treatments-xml/data/18/90/A5/1890A560C11158B8BCA96992E847D0AA.xml

<document ID-DOI="http://dx.doi.org/10.3897/zse.99.90875" ID-Pensoft-Pub="1860-0743-1-45" ID-Pensoft-UUID="088FB3FF68605537BB17D8F33EB98F01" ID-ZooBank="B9F5B3C59F014D66B5422850719DEBEA" ModsDocID="1860-0743-99-1-45" checkinTime="1673085750920" checkinUser="pensoft" docAuthor="Siyam, Manal, Dunlop, Jason A., Kovarik, Frantisek &amp; Mohammad, Abubakr" docDate="2023" docId="1890A560C11158B8BCA96992E847D0AA" docLanguage="en" docName="ZoosystEvol 99(1): 45-53" docOrigin="Zoosystematics and Evolution 99 (1)" docPubDate="2023-01-06" docSource="http://dx.doi.org/10.3897/zse.99.90875" docTitle="Compsobuthus werneri" docType="treatment" docVersion="1" id="088FB3FF68605537BB17D8F33EB98F01" lastPageNumber="45" masterDocId="088FB3FF68605537BB17D8F33EB98F01" masterDocTitle="Additions to the distribution of Sudanese scorpions" masterLastPageNumber="53" masterPageNumber="45" pageNumber="45" updateTime="1673085750920" updateUser="pensoft">
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<mods:title>Additions to the distribution of Sudanese scorpions</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Siyam, Manal</mods:namePart>
<mods:affiliation>University of Khartoum, Faculty of Science, Sudan Natural History Museum, Al-Jamaa Avenue, P. O Box 321, Khartoum, Sudan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dunlop, Jason A.</mods:namePart>
<mods:affiliation>Museum fuer Naturkunde, Leibniz Institute for Research on Evolution and Biodiversity, Invalidenstrasse 43, D- 10115 Berlin, Germany</mods:affiliation>
<mods:nameIdentifier type="email">jason.dunlop@mfn.berlin</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kovarik, Frantisek</mods:namePart>
<mods:affiliation>Charles University, Department of Zoology, Vinicna 7, CZ- 128 44 Praha 2, Czech Republic</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mohammad, Abubakr</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7297-9284</mods:nameIdentifier>
<mods:affiliation>Independent researcher, Natural Heritage, Sudan &amp; Current address: Flat 4, 19 Byram Street, Huddersfield, HD 1 1 DR, UK</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Zoosystematics and Evolution</mods:title>
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<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-06</mods:number>
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<mods:detail type="volume">
<mods:number>99</mods:number>
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<mods:detail type="issue">
<mods:number>1</mods:number>
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<mods:start>45</mods:start>
<mods:end>53</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zse.99.90875</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.99.90875</mods:identifier>
<mods:identifier type="Pensoft-Pub">1860-0743-1-45</mods:identifier>
<mods:identifier type="ZooBank">B9F5B3C59F014D66B5422850719DEBEA</mods:identifier>
<mods:identifier type="Pensoft-UUID">088FB3FF68605537BB17D8F33EB98F01</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:1890A560C11158B8BCA96992E847D0AA" httpUri="http://treatment.plazi.org/id/1890A560C11158B8BCA96992E847D0AA" lastPageNumber="45" pageId="0" pageNumber="45">
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<paragraph pageId="0" pageNumber="45">
<taxonomicName LSID="1890A560-C111-58B8-BCA9-6992E847D0AA" authority="(Birula, 1908)" baseAuthorityName="Birula" baseAuthorityYear="1908" class="Arachnida" family="Buthidae" genus="Compsobuthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Compsobuthus werneri" order="Scorpiones" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="werneri">Compsobuthus werneri (Birula, 1908)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="description">
<paragraph pageId="0" pageNumber="45">
