treatments-xml/data/2D/0A/57/2D0A576DFF9C3127A895B737FCD8FCF7.xml

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<mods:title id="E7B3276EED32126EAD3AAAF28361099B">Two new species of Stenostygnus from South America (Opiliones: Biantidae Stenostygninae)</mods:title>
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<mods:namePart id="ED055D7BE4BC83D771784F04BF8A7A07">Mamani, Claudia Vanesa</mods:namePart>
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<mods:namePart id="8C5602DBC214E0371078633E3FAC86A2">Porto, Willians</mods:namePart>
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<mods:namePart id="9FCAF2C41E2255EFAA90771B8D1CFE5E">Iglesias, Patricia P.</mods:namePart>
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<mods:affiliation id="60C500EF311473EBD9A4FD80131E9AEF">Current address: Laboratorio de Genética Evolutiva, Universidad Nacional de Misiones - CONICET, Félix de Azara 1552, N 3300 LQH, Misiones, Argentina; https: // orcid. org / 0000 - 0002 - 3979 - 4253</mods:affiliation>
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<mods:namePart id="AFC4B7ACD08366985A5A9C8B0B23C6D4">González, Abel Pérez-</mods:namePart>
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<taxonomicName id="62A39DF8FF9C3122A895B737FE25F9B6" ID-CoL="9X5HN" authority="Mamani &amp; Porto &amp; Iglesias &amp; González, 2021" authorityName="Mamani &amp; Porto &amp; Iglesias &amp; González" authorityYear="2021" box="[151,424,1541,1567]" class="Arachnida" family="Biantidae" genus="Stenostygnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="185" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov.">
<emphasis id="97D73A69FF9C3122A895B737FE25F9B6" bold="true" box="[151,424,1541,1567]" italics="true" pageId="3" pageNumber="185">Stenostygnus martensi</emphasis>
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<figureCitation id="3D98FAFEFF9C3122A895B71BFF75F9EA" box="[151,248,1577,1603]" captionStart-0="FIGURES 1" captionStart-1="FIGURES 2" captionStart-2="FIGURES 3" captionStart-3="FIGURES 4" captionStart-4="FIGURES 5" captionStart-5="FIGURES 6" captionStartId-0="2.[151,264,1943,1967]" captionStartId-1="3.[151,264,1399,1423]" captionStartId-2="4.[151,264,1624,1648]" captionStartId-3="5.[151,264,1295,1319]" captionStartId-4="8.[151,264,1283,1307]" captionStartId-5="9.[151,264,1943,1967]" captionTargetBox-0="[172,1415,179,1917]" captionTargetBox-1="[151,1436,181,1375]" captionTargetBox-2="[181,1407,561,1601]" captionTargetBox-3="[173,1414,181,1271]" captionTargetBox-4="[148,1440,181,1259]" captionTargetBox-5="[208,1379,181,1919]" captionTargetId-0="figure-17@2.[172,1415,179,1917]" captionTargetId-1="figure-19@3.[151,1436,181,1375]" captionTargetId-2="figure-240@4.[181,1407,561,1601]" captionTargetId-3="figure-19@5.[173,1414,181,1271]" captionTargetId-4="figure-17@8.[148,1440,181,1259]" captionTargetId-5="figure-19@9.[208,1379,181,1919]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="8" captionTargetPageId-5="9" captionText-0="FIGURES 1A–C. Stenostygnus martensi spec. nov., male (holotype, MIZA 0103460), habitus (A dorsal, B ventral, C lateral). Scale bars: 1 mm." captionText-1="FIGURES 2A–F. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A, B, E= 500µm; C= 200 µm; D= 50 µm; F= 100µm." captionText-2="FIGURES 3A–F. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692). A–C Right chelicera (A ectal, B frontal, C detail of fingers). D–F Right pedipalp (D mesal, E detail of tibia and tarsus, F detail of femur). Scale bars: A, B, E= 200 µm; C, F= 100 µm; D= 500 µm." captionText-3="FIGURES 4A–E. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), right leg IV (A retrolateral, B detail of patella and tibia, C detail of tarsus, D detail of frontal scopula, E detail of lateral scopula). Scale bars: A= 1mm; B= 500µm; C= 200µm; D= 100µm; E= 50µm." captionText-4="FIGURES 5A–G. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), penis (A, E dorsal, B, F lateral, C, G ventral, D detail of conductor and stylus). CO—conductor. LA—lamina apicalis. SO—stylus opening of ductus ejaculatorius. ST—stylus. Ti—Titillator. Scale bars:A= 50 µm; B, C, E-G= 100 µm; D= 20 µm. Stylus in green, conductors in pink and capsula externa in blue (For interpretation of the references to colour the reader is referred to the pdf version of this article)." captionText-5="FIGURES 6A–D. Stenostygnus martensi spec. nov., sexual dimorphism (A–B female paratype, MNRJ 2692, C–D holotype male, MIZA 0103460). A, C Dorsal view. B, D Detail of tibia. Scale bars: A, C= 2 mm; B, D= 500 µm." figureDoi-0="http://doi.org/10.5281/zenodo.4926891" figureDoi-1="http://doi.org/10.5281/zenodo.4926893" figureDoi-2="http://doi.org/10.5281/zenodo.4926895" figureDoi-3="http://doi.org/10.5281/zenodo.4926897" figureDoi-4="http://doi.org/10.5281/zenodo.4926899" figureDoi-5="http://doi.org/10.5281/zenodo.4926903" httpUri-0="https://zenodo.org/record/4926891/files/figure.png" httpUri-1="https://zenodo.org/record/4926893/files/figure.png" httpUri-2="https://zenodo.org/record/4926895/files/figure.png" httpUri-3="https://zenodo.org/record/4926897/files/figure.png" httpUri-4="https://zenodo.org/record/4926899/files/figure.png" httpUri-5="https://zenodo.org/record/4926903/files/figure.png" pageId="3" pageNumber="185">Figs 1–6</figureCitation>
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<emphasis id="97D73A69FF9C3122A895B7BDFEC4F900" bold="true" box="[151,329,1679,1705]" pageId="3" pageNumber="185">Type material.</emphasis>
<materialsCitation id="15CBEC26FF9C3122A950B7BDFEF0F958" ID-GBIF-Occurrence="3335045308" collectionCode="MIZA" collectorName="Osvaldo Villareal &amp; Néstor Sánchez &amp;  Andrés García" country="Venezuela" location="Andres Bello" pageId="3" pageNumber="185" specimenCode="MIZA 0103460" specimenCount="1" stateProvince="Merida" typeStatus="holotype">
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<collectingCountry id="DDB4A6EBFF9C3122A950B7BDFD8FF900" box="[338,514,1679,1705]" name="Venezuela" pageId="3" pageNumber="185">VENEZUELA</collectingCountry>
, 
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: 
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male
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(
<specimenCode id="F5054E00FF9C3122AB3FB7BDFC7AF903" box="[829,1015,1679,1706]" collectionCode="MIZA" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="3" pageNumber="185" type="Museum">MIZA 0103460</specimenCode>
): 
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, vía a 
<collectorName id="085683ADFF9C3122ACFBB7BDFAF9F900" box="[1273,1396,1679,1705]" pageId="3" pageNumber="185">La Azulita</collectorName>
, 
<collectorName id="085683ADFF9C3122AD81B7BDFF71F964" pageId="3" pageNumber="185">El Chorotal</collectorName>
, 
<collectorName id="085683ADFF9C3122A90BB781FE53F964" box="[265,478,1715,1741]" pageId="3" pageNumber="185">Sector El Frontino</collectorName>
, 8.663598°; -71.419705°, 
<date id="D11DC0BBFF9C3122AB09B781FC1DF964" box="[779,912,1715,1741]" pageId="3" pageNumber="185" value="2017-02-03">3 Feb 2017</date>
, 
<quantity id="625B4B9EFF9C3122AB9FB781FC7EF964" box="[925,1011,1715,1741]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="185" unit="m" value="2200.0">2200 m</quantity>
a.s.l., Osvaldo Villareal, Néstor Sánchez, Andrés García
</materialsCitation>
. 
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:
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(
<collectionCode id="C3B27EBEFF9C3122AA69B7E5FD2BF958" box="[619,678,1751,1777]" pageId="3" pageNumber="185">SEM</collectionCode>
voucher), 
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(
<specimenCode id="F5054E00FF9C3122AB98B7E5FBA7F95B" box="[922,1066,1751,1778]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="3" pageNumber="185">MNRJ 2692</specimenCode>
), with same data as for holotype
</materialsCitation>
. 
<materialsCitation id="15CBEC26FF9C3122A895B7C9FCC3F8BC" ID-GBIF-Occurrence="3335045305" box="[151,846,1787,1813]" collectionCode="MIZA" collectorName="Osvaldo Villareal &amp; Néstor Sánchez &amp;  Andrés García" country="Venezuela" location="Andres Bello" pageId="3" pageNumber="185" specimenCount="2" specimenCount-female="2" stateProvince="Paratypes" typeStatus="paratype">
<specimenCount id="B3A52DF2FF9C3122A895B7C9FE8EF8BC" box="[151,259,1787,1813]" pageId="3" pageNumber="185" type="female">2 females</specimenCount>
(MACN-Ar 41022), with same data as for holotype
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.
