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<mods:title id="2C513743E713B0D1207065AC23AA2A3F">A monographic catalogue on the systematics and phylogeny of the South American iguanian lizard family Liolaemidae (Squamata, Iguania)</mods:title>
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<taxonomicName id="7BB630F1FF85612D6CFBEA58FDAA8838" ID-CoL="6QHKX" authority="(Donoso-Barros)" baseAuthorityName="Donoso-Barros" box="[151,613,327,353]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="cranwelli">
<emphasis id="8EC29760FF85612D6CFBEA58FDAA8838" bold="true" box="[151,613,327,353]" pageId="53" pageNumber="54">
<emphasis id="8EC29760FF85612D6CFBEA58FE428838" bold="true" box="[151,397,327,353]" italics="true" pageId="53" pageNumber="54">Liolaemus cranwelli</emphasis>
(Donoso-Barros)
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<taxonomicName id="7BB630F1FF85612D6CFBEA6FFD6C88DE" ID-CoL="76FLN" authority="Donoso-Barros 1973 a: 133" authorityName="Donoso-Barros" authorityYear="1973" box="[151,675,368,391]" class="Reptilia" family="Liolaemidae" genus="Pelusaurus" kingdom="Animalia" order="Squamata" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="cranwelli">
<emphasis id="8EC29760FF85612D6CFBEA6FFEB488DE" box="[151,379,368,391]" italics="true" pageId="53" pageNumber="54">Pelusaurus cranwelli</emphasis>
<bibRefCitation id="D8273683FF85612D6DEEEA6FFD6C88DE" author="Donoso-Barros" box="[386,675,368,391]" firstAuthor="Donoso-Barros" page="133" pageId="53" pageNumber="54" pagination="132 - 134" refId="ref58210" refString="Donoso-Barros, R. (1973 a) Un nuevo saurio de Bolivia (Lacertilia, Iguanidae). Neotropica, 19, 132 - 134." type="journal article" year="1973" yearSuffix="a">Donoso-Barros 1973a: 133</bibRefCitation>
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(
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locality: Nueva Moka, 
<collectingRegion id="7E728590FF85612D6FB3EA6FFB9888DE" box="[991,1111,368,391]" country="Bolivia" name="Santa Cruz" pageId="53" pageNumber="54">Santa Cruz</collectingRegion>
, 
<collectingCountry id="C4A10BE2FF85612D680FEA6FFB7988DE" box="[1123,1206,368,391]" name="Bolivia" pageId="53" pageNumber="54">Bolivia</collectingCountry>
); 
<treatmentCitation id="3D176D63FF85612D68A5EA6FFA5888DE" author="Vanzolini, P. E." box="[1225,1431,368,391]" page="13" pageId="53" pageNumber="54" year="1986">
<bibRefCitation id="D8273683FF85612D68A5EA6FFA5888DE" author="Vanzolini" box="[1225,1431,368,391]" firstAuthor="Vanzolini" page="13" pageId="53" pageNumber="54" pagination="1 - 25" refId="ref73558" refString="Vanzolini, P. E. (1986) Addenda and corrigenda to the Catalogue of Neotropical Squamata. Smithsonian Herpetological Information Service, 70, 1 - 25." type="journal article" year="1986">Vanzolini 1986: 13</bibRefCitation>
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;
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<bibRefCitation id="D8273683FF85612D6CABEA8FFEBC88FE" author="Fugler" box="[199,371,400,423]" firstAuthor="Fugler" page="61" pageId="53" pageNumber="54" pagination="57 - 75" refId="ref60096" refString="Fugler, C. M. (1989) Lista preliminar de los saurios. Ecologia en Bolivia 13, 57 - 75." type="journal article" year="1989">Fugler 1989: 61</bibRefCitation>
; 