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus photographs of several Sudanese scorpion species, all in the family Buthidae, for which we can offer new distribution data here. a. Androctonus amoreuxi (Audouin, 1826); b. Buthacus leptochelys (Ehrenberg, 1829); c. Compsobuthus werneri (Birula, 1908); d. Leiurus quinquestriatus (Ehrenberg, 1829); e. Orthochirus olivaceus (Karsch, 1881); f. Parabuthus abyssinicus (Pocock, 1901). Of these, A. amoreuxi, L. quinquestriatus and P. abyssinicus have a potent venom and are regarded as medically significant." figureDoi="10.3897/zse.99.90875.figure2" httpUri="https://binary.pensoft.net/fig/792292" pageId="0" pageNumber="45">Figure 2c</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="materials_examined">
<paragraph pageId="0" pageNumber="45">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="45">
<materialsCitation accessionNumber="ZMB 48709" collectingDate="2011-10-06" collectorName="M. Siyam" country="Sudan" county="The Republic" determinerName="F. Kovarik." elevation="1084" latitude="18.796944" location="Jebel El-sitt" longLatPrecision="1" longitude="37.11222" municipality="Erkweet" specimenCount="1" specimenCount-female="1" stateProvince="Red Sea State">
<specimenCount type="female">1♀</specimenCount>
; 
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/ZMB48709">ZMB 48709</accessionNumber>
, 
<collectingCounty>The Republic</collectingCounty>
of 
<collectingCountry name="Sudan">Sudan</collectingCountry>
, 
<collectingRegion country="Sudan" name="Al Bahr al Ahmar">Red Sea State</collectingRegion>
, 
<collectingMunicipality>Erkweet</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:1890A560C11158B8BCA96992E847D0AA:B261B58B1E754983A1615CA1F3C61941" country="Sudan" county="The Republic" latitude="18.796944" longLatPrecision="1" longitude="37.11222" municipality="Erkweet" name="Jebel El-sitt" stateProvince="Red Sea State">Jebel El-sitt</location>
(
<geoCoordinate degrees="18" direction="north" minutes="47" orientation="latitude" precision="1" seconds="49.0" value="18.796944">18°47'49.0&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="37" direction="east" minutes="06" orientation="longitude" precision="1" seconds="44.0" value="37.11222">37°06'44.0&quot;E</geoCoordinate>
, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.084" unit="m" value="1084.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.084" unit="m" value="1084.0">1084 m</elevation>
</quantity>
a.s.l), leg. 
<collectorName>M. Siyam</collectorName>
, 
<collectingDate value="2011-10-06">06.10.2011</collectingDate>
, det. 
<determinerName>
F. 
<normalizedToken originalValue="Kovařík">Kovarik</normalizedToken>
.
</determinerName>
</materialsCitation>
<materialsCitation collectingDate="2021-08-10" collectorName="H. Marzoug" country="Sudan" determinerName="M. Siyam." elevation="685" latitude="20.34042" location="El Madeqeen - Kouya" longLatPrecision="1" longitude="30.40493" specimenCount="1" specimenCount-female="1" stateProvince="Northern State">
<specimenCount type="female">1♀</specimenCount>
; 2.550, 
<collectingRegion inferredBySuffix="true">Northern State</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:1890A560C11158B8BCA96992E847D0AA:D34EE0BDB8B3476111519C49A35780A3" country="Sudan" latitude="20.34042" longLatPrecision="1" longitude="30.40493" name="El Madeqeen - Kouya" stateProvince="Northern State">El Madeqeen - Kouya</location>
(
<geoCoordinate degrees="20" direction="north" minutes="20" orientation="latitude" precision="1" seconds="25.512" value="20.34042">20°20'25.512&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="30" direction="east" minutes="24" orientation="longitude" precision="1" seconds="17.748" value="30.40493">30°24'17.748&quot;E</geoCoordinate>
, 
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.85" unit="m" value="685.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.85" unit="m" value="685.0">685 m</elevation>
</quantity>
a.s.l), 
<collectingDate value="2021-08-10">10.08.2021</collectingDate>
, leg. 
<collectorName>H. Marzoug</collectorName>
, det. 
