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<mods:title id="75A1131CCA6F4ED046663C2839668ABA">Redescriptions of three species of freshwater crabs from Angola, southern Africa (Brachyura: Potamoidea: Potamonautidae)</mods:title>
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<mods:namePart id="BF3DAC7F2C3E0AAD27ECDF2702C965CC">Ahles, Katelyn M.</mods:namePart>
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<mods:namePart id="7CC2DF1899F30CCD1EB1B51DC4D3C182">Daniels, Savel R.</mods:namePart>
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<taxonomicName id="88C94D036040FFCA6B83FA72FDB0FA3E" ID-CoL="4M4J6" authority="Bott, 1953" authorityName="Bott" authorityYear="1953" box="[151,629,1410,1437]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="521" phylum="Arthropoda" rank="species" species="macrobrachii">
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<emphasis id="7DBDEA926040FFCA6B83FA72FE2BFA3F" bold="true" box="[151,494,1410,1437]" italics="true" pageId="5" pageNumber="521">Potamonautes macrobrachii</emphasis>
<bibRefCitation id="2B584B716040FFCA6AE2FA72FDB0FA3E" author="Bott, R." box="[502,629,1410,1437]" pageId="5" pageNumber="521" pagination="133 - 148" refId="ref9019" refString="Bott, R. (1953) Potamoniden (Crust. Decap.) von Angola. Publicacoes culturais da Companhia de Diamantes de Angola, Lisboa, 16, 133 - 148." type="journal article" year="1953">Bott, 1953</bibRefCitation>
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<paragraph id="4F7636806040FFCA6B83FA56FD86FA63" blockId="5.[151,629,1410,1473]" box="[151,579,1446,1473]" pageId="5" pageNumber="521">
(
<figureCitation id="D7F22A056040FFCA6B8BFA56FF2BFA63" box="[159,238,1447,1473]" captionStart="FIGURE 3" captionStartId="9.[151,250,1957,1981]" captionTargetBox="[191,1396,181,1932]" captionTargetId="figure-21@9.[191,1396,181,1932]" captionTargetPageId="9" captionText="FIGURE 3. Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Whole animal, dorsal view (A), whole animal, ventral view (B). Scale bars: A = 12.5 mm, B = 5.38 mm." figureDoi="http://doi.org/10.5281/zenodo.5497105" httpUri="https://zenodo.org/record/5497105/files/figure.png" pageId="5" pageNumber="521">Figs. 3</figureCitation>
, 
<figureCitation id="D7F22A056040FFCA6BEFFA56FEDBFA63" box="[251,286,1447,1473]" captionStart="FIGURE 4" captionStartId="10.[151,250,1747,1771]" captionTargetBox="[152,1436,309,1723]" captionTargetId="figure-93@10.[152,1436,309,1723]" captionTargetPageId="10" captionText="FIGURE 4. Frontal view of carapace. (A) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (B) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (C) Potamonautes macrobrachii Bott, 1953, paratype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Photograph B by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A = 10.4 mm; B = 5.7 mm; C = 4.7 mm." figureDoi="http://doi.org/10.5281/zenodo.5497107" httpUri="https://zenodo.org/record/5497107/files/figure.png" pageId="5" pageNumber="521">4C</figureCitation>
, 
<figureCitation id="D7F22A056040FFCA6A3DFA57FE62FA63" box="[297,423,1446,1473]" captionStart="FIGURE 5" captionStartId="11.[151,250,1588,1612]" captionTargetBox="[159,1428,181,1565]" captionTargetId="figure-21@11.[159,1428,181,1565]" captionTargetPageId="11" captionText="FIGURE 5. (A–F) Frontal view of right and left chela; (G–L) cheliped merus and carpus. (A, B) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (C, D) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (E, F) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892); (G, H) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (G) ventral view of right chela merus; (H) dorsal view of right chela carpus and merus; (I, J) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province,Angola (AMG GEN 1887A) (I) ventral view of right chela merus; (J) dorsal view of right chela merus and carpus; (K, L) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892): (K) ventral view of right chela merus; (L) dorsal view of right chela merus and carpus;. Photographs C, D, I, J by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A = 18.4 mm; B = 17.9 mm; C, D = 8.9 mm; E = 8.33 mm; F = 5.81 mm; G = 17.9 mm; H = 7.3 mm; I = 8.9 mm; J = 5.9 mm; K = 5.81 mm; L = 5.6 mm." figureDoi="http://doi.org/10.5281/zenodo.5497109" httpUri="https://zenodo.org/record/5497109/files/figure.png" pageId="5" pageNumber="521">5E, F, K, L</figureCitation>