<bibRefCitation id="D8273683FF85612D6D13EA8FFD8188FE" author="Etheridge" box="[383,590,400,423]" firstAuthor="Etheridge" page="34" pageId="53" pageNumber="54" pagination="1 - 34" refId="ref59230" refString="Etheridge, R. (1995) Redescription of Ctenoblepharys adspersa Tschudi, 1845, and the taxonomy of Liolaeminae (Reptilia: Squamata: Tropiduridae). American Museum Novitates, 3142, 1 - 34." type="journal article" year="1995">Etheridge 1995: 34</bibRefCitation>
; 
<bibRefCitation id="D8273683FF85612D6E36EA8FFC5188FE" author="Etheridge &amp; Espinoza" box="[602,926,400,423]" firstAuthor="Etheridge" page="5" pageId="53" pageNumber="54" pagination="1 - 64" refId="ref59521" refString="Etheridge, R &amp; Espinoza, R. E. (2000) Taxonomy of the Liolaeminae (Squamata: Iguania: Tropiduridae) and a semiannotated bibliography. Smithsonian Herpetological Information Service, 126, 1 - 64." type="journal article" year="2000">Etheridge &amp; Espinoza 2000: 5</bibRefCitation>
; 
<bibRefCitation id="D8273683FF85612D6FC6EA8FFB1088FE" author="Pincheira-Donoso" box="[938,1247,400,423]" firstAuthor="Pincheira-Donoso" page="21" pageId="53" pageNumber="54" pagination="18 - 27" refId="ref69329" refString="Pincheira-Donoso, D. (2002 b) Lista Sistematica de los reptiles de Bolivia. Noticiario Mensual Museo Nacional de Historia Natural de Chile, 350, 18 - 27." type="journal article" year="2002" yearSuffix="b">Pincheira-Donoso 2002b: 21</bibRefCitation>
.
</paragraph>
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<taxonomicName id="7BB630F1FF85612D6CFBEAAFFDFF889E" authority="Laurent 1982 c: 9" authorityName="Laurent" authorityPageNumber="9" authorityYear="1982" box="[151,560,432,455]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="cranwelli">
<emphasis id="8EC29760FF85612D6CFBEAAFFEBE889E" box="[151,369,432,455]" italics="true" pageId="53" pageNumber="54">Liolaemus cranwelli</emphasis>
<bibRefCitation id="D8273683FF85612D6D14EAAFFDFF889E" author="Laurent" box="[376,560,432,455]" firstAuthor="Laurent" page="9" pageId="53" pageNumber="54" pagination="9 - 10" refId="ref63673" refString="Laurent, R. F. (1982 c) Sinonimia del genero Pelusaurus Donoso-Barros con Liolaemus (Sauria, Iguanidae). Boletin de la Asociacion Herpetologica Argentina, 1, 9 - 10." type="journal article" year="1982" yearSuffix="c">Laurent 1982c: 9</bibRefCitation>
</taxonomicName>
; 
<bibRefCitation id="D8273683FF85612D6E50EAAFFC80889E" author="Verrastro" box="[572,847,432,455]" etAl="et al." firstAuthor="Verrastro" page="109" pageId="53" pageNumber="54" pagination="105 - 118" refId="ref74055" refString="Verrastro, L., Veronese, L., Bujes, C. &amp; Martins Dias Filho, M. (2003) A new species of Liolaemus from southern Brazil (Iguania: Tropiduridae). Herpetologica, 59, 105 - 118." type="journal article" year="2003">
Verrastro 
<emphasis id="8EC29760FF85612D6EC8EAAFFD15889E" box="[676,730,432,455]" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
2003: 109
</bibRefCitation>
; 
<bibRefCitation id="D8273683FF85612D6F36EAAFFBCF889E" author="Sura" box="[858,1024,432,455]" firstAuthor="Sura" page="428" pageId="53" pageNumber="54" refId="ref73002" refString="Sura, P. 2005. Encyclopedia of extant amphibians and reptiles, Wydawnictwo Fundacja, Nowy Sacz, 544 pp." type="book" year="2005">Sura 2005: 428</bibRefCitation>
.
