Novitates 2018 (3916)" docSource="https://bioone.org/journals/american-museum-novitates/volume-2018/issue-3916/3916.1/The-Cuban-Crocodile-Crocodylus-rhombifer-from-Late-Quaternary-Underwater-Cave/10.1206/3916.1.full" docStyle="DocumentStyle:26C3580A17D4BBE58BD8D8AC6AE09AC4.6:AmMusNovit.2010-.journal_article" docStyleId="26C3580A17D4BBE58BD8D8AC6AE09AC4" docStyleName="AmMusNovit.2010-.journal_article" docStyleVersion="6" docTitle="Crocodylus rhombifer Cuvier 1807" docType="treatment" docVersion="3" masterDocId="444887337E38D345FFC7FF83FB46975C" masterDocTitle="The Cuban Crocodile (Crocodylus rhombifer) from Late Quaternary Underwater Cave Deposits in the Dominican Republic" masterLastPageNumber="1" masterPageNumber="1" pageNumber="1" updateTime="1699204417027" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="03093570F180F25FCF25ED759C6B6D4A">
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<mods:title id="0922900E08D43A6B74518E7322739484">The Cuban Crocodile (Crocodylus rhombifer) from Late Quaternary Underwater Cave Deposits in the Dominican Republic</mods:title>
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<mods:namePart id="72D2BD35670B9D3D2E1EB4CF56E6817F">Morgan, Gary S.</mods:namePart>
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<mods:namePart id="F0756FDCE73889297C2EF02187BB63B4">Albury, Nancy A.</mods:namePart>
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<mods:namePart id="2DA436D7BDDA98C0869A04BC53C36A0D">Rímoli, Renato</mods:namePart>
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<mods:namePart id="CDB6F2C982277DA07E073157CED87374">Lehman, Phillip</mods:namePart>
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<mods:namePart id="9B744453055026F9CCFD0E6543C4DB73">Rosenberger, Alfred L.</mods:namePart>
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<mods:namePart id="1DD5FB4996D15108FE467E61332E3FD8">Cooke, Siobhán B.</mods:namePart>
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<mods:typeOfResource id="EAF92DE427E9BFD53BB6799294D863AD">text</mods:typeOfResource>
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<mods:title id="11B906E335B9FDA0D8462F210F82BF58">American Museum Novitates</mods:title>
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</mods:titleInfo>
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<mods:part id="7A185571C1E91F667B42D00ACB371505">
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<mods:date id="522D3D89C5D19B402E95A5123F1256DF">2018</mods:date>
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<mods:number id="48A266EC2EF155AB2D522C2A35DB312E">2018-12-21</mods:number>
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<mods:detail id="CB5AA3907F59C286F8E0EBB01FD4318D" type="volume">
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<mods:number id="83BF4AF9A5D934F27AA7A6EA5E23C69E">2018</mods:number>
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<mods:detail id="76C8610E58A71E3661A36515467DC6D8" type="issue">
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<mods:number id="6C709DFBA88A73A7E10E0202A3EFD5C0">3916</mods:number>
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<mods:start id="478FA039108F32ECF563C4623A0ACC84">1</mods:start>
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<mods:location id="2817249D9A06972DA2C7F280EAF8003E">
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<mods:url id="B199E52A876160F701A94E757609D9E9">https://bioone.org/journals/american-museum-novitates/volume-2018/issue-3916/3916.1/The-Cuban-Crocodile-Crocodylus-rhombifer-from-Late-Quaternary-Underwater-Cave/10.1206/3916.1.full</mods:url>
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<mods:classification id="1EF4C7587457BE75B95A9DEBDAEA3594">journal article</mods:classification>
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<mods:identifier id="5A12122B31A7CC3CD45B1DF58F381E48" type="DOI">10.1206/3916.1</mods:identifier>
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<mods:identifier id="355D5D8EB1A04171DCC9BA1FDF9C3948" type="ISSN">0003-0082</mods:identifier>
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<mods:identifier id="6F5862BAECCB913E46E69B4137DB9541" type="Zenodo-Dep">5369804</mods:identifier>
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<treatment id="B871FF4B7E33D34EFE26FD24F95E91B5" LSID="urn:lsid:plazi:treatment:B871FF4B7E33D34EFE26FD24F95E91B5" httpUri="http://treatment.plazi.org/id/B871FF4B7E33D34EFE26FD24F95E91B5" pageId="11">
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<subSubSection id="78C21DD67E33D34EFE26FD24F85795A5" pageId="11" type="nomenclature">
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<paragraph id="30674E5D7E33D34EFE26FD24F831959D" blockId="11.[481,887,679,705]" box="[481,887,679,705]" pageId="11">