, 
<figureCitation id="D7F22A056040FFCA6AA7FA56FE37FA63" box="[435,498,1447,1473]" captionStart="FIGURE 6" captionStartId="12.[151,250,1828,1852]" captionTargetBox="[197,1391,181,1805]" captionTargetId="figure-17@12.[197,1391,181,1805]" captionTargetPageId="12" captionText="FIGURE 6. Right G1, ventral view (A–C) and dorsal view (D, F). (A, D) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (B, E) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (C, F) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Photo- graphs B and E by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A= 6.7 mm; B = 2.3 mm; C = 1.13 mm; D = 2.7 mm; E = 2.3 mm; F = 1.2." figureDoi="http://doi.org/10.5281/zenodo.5497111" httpUri="https://zenodo.org/record/5497111/files/figure.png" pageId="5" pageNumber="521">6C, F</figureCitation>
, 
<figureCitation id="D7F22A056040FFCA6AE9FA56FDE5FA63" box="[509,544,1447,1473]" captionStart="FIGURE 7" captionStartId="13.[151,250,1855,1879]" captionTargetBox="[264,1323,181,1831]" captionTargetId="figure-21@13.[264,1323,181,1831]" captionTargetPageId="13" captionText="FIGURE 7. Right G2 ventral view (A–C); (A) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (B) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (C) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Photograph C by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A = 5.6 mm; B = 2.3 mm; C = 1.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5497113" httpUri="https://zenodo.org/record/5497113/files/figure.png" pageId="5" pageNumber="521">7C</figureCitation>
, 
<figureCitation id="D7F22A056040FFCA6938FA56FDFFFA63" box="[556,570,1447,1473]" captionStart="FIGURE 8" captionStartId="14.[151,250,1336,1360]" captionTargetBox="[159,1428,189,1302]" captionTargetId="figure-17@14.[151,1436,181,1312]" captionTargetPageId="14" captionText="FIGURE 8. Map of the Angola showing the known localities of Potamonautes adeleae (black square, not shown to scale), P. kensleyi (black circles, not shown to scale), and P. macrobrachii (black diamond, not shown to scale). Scale bar = 520 km. See text for exact locality coordinates." figureDoi="http://doi.org/10.5281/zenodo.5497115" httpUri="https://zenodo.org/record/5497115/files/figure.png" pageId="5" pageNumber="521">8</figureCitation>
)
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<treatmentCitation id="CE6810916040FFCA6B83FA1DFC93F9A1" author="Bott, R." box="[151,854,1515,1539]" page="139" pageId="5" pageNumber="521" year="1953">
<taxonomicName id="88C94D036040FFCA6B83FA1DFC93F9A1" ID-CoL="4M4J6" authority="Bott, 1953: 138 - 139" authorityName="Bott" authorityPageNumber="138 - 139" authorityYear="1953" box="[151,854,1515,1539]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="521" phylum="Arthropoda" rank="species" species="macrobrachii" subGenus="Isopotamonautes">
<emphasis id="7DBDEA926040FFCA6B83FA1DFDB9F9A1" box="[151,636,1515,1539]" italics="true" pageId="5" pageNumber="521">Potamonautes (Isopotamonautes) macrobrachii</emphasis>
<bibRefCitation id="2B584B716040FFCA6996FA1DFC93F9A1" author="Bott, R." box="[642,854,1515,1539]" pageId="5" pageNumber="521" pagination="133 - 148" refId="ref9019" refString="Bott, R. (1953) Potamoniden (Crust. Decap.) von Angola. Publicacoes culturais da Companhia de Diamantes de Angola, Lisboa, 16, 133 - 148." type="journal article" year="1953">Bott, 1953: 138–139</bibRefCitation>