</paragraph>
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<taxonomicName id="7BB630F1FF85612D6CFBEACFFDA688BE" authority="Etheridge 2000: 310" authorityName="Etheridge" authorityPageNumber="310" authorityYear="2000" box="[151,617,464,487]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="wiegmannii">
<emphasis id="8EC29760FF85612D6CFBEACFFE4688BE" box="[151,393,464,487]" italics="true" pageId="53" pageNumber="54">Liolaemus wiegmannii</emphasis>
<bibRefCitation id="D8273683FF85612D6DFCEACFFDA688BE" author="Etheridge, R." box="[400,617,464,487]" pageId="53" pageNumber="54" pagination="293 - 352" refId="ref59302" refString="Etheridge, R. (2000) A review of lizards of the Liolaemus wiegmannii group (Squamata, Iguania, Tropiduridae), and a history of morphological change in the sand-dwelling species. Herpetological Monographs, 14, 293 - 352." type="journal article" year="2000">Etheridge 2000: 310</bibRefCitation>
</taxonomicName>
(
<emphasis id="8EC29760FF85612D6E15EACFFCC388BE" box="[633,780,464,487]" italics="true" pageId="53" pageNumber="54">nomina dubia</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="F4AC18F9FF85612D6CFBE908FE548D83" pageId="53" pageNumber="54" type="description">
<paragraph id="BC094B72FF85612D6CFBE908FAEA8A7B" blockId="53.[151,1436,535,1242]" pageId="53" pageNumber="54">
<emphasis id="8EC29760FF85612D6CFBE908FE898B68" bold="true" box="[151,326,535,561]" pageId="53" pageNumber="54">Observations:</emphasis>
<taxonomicName id="7BB630F1FF85612D6D23E907FD8D8B68" authorityName="Laurent" authorityYear="1982" box="[335,578,536,561]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="cranwelli">
<emphasis id="8EC29760FF85612D6D23E907FD8D8B68" box="[335,578,536,561]" italics="true" pageId="53" pageNumber="54">Liolaemus cranwelli</emphasis>
</taxonomicName>
is one of the most controversial species of the genus, mainly because it was established on the basis of a single male museum specimen (
<bibRefCitation id="D8273683FF85612D6FD5E95FFB0B8B03" author="Donoso-Barros" box="[953,1220,576,602]" firstAuthor="Donoso-Barros" pageId="53" pageNumber="54" pagination="132 - 134" refId="ref58210" refString="Donoso-Barros, R. (1973 a) Un nuevo saurio de Bolivia (Lacertilia, Iguanidae). Neotropica, 19, 132 - 134." type="journal article" year="1973" yearSuffix="a">Donoso-Barros 1973a</bibRefCitation>
; 
<bibRefCitation id="D8273683FF85612D68BFE95FFA5F8B03" author="Etheridge, R." box="[1235,1424,576,602]" pageId="53" pageNumber="54" pagination="293 - 352" refId="ref59302" refString="Etheridge, R. (2000) A review of lizards of the Liolaemus wiegmannii group (Squamata, Iguania, Tropiduridae), and a history of morphological change in the sand-dwelling species. Herpetological Monographs, 14, 293 - 352." type="journal article" year="2000">Etheridge 2000</bibRefCitation>
). There is no information on the living colour pattern of the species and the female is entirely unknown. Over the last decade, the debate on the validity of this species has increased (
<bibRefCitation id="D8273683FF85612D6F8DE98FFB558BF3" author="Etheridge, R." box="[993,1178,656,682]" pageId="53" pageNumber="54" pagination="293 - 352" refId="ref59302" refString="Etheridge, R. (2000) A review of lizards of the Liolaemus wiegmannii group (Squamata, Iguania, Tropiduridae), and a history of morphological change in the sand-dwelling species. Herpetological Monographs, 14, 293 - 352." type="journal article" year="2000">Etheridge 2000</bibRefCitation>
; see also 
<bibRefCitation id="D8273683FF85612D6967E98FFE868B8B" author="Etheridge &amp; Espinoza" firstAuthor="Etheridge" pageId="53" pageNumber="54" pagination="1 - 64" refId="ref59521" refString="Etheridge, R &amp; Espinoza, R. E. (2000) Taxonomy of the Liolaeminae (Squamata: Iguania: Tropiduridae) and a semiannotated bibliography. Smithsonian Herpetological Information Service, 126, 1 - 64." type="journal article" year="2000">Etheridge &amp; Espinoza 2000</bibRefCitation>
). 