<determinerName>M. Siyam.</determinerName>
</materialsCitation>
<materialsCitation accessionNumber="ZMB 49494" collectingDate="2016-06-07" collectorName="M. Siyam" country="Sudan" determinerName="F. Kovarik." elevation="228" latitude="19.144112" location="Dongola" longLatPrecision="1" longitude="30.481" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Northern State">
<specimenCount type="male">1♂</specimenCount>
<specimenCount type="female">1♀</specimenCount>
; 
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/ZMB49494">ZMB 49494</accessionNumber>
, 
<collectingRegion inferredBySuffix="true">Northern State</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:1890A560C11158B8BCA96992E847D0AA:5730E0D99C56A4D0ABA08D0D9F61E811" country="Sudan" latitude="19.144112" longLatPrecision="1" longitude="30.481" name="Dongola" stateProvince="Northern State">Dongola</location>
(
<geoCoordinate degrees="19" direction="north" minutes="08" orientation="latitude" precision="1" seconds="38.8" value="19.144112">19°08'38.8&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="30" direction="east" minutes="28" orientation="longitude" precision="1" seconds="51.6" value="30.481">30°28'51.6&quot;E</geoCoordinate>
, 
<quantity metricMagnitude="2" metricUnit="m" metricValue="2.28" unit="m" value="228.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="2.28" unit="m" value="228.0">228 m</elevation>
</quantity>
a.s.l), leg. 
<collectorName>M. Siyam</collectorName>
, 
<collectingDate value="2016-06-07">07.06.2016</collectingDate>
, det. 
<determinerName>
F. 
<normalizedToken originalValue="Kovařík">Kovarik</normalizedToken>
.
</determinerName>
</materialsCitation>
<materialsCitation accessionNumber="ZMB 49448, ZMB 49493" collectingDate="2019-09-15" collectorName="Al-Khidir" country="Sudan" determinerName="M. Siyam." elevation="219" latitude="19.463196" location="Wadi" longLatPrecision="1" longitude="30.256472" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Northern State">
<specimenCount type="male">1♂</specimenCount>
<specimenCount type="female">1♀</specimenCount>
; 
<accessionNumber isEnumeration="true">ZMB 49448, ZMB 49493</accessionNumber>
, 
<collectingRegion inferredBySuffix="true">Northern State</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:1890A560C11158B8BCA96992E847D0AA:B227637AC339535051B5BA652302FB8C" country="Sudan" latitude="19.463196" longLatPrecision="1" longitude="30.256472" name="Wadi" stateProvince="Northern State">Wadi</location>
<normalizedToken originalValue="El-Ga’ab">El-Ga'ab</normalizedToken>
(
<geoCoordinate degrees="19" direction="north" minutes="27" orientation="latitude" precision="1" seconds="47.5" value="19.463196">19°27'47.5&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="30" direction="east" minutes="15" orientation="longitude" precision="1" seconds="23.3" value="30.256472">30°15'23.3&quot;E</geoCoordinate>
, 
<quantity metricMagnitude="2" metricUnit="m" metricValue="2.19" unit="m" value="219.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="2.19" unit="m" value="219.0">219 m</elevation>
</quantity>
a.s.l), 
<collectingDate value="2019-09-15">15.09.2019</collectingDate>
, leg. I, 
<collectorName>Al-Khidir</collectorName>
, det. 
<determinerName>M. Siyam.</determinerName>
</materialsCitation>
<materialsCitation collectingDate="2018-09-14" collectorName="M. Siyam" country="Sudan" determinerName="F. Kovarik." elevation="600" latitude="13.073889" location="El-Obeid - Tagat" longLatPrecision="1" longitude="30.345612" specimenCount="1" specimenCount-female="1" stateProvince="North Kordofan State">
<specimenCount type="female">1♀</specimenCount>
; SNHMK 2.306, 
<collectingRegion inferredBySuffix="true">North Kordofan State</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:1890A560C11158B8BCA96992E847D0AA:23A24E9496A2CAD5E18133DF2F20B7A2" country="Sudan" latitude="13.073889" longLatPrecision="1" longitude="30.345612" name="El-Obeid - Tagat" stateProvince="North Kordofan State">El-Obeid - Tagat</location>
(
<geoCoordinate degrees="13" direction="north" minutes="04" orientation="latitude" precision="1" seconds="26.0" value="13.073889">13°04'26.0&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="30" direction="east" minutes="20" orientation="longitude" precision="1" seconds="44.2" value="30.345612">30°20'44.2&quot;E</geoCoordinate>
, 
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">600 m</elevation>
</quantity>
a.s.l), leg. 