</paragraph>
</subSubSection>
<subSubSection id="EDB9B5F0FF9C3122A8C5B62DFE37F80C" pageId="3" pageNumber="185" type="etymology">
<paragraph id="A51CE67BFF9C3122A8C5B62DFE37F80C" blockId="3.[151,1437,1678,2029]" pageId="3" pageNumber="185">
<emphasis id="97D73A69FF9C3122A8C5B62DFEC3F890" bold="true" box="[199,334,1823,1849]" pageId="3" pageNumber="185">Etymology.</emphasis>
Patronymic in honor to the German zoologist Jochen Martens, in recognition of his enormous contribution to widen our knowledge of harvestmen. His work helped to found the basis of the new opilionology—as opposed to the old typological approach—and still exerts great influence on new generations of opilionologists; name in the genitive case.
</paragraph>
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<paragraph id="A51CE67BFF9C3125A8C5B69DFCA2FDB0" blockId="3.[151,1437,1678,2029]" lastBlockId="4.[151,1437,151,537]" lastPageId="4" lastPageNumber="186" pageId="3" pageNumber="185">
<emphasis id="97D73A69FF9C3122A8C5B69DFECFF860" bold="true" box="[199,322,1967,1993]" pageId="3" pageNumber="185">Diagnosis.</emphasis>
Differs from all the species of 
<taxonomicName id="62A39DF8FF9C3122AA9BB69DFCA6F860" authorityName="Simon" authorityYear="1879" box="[665,811,1967,1993]" class="Arachnida" family="Biantidae" genus="Stenostygnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="185" phylum="Arthropoda" rank="genus">
<emphasis id="97D73A69FF9C3122AA9BB69DFCA6F860" box="[665,811,1967,1993]" italics="true" pageId="3" pageNumber="185">Stenostygnus</emphasis>
</taxonomicName>
by the following characteristics: absence of granules on mesotergal areas, posterior border of the 
<emphasis id="97D73A69FF9C3122AA6BB6E6FD35F844" box="[617,696,2004,2029]" italics="true" pageId="3" pageNumber="185">scutum</emphasis>
(
<figureCitation id="3D98FAFEFF9C3122AACBB6E1FCA4F844" box="[713,809,2003,2029]" captionStart="FIGURES 2" captionStartId="3.[151,264,1399,1423]" captionTargetBox="[151,1436,181,1375]" captionTargetId="figure-19@3.[151,1436,181,1375]" captionTargetPageId="3" captionText="FIGURES 2A–F. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A, B, E= 500µm; C= 200 µm; D= 50 µm; F= 100µm." figureDoi="http://doi.org/10.5281/zenodo.4926893" httpUri="https://zenodo.org/record/4926893/files/figure.png" pageId="3" pageNumber="185">Fig. 2 A</figureCitation>
) and ventral surface of the coxae III and IV (
<figureCitation id="3D98FAFEFF9C3122AD2FB6E1FA00F844" box="[1325,1421,2003,2029]" captionStart="FIGURES 2" captionStartId="3.[151,264,1399,1423]" captionTargetBox="[151,1436,181,1375]" captionTargetId="figure-19@3.[151,1436,181,1375]" captionTargetPageId="3" captionText="FIGURES 2A–F. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A, B, E= 500µm; C= 200 µm; D= 50 µm; F= 100µm." figureDoi="http://doi.org/10.5281/zenodo.4926893" httpUri="https://zenodo.org/record/4926893/files/figure.png" pageId="3" pageNumber="185">Fig. 2 B</figureCitation>
); males with a retrolateral row of pointed strong tubercles on tibia IV (
<figureCitation id="3D98FAFEFF9B3125ABAEB1A5FBB9FF18" box="[940,1076,151,177]" captionStart="FIGURES 4" captionStartId="5.[151,264,1295,1319]" captionTargetBox="[173,1414,181,1271]" captionTargetId="figure-19@5.[173,1414,181,1271]" captionTargetPageId="5" captionText="FIGURES 4A–E. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), right leg IV (A retrolateral, B detail of patella and tibia, C detail of tarsus, D detail of frontal scopula, E detail of lateral scopula). Scale bars: A= 1mm; B= 500µm; C= 200µm; D= 100µm; E= 50µm." figureDoi="http://doi.org/10.5281/zenodo.4926897" httpUri="https://zenodo.org/record/4926897/files/figure.png" pageId="4" pageNumber="186">Figs 4 A, B</figureCitation>
; 
<figureCitation id="3D98FAFEFF9B3125AC40B1A5FBE0FF18" box="[1090,1133,151,177]" captionStart="FIGURES 6" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[208,1379,181,1919]" captionTargetId="figure-19@9.[208,1379,181,1919]" captionTargetPageId="9" captionText="FIGURES 6A–D. Stenostygnus martensi spec. nov., sexual dimorphism (A–B female paratype, MNRJ 2692, C–D holotype male, MIZA 0103460). A, C Dorsal view. B, D Detail of tibia. Scale bars: A, C= 2 mm; B, D= 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4926903" httpUri="https://zenodo.org/record/4926903/files/figure.png" pageId="4" pageNumber="186">6 D</figureCitation>
)
<subScript id="3927E43EFF9B3125AC74B19BFBF6FF1E" attach="left" box="[1142,1147,169,183]" fontSize="6" pageId="4" pageNumber="186">;</subScript>
<emphasis id="97D73A69FF9B3125AC81B1A5FABFFF18" box="[1155,1330,151,177]" italics="true" pageId="4" pageNumber="186">lamina apicalis</emphasis>
medially divided with both halves medially in contact (
<figureCitation id="3D98FAFEFF9B3125AA90B189FCD8FF7C" box="[658,853,186,213]" captionStart="FIGURES 5" captionStartId="8.[151,264,1283,1307]" captionTargetBox="[148,1440,181,1259]" captionTargetId="figure-17@8.[148,1440,181,1259]" captionTargetPageId="8" captionText="FIGURES 5A–G. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), penis (A, E dorsal, B, F lateral, C, G ventral, D detail of conductor and stylus). CO—conductor. LA—lamina apicalis. SO—stylus opening of ductus ejaculatorius. ST—stylus. Ti—Titillator. Scale bars:A= 50 µm; B, C, E-G= 100 µm; D= 20 µm. Stylus in green, conductors in pink and capsula externa in blue (For interpretation of the references to colour the reader is referred to the pdf version of this article)." figureDoi="http://doi.org/10.5281/zenodo.4926899" httpUri="https://zenodo.org/record/4926899/files/figure.png" pageId="4" pageNumber="186">Figs 5 A, C, E, G</figureCitation>
). It can be distinguished from 
<taxonomicName id="62A39DF8FF9B3125ACA7B189FAF7FF7D" authorityName="Simon" authorityYear="1879" box="[1189,1402,187,213]" class="Arachnida" family="Biantidae" genus="Stenostygnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="186" phylum="Arthropoda" rank="species" species="pusio">
<emphasis id="97D73A69FF9B3125ACA7B189FAF7FF7D" box="[1189,1402,187,213]" italics="true" pageId="4" pageNumber="186">Stenostygnus pusio</emphasis>
</taxonomicName>
by the lateral borders of the scutum almost parallel, and mesotergal areas almost equally wide (
<figureCitation id="3D98FAFEFF9B3125AC91B1EDFB7DFF50" box="[1171,1264,223,249]" captionStart="FIGURES 2" captionStartId="3.[151,264,1399,1423]" captionTargetBox="[151,1436,181,1375]" captionTargetId="figure-19@3.[151,1436,181,1375]" captionTargetPageId="3" captionText="FIGURES 2A–F. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A, B, E= 500µm; C= 200 µm; D= 50 µm; F= 100µm." figureDoi="http://doi.org/10.5281/zenodo.4926893" httpUri="https://zenodo.org/record/4926893/files/figure.png" pageId="4" pageNumber="186">Fig. 2 A</figureCitation>
) 
<emphasis id="97D73A69FF9B3125ACFCB1EDFACBFF51" box="[1278,1350,223,248]" italics="true" pageId="4" pageNumber="186">versus</emphasis>
scutum lateral borders slightly arched (giving an oval appearance to the habitus) (
<bibRefCitation id="C1329B8AFF9B3125ABE0B031FB5AFEB4" author="Pinto-da-Rocha, R." box="[994,1239,258,285]" pageId="4" pageNumber="186" pagination="194 - 198" refId="ref8901" refString="Pinto-da-Rocha, R. (1995) Redescription of Stenostygnus pusio Simon and synonymy of Caribbiantinae with Stenostygninae (Opiliones, Laniatores, Biantidae). The Journal of Arachnology, 23, 194 - 198." type="journal article" year="1995">Pinto-da-Rocha 1995</bibRefCitation>
, fig. 1); presence of ventral armature on the femur of the pedipalp 
<emphasis id="97D73A69FF9B3125AAC2B015FC85FEE9" box="[704,776,295,320]" italics="true" pageId="4" pageNumber="186">versus</emphasis>
unarmed in 
<taxonomicName id="62A39DF8FF9B3125AB94B015FC7DFEE9" authorityName="Simon" authorityYear="1879" box="[918,1008,295,321]" class="Arachnida" family="Biantidae" genus="Stenostygnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="186" phylum="Arthropoda" rank="species" species="pusio">
<emphasis id="97D73A69FF9B3125AB94B015FC7DFEE9" box="[918,1008,295,321]" italics="true" pageId="4" pageNumber="186">S. pusio</emphasis>