<bibRefCitation id="D8273683FF85612D6D31E9A7FDEC8B8B" author="Etheridge, R." box="[349,547,696,722]" pageId="53" pageNumber="54" pagination="293 - 352" refId="ref59302" refString="Etheridge, R. (2000) A review of lizards of the Liolaemus wiegmannii group (Squamata, Iguania, Tropiduridae), and a history of morphological change in the sand-dwelling species. Herpetological Monographs, 14, 293 - 352." type="journal article" year="2000">Etheridge (2000)</bibRefCitation>
suggested that 
<taxonomicName id="7BB630F1FF85612D6EB5E9A7FCA98B88" authorityName="Laurent" authorityYear="1982" box="[729,870,696,721]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="cranwelli">
<emphasis id="8EC29760FF85612D6EB5E9A7FCA98B88" box="[729,870,696,721]" italics="true" pageId="53" pageNumber="54">L. cranwelli</emphasis>
</taxonomicName>
may be a synonym of 
<taxonomicName id="7BB630F1FF85612D681DE9A7FAD78B88" authorityName="Etheridge" authorityYear="2000" box="[1137,1304,696,721]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="wiegmannii">
<emphasis id="8EC29760FF85612D681DE9A7FAD78B88" box="[1137,1304,696,721]" italics="true" pageId="53" pageNumber="54">L. wiegmannii</emphasis>
</taxonomicName>
. Nevertheless, further studies are needed to clarify the problem. Unfortunately, because it is currently unfeasible to gather information on the variation of 
<taxonomicName id="7BB630F1FF85612D6E34E817FD2A8A78" authorityName="Laurent" authorityYear="1982" box="[600,741,776,801]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="cranwelli">
<emphasis id="8EC29760FF85612D6E34E817FD2A8A78" box="[600,741,776,801]" italics="true" pageId="53" pageNumber="54">L. cranwelli</emphasis>
</taxonomicName>
, powerful conclusions might not be possible yet.
</paragraph>
<paragraph id="BC094B72FF85612D6CABE82FFE548D83" blockId="53.[151,1436,535,1242]" pageId="53" pageNumber="54">
Additionally, it is necessary to clarify some aspects concerning the exact type locality of this species. In his monograph, 
<bibRefCitation id="D8273683FF85612D6D3BE847FDEF8A2B" author="Etheridge, R." box="[343,544,856,882]" pageId="53" pageNumber="54" pagination="293 - 352" refId="ref59302" refString="Etheridge, R. (2000) A review of lizards of the Liolaemus wiegmannii group (Squamata, Iguania, Tropiduridae), and a history of morphological change in the sand-dwelling species. Herpetological Monographs, 14, 293 - 352." type="journal article" year="2000">Etheridge (2000)</bibRefCitation>
pointed out that the only known specimen of 
<taxonomicName id="7BB630F1FF85612D6829E847FB1B8A28" authorityName="Laurent" authorityYear="1982" box="[1093,1236,856,881]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="cranwelli">
<emphasis id="8EC29760FF85612D6829E847FB1B8A28" box="[1093,1236,856,881]" italics="true" pageId="53" pageNumber="54">L. cranwelli</emphasis>
</taxonomicName>
was collected in “Macho Moka, 
<collectingRegion id="7E728590FF85612D6D3DE89FFE198AC3" box="[337,470,896,922]" inferredBySuffix="true" pageId="53" pageNumber="54">Santa Cruz</collectingRegion>
, 
<collectingCountry id="C4A10BE2FF85612D6D8EE89FFDF18AC3" box="[482,574,896,922]" name="Bolivia" pageId="53" pageNumber="54">Bolivia</collectingCountry>
”, and that the distribution of this taxon would be “about 
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north of the Sierra de Santa Bárbara....., from Macho, Nueva Moka, in the department of 
<collectingRegion id="7E728590FF85612D681FE8B7FB348A9B" box="[1139,1275,936,962]" country="Bolivia" name="Santa Cruz" pageId="53" pageNumber="54">Santa Cruz</collectingRegion>
, southeastern 