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<heading id="6B2FF9317E33D34EFE26FD24F831959D" box="[481,887,679,705]" centered="true" fontSize="11" level="2" pageId="11" reason="5">
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<taxonomicName id="F7D835DE7E33D34EFE26FD24F831959D" authority="Cuvier, 1807" authorityName="Cuvier" authorityYear="1807" box="[481,887,679,705]" class="Reptilia" family="Crocodylidae" genus="Crocodylus" kingdom="Animalia" order="Crocodylia" pageId="11" phylum="Chordata" rank="species" species="rhombifer">
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<emphasis id="02AC924F7E33D34EFE26FD24F99F959D" box="[481,729,679,705]" italics="true" pageId="11">Crocodylus rhombifer</emphasis>
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Cuvier, 1807
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="30674E5D7E33D34EFD80FD5CF85795A5" blockId="11.[583,785,735,761]" box="[583,785,735,761]" pageId="11">Cuban Crocodile</paragraph>
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</subSubSection>
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<subSubSection id="78C21DD67E33D34EFDA7FC94F95E91B5" pageId="11" type="description">
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<paragraph id="30674E5D7E33D34EFDA7FC94F9BE946D" blockId="11.[608,760,791,817]" box="[608,760,791,817]" pageId="11">
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<heading id="6B2FF9317E33D34EFDA7FC94F9BE946D" box="[608,760,791,817]" centered="true" fontSize="36" level="3" pageId="11" reason="7">
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<figureCitation id="A8E352D87E33D34EFDA7FC94F9BE946D" box="[608,760,791,817]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStart-3="FIGURE 9" captionStart-4="FIGURE 10" captionStartId-0="13.[132,218,1622,1643]" captionStartId-1="14.[132,218,1687,1708]" captionStartId-2="15.[132,218,1701,1722]" captionStartId-3="25.[132,218,1476,1497]" captionStartId-4="26.[132,218,953,974]" captionTargetBox-0="[147,1227,248,1592]" captionTargetBox-1="[132,1182,248,1661]" captionTargetBox-2="[132,1236,248,1675]" captionTargetBox-3="[132,1236,248,1450]" captionTargetBox-4="[172,1195,248,927]" captionTargetId-0="figure-6@13.[147,1227,248,1592]" captionTargetId-1="figure-10@14.[132,1182,248,1661]" captionTargetId-2="figure-6@15.[132,1236,248,1675]" captionTargetId-3="figure-6@25.[132,1236,248,1450]" captionTargetId-4="figure-314@26.[172,1195,248,927]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="15" captionTargetPageId-3="25" captionTargetPageId-4="26" captionText-0="FIGURE 6. Skull and articulated right and left mandibles of a Cuban crocodile (Crocodylus rhombifer) of Late Quaternary age from Ni-Rahu (= Cueva de Lynn), Dominican Republic (MHD 414). The skull and mandibles of this specimen are partially covered with a thick layer of calcite flowstone, obscuring many morphological features. A. Dorsal, B. ventral, C. left lateral, D. right lateral, and E. posterior views." captionText-1="FIGURE 7. Skull of a Cuban crocodile (Crocodylus rhombifer) of Late Quaternary age from Oleg’s Bat Cave, Dominican Republic (MHD 572) in A. dorsal, B. ventral, C. left lateral, and D. posterior views." captionText-2="FIGURE 8. Skull and left mandible of a Cuban crocodile (Crocodylus rhombifer) of Late Quaternary age from Oleg’s Bat Cave, Dominican Republic (MHD 574). Skull in A. dorsal, B. ventral, C. left lateral, and D. posterior views. Left mandible in E. medial and F. lateral views." captionText-3="FIGURE 9. Limb bones of the Cuban crocodile (Crocodylus rhombifer) of Late Quaternary age from Oleg’s Bat Cave, Dominican Republic. All limb bones are from the left side and are from the same associated individual (MHD 576). A. Humerus, B. radius, C. ulna, D. femur, E. tibia, F. fibula. In each pair of photographs, the anterior view is on the left and posterior view on the right." captionText-4="FIGURE 10. Ilia of the Cuban crocodile (Crocodylus rhombifer) of Late Quaternary age from Oleg’s Bat Cave, Dominican Republic (MHD 575). A. External and B. internal views of the right ilium; C. external and D. internal views of the left ilium." figureDoi-0="http://doi.org/10.5281/zenodo.5369816" figureDoi-1="http://doi.org/10.5281/zenodo.5369818" figureDoi-2="http://doi.org/10.5281/zenodo.5369820" figureDoi-3="http://doi.org/10.5281/zenodo.5369822" figureDoi-4="http://doi.org/10.5281/zenodo.5369824" httpUri-0="https://zenodo.org/record/5369816/files/figure.png" httpUri-1="https://zenodo.org/record/5369818/files/figure.png" httpUri-2="https://zenodo.org/record/5369820/files/figure.png" httpUri-3="https://zenodo.org/record/5369822/files/figure.png" httpUri-4="https://zenodo.org/record/5369824/files/figure.png" pageId="11">Figures 6–10</figureCitation>