</taxonomicName>
; 
<emphasis id="97D73A69FF9B3125ABFDB015FB04FEE8" box="[1023,1161,295,321]" italics="true" pageId="4" pageNumber="186">pars distalis</emphasis>
ventrally convex 
<emphasis id="97D73A69FF9B3125AD57B015FA10FEE9" box="[1365,1437,295,320]" italics="true" pageId="4" pageNumber="186">versus</emphasis>
concave in 
<taxonomicName id="62A39DF8FF9B3125A917B079FEE0FECD" authorityName="Simon" authorityYear="1879" box="[277,365,331,357]" class="Arachnida" family="Biantidae" genus="Stenostygnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="186" phylum="Arthropoda" rank="species" species="pusio">
<emphasis id="97D73A69FF9B3125A917B079FEE0FECD" box="[277,365,331,357]" italics="true" pageId="4" pageNumber="186">S. pusio</emphasis>
</taxonomicName>
(
<bibRefCitation id="C1329B8AFF9B3125A978B079FDE0FECC" author="Pinto-da-Rocha, R." box="[378,621,330,357]" pageId="4" pageNumber="186" pagination="194 - 198" refId="ref8901" refString="Pinto-da-Rocha, R. (1995) Redescription of Stenostygnus pusio Simon and synonymy of Caribbiantinae with Stenostygninae (Opiliones, Laniatores, Biantidae). The Journal of Arachnology, 23, 194 - 198." type="journal article" year="1995">Pinto-da-Rocha 1995</bibRefCitation>
, fig. 7). It differs from 
<taxonomicName id="62A39DF8FF9B3125AB68B079FC43FECC" authority="Mamani &amp; Porto &amp; Iglesias &amp; González, 2021" authorityName="Mamani &amp; Porto &amp; Iglesias &amp; González" authorityYear="2021" box="[874,974,331,357]" class="Arachnida" family="Biantidae" genus="Stenostygnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="186" phylum="Arthropoda" rank="species" species="huberi" status="sp. nov.">
<emphasis id="97D73A69FF9B3125AB68B079FC43FECC" box="[874,974,331,357]" italics="true" pageId="4" pageNumber="186">S. huberi</emphasis>
</taxonomicName>
<emphasis id="97D73A69FF9B3125ABD6B079FBC8FECC" bold="true" box="[980,1093,331,357]" pageId="4" pageNumber="186">
<taxonomicNameLabel id="8CE48712FF9B3125ABD6B079FBC8FECC" box="[980,1093,331,357]" pageId="4" pageNumber="186" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
by the smooth interocular area and mesotergal areas I to IV smooth (
<figureCitation id="3D98FAFEFF9B3125AA3EB05DFD17FE20" box="[572,666,367,393]" captionStart="FIGURES 2" captionStartId="3.[151,264,1399,1423]" captionTargetBox="[151,1436,181,1375]" captionTargetId="figure-19@3.[151,1436,181,1375]" captionTargetPageId="3" captionText="FIGURES 2A–F. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A, B, E= 500µm; C= 200 µm; D= 50 µm; F= 100µm." figureDoi="http://doi.org/10.5281/zenodo.4926893" httpUri="https://zenodo.org/record/4926893/files/figure.png" pageId="4" pageNumber="186">Fig. 2 A</figureCitation>
) 
<emphasis id="97D73A69FF9B3125AAAAB05DFD7DFE21" box="[680,752,367,392]" italics="true" pageId="4" pageNumber="186">versus</emphasis>
presence of scattered granules in 
<taxonomicName id="62A39DF8FF9B3125AC6EB05DFB5CFE20" authority="Mamani &amp; Porto &amp; Iglesias &amp; González, 2021" authorityName="Mamani &amp; Porto &amp; Iglesias &amp; González" authorityYear="2021" box="[1132,1233,367,393]" class="Arachnida" family="Biantidae" genus="Stenostygnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="186" phylum="Arthropoda" rank="species" species="huberi" status="sp. nov.">
<emphasis id="97D73A69FF9B3125AC6EB05DFB5CFE20" box="[1132,1233,367,393]" italics="true" pageId="4" pageNumber="186">S. huberi</emphasis>
</taxonomicName>
<emphasis id="97D73A69FF9B3125ACDAB05DFAC7FE20" bold="true" box="[1240,1354,367,393]" pageId="4" pageNumber="186">
<taxonomicNameLabel id="8CE48712FF9B3125ACDAB05DFAC7FE20" box="[1240,1354,367,393]" pageId="4" pageNumber="186" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="3D98FAFEFF9B3125AD58B05DFF20FE04" captionStart="FIGURES 8" captionStartId="11.[151,264,1351,1375]" captionTargetBox="[151,1436,181,1328]" captionTargetId="figure-19@11.[151,1436,181,1328]" captionTargetPageId="11" captionText="FIGURES 8A–F. Stenostygnus huberi spec. nov., male (ss) (paratype, MNRJ 2429), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A = 1 mm; B, E= 500 µm; C, F= 100 µm; D= 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4926909" httpUri="https://zenodo.org/record/4926909/files/figure.png" pageId="4" pageNumber="186">Fig. 8 A</figureCitation>
)
<subScript id="3927E43EFF9B3125A8B7B097FF37FE1A" attach="left" box="[181,186,421,435]" fontSize="6" pageId="4" pageNumber="186">;</subScript>
absence of granules on III and IV coxae (
<figureCitation id="3D98FAFEFF9B3125AA8DB0A1FD61FE04" box="[655,748,403,429]" captionStart="FIGURES 2" captionStartId="3.[151,264,1399,1423]" captionTargetBox="[151,1436,181,1375]" captionTargetId="figure-19@3.[151,1436,181,1375]" captionTargetPageId="3" captionText="FIGURES 2A–F. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A, B, E= 500µm; C= 200 µm; D= 50 µm; F= 100µm." figureDoi="http://doi.org/10.5281/zenodo.4926893" httpUri="https://zenodo.org/record/4926893/files/figure.png" pageId="4" pageNumber="186">Fig. 2 B</figureCitation>
) 
<emphasis id="97D73A69FF9B3125AAFEB0A1FCC9FE05" box="[764,836,403,428]" italics="true" pageId="4" pageNumber="186">versus</emphasis>
presence of granules on III and IV coxae in 
<taxonomicName id="62A39DF8FF9B3125AD35B0A1FA11FE04" authority="Mamani &amp; Porto &amp; Iglesias &amp; González, 2021" authorityName="Mamani &amp; Porto &amp; Iglesias &amp; González" authorityYear="2021" box="[1335,1436,403,429]" class="Arachnida" family="Biantidae" genus="Stenostygnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="186" phylum="Arthropoda" rank="species" species="huberi" status="sp. nov.">
<emphasis id="97D73A69FF9B3125AD35B0A1FA11FE04" box="[1335,1436,403,429]" italics="true" pageId="4" pageNumber="186">S. huberi</emphasis>
</taxonomicName>
<emphasis id="97D73A69FF9B3125A895B085FE84FE78" bold="true" box="[151,265,439,465]" pageId="4" pageNumber="186">
<taxonomicNameLabel id="8CE48712FF9B3125A895B085FE84FE78" box="[151,265,439,465]" pageId="4" pageNumber="186" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="3D98FAFEFF9B3125A915B085FEF9FE78" box="[279,372,439,465]" captionStart="FIGURES 8" captionStartId="11.[151,264,1351,1375]" captionTargetBox="[151,1436,181,1328]" captionTargetId="figure-19@11.[151,1436,181,1328]" captionTargetPageId="11" captionText="FIGURES 8A–F. Stenostygnus huberi spec. nov., male (ss) (paratype, MNRJ 2429), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A = 1 mm; B, E= 500 µm; C, F= 100 µm; D= 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4926909" httpUri="https://zenodo.org/record/4926909/files/figure.png" pageId="4" pageNumber="186">Fig. 8 B</figureCitation>
); femur with two small ectal spines (
<figureCitation id="3D98FAFEFF9B3125AB0FB085FCE8FE78" box="[781,869,439,465]" captionStart="FIGURES 3" captionStartId="4.[151,264,1624,1648]" captionTargetBox="[181,1407,561,1601]" captionTargetId="figure-240@4.[181,1407,561,1601]" captionTargetPageId="4" captionText="FIGURES 3A–F. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692). A–C Right chelicera (A ectal, B frontal, C detail of fingers). D–F Right pedipalp (D mesal, E detail of tibia and tarsus, F detail of femur). Scale bars: A, B, E= 200 µm; C, F= 100 µm; D= 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4926895" httpUri="https://zenodo.org/record/4926895/files/figure.png" pageId="4" pageNumber="186">Fig. 3 F</figureCitation>
) 
<emphasis id="97D73A69FF9B3125AB71B085FC36FE79" box="[883,955,439,464]" italics="true" pageId="4" pageNumber="186">versus</emphasis>
without ectal spines in 
<taxonomicName id="62A39DF8FF9B3125ACC2B085FAA9FE78" authority="Mamani &amp; Porto &amp; Iglesias &amp; González, 2021" authorityName="Mamani &amp; Porto &amp; Iglesias &amp; González" authorityYear="2021" box="[1216,1316,439,465]" class="Arachnida" family="Biantidae" genus="Stenostygnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="186" phylum="Arthropoda" rank="species" species="huberi" status="sp. nov.">
<emphasis id="97D73A69FF9B3125ACC2B085FAA9FE78" box="[1216,1316,439,465]" italics="true" pageId="4" pageNumber="186">S. huberi</emphasis>