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, pl. 4, 5; 
<bibRefCitation id="2B584B716040FFCA68A0FA1DFBD9F9A1" author="Bott, R." box="[948,1052,1515,1539]" pageId="5" pageNumber="521" pagination="209 - 352" refId="ref9053" refString="Bott, R. (1955) Die Susswasserkrabben von Afrika (Crust. Decap.) und Ihre Stammesgeschichte. Annales du Musee Du Congo Belge, Tervuren, Belgique), C Zoologie, (3, 3), 3 (1), 209 - 352." type="journal article" year="1955">Bott 1955</bibRefCitation>
: table IX, fig. 2 a–d, pl. 25, 78.b 
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<taxonomicName id="88C94D036040FFCA6B83F9FAFCF7F980" ID-CoL="4M4J6" authority=", Cumberlidge &amp; Tavares, 2006: 248" authorityName=", Cumberlidge &amp; Tavares" authorityPageNumber="248" authorityYear="2006" box="[151,818,1546,1570]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="521" phylum="Arthropoda" rank="species" species="macrobrachii">
<emphasis id="7DBDEA926040FFCA6B83F9FAFE7AF980" box="[151,447,1546,1570]" italics="true" pageId="5" pageNumber="521">Potamonautes macrobrachii,</emphasis>
<bibRefCitation id="2B584B716040FFCA6AD2F9FBFCF7F980" author="Cumberlidge, N. &amp; Tavares, M." box="[454,818,1546,1570]" pageId="5" pageNumber="521" pagination="248 - 257" refId="ref9684" refString="Cumberlidge, N. &amp; Tavares, M. (2006) Remarks on the freshwater crabs of Angola, southwestern Africa, with the description of Potamonautes kensleyi, new species (Brachyura: Potamoidea: Potamonautidae). Journal of Crustacean Biology, 26 (2), 248 - 257. https: // doi. org / 10.1651 / s- 2653.1" type="journal article" year="2006">Cumberlidge &amp; Tavares, 2006: 248</bibRefCitation>
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, 250, 253–256, table 1; 
<treatmentCitation id="CE6810916040FFCA6F3FF9FBFAE9F980" author="Cumberlidge, N." box="[1067,1324,1546,1570]" page="1" pageId="5" pageNumber="521" year="2008">
<bibRefCitation id="2B584B716040FFCA6F3FF9FBFAE9F980" author="Cumberlidge, N." box="[1067,1324,1546,1570]" pageId="5" pageNumber="521" pagination="64384" refId="ref9510" refString="Cumberlidge, N. (2008 b) Potamonautes kensleyi. The IUCN Red List of Threatened Species, 2008, e. T 64384 A 12765673. https: // doi. org / 10.2305 / IUCN. UK. 2008. RLTS. T 64384 A 12765673. en" type="journal article" year="2008">Cumberlidge 2008b: 1–8</bibRefCitation>
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, fig. 1; 
<bibRefCitation id="2B584B716040FFCA6E69F9FBFA59F980" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." box="[1405,1436,1546,1570]" pageId="5" pageNumber="521" pagination="1 - 286" refId="ref10320" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyuorum: part I. An annotated checklist of extant Brachyuran crabs of the world. Raffles Bulletin of Zoology, 17, 1 - 286." type="journal article" year="2008">Ng</bibRefCitation>
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</paragraph>
<paragraph id="4F7636806040FFCA6BD3F9D8FEA8F9E3" blockId="5.[151,1436,1515,1601]" box="[199,365,1576,1601]" pageId="5" pageNumber="521">
<emphasis id="7DBDEA926040FFCA6BD3F9D8FF3FF9E2" box="[199,250,1576,1600]" italics="true" pageId="5" pageNumber="521">et al.</emphasis>
2008: 171.
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<materialsCitation id="FFA13CDD6040FFCA6B83F99EFA5DF973" collectingDate="1949-09-05" collectionCode="SMF" collectorName="Kuketa &amp; A. de Barros Machado." country="Angola" county="Benguala" elevation="1800" latitude="-12.466667" location="Serra do Moco" longLatPrecision="1" longitude="15.166667" municipality="Luimbale" pageId="5" pageNumber="521" specimenCode="SMF 1892" specimenCount="1" stateProvince="Huambo" typeStatus="holotype">
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<typeStatus id="907288226040FFCA6B83F99EFEC3F92B" box="[151,262,1647,1673]" pageId="5" pageNumber="521" type="holotype">Holotype</typeStatus>
.