<collectingCountry id="C4A10BE2FF85612D6CFBE8CFFF3C8AB3" box="[151,243,976,1002]" name="Bolivia" pageId="53" pageNumber="54">Bolivia</collectingCountry>
”. However, this information is partially misleading, because “Macho” is not a geographical point, but the Spanish word “male”. In the original publication, 
<bibRefCitation id="D8273683FF85612D6F2DE8E7FB928D4B" author="Donoso-Barros" box="[833,1117,1016,1042]" firstAuthor="Donoso-Barros" pageId="53" pageNumber="54" pagination="132 - 134" refId="ref58210" refString="Donoso-Barros, R. (1973 a) Un nuevo saurio de Bolivia (Lacertilia, Iguanidae). Neotropica, 19, 132 - 134." type="journal article" year="1973" yearSuffix="a">Donoso-Barros (1973a)</bibRefCitation>
detailed “Holotipo: 3632 M.A.C.N. Macho, Nueva Moka, 
<collectingRegion id="7E728590FF85612D6E73EF3FFD6A8D63" box="[543,677,1056,1082]" country="Bolivia" name="Santa Cruz" pageId="53" pageNumber="54">Santa Cruz</collectingRegion>
, 
<collectingCountry id="C4A10BE2FF85612D6EDEEF3FFCC18D63" box="[690,782,1056,1082]" name="Bolivia" pageId="53" pageNumber="54">Bolivia</collectingCountry>
”, which really means “ 
<typeStatus id="630DF5D0FF85612D6872EF3FFB5D8D63" box="[1054,1170,1056,1082]" pageId="53" pageNumber="54" type="holotype">Holotype</typeStatus>
: 3632 M.A.C.N. Male [individual], [collected in] Nueva Moka, 
<collectingRegion id="7E728590FF85612D6E16EF57FD318D3B" box="[634,766,1096,1122]" country="Bolivia" name="Santa Cruz" pageId="53" pageNumber="54">Santa Cruz</collectingRegion>
, 
<collectingCountry id="C4A10BE2FF85612D6F66EF57FCA98D3B" box="[778,870,1096,1122]" name="Bolivia" pageId="53" pageNumber="54">Bolivia</collectingCountry>
”. Consequently, the type locality of 
<taxonomicName id="7BB630F1FF85612D6962EF57FA538D38" authorityName="Laurent" authorityYear="1982" box="[1294,1436,1096,1121]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="cranwelli">
<emphasis id="8EC29760FF85612D6962EF57FA538D38" box="[1294,1436,1096,1121]" italics="true" pageId="53" pageNumber="54">L. cranwelli</emphasis>
</taxonomicName>
is “Nueva Moka, 
<collectingRegion id="7E728590FF85612D6D0AEF6FFE258DD3" box="[358,490,1136,1162]" country="Bolivia" name="Santa Cruz" pageId="53" pageNumber="54">Santa Cruz</collectingRegion>
, 
<collectingCountry id="C4A10BE2FF85612D6D9BEF6FFD9C8DD3" box="[503,595,1136,1162]" name="Bolivia" pageId="53" pageNumber="54">Bolivia</collectingCountry>
”, and the proposed distribution provided by 
<bibRefCitation id="D8273683FF85612D6830EF6FFAEC8DD3" author="Etheridge, R." box="[1116,1315,1136,1162]" pageId="53" pageNumber="54" pagination="293 - 352" refId="ref59302" refString="Etheridge, R. (2000) A review of lizards of the Liolaemus wiegmannii group (Squamata, Iguania, Tropiduridae), and a history of morphological change in the sand-dwelling species. Herpetological Monographs, 14, 293 - 352." type="journal article" year="2000">Etheridge (2000)</bibRefCitation>
should be “about 
<quantity id="7B4EE697FF85612D6C81EF87FE898DEB" box="[237,326,1176,1202]" metricMagnitude="5" metricUnit="m" metricValue="4.7" pageId="53" pageNumber="54" unit="km" value="470.0">470 km</quantity>
north of the Sierra de Santa Bárbara....., from Nueva Moka, in the department of 
<collectingRegion id="7E728590FF85612D697EEF87FA578DEB" box="[1298,1432,1176,1202]" country="Bolivia" name="Santa Cruz" pageId="53" pageNumber="54">Santa Cruz</collectingRegion>
, southeastern 
<collectingCountry id="C4A10BE2FF85612D6D5DEFDFFE428D83" box="[305,397,1216,1242]" name="Bolivia" pageId="53" pageNumber="54">Bolivia</collectingCountry>
”.
</paragraph>
</subSubSection>
</treatment>
</document>