</taxonomicName>
<emphasis id="97D73A69FF9B3125AD29B085FA11FE78" bold="true" box="[1323,1436,439,465]" pageId="4" pageNumber="186">
<taxonomicNameLabel id="8CE48712FF9B3125AD29B085FA11FE78" box="[1323,1436,439,465]" pageId="4" pageNumber="186" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="3D98FAFEFF9B3125A8A2B0E9FF70FE5C" box="[160,253,475,501]" captionStart="FIGURES 9" captionStartId="12.[151,264,1864,1888]" captionTargetBox="[151,1436,957,1841]" captionTargetId="figure-420@12.[151,1436,957,1841]" captionTargetPageId="12" captionText="FIGURES 9A–E. Stenostygnus huberi spec. nov., male (ss) (paratype, MNRJ 2429). A–C Left chelicera (A ectal, B frontal, C detail of fingers), D–E Left pedipalp (D ectal, E detail of femur). Scale bars: A–C= 200 µm; D= 1 mm; E= 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4926911" httpUri="https://zenodo.org/record/4926911/files/figure.png" pageId="4" pageNumber="186">Fig. 9 E</figureCitation>
)
<subScript id="3927E43EFF9B3125A907B0DFFE87FE52" attach="left" box="[261,266,493,507]" fontSize="6" pageId="4" pageNumber="186">;</subScript>
<emphasis id="97D73A69FF9B3125A910B0E9FE4CFE5C" box="[274,449,475,501]" italics="true" pageId="4" pageNumber="186">lamina apicalis</emphasis>
divided in two halves medially in contact (
<figureCitation id="3D98FAFEFF9B3125ABAFB0E9FB81FE5C" box="[941,1036,474,501]" captionStart="FIGURES 5" captionStartId="8.[151,264,1283,1307]" captionTargetBox="[148,1440,181,1259]" captionTargetId="figure-17@8.[148,1440,181,1259]" captionTargetPageId="8" captionText="FIGURES 5A–G. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), penis (A, E dorsal, B, F lateral, C, G ventral, D detail of conductor and stylus). CO—conductor. LA—lamina apicalis. SO—stylus opening of ductus ejaculatorius. ST—stylus. Ti—Titillator. Scale bars:A= 50 µm; B, C, E-G= 100 µm; D= 20 µm. Stylus in green, conductors in pink and capsula externa in blue (For interpretation of the references to colour the reader is referred to the pdf version of this article)." figureDoi="http://doi.org/10.5281/zenodo.4926899" httpUri="https://zenodo.org/record/4926899/files/figure.png" pageId="4" pageNumber="186">Fig. 5 C</figureCitation>
) 
<emphasis id="97D73A69FF9B3125AC1EB0E9FBE9FE5D" box="[1052,1124,475,500]" italics="true" pageId="4" pageNumber="186">versus</emphasis>
medially divided by a cleft that creates a “V” shape in 
<taxonomicName id="62A39DF8FF9B3125A9CAB0CDFDA0FDB0" authority="Mamani &amp; Porto &amp; Iglesias &amp; González, 2021" authorityName="Mamani &amp; Porto &amp; Iglesias &amp; González" authorityYear="2021" box="[456,557,511,537]" class="Arachnida" family="Biantidae" genus="Stenostygnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="186" phylum="Arthropoda" rank="species" species="huberi" status="sp. nov.">
<emphasis id="97D73A69FF9B3125A9CAB0CDFDA0FDB0" box="[456,557,511,537]" italics="true" pageId="4" pageNumber="186">S. huberi</emphasis>
</taxonomicName>
<emphasis id="97D73A69FF9B3125AA36B0CDFD2BFDB0" bold="true" box="[564,678,511,537]" pageId="4" pageNumber="186">
<taxonomicNameLabel id="8CE48712FF9B3125AA36B0CDFD2BFDB0" box="[564,678,511,537]" pageId="4" pageNumber="186" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="3D98FAFEFF9B3125AAB4B0CDFCAFFDB0" box="[694,802,511,537]" captionStart="FIGURES 11" captionStartId="16.[151,264,1327,1351]" captionTargetBox="[158,1429,181,1303]" captionTargetId="figure-17@16.[158,1429,181,1304]" captionTargetPageId="16" captionText="FIGURES 11A–G. Stenostygnus huberi spec. nov., penis (A–B male (ss) paratype, MNRJ 2429; C male (ss), paratype, MACN 41020; D–G holotype male (α), MIZA 0103461). A, D Dorsal view. B, E, G Lateral view (G with detail of stylus and conduc- tors). C, F Ventral view. LA—lamina apicalis. Ti—Titillator. Scale bars: 100 µm. Stylus in green, conductors in pink and capsula externa in blue (For interpretation of the references to colour the reader is referred to the pdf version of this article)." figureDoi="http://doi.org/10.5281/zenodo.4926915" httpUri="https://zenodo.org/record/4926915/files/figure.png" pageId="4" pageNumber="186">Fig. 11 A</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="F1DCB6F3FF9B3125A895B76AFE1FF910" ID-DOI="http://doi.org/10.5281/zenodo.4926895" ID-Zenodo-Dep="4926895" httpUri="https://zenodo.org/record/4926895/files/figure.png" pageId="4" pageNumber="186" startId="4.[151,264,1624,1648]" targetBox="[181,1407,561,1601]" targetPageId="4">
<paragraph id="A51CE67BFF9B3125A895B76AFE1FF910" blockId="4.[151,1436,1624,1721]" pageId="4" pageNumber="186">
<emphasis id="97D73A69FF9B3125A895B76AFEC0F9D9" bold="true" box="[151,333,1624,1648]" pageId="4" pageNumber="186">FIGURES 3A–F.</emphasis>
<taxonomicName id="62A39DF8FF9B3125A957B76BFDB4F9D9" authority="Mamani &amp; Porto &amp; Iglesias &amp; González, 2021" authorityName="Mamani &amp; Porto &amp; Iglesias &amp; González" authorityYear="2021" box="[341,569,1625,1648]" class="Arachnida" family="Biantidae" genus="Stenostygnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="186" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov.">
<emphasis id="97D73A69FF9B3125A957B76BFDB4F9D9" box="[341,569,1625,1648]" italics="true" pageId="4" pageNumber="186">Stenostygnus martensi</emphasis>
</taxonomicName>
<emphasis id="97D73A69FF9B3125AA42B76BFD25F9D8" bold="true" box="[576,680,1625,1649]" pageId="4" pageNumber="186">
<taxonomicNameLabel id="8CE48712FF9B3125AA42B76BFD25F9D8" box="[576,680,1625,1649]" pageId="4" pageNumber="186" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, male (paratype, MNRJ 2692). A–C Right chelicera (A ectal, B frontal, C detail of fingers). D–F Right pedipalp (D mesal, E detail of tibia and tarsus, F detail of femur). Scale bars: A, B, E= 200 µm; C, F= 100 µm; D= 500 µm.
</paragraph>
</caption>
<subSubSection id="EDB9B5F0FF9B3127A8C5B7D3FB81FC90" lastPageId="6" lastPageNumber="188" pageId="4" pageNumber="186" type="description">
<paragraph id="A51CE67BFF9B3125A8C5B7D3FAE1F819" blockId="4.[151,1437,1761,2040]" pageId="4" pageNumber="186">
<emphasis id="97D73A69FF9B3125A8C5B7D3FE1AF952" bold="true" box="[199,407,1761,1787]" pageId="4" pageNumber="186">Description. Male</emphasis>
(
<typeStatus id="7A1858D9FF9B3125A9A5B7D3FD81F955" box="[423,524,1761,1788]" pageId="4" pageNumber="186" type="holotype">holotype</typeStatus>
): Measurements in 
<tableCitation id="E821D3C0FF9B3125AAE0B7D3FCBEF952" box="[738,819,1761,1787]" captionStart="TABLE 1" captionStartId="6.[151,239,908,932]" captionTargetPageId="6" captionText="TABLE 1. Measurements (in mm) of the type series of Stenostygnus martensi spec. nov. Holotype in bold; TBL—total body length, CL—carapace length, MSL—mesotergal scutum length, CMW—carapace maximum width, MSMW—mesotergal scutum maximum width, Tr—Trochanter, Fe—Femur, Pa—Patella, Ti—Tibia, Ta—Tarsus, T—Total." httpUri="http://table.plazi.org/id/F1DCB6F3FF993127A895B2BEFAA0FC45" pageId="4" pageNumber="186" tableUuid="F1DCB6F3FF993127A895B2BEFAA0FC45">Table 1</tableCitation>
. Dorsum: Habitus oval (
<figureCitation id="3D98FAFEFF9B3125AC41B7D3FB2BF955" box="[1091,1190,1761,1788]" captionStart="FIGURES 1" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[172,1415,179,1917]" captionTargetId="figure-17@2.[172,1415,179,1917]" captionTargetPageId="2" captionText="FIGURES 1A–C. Stenostygnus martensi spec. nov., male (holotype, MIZA 0103460), habitus (A dorsal, B ventral, C lateral). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4926891" httpUri="https://zenodo.org/record/4926891/files/figure.png" pageId="4" pageNumber="186">Figs 1 A</figureCitation>
; 
<figureCitation id="3D98FAFEFF9B3125ACB0B7D3FB50F955" box="[1202,1245,1761,1788]" captionStart="FIGURES 2" captionStartId="3.[151,264,1399,1423]" captionTargetBox="[151,1436,181,1375]" captionTargetId="figure-19@3.[151,1436,181,1375]" captionTargetPageId="3" captionText="FIGURES 2A–F. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A, B, E= 500µm; C= 200 µm; D= 50 µm; F= 100µm." figureDoi="http://doi.org/10.5281/zenodo.4926893" httpUri="https://zenodo.org/record/4926893/files/figure.png" pageId="4" pageNumber="186">2 A</figureCitation>