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<specimenCode id="1F6F9EFB6040FFCA6A06F99EFE49F92B" box="[274,396,1647,1673]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="521">SMF 1892</specimenCode>
, adult male (CW 32.91, CL 22.3, CH 10.9, FW 
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), 
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<collectingRegion id="8D0DF8626040FFCA6FE2F99EFF0DF90F" country="Angola" name="Huambo" pageId="5" pageNumber="521">Huambo Province</collectingRegion>
, 
<collectingMunicipality id="AF12ACFA6040FFCA6BC7F962FE84F90F" box="[211,321,1683,1709]" pageId="5" pageNumber="521">Luimbale</collectingMunicipality>
, 
<location id="4A16605B6040FFCA6A5FF962FE2BF90F" LSID="urn:lsid:plazi:treatment:C76087966040FFC16B83FA72FE1EF8A3:4A16605B6040FFCA6A5FF962FE2BF90F" box="[331,494,1683,1709]" country="Angola" county="Benguala" latitude="-12.466667" longLatPrecision="1" longitude="15.166667" municipality="Luimbale" name="Serra do Moco" pageId="5" pageNumber="521" stateProvince="Huambo">Serra do Moco</location>
(escarpment of 
<location id="4A16605B6040FFCA69B4F962FD26F90F" LSID="urn:lsid:plazi:treatment:C76087966040FFC16B83FA72FE1EF8A3:4A16605B6040FFCA69B4F962FD26F90F" box="[672,739,1683,1709]" country="Angola" county="Benguala" latitude="-12.466667" longLatPrecision="1" longitude="15.166667" municipality="Luimbale" name="Moco" pageId="5" pageNumber="521" stateProvince="Huambo">Moco</location>
) (
<geoCoordinate id="2AFD50476040FFCA69ECF962FCB8F90F" box="[760,893,1683,1709]" degrees="12.466667" direction="south" orientation="latitude" pageId="5" pageNumber="521" precision="1" value="-12.466667">-12.466667</geoCoordinate>
, 
<geoCoordinate id="2AFD50476040FFCA6893F963FBC5F90F" box="[903,1024,1682,1709]" degrees="15.166667" direction="east" orientation="longitude" pageId="5" pageNumber="521" precision="1" value="15.166667">15.166667</geoCoordinate>
) 
<quantity id="88319B656040FFCA6F19F962FBADF90F" box="[1037,1128,1683,1709]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="521" unit="m" value="1800.0">
<elevation id="C4E4D1B36040FFCA6F19F962FBADF90F" box="[1037,1128,1683,1709]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="521" unit="m" value="1800.0">1,800 m</elevation>
</quantity>
ASL, 
<collectorName id="E23C53566040FFCA6FBAF962FB3AF90F" box="[1198,1279,1683,1709]" pageId="5" pageNumber="521">Kuketa</collectorName>
stream (tributary of the 
<location id="4A16605B6040FFCA6A05F946FE17F970" LSID="urn:lsid:plazi:treatment:C76087966040FFC16B83FA72FE1EF8A3:4A16605B6040FFCA6A05F946FE17F970" box="[273,466,1719,1746]" country="Angola" county="Benguala" latitude="-12.466667" longLatPrecision="1" longitude="15.166667" municipality="Luimbale" name="Catumbela River" pageId="5" pageNumber="521" stateProvince="Huambo">Catumbela River</location>
) draining the 
<location id="4A16605B6040FFCA697AF946FC7BF973" LSID="urn:lsid:plazi:treatment:C76087966040FFC16B83FA72FE1EF8A3:4A16605B6040FFCA697AF946FC7BF973" box="[622,958,1719,1745]" country="Angola" county="Benguala" latitude="-12.466667" longLatPrecision="1" longitude="15.166667" municipality="Luimbale" name="Mount Moco Special Reserve" pageId="5" pageNumber="521" stateProvince="Huambo">Mount Moco Special Reserve</location>
, 
<date id="3B7710406040FFCA68DDF947FB94F973" box="[969,1105,1718,1745]" pageId="5" pageNumber="521" value="1949-09-05">
<collectingDate id="2B33E9A86040FFCA68DDF947FB94F973" box="[969,1105,1718,1745]" pageId="5" pageNumber="521" value="1949-09-05">5 Sep. 1949</collectingDate>
</date>
, coll. 