<collectorName>M. Siyam</collectorName>
, 
<collectingDate value="2018-09-14">14.9.2018</collectingDate>
, det. 
<determinerName>
F. 
<normalizedToken originalValue="Kovařík">Kovarik</normalizedToken>
.
</determinerName>
</materialsCitation>
<materialsCitation collectingDate="2018-09-17" collectorName="M. Siyam" country="Sudan" determinerName="F. Kovarik." elevation="385" latitude="13.160944" location="Kosti" longLatPrecision="1" longitude="32.683113" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="White Nile State">
<specimenCount type="male">1♂</specimenCount>
<specimenCount type="female">1♀</specimenCount>
; SNHMK 2.311, SNHMK 2.310, 
<collectingRegion country="Sudan" name="An Nil al Abyad">White Nile State</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:1890A560C11158B8BCA96992E847D0AA:573CD077E5618E5AB1D3EFAEA7F91959" country="Sudan" latitude="13.160944" longLatPrecision="1" longitude="32.683113" name="Kosti" stateProvince="White Nile State">Kosti</location>
- a farm (
<geoCoordinate degrees="13" direction="north" minutes="09" orientation="latitude" precision="1" seconds="39.4" value="13.160944">13°09'39.4&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="32" direction="east" minutes="40" orientation="longitude" precision="1" seconds="59.2" value="32.683113">32°40'59.2&quot;E</geoCoordinate>
, 
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.85" unit="m" value="385.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="3.85" unit="m" value="385.0">385 m</elevation>
</quantity>
a.s.l), leg. 
<collectorName>M. Siyam</collectorName>
, 
<collectingDate value="2018-09-17">17.09.2018</collectingDate>
, det. 
<determinerName>
F. 
<normalizedToken originalValue="Kovařík">Kovarik</normalizedToken>
.
</determinerName>
</materialsCitation>
<materialsCitation accessionNumber="ZMB 49440" collectingDate="2019-02-19" collectorName="M. Siyam, O. Khalil" country="Sudan" determinerName="M. Siyam." elevation="502" latitude="14.279388" location="Muraba'a village - Berno Wadi" longLatPrecision="1" longitude="35.876694" specimenCount="4" specimenCount-female="4" stateProvince="Kassala State">
<specimenCount type="female">4♀</specimenCount>
; 
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/ZMB49440">ZMB 49440</accessionNumber>
-443, 
<collectingRegion country="Sudan" name="Kassala">Kassala State</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:1890A560C11158B8BCA96992E847D0AA:881E0A9A34DFCBF430352485C6EACB4D" country="Sudan" latitude="14.279388" longLatPrecision="1" longitude="35.876694" name="Muraba'a village - Berno Wadi" stateProvince="Kassala State">
<normalizedToken originalValue="Muraba’a">Muraba'a</normalizedToken>
village - Berno Wadi
</location>
(
<geoCoordinate degrees="14" direction="north" minutes="16" orientation="latitude" precision="1" seconds="45.8" value="14.279388">14°16'45.8&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="35" direction="east" minutes="52" orientation="longitude" precision="1" seconds="36.1" value="35.876694">35°52'36.1&quot;E</geoCoordinate>
, 
<quantity metricMagnitude="2" metricUnit="m" metricValue="5.02" unit="m" value="502.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="5.02" unit="m" value="502.0">502 m</elevation>
</quantity>
a.s.l), leg. 
<collectorName>M. Siyam</collectorName>
and 
<collectorName>O. Khalil</collectorName>
, 14- 
<collectingDate value="2019-02-19">19.02.2019</collectingDate>
, det. 