). Carapace wider than long with the anterior border straight and smooth. Cheliceral sockets not marked. Eyes separated; eye mounds high and smooth, near sulcus I; interocular area completely smooth (
<figureCitation id="3D98FAFEFF9B3125AB43B61BFC11F8ED" box="[833,924,1833,1860]" captionStart="FIGURES 2" captionStartId="3.[151,264,1399,1423]" captionTargetBox="[151,1436,181,1375]" captionTargetId="figure-19@3.[151,1436,181,1375]" captionTargetPageId="3" captionText="FIGURES 2A–F. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A, B, E= 500µm; C= 200 µm; D= 50 µm; F= 100µm." figureDoi="http://doi.org/10.5281/zenodo.4926893" httpUri="https://zenodo.org/record/4926893/files/figure.png" pageId="4" pageNumber="186">Fig. 2 A</figureCitation>
). Carapace in lateral view slightly convex (
<figureCitation id="3D98FAFEFF9B3125AD6EB61BFF32F8CE" captionStart="FIGURES 1" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[172,1415,179,1917]" captionTargetId="figure-17@2.[172,1415,179,1917]" captionTargetPageId="2" captionText="FIGURES 1A–C. Stenostygnus martensi spec. nov., male (holotype, MIZA 0103460), habitus (A dorsal, B ventral, C lateral). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4926891" httpUri="https://zenodo.org/record/4926891/files/figure.png" pageId="4" pageNumber="186">Figs 1 C</figureCitation>
; 
<figureCitation id="3D98FAFEFF9B3125A8CBB67FFF7DF8C1" box="[201,240,1869,1896]" captionStart="FIGURES 2" captionStartId="3.[151,264,1399,1423]" captionTargetBox="[151,1436,181,1375]" captionTargetId="figure-19@3.[151,1436,181,1375]" captionTargetPageId="3" captionText="FIGURES 2A–F. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A, B, E= 500µm; C= 200 µm; D= 50 µm; F= 100µm." figureDoi="http://doi.org/10.5281/zenodo.4926893" httpUri="https://zenodo.org/record/4926893/files/figure.png" pageId="4" pageNumber="186">2 E</figureCitation>
). Mesotergum in lateral view convex with areas well defined, with sulcus well marked (
<figureCitation id="3D98FAFEFF9B3125AC94B67FFB63F8C1" box="[1174,1262,1869,1896]" captionStart="FIGURES 2" captionStartId="3.[151,264,1399,1423]" captionTargetBox="[151,1436,181,1375]" captionTargetId="figure-19@3.[151,1436,181,1375]" captionTargetPageId="3" captionText="FIGURES 2A–F. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A, B, E= 500µm; C= 200 µm; D= 50 µm; F= 100µm." figureDoi="http://doi.org/10.5281/zenodo.4926893" httpUri="https://zenodo.org/record/4926893/files/figure.png" pageId="4" pageNumber="186">Fig. 2 E</figureCitation>
). Mesotergal areas I to IV unarmed. Lateral margins of scutum dorsally with a row of small granules (
<figureCitation id="3D98FAFEFF9B3125AC29B643FB28F825" box="[1067,1189,1905,1932]" captionStart="FIGURES 2" captionStartId="3.[151,264,1399,1423]" captionTargetBox="[151,1436,181,1375]" captionTargetId="figure-19@3.[151,1436,181,1375]" captionTargetPageId="3" captionText="FIGURES 2A–F. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A, B, E= 500µm; C= 200 µm; D= 50 µm; F= 100µm." figureDoi="http://doi.org/10.5281/zenodo.4926893" httpUri="https://zenodo.org/record/4926893/files/figure.png" pageId="4" pageNumber="186">Figs 2 E–F</figureCitation>
). Ozopore with an oval orifice with a descending channel toward the posterior region (
<figureCitation id="3D98FAFEFF9B3125AB3EB6A7FC19F819" box="[828,916,1941,1968]" captionStart="FIGURES 2" captionStartId="3.[151,264,1399,1423]" captionTargetBox="[151,1436,181,1375]" captionTargetId="figure-19@3.[151,1436,181,1375]" captionTargetPageId="3" captionText="FIGURES 2A–F. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A, B, E= 500µm; C= 200 µm; D= 50 µm; F= 100µm." figureDoi="http://doi.org/10.5281/zenodo.4926893" httpUri="https://zenodo.org/record/4926893/files/figure.png" pageId="4" pageNumber="186">Fig. 2 F</figureCitation>
). Free tergites with few granules (
<figureCitation id="3D98FAFEFF9B3125AD01B6A7FAD2F819" box="[1283,1375,1941,1968]" captionStart="FIGURES 2" captionStartId="3.[151,264,1399,1423]" captionTargetBox="[151,1436,181,1375]" captionTargetId="figure-19@3.[151,1436,181,1375]" captionTargetPageId="3" captionText="FIGURES 2A–F. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A, B, E= 500µm; C= 200 µm; D= 50 µm; F= 100µm." figureDoi="http://doi.org/10.5281/zenodo.4926893" httpUri="https://zenodo.org/record/4926893/files/figure.png" pageId="4" pageNumber="186">Fig. 2 A</figureCitation>
).
</paragraph>
<paragraph id="A51CE67BFF9B3125A8C5B688FD61F851" blockId="4.[151,1437,1761,2040]" pageId="4" pageNumber="186">
Venter: Without any remarkable features (
<figureCitation id="3D98FAFEFF9B3125AAAAB68BFCBEF87A" box="[680,819,1977,2003]" captionStart="FIGURES 2" captionStartId="3.[151,264,1399,1423]" captionTargetBox="[151,1436,181,1375]" captionTargetId="figure-19@3.[151,1436,181,1375]" captionTargetPageId="3" captionText="FIGURES 2A–F. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A, B, E= 500µm; C= 200 µm; D= 50 µm; F= 100µm." figureDoi="http://doi.org/10.5281/zenodo.4926893" httpUri="https://zenodo.org/record/4926893/files/figure.png" pageId="4" pageNumber="186">Figs 2 B–C</figureCitation>
). Spiracles visible (
<figureCitation id="3D98FAFEFF9B3125AC16B68BFB2FF87D" box="[1044,1186,1977,2004]" captionStart="FIGURES 2" captionStartId="3.[151,264,1399,1423]" captionTargetBox="[151,1436,181,1375]" captionTargetId="figure-19@3.[151,1436,181,1375]" captionTargetPageId="3" captionText="FIGURES 2A–F. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A, B, E= 500µm; C= 200 µm; D= 50 µm; F= 100µm." figureDoi="http://doi.org/10.5281/zenodo.4926893" httpUri="https://zenodo.org/record/4926893/files/figure.png" pageId="4" pageNumber="186">Figs 2 B, D</figureCitation>
). Coxae I and II with minute granules. Coxae III and IV smooth. (
<figureCitation id="3D98FAFEFF9B3125AA85B6EFFD6EF851" box="[647,739,2013,2040]" captionStart="FIGURES 2" captionStartId="3.[151,264,1399,1423]" captionTargetBox="[151,1436,181,1375]" captionTargetId="figure-19@3.[151,1436,181,1375]" captionTargetPageId="3" captionText="FIGURES 2A–F. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A, B, E= 500µm; C= 200 µm; D= 50 µm; F= 100µm." figureDoi="http://doi.org/10.5281/zenodo.4926893" httpUri="https://zenodo.org/record/4926893/files/figure.png" pageId="4" pageNumber="186">Fig. 2 B</figureCitation>
)
</paragraph>
<caption id="F1DCB6F3FF9A3124A895B43DFE5DFAC6" ID-DOI="http://doi.org/10.5281/zenodo.4926897" ID-Zenodo-Dep="4926897" httpUri="https://zenodo.org/record/4926897/files/figure.png" pageId="5" pageNumber="187" startId="5.[151,264,1295,1319]" targetBox="[173,1414,181,1271]" targetPageId="5">
<paragraph id="A51CE67BFF9A3124A895B43DFE5DFAC6" blockId="5.[151,1436,1295,1391]" pageId="5" pageNumber="187">
<emphasis id="97D73A69FF9A3124A895B43DFEDFFA8E" bold="true" box="[151,338,1295,1319]" pageId="5" pageNumber="187">FIGURES 4A–E.</emphasis>
<taxonomicName id="62A39DF8FF9A3124A95AB43DFDB1FA8E" authority="Mamani &amp; Porto &amp; Iglesias &amp; González, 2021" authorityName="Mamani &amp; Porto &amp; Iglesias &amp; González" authorityYear="2021" box="[344,572,1295,1319]" class="Arachnida" family="Biantidae" genus="Stenostygnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="187" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov.">
<emphasis id="97D73A69FF9A3124A95AB43DFDB1FA8E" box="[344,572,1295,1319]" italics="true" pageId="5" pageNumber="187">Stenostygnus martensi</emphasis>
</taxonomicName>
<emphasis id="97D73A69FF9A3124AA41B43DFD27FA8E" bold="true" box="[579,682,1295,1319]" pageId="5" pageNumber="187">
<taxonomicNameLabel id="8CE48712FF9A3124AA41B43DFD27FA8E" box="[579,682,1295,1319]" pageId="5" pageNumber="187" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, male (paratype, MNRJ 2692), right leg IV (A retrolateral, B detail of patella and tibia, C detail of tarsus, D detail of frontal scopula, E detail of lateral scopula). Scale bars: A= 1mm; B= 500µm; C= 200µm; D= 100µm; E= 50µm.