<collectorName id="E23C53566040FFCA6F87F946FA5DF973" box="[1171,1432,1719,1745]" pageId="5" pageNumber="521">A. de Barros Machado.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="4F7636806040FFC96BD3F92BFC35FF77" blockId="5.[151,1437,1647,2033]" lastBlockId="6.[151,1437,151,2013]" lastPageId="6" lastPageNumber="522" pageId="5" pageNumber="521">
<emphasis id="7DBDEA926040FFCA6BD3F92BFE9BF956" bold="true" box="[199,350,1754,1780]" pageId="5" pageNumber="521">Rediagnosis.</emphasis>
Surface of carapace completely smooth with no deep grooves, granules, or carinae; postfrontal crest completely crossing carapace, low in middle, sharp laterally, extending to meet anterolateral margins (
<figureCitation id="D7F22A056040FFCA6E70F90EFF7EF89F" captionStart="FIGURE 3" captionStartId="9.[151,250,1957,1981]" captionTargetBox="[191,1396,181,1932]" captionTargetId="figure-21@9.[191,1396,181,1932]" captionTargetPageId="9" captionText="FIGURE 3. Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Whole animal, dorsal view (A), whole animal, ventral view (B). Scale bars: A = 12.5 mm, B = 5.38 mm." figureDoi="http://doi.org/10.5281/zenodo.5497105" httpUri="https://zenodo.org/record/5497105/files/figure.png" pageId="5" pageNumber="521">Figs. 3A</figureCitation>
, 
<figureCitation id="D7F22A056040FFCA6BDCF8D2FF28F89F" box="[200,237,1827,1853]" captionStart="FIGURE 4" captionStartId="10.[151,250,1747,1771]" captionTargetBox="[152,1436,309,1723]" captionTargetId="figure-93@10.[152,1436,309,1723]" captionTargetPageId="10" captionText="FIGURE 4. Frontal view of carapace. (A) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (B) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (C) Potamonautes macrobrachii Bott, 1953, paratype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Photograph B by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A = 10.4 mm; B = 5.7 mm; C = 4.7 mm." figureDoi="http://doi.org/10.5281/zenodo.5497107" httpUri="https://zenodo.org/record/5497107/files/figure.png" pageId="5" pageNumber="521">4C</figureCitation>
). Exorbital tooth small, low, blunt; epibranchial tooth reduced to granule; anterolateral margin posterior to epibranchial tooth smooth, posterior end continuous with posterolateral margin (
<figureCitation id="D7F22A056040FFCA6F34F8B6FB5EF8C3" box="[1056,1179,1863,1889]" captionStart="FIGURE 3" captionStartId="9.[151,250,1957,1981]" captionTargetBox="[191,1396,181,1932]" captionTargetId="figure-21@9.[191,1396,181,1932]" captionTargetPageId="9" captionText="FIGURE 3. Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Whole animal, dorsal view (A), whole animal, ventral view (B). Scale bars: A = 12.5 mm, B = 5.38 mm." figureDoi="http://doi.org/10.5281/zenodo.5497105" httpUri="https://zenodo.org/record/5497105/files/figure.png" pageId="5" pageNumber="521">Fig. 3A, B</figureCitation>
). S3/4 complete, deep, v-shaped, meeting anterior margin of sternopleonal cavity; outer margins of S4 thickened, raised (
<figureCitation id="D7F22A056040FFCA6FEDF89AFA9BF827" box="[1273,1374,1899,1925]" captionStart="FIGURE 3" captionStartId="9.[151,250,1957,1981]" captionTargetBox="[191,1396,181,1932]" captionTargetId="figure-21@9.[191,1396,181,1932]" captionTargetPageId="9" captionText="FIGURE 3. Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Whole animal, dorsal view (A), whole animal, ventral view (B). Scale bars: A = 12.5 mm, B = 5.38 mm." figureDoi="http://doi.org/10.5281/zenodo.5497105" httpUri="https://zenodo.org/record/5497105/files/figure.png" pageId="5" pageNumber="521">Figs. 3B</figureCitation>
, 