<determinerName>M. Siyam.</determinerName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="description">
<paragraph pageId="0" pageNumber="45">Description.</paragraph>
<paragraph pageId="0" pageNumber="45">Male / female total body length (5-6 cm). General coloration pale brown. Prosoma light brown with darker carinae. Mesosoma brown with granulated tergites except VII with lighter coloration. Metasoma yellowish brown, thin and ventral surface granulated with vertical black bands. Telson bulbous, brown, aculeus shorter than vesicle; tip black. Legs yellow. Pedialps medium yellowish brown, manus slightly swollen. Pectines with 17 teeth in females and 20 teeth in males.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="remarks">
<paragraph pageId="0" pageNumber="45">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="45">
<taxonomicName authorityName="Vachon" authorityYear="1949" class="Arachnida" family="Buthidae" genus="Compsobuthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Compsobuthus" order="Scorpiones" pageId="0" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="45">Compsobuthus</emphasis>
</taxonomicName>
is one of the most widely distributed genera of the family 
<taxonomicName authorityName="C. L. Koch" authorityYear="1837" class="Arachnida" family="Buthidae" kingdom="Animalia" lsidName="" order="Scorpiones" pageId="0" pageNumber="45" phylum="Arthropoda" rank="family">Buthidae</taxonomicName>
with records from the Republic of Sudan, Ethiopia and Somaliland as the south-eastern limits of distribution for this genus (
<bibRefCitation DOI="https://doi.org/10.18590/euscorpius.2016.vol2016.iss226.1" author="Kovarik, F" journalOrPublisher="Euscorpius" pageId="0" pageNumber="45" pagination="1 - 21" refId="B25" refString="Kovarik, F, Lowe, G, Pliskova, J, Stahlavsky, F, 2016a. Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part VI. Compsobuthus Vachon, 1949 (Buthidae), with a description of C. eritreaensis sp. n. Euscorpius 226 (226): 1 - 21, DOI: https://doi.org/10.18590/euscorpius.2016.vol2016.iss226.1" title="Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part VI. Compsobuthus Vachon, 1949 (Buthidae), with a description of C. eritreaensis sp. n." url="https://doi.org/10.18590/euscorpius.2016.vol2016.iss226.1" volume="226" year="2016 a">
<normalizedToken originalValue="Kovařík">Kovarik</normalizedToken>
et al. 2016a
</bibRefCitation>
). Despite its small body size, 
<taxonomicName baseAuthorityName="Birula" baseAuthorityYear="1908" class="Arachnida" family="Buthidae" genus="Compsobuthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Compsobuthus werneri" order="Scorpiones" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="werneri">
<emphasis italics="true" pageId="0" pageNumber="45">Compsobuthus werneri</emphasis>
</taxonomicName>
is a fairly toxic species, usually found under stones or embedded in sand or underground. It was previously documented from several localities in the Republic of Sudan (see 
<bibRefCitation DOI="https://doi.org/10.13156/arac.2018.17.9.449" author="Dunlop, JA" journalOrPublisher="Arachnology" pageId="0" pageNumber="45" pagination="449 - 457" refId="B10" refString="Dunlop, JA, Siyam, M, Kovarik, F, 2018. Smaller orders of Arachnida in Sudan: A literature review. Arachnology 17 (9): 449 - 457, DOI: https://doi.org/10.13156/arac.2018.17.9.449" title="Smaller orders of Arachnida in Sudan: A literature review." url="https://doi.org/10.13156/arac.2018.17.9.449" volume="17" year="2018">Dunlop et al. 2018</bibRefCitation>
) and its range is here expanded to include four new localities representing three new states (North Kordofan, White Nile and Kassala).
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
Genus 
<taxonomicName authorityName="Ehrenberg" authorityYear="1828" class="Arachnida" family="Buthidae" genus="Leiurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leiurus" order="Scorpiones" pageId="0" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="45">Leiurus</emphasis>
</taxonomicName>
Ehrenberg, 1828
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>