</paragraph>
</caption>
<paragraph id="A51CE67BFF9A3124A8C5B4AFFA17FA72" blockId="5.[151,1437,1437,2033]" pageId="5" pageNumber="187">
Chelicerae: Without remarkable armature. Basichelicerite smooth with a short bulla. Cheliceral hand unarmed, covered with sparse setae (
<figureCitation id="3D98FAFEFF9A3124A9C1B4F3FDC4FA75" box="[451,585,1473,1500]" captionStart="FIGURES 3" captionStartId="4.[151,264,1624,1648]" captionTargetBox="[181,1407,561,1601]" captionTargetId="figure-240@4.[181,1407,561,1601]" captionTargetPageId="4" captionText="FIGURES 3A–F. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692). A–C Right chelicera (A ectal, B frontal, C detail of fingers). D–F Right pedipalp (D mesal, E detail of tibia and tarsus, F detail of femur). Scale bars: A, B, E= 200 µm; C, F= 100 µm; D= 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4926895" httpUri="https://zenodo.org/record/4926895/files/figure.png" pageId="5" pageNumber="187">Figs 3 A–B</figureCitation>
). Fingers thin, each with four teeth spaced one from the other (
<figureCitation id="3D98FAFEFF9A3124AD0AB4F3FA00FA72" box="[1288,1421,1473,1499]" captionStart="FIGURES 3" captionStartId="4.[151,264,1624,1648]" captionTargetBox="[181,1407,561,1601]" captionTargetId="figure-240@4.[181,1407,561,1601]" captionTargetPageId="4" captionText="FIGURES 3A–F. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692). A–C Right chelicera (A ectal, B frontal, C detail of fingers). D–F Right pedipalp (D mesal, E detail of tibia and tarsus, F detail of femur). Scale bars: A, B, E= 200 µm; C, F= 100 µm; D= 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4926895" httpUri="https://zenodo.org/record/4926895/files/figure.png" pageId="5" pageNumber="187">Figs 3 B–C</figureCitation>
).
</paragraph>
<paragraph id="A51CE67BFF9A3124A8C5B4D7FBBFF91D" blockId="5.[151,1437,1437,2033]" pageId="5" pageNumber="187">
Pedipalps: Raptorial, with spines concentrated on tibia and tarsus (
<figureCitation id="3D98FAFEFF9A3124ABC2B4D7FBA8F9A9" box="[960,1061,1509,1536]" captionStart="FIGURES 3" captionStartId="4.[151,264,1624,1648]" captionTargetBox="[181,1407,561,1601]" captionTargetId="figure-240@4.[181,1407,561,1601]" captionTargetPageId="4" captionText="FIGURES 3A–F. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692). A–C Right chelicera (A ectal, B frontal, C detail of fingers). D–F Right pedipalp (D mesal, E detail of tibia and tarsus, F detail of femur). Scale bars: A, B, E= 200 µm; C, F= 100 µm; D= 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4926895" httpUri="https://zenodo.org/record/4926895/files/figure.png" pageId="5" pageNumber="187">Fig. 3 D</figureCitation>
). Coxa elongated, surpassing the anterior border of the carapace (
<emphasis id="97D73A69FF9A3124AA01B738FDABF98A" box="[515,550,1546,1571]" italics="true" pageId="5" pageNumber="187">i.e.</emphasis>
can be seen in dorsal view) (
<figureCitation id="3D98FAFEFF9A3124AB73B73BFC5BF98D" box="[881,982,1545,1572]" captionStart="FIGURES 1" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[172,1415,179,1917]" captionTargetId="figure-17@2.[172,1415,179,1917]" captionTargetPageId="2" captionText="FIGURES 1A–C. Stenostygnus martensi spec. nov., male (holotype, MIZA 0103460), habitus (A dorsal, B ventral, C lateral). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4926891" httpUri="https://zenodo.org/record/4926891/files/figure.png" pageId="5" pageNumber="187">Figs 1 A</figureCitation>
; 
<figureCitation id="3D98FAFEFF9A3124ABE1B73BFB83F98D" box="[995,1038,1545,1572]" captionStart="FIGURES 2" captionStartId="3.[151,264,1399,1423]" captionTargetBox="[151,1436,181,1375]" captionTargetId="figure-19@3.[151,1436,181,1375]" captionTargetPageId="3" captionText="FIGURES 2A–F. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A, B, E= 500µm; C= 200 µm; D= 50 µm; F= 100µm." figureDoi="http://doi.org/10.5281/zenodo.4926893" httpUri="https://zenodo.org/record/4926893/files/figure.png" pageId="5" pageNumber="187">2 A</figureCitation>
), proximally with one small dorsoectal protuberance; trochanter small and smooth; Femur with tubular shape, slightly narrowed distally, and with a meso-medial spine and with two small ectal spines (
<figureCitation id="3D98FAFEFF9A3124AAD4B763FCBDF9C5" box="[726,816,1617,1644]" captionStart="FIGURES 3" captionStartId="4.[151,264,1624,1648]" captionTargetBox="[181,1407,561,1601]" captionTargetId="figure-240@4.[181,1407,561,1601]" captionTargetPageId="4" captionText="FIGURES 3A–F. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692). A–C Right chelicera (A ectal, B frontal, C detail of fingers). D–F Right pedipalp (D mesal, E detail of tibia and tarsus, F detail of femur). Scale bars: A, B, E= 200 µm; C, F= 100 µm; D= 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4926895" httpUri="https://zenodo.org/record/4926895/files/figure.png" pageId="5" pageNumber="187">Fig. 3 F</figureCitation>
). Patella is short, clavate, strongly swollen distally, with a strong meso-distal spine. Tibia armed with one ventro-basal spine and with three ectal and three mesal ventral spines (
<figureCitation id="3D98FAFEFF9A3124A8EFB7ABFEC6F91D" box="[237,331,1689,1716]" captionStart="FIGURES 3" captionStartId="4.[151,264,1624,1648]" captionTargetBox="[181,1407,561,1601]" captionTargetId="figure-240@4.[181,1407,561,1601]" captionTargetPageId="4" captionText="FIGURES 3A–F. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692). A–C Right chelicera (A ectal, B frontal, C detail of fingers). D–F Right pedipalp (D mesal, E detail of tibia and tarsus, F detail of femur). Scale bars: A, B, E= 200 µm; C, F= 100 µm; D= 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4926895" httpUri="https://zenodo.org/record/4926895/files/figure.png" pageId="5" pageNumber="187">Fig. 3 E</figureCitation>
). Tarsus mesal and ectally with two spines on each side (
<figureCitation id="3D98FAFEFF9A3124ABC5B7ABFBABF91D" box="[967,1062,1689,1716]" captionStart="FIGURES 3" captionStartId="4.[151,264,1624,1648]" captionTargetBox="[181,1407,561,1601]" captionTargetId="figure-240@4.[181,1407,561,1601]" captionTargetPageId="4" captionText="FIGURES 3A–F. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692). A–C Right chelicera (A ectal, B frontal, C detail of fingers). D–F Right pedipalp (D mesal, E detail of tibia and tarsus, F detail of femur). Scale bars: A, B, E= 200 µm; C, F= 100 µm; D= 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4926895" httpUri="https://zenodo.org/record/4926895/files/figure.png" pageId="5" pageNumber="187">Fig. 3 E</figureCitation>
).
</paragraph>
<paragraph id="A51CE67BFF9A3124A8C5B78CFC14F889" blockId="5.[151,1437,1437,2033]" pageId="5" pageNumber="187">
Legs: Tubular (
<figureCitation id="3D98FAFEFF9A3124A970B78FFE5DF971" box="[370,464,1725,1752]" captionStart="FIGURES 4" captionStartId="5.[151,264,1295,1319]" captionTargetBox="[173,1414,181,1271]" captionTargetId="figure-19@5.[173,1414,181,1271]" captionTargetPageId="5" captionText="FIGURES 4A–E. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), right leg IV (A retrolateral, B detail of patella and tibia, C detail of tarsus, D detail of frontal scopula, E detail of lateral scopula). Scale bars: A= 1mm; B= 500µm; C= 200µm; D= 100µm; E= 50µm." figureDoi="http://doi.org/10.5281/zenodo.4926897" httpUri="https://zenodo.org/record/4926897/files/figure.png" pageId="5" pageNumber="187">Fig. 4 A</figureCitation>
). Coxa IV unarmed. Femur IV with a row of tubercles (
<figureCitation id="3D98FAFEFF9A3124AC3AB78FFB1AF971" box="[1080,1175,1725,1752]" captionStart="FIGURES 4" captionStartId="5.[151,264,1295,1319]" captionTargetBox="[173,1414,181,1271]" captionTargetId="figure-19@5.[173,1414,181,1271]" captionTargetPageId="5" captionText="FIGURES 4A–E. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), right leg IV (A retrolateral, B detail of patella and tibia, C detail of tarsus, D detail of frontal scopula, E detail of lateral scopula). Scale bars: A= 1mm; B= 500µm; C= 200µm; D= 100µm; E= 50µm." figureDoi="http://doi.org/10.5281/zenodo.4926897" httpUri="https://zenodo.org/record/4926897/files/figure.png" pageId="5" pageNumber="187">Fig. 4 A</figureCitation>
); patella I–III short and swollen; patella IV armed with granules; tibia IV with a retrolateral row of pointed strong tubercles (
<figureCitation id="3D98FAFEFF9A3124ACFBB7D3FADAF955" box="[1273,1367,1761,1788]" captionStart="FIGURES 4" captionStartId="5.[151,264,1295,1319]" captionTargetBox="[173,1414,181,1271]" captionTargetId="figure-19@5.[173,1414,181,1271]" captionTargetPageId="5" captionText="FIGURES 4A–E. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), right leg IV (A retrolateral, B detail of patella and tibia, C detail of tarsus, D detail of frontal scopula, E detail of lateral scopula). Scale bars: A= 1mm; B= 500µm; C= 200µm; D= 100µm; E= 50µm." figureDoi="http://doi.org/10.5281/zenodo.4926897" httpUri="https://zenodo.org/record/4926897/files/figure.png" pageId="5" pageNumber="187">Fig. 4 B</figureCitation>
); tarsi III and IV with scopula (
<figureCitation id="3D98FAFEFF9A3124A9A9B637FDA2F8B6" box="[427,559,1797,1823]" captionStart="FIGURES 4" captionStartId="5.[151,264,1295,1319]" captionTargetBox="[173,1414,181,1271]" captionTargetId="figure-19@5.[173,1414,181,1271]" captionTargetPageId="5" captionText="FIGURES 4A–E. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), right leg IV (A retrolateral, B detail of patella and tibia, C detail of tarsus, D detail of frontal scopula, E detail of lateral scopula). Scale bars: A= 1mm; B= 500µm; C= 200µm; D= 100µm; E= 50µm." figureDoi="http://doi.org/10.5281/zenodo.4926897" httpUri="https://zenodo.org/record/4926897/files/figure.png" pageId="5" pageNumber="187">Figs 4 C–E</figureCitation>
). Tarsal formula: 7(3):15(3):6:6.