<figureCitation id="D7F22A056040FFCA6E7FF89AFA55F827" box="[1387,1424,1899,1925]" captionStart="FIGURE 4" captionStartId="10.[151,250,1747,1771]" captionTargetBox="[152,1436,309,1723]" captionTargetId="figure-93@10.[152,1436,309,1723]" captionTargetPageId="10" captionText="FIGURE 4. Frontal view of carapace. (A) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (B) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (C) Potamonautes macrobrachii Bott, 1953, paratype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Photograph B by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A = 10.4 mm; B = 5.7 mm; C = 4.7 mm." figureDoi="http://doi.org/10.5281/zenodo.5497107" httpUri="https://zenodo.org/record/5497107/files/figure.png" pageId="5" pageNumber="521">4C</figureCitation>
). Third maxilliped ischium vertical sulcus either lacking, or very faint (
<figureCitation id="D7F22A056040FFCA68BAF87EFBC3F80B" box="[942,1030,1935,1961]" captionStart="FIGURE 3" captionStartId="9.[151,250,1957,1981]" captionTargetBox="[191,1396,181,1932]" captionTargetId="figure-21@9.[191,1396,181,1932]" captionTargetPageId="9" captionText="FIGURE 3. Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Whole animal, dorsal view (A), whole animal, ventral view (B). Scale bars: A = 12.5 mm, B = 5.38 mm." figureDoi="http://doi.org/10.5281/zenodo.5497105" httpUri="https://zenodo.org/record/5497105/files/figure.png" pageId="5" pageNumber="521">Fig. 3B</figureCitation>
). Distal, proximal teeth on cheliped carpus inner margin both reduced to granules (
<figureCitation id="D7F22A056040FFCA69B0F842FCC2F86F" box="[676,775,1971,1997]" captionStart="FIGURE 3" captionStartId="9.[151,250,1957,1981]" captionTargetBox="[191,1396,181,1932]" captionTargetId="figure-21@9.[191,1396,181,1932]" captionTargetPageId="9" captionText="FIGURE 3. Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Whole animal, dorsal view (A), whole animal, ventral view (B). Scale bars: A = 12.5 mm, B = 5.38 mm." figureDoi="http://doi.org/10.5281/zenodo.5497105" httpUri="https://zenodo.org/record/5497105/files/figure.png" pageId="5" pageNumber="521">Figs. 3A</figureCitation>
, 
<figureCitation id="D7F22A056040FFCA6800F843FCF3F86F" box="[788,822,1970,1997]" captionStart="FIGURE 5" captionStartId="11.[151,250,1588,1612]" captionTargetBox="[159,1428,181,1565]" captionTargetId="figure-21@11.[159,1428,181,1565]" captionTargetPageId="11" captionText="FIGURE 5. (A–F) Frontal view of right and left chela; (G–L) cheliped merus and carpus. (A, B) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (C, D) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (E, F) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892); (G, H) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (G) ventral view of right chela merus; (H) dorsal view of right chela carpus and merus; (I, J) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province,Angola (AMG GEN 1887A) (I) ventral view of right chela merus; (J) dorsal view of right chela merus and carpus; (K, L) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892): (K) ventral view of right chela merus; (L) dorsal view of right chela merus and carpus;. Photographs C, D, I, J by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A = 18.4 mm; B = 17.9 mm; C, D = 8.9 mm; E = 8.33 mm; F = 5.81 mm; G = 17.9 mm; H = 7.3 mm; I = 8.9 mm; J = 5.9 mm; K = 5.81 mm; L = 5.6 mm." figureDoi="http://doi.org/10.5281/zenodo.5497109" httpUri="https://zenodo.org/record/5497109/files/figure.png" pageId="5" pageNumber="521">5L</figureCitation>
). G1 
<collectionCode id="29D8AE456040FFCA6860F842FC53F86F" box="[884,918,1971,1997]" country="0" name="Timescale Adventures Research and Interpretive Center" pageId="5" pageNumber="521">TA</collectionCode>
short (
<collectionCode id="29D8AE456040FFCA68F7F842FBCDF86F" box="[995,1032,1971,1997]" country="0" name="Timescale Adventures Research and Interpretive Center" pageId="5" pageNumber="521">TA</collectionCode>