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</heading>
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</paragraph>
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<paragraph id="30674E5D7E33D34EFF73FCCCF87D9382" blockId="11.[132,1237,847,1769]" pageId="11">
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REFERRED SPECIMENS: The sample of Quaternary crocodile fossils from
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<collectingCountry id="48CF0ECD7E33D34EFBD8FCCCFF939435" box="[1055,1237,847,873]" name="Dominican Republic" pageId="11">the Dominican</collectingCountry>
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Republic consists of about
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<specimenCount id="26DE85D47E33D34EFE00FCF7F93194D2" box="[455,631,884,910]" pageId="11" type="generic">100 specimens</specimenCount>
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, five of which consist of partial to complete skulls with partial associated postcranial skeletons. The following list of specimens includes only the most complete fossil skulls and mandibles of
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<taxonomicName id="F7D835DE7E33D34EFD65FC3CF8DD9485" authorityName="Cuvier" authorityYear="1807" box="[674,923,959,985]" class="Reptilia" family="Crocodylidae" genus="Crocodylus" kingdom="Animalia" order="Crocodylia" pageId="11" phylum="Chordata" rank="species" species="rhombifer">
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<emphasis id="02AC924F7E33D34EFD65FC3CF8DD9485" box="[674,923,959,985]" italics="true" pageId="11">Crocodylus rhombifer</emphasis>
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</taxonomicName>
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from Oleg’s Bat Cave and Ni-Rahu used in the cranial descriptions and comparisons. The appendix provides a more complete list of the fossil sample of
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<taxonomicName id="F7D835DE7E33D34EFD86FB89F99E937F" authorityName="Cuvier" authorityYear="1807" box="[577,728,1033,1060]" class="Reptilia" family="Crocodylidae" genus="Crocodylus" kingdom="Animalia" order="Crocodylia" pageId="11" phylum="Chordata" rank="species" species="rhombifer">
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<emphasis id="02AC924F7E33D34EFD86FB89F9129378" box="[577,596,1034,1060]" italics="true" pageId="11">C</emphasis>
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.
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<emphasis id="02AC924F7E33D34EFDA1FB8AF99E937F" box="[614,728,1033,1059]" italics="true" pageId="11">rhombifer</emphasis>
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</taxonomicName>
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from
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<collectingCountry id="48CF0ECD7E33D34EFCEFFB8AF898937F" box="[808,990,1033,1059]" name="Dominican Republic" pageId="11">the Dominican</collectingCountry>
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Republic, including postcranial material associated with several of the skulls listed here. Isolated postcranial elements from Oleg’s Bat Cave are listed as
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<taxonomicName id="F7D835DE7E33D34EFD9AFBD6F9A99332" authorityName="Cuvier" authorityYear="1807" box="[605,751,1108,1135]" class="Reptilia" family="Crocodylidae" genus="Crocodylus" kingdom="Animalia" order="Crocodylia" pageId="11" phylum="Chordata" rank="species" species="rhombifer">
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<emphasis id="02AC924F7E33D34EFD9AFBD6F9A99332" box="[605,751,1108,1135]" italics="true" pageId="11">C. rhombifer</emphasis>
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</taxonomicName>
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in the appendix. Although most postcranial elements of crocodiles cannot be confidently identified to the species level, it seems reasonable to tentatively refer these specimens to
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<taxonomicName id="F7D835DE7E33D34EFD15FB1CF82393E5" authorityName="Cuvier" authorityYear="1807" box="[722,869,1183,1209]" class="Reptilia" family="Crocodylidae" genus="Crocodylus" kingdom="Animalia" order="Crocodylia" pageId="11" phylum="Chordata" rank="species" species="rhombifer">
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<emphasis id="02AC924F7E33D34EFD15FB1CF82393E5" box="[722,869,1183,1209]" italics="true" pageId="11">C. rhombifer</emphasis>
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</taxonomicName>
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considering that all diagnostic cranial material from this same site represents this species.