</paragraph>
<paragraph id="A51CE67BFF9A3124A8C5B61BFF7FF8C8" blockId="5.[151,1437,1437,2033]" pageId="5" pageNumber="187">
Color (specimen preserved in 80% ethanol): Body yellowish. Mesotergal areas, with faint brown reticulations (
<figureCitation id="3D98FAFEFF9A3124A8A2B675FF6CF8C8" box="[160,225,1863,1889]" captionStart="FIGURES 1" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[172,1415,179,1917]" captionTargetId="figure-17@2.[172,1415,179,1917]" captionTargetPageId="2" captionText="FIGURES 1A–C. Stenostygnus martensi spec. nov., male (holotype, MIZA 0103460), habitus (A dorsal, B ventral, C lateral). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4926891" httpUri="https://zenodo.org/record/4926891/files/figure.png" pageId="5" pageNumber="187">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="A51CE67BFF9A3127A8C5B659FDCBFE5C" blockId="5.[151,1437,1437,2033]" lastBlockId="6.[151,1437,150,862]" lastPageId="6" lastPageNumber="188" pageId="5" pageNumber="187">
Genitalia: Penis with 
<emphasis id="97D73A69FF9A3124A9BEB659FDC8F82C" box="[444,581,1899,1925]" italics="true" pageId="5" pageNumber="187">pars basalis</emphasis>
and 
<emphasis id="97D73A69FF9A3124AA7CB659FC84F82C" box="[638,777,1899,1925]" italics="true" pageId="5" pageNumber="187">pars distalis</emphasis>
easily distinguishable. 
<emphasis id="97D73A69FF9A3124AC11B659FB12F82C" box="[1043,1183,1899,1925]" italics="true" pageId="5" pageNumber="187">Pars basalis</emphasis>
tubular slightly thickened distally while 
<emphasis id="97D73A69FF9A3124A971B6BDFE73F800" box="[371,510,1935,1961]" italics="true" pageId="5" pageNumber="187">pars distalis</emphasis>
is slightly flattened dorsoventrally (
<figureCitation id="3D98FAFEFF9A3124AB97B6BDFBD4F800" box="[917,1113,1934,1961]" captionStart="FIGURES 5" captionStartId="8.[151,264,1283,1307]" captionTargetBox="[148,1440,181,1259]" captionTargetId="figure-17@8.[148,1440,181,1259]" captionTargetPageId="8" captionText="FIGURES 5A–G. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), penis (A, E dorsal, B, F lateral, C, G ventral, D detail of conductor and stylus). CO—conductor. LA—lamina apicalis. SO—stylus opening of ductus ejaculatorius. ST—stylus. Ti—Titillator. Scale bars:A= 50 µm; B, C, E-G= 100 µm; D= 20 µm. Stylus in green, conductors in pink and capsula externa in blue (For interpretation of the references to colour the reader is referred to the pdf version of this article)." figureDoi="http://doi.org/10.5281/zenodo.4926899" httpUri="https://zenodo.org/record/4926899/files/figure.png" pageId="5" pageNumber="187">Figs 5 A–B, E–F</figureCitation>
). 
<emphasis id="97D73A69FF9A3124AC6EB6BDFB77F800" box="[1132,1274,1935,1961]" italics="true" pageId="5" pageNumber="187">Pars distalis</emphasis>
with a 
<emphasis id="97D73A69FF9A3124AD4CB6BDFF7DF864" italics="true" pageId="5" pageNumber="187">lamina apicalis</emphasis>
ending in two flattened triangular halves. The halves are apically rounded and separated by a deep and narrow groove (
<figureCitation id="3D98FAFEFF9A3124A949B6E5FE5EF858" box="[331,467,2006,2033]" captionStart="FIGURES 5" captionStartId="8.[151,264,1283,1307]" captionTargetBox="[148,1440,181,1259]" captionTargetId="figure-17@8.[148,1440,181,1259]" captionTargetPageId="8" captionText="FIGURES 5A–G. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), penis (A, E dorsal, B, F lateral, C, G ventral, D detail of conductor and stylus). CO—conductor. LA—lamina apicalis. SO—stylus opening of ductus ejaculatorius. ST—stylus. Ti—Titillator. Scale bars:A= 50 µm; B, C, E-G= 100 µm; D= 20 µm. Stylus in green, conductors in pink and capsula externa in blue (For interpretation of the references to colour the reader is referred to the pdf version of this article)." figureDoi="http://doi.org/10.5281/zenodo.4926899" httpUri="https://zenodo.org/record/4926899/files/figure.png" pageId="5" pageNumber="187">Figs 5 C, G</figureCitation>
). Latero-ventral border of 
<emphasis id="97D73A69FF9A3124AAFCB6E5FC04F858" box="[766,905,2007,2033]" italics="true" pageId="5" pageNumber="187">pars distalis</emphasis>
armed with 8 pairs of leaf-like macrosetae and 2 subapical pairs of truncated setae, also the cuticle has a rugose (folding) appearance (
<figureCitation id="3D98FAFEFF993127AC71B1A5FB7AFF18" box="[1139,1271,150,177]" captionStart="FIGURES 5" captionStartId="8.[151,264,1283,1307]" captionTargetBox="[148,1440,181,1259]" captionTargetId="figure-17@8.[148,1440,181,1259]" captionTargetPageId="8" captionText="FIGURES 5A–G. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), penis (A, E dorsal, B, F lateral, C, G ventral, D detail of conductor and stylus). CO—conductor. LA—lamina apicalis. SO—stylus opening of ductus ejaculatorius. ST—stylus. Ti—Titillator. Scale bars:A= 50 µm; B, C, E-G= 100 µm; D= 20 µm. Stylus in green, conductors in pink and capsula externa in blue (For interpretation of the references to colour the reader is referred to the pdf version of this article)." figureDoi="http://doi.org/10.5281/zenodo.4926899" httpUri="https://zenodo.org/record/4926899/files/figure.png" pageId="6" pageNumber="188">Figs 5 B, F</figureCitation>
). Dorsally, the capsula externa has two small titillators, which present a small crease towards the dorsal region. Inner surface of titillators covered by digitiform projections (
<figureCitation id="3D98FAFEFF993127AA8EB1EDFC9EFF50" box="[652,787,222,249]" captionStart="FIGURES 5" captionStartId="8.[151,264,1283,1307]" captionTargetBox="[148,1440,181,1259]" captionTargetId="figure-17@8.[148,1440,181,1259]" captionTargetPageId="8" captionText="FIGURES 5A–G. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), penis (A, E dorsal, B, F lateral, C, G ventral, D detail of conductor and stylus). CO—conductor. LA—lamina apicalis. SO—stylus opening of ductus ejaculatorius. ST—stylus. Ti—Titillator. Scale bars:A= 50 µm; B, C, E-G= 100 µm; D= 20 µm. Stylus in green, conductors in pink and capsula externa in blue (For interpretation of the references to colour the reader is referred to the pdf version of this article)." figureDoi="http://doi.org/10.5281/zenodo.4926899" httpUri="https://zenodo.org/record/4926899/files/figure.png" pageId="6" pageNumber="188">Figs 5 A–B</figureCitation>
). Capsula interna with two laminar conductors bearing an also laminar and pointed stylus with a ventral subapical opening of 
<emphasis id="97D73A69FF993127AB9BB031FBF3FEB4" box="[921,1150,259,285]" italics="true" pageId="6" pageNumber="188">ductus ejaculatorius</emphasis>
(
<figureCitation id="3D98FAFEFF993127AC8DB031FB7FFEB4" box="[1167,1266,258,285]" captionStart="FIGURES 5" captionStartId="8.[151,264,1283,1307]" captionTargetBox="[148,1440,181,1259]" captionTargetId="figure-17@8.[148,1440,181,1259]" captionTargetPageId="8" captionText="FIGURES 5A–G. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), penis (A, E dorsal, B, F lateral, C, G ventral, D detail of conductor and stylus). CO—conductor. LA—lamina apicalis. SO—stylus opening of ductus ejaculatorius. ST—stylus. Ti—Titillator. Scale bars:A= 50 µm; B, C, E-G= 100 µm; D= 20 µm. Stylus in green, conductors in pink and capsula externa in blue (For interpretation of the references to colour the reader is referred to the pdf version of this article)." figureDoi="http://doi.org/10.5281/zenodo.4926899" httpUri="https://zenodo.org/record/4926899/files/figure.png" pageId="6" pageNumber="188">Fig. 5 D</figureCitation>
). The cuticular area of the lateral macrosetae has a rugose (folding) appearance (
<figureCitation id="3D98FAFEFF993127AB85B015FC67FEE8" box="[903,1002,294,321]" captionStart="FIGURES 5" captionStartId="8.[151,264,1283,1307]" captionTargetBox="[148,1440,181,1259]" captionTargetId="figure-17@8.[148,1440,181,1259]" captionTargetPageId="8" captionText="FIGURES 5A–G. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), penis (A, E dorsal, B, F lateral, C, G ventral, D detail of conductor and stylus). CO—conductor. LA—lamina apicalis. SO—stylus opening of ductus ejaculatorius. ST—stylus. Ti—Titillator. Scale bars:A= 50 µm; B, C, E-G= 100 µm; D= 20 µm. Stylus in green, conductors in pink and capsula externa in blue (For interpretation of the references to colour the reader is referred to the pdf version of this article)." figureDoi="http://doi.org/10.5281/zenodo.4926899" httpUri="https://zenodo.org/record/4926899/files/figure.png" pageId="6" pageNumber="188">Fig. 5 B</figureCitation>
), this could be an effect of thinner or less chitinous cuticular layer in this region, that could be (hypothetically) related to the region of 
<emphasis id="97D73A69FF993127ACE2B079FAE6FECC" box="[1248,1387,331,357]" italics="true" pageId="6" pageNumber="188">pars distalis</emphasis>
that became swollen during the turgescent stage in the hydraulic penial functioning, this inflation could be responsible for the macrosetae erection necessary for functional interaction with the ovipositor during the copulation. These putative expansible areas could functionally correspond to the spongy body (‘Schwellkörper’ in 
<bibRefCitation id="C1329B8AFF993127ACF7B085FA19FE78" author="Martens, J." box="[1269,1428,439,465]" pageId="6" pageNumber="188" pagination="347 - 414" refId="ref8555" refString="Martens, J. (1978) Opiliones aus dem Nepal-Himalaya. IV. Biantidae (Arachnida). Senckenbergiana biologica, 58 (5 / 6), 347 - 414." type="journal article" year="1978">Martens 1978</bibRefCitation>
) commonly observed in Asian biantids.
</paragraph>
<paragraph id="A51CE67BFF993127A8C5B0CDFF51FDC8" blockId="6.[151,1437,150,862]" pageId="6" pageNumber="188">
<emphasis id="97D73A69FF993127A8C5B0CDFE93FDB0" bold="true" box="[199,286,511,537]" pageId="6" pageNumber="188">Female</emphasis>
(
<typeStatus id="7A1858D9FF993127A92FB0CDFE1FFDB0" box="[301,402,511,537]" pageId="6" pageNumber="188" type="paratype">paratype</typeStatus>
, MNRJ 2692): measurements as in 
<tableCitation id="E821D3C0FF993127AB1DB0CDFCFEFDB0" box="[799,883,511,537]" captionStart="TABLE 1" captionStartId="6.[151,239,908,932]" captionTargetPageId="6" captionText="TABLE 1. Measurements (in mm) of the type series of Stenostygnus martensi spec. nov. Holotype in bold; TBL—total body length, CL—carapace length, MSL—mesotergal scutum length, CMW—carapace maximum width, MSMW—mesotergal scutum maximum width, Tr—Trochanter, Fe—Femur, Pa—Patella, Ti—Tibia, Ta—Tarsus, T—Total." httpUri="http://table.plazi.org/id/F1DCB6F3FF993127A895B2BEFAA0FC45" pageId="6" pageNumber="188" tableUuid="F1DCB6F3FF993127A895B2BEFAA0FC45">Table 1</tableCitation>
. Similar to male in relation to the armature of the dorsal scutum (
<figureCitation id="3D98FAFEFF993127A946B311FE46FD94" box="[324,459,547,573]" captionStart="FIGURES 6" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[208,1379,181,1919]" captionTargetId="figure-19@9.[208,1379,181,1919]" captionTargetPageId="9" captionText="FIGURES 6A–D. Stenostygnus martensi spec. nov., sexual dimorphism (A–B female paratype, MNRJ 2692, C–D holotype male, MIZA 0103460). A, C Dorsal view. B, D Detail of tibia. Scale bars: A, C= 2 mm; B, D= 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4926903" httpUri="https://zenodo.org/record/4926903/files/figure.png" pageId="6" pageNumber="188">Figs 6 A, C</figureCitation>
). Sexual dimorphism evident in tibia IV: females without retrolateral granules. (
<figureCitation id="3D98FAFEFF993127AD57B311FF5DFDC8" captionStart="FIGURES 6" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[208,1379,181,1919]" captionTargetId="figure-19@9.[208,1379,181,1919]" captionTargetPageId="9" captionText="FIGURES 6A–D. Stenostygnus martensi spec. nov., sexual dimorphism (A–B female paratype, MNRJ 2692, C–D holotype male, MIZA 0103460). A, C Dorsal view. B, D Detail of tibia. Scale bars: A, C= 2 mm; B, D= 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4926903" httpUri="https://zenodo.org/record/4926903/files/figure.png" pageId="6" pageNumber="188">Figs 6 B, D</figureCitation>
).
</paragraph>
<paragraph id="A51CE67BFF993127A8C5B358FB81FC90" blockId="6.[151,1437,150,862]" pageId="6" pageNumber="188">
<emphasis id="97D73A69FF993127A8C5B358FEB2FD2D" bold="true" box="[199,319,618,644]" pageId="6" pageNumber="188">Variation.</emphasis>
Measurement (minimum-maximum) (
<tableCitation id="E821D3C0FF993127AAE8B359FCB0FD2C" box="[746,829,619,645]" captionStart="TABLE 1" captionStartId="6.[151,239,908,932]" captionTargetPageId="6" captionText="TABLE 1. Measurements (in mm) of the type series of Stenostygnus martensi spec. nov. Holotype in bold; TBL—total body length, CL—carapace length, MSL—mesotergal scutum length, CMW—carapace maximum width, MSMW—mesotergal scutum maximum width, Tr—Trochanter, Fe—Femur, Pa—Patella, Ti—Tibia, Ta—Tarsus, T—Total." httpUri="http://table.plazi.org/id/F1DCB6F3FF993127A895B2BEFAA0FC45" pageId="6" pageNumber="188" tableUuid="F1DCB6F3FF993127A895B2BEFAA0FC45">Table 1</tableCitation>
): Males (n=2): total body length (2.15–2.39), carapace length (0.72–0.89), mesotergal length (2.05–2.27), carapace maximum width (1.24–1.27), mesotergal 
<emphasis id="97D73A69FF993127AD07B3BDFAD9FD01" box="[1285,1364,655,680]" italics="true" pageId="6" pageNumber="188">scutum</emphasis>
maximum width (1.65–1.70), leg I (5.89–6.24), leg II (11.27–11.73), leg III (7.70–7.76), leg IV (9.22–9.58); Females (n=5): total body length (2.39–2.75), carapace length (0.70–0.78), mesotergal length (1.90–2.13), carapace maximum width (1.12–1.20), mesotergal 
<emphasis id="97D73A69FF993127AA30B3C9FD0CFCBD" box="[562,641,763,788]" italics="true" pageId="6" pageNumber="188">scutum</emphasis>
maximum width (1.55–1.60), leg I (4.51–5.21), leg II (9.19–9.54), leg III (6.02–6.26), leg IV (7.51–8.10). Tarsal formula (n=7): 6–7(3):10–15(3):6:6.
</paragraph>
</subSubSection>
<subSubSection id="EDB9B5F0FF993127A8C5B270FCD8FCF7" box="[199,853,834,862]" pageId="6" pageNumber="188" type="distribution">
<paragraph id="A51CE67BFF993127A8C5B270FCD8FCF7" blockId="6.[151,1437,150,862]" box="[199,853,834,862]" pageId="6" pageNumber="188">
<emphasis id="97D73A69FF993127A8C5B270FEECFCF5" bold="true" box="[199,353,834,860]" pageId="6" pageNumber="188">Distribution.</emphasis>
Known only from the 
<typeStatus id="7A1858D9FF993127AA62B271FD1DFCF4" box="[608,656,835,861]" pageId="6" pageNumber="188">type</typeStatus>
locality (
<figureCitation id="3D98FAFEFF993127AAF9B271FCCAFCF7" box="[763,839,835,862]" captionStart="FIGURE 13" captionStartId="18.[151,250,1113,1137]" captionTargetBox="[151,1436,181,1090]" captionTargetId="figure-17@18.[151,1436,181,1090]" captionTargetPageId="18" captionText="FIGURE 13. Geographical distribution of the genus Stenostygnus in South America. Stenostygnus pusio (circles) in Colombia, Ecuador and Brazil, Stenostygnus huberi spec. nov. (square) in Lara State, Venezuela and Stenostygnus martensi spec. nov. (triangle) in Mérida State, Venezuela." figureDoi="http://doi.org/10.5281/zenodo.4926919" httpUri="https://zenodo.org/record/4926919/files/figure.png" pageId="6" pageNumber="188">Fig.13</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>