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</paragraph>
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<paragraph id="30674E5D7E33D34EFF73FB69F99F9172" blockId="11.[132,1237,847,1769]" pageId="11">
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NI-RAHU: MHD 414. Complete skull and articulated mandibles, and two associated vertebrae. Collected by Phillip Lehman and Cristian Pittaro of the DRSS on
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<date id="4466689D7E33D34EFC1AFA8CFFD89275" box="[989,1182,1295,1321]" pageId="11" value="2013-08-10">August 10, 2013</date>
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. The skull and mandibles from Ni-Rahu are illustrated in figure 6. The left half of the skull from the premaxilla to the quadrate is covered with a thick layer of calcite flowstone. No observations could be made on the palate of this specimen because of the flowstone coating. The mandibles are also covered with flowstone, with only the teeth on the right side visible. Several cranial measurements either could not be taken or are estimates (table 1). The mandibles are firmly attached to the skull, and are so completely covered in flowstone that little of their anatomy is evident, except for the teeth in the right mandible.
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</paragraph>
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<paragraph id="30674E5D7E33D34EFF73F9B9F95E91B5" blockId="11.[132,1237,847,1769]" pageId="11">
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<materialsCitation id="80B044007E33D34EFF73F9B9F95E91B5" collectingDate="2012-08-12" collectionCode="CAVE, DRSS" location="This" municipality="Oleg's Bat Cave" pageId="11" specimenCode="MHD 572" specimenCount="1">
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OLEG’S BAT
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<collectionCode id="56C9D6987E33D34EFE84F9B9FAC59108" box="[323,387,1594,1620]" country="USA" httpUri="http://grbio.org/cool/wvdq-gi6n" name="DOI/NPS, Carlsbad Caverns National Park" pageId="11">CAVE</collectionCode>
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:
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<specimenCode id="607EE6267E33D34EFE57F9B9F94F9108" box="[400,521,1593,1620]" collectionCode="CAVE" country="USA" httpUri="http://grbio.org/cool/wvdq-gi6n" name="DOI/NPS, Carlsbad Caverns National Park" pageId="11">MHD 572</specimenCode>
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. A nearly complete skull lacking only the left premaxilla, right nasal, and the ectopterygoids and palatines (fig. 7).
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<location id="350718867E33D34EFD39F9DCF8769125" LSID="urn:lsid:plazi:treatment:B871FF4B7E33D34EFE26FD24F95E91B5:350718867E33D34EFD39F9DCF8769125" box="[766,816,1631,1657]" municipality="Oleg's Bat Cave" name="This" pageId="11">This</location>
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specimen also includes associated postcranial elements (see appendix). Collected “under mud” on the floor of
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<collectingMunicipality id="D003D4277E33D34EFC3AF907FFF791C2" box="[1021,1201,1668,1695]" pageId="11">Oleg’s Bat Cave</collectingMunicipality>
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by members of the
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<collectionCode id="56C9D6987E33D34EFE83F929FAC19198" box="[324,391,1706,1732]" pageId="11">DRSS</collectionCode>
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on
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<date id="4466689D7E33D34EFE71F929F9319198" box="[438,631,1706,1732]" pageId="11" value="2012-08-12">
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<collectingDate id="542291757E33D34EFE71F929F9319198" box="[438,631,1706,1732]" pageId="11" value="2012-08-12">August 12, 2012</collectingDate>
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</date>
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. A handwritten note in the MHD collection identifies this specimen as “crocodile 1.”
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</materialsCitation>
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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