/ 
<collectionCode id="29D8AE456040FFCA6F04F842FBF7F86F" box="[1040,1074,1971,1997]" country="France" name="Museum national d'Histoire Naturelle, Laboratiore de Paleontologie" pageId="5" pageNumber="521">SA</collectionCode>
0.33), basal half angled slightly outward at 50 
<emphasis id="7DBDEA926040FFCA6A26F827FEF8F852" bold="true" box="[306,317,2006,2032]" pageId="5" pageNumber="521">°</emphasis>
to longitudinal axis of G1 
<collectionCode id="29D8AE456040FFCA6979F826FD56F853" box="[621,659,2007,2033]" country="France" name="Museum national d'Histoire Naturelle, Laboratiore de Paleontologie" pageId="5" pageNumber="521">SA</collectionCode>
, distal third tapering to pointed tip; G1 
<collectionCode id="29D8AE456040FFCA6F46F826FBB1F853" box="[1106,1140,2007,2033]" country="0" name="Timescale Adventures Research and Interpretive Center" pageId="5" pageNumber="521">TA</collectionCode>
widened in midsection by distinctly raised rounded mesial, lateral folds, equally high, (
<figureCitation id="D7F22A056043FFC96823FF66FC6EFF13" box="[823,939,151,177]" captionStart="FIGURE 6" captionStartId="12.[151,250,1828,1852]" captionTargetBox="[197,1391,181,1805]" captionTargetId="figure-17@12.[197,1391,181,1805]" captionTargetPageId="12" captionText="FIGURE 6. Right G1, ventral view (A–C) and dorsal view (D, F). (A, D) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (B, E) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (C, F) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Photo- graphs B and E by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A= 6.7 mm; B = 2.3 mm; C = 1.13 mm; D = 2.7 mm; E = 2.3 mm; F = 1.2." figureDoi="http://doi.org/10.5281/zenodo.5497111" httpUri="https://zenodo.org/record/5497111/files/figure.png" pageId="6" pageNumber="522">Fig. 6C, F</figureCitation>
); longitudinal groove of G1 
<collectionCode id="29D8AE456043FFC96FF2FF66FACDFF13" box="[1254,1288,151,177]" country="0" name="Timescale Adventures Research and Interpretive Center" pageId="6" pageNumber="522">TA</collectionCode>
visible along proximal two-thirds. G2 
<collectionCode id="29D8AE456043FFC96AB9FF4AFE0AFF77" box="[429,463,187,213]" country="0" name="Timescale Adventures Research and Interpretive Center" pageId="6" pageNumber="522">TA</collectionCode>
long, flagellum-like (G2 
<collectionCode id="29D8AE456043FFC969F9FF4AFCD4FF77" box="[749,785,187,213]" country="0" name="Timescale Adventures Research and Interpretive Center" pageId="6" pageNumber="522">TA</collectionCode>
/ 
<collectionCode id="29D8AE456043FFC9680DFF4AFCFFFF77" box="[793,826,187,213]" country="France" name="Museum national d'Histoire Naturelle, Laboratiore de Paleontologie" pageId="6" pageNumber="522">SA</collectionCode>
0.33) (
<figureCitation id="D7F22A056043FFC9689FFF4AFC26FF77" box="[907,995,187,213]" captionStart="FIGURE 7" captionStartId="13.[151,250,1855,1879]" captionTargetBox="[264,1323,181,1831]" captionTargetId="figure-21@13.[264,1323,181,1831]" captionTargetPageId="13" captionText="FIGURE 7. Right G2 ventral view (A–C); (A) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (B) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (C) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Photograph C by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A = 5.6 mm; B = 2.3 mm; C = 1.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5497113" httpUri="https://zenodo.org/record/5497113/files/figure.png" pageId="6" pageNumber="522">Fig. 7C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="07D3650B6043FFC96BD3FF2FFA98FA93" pageId="6" pageNumber="522" type="description">
<paragraph id="4F7636806043FFC96BD3FF2FFA4AFD27" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="522">
<emphasis id="7DBDEA926043FFC96BD3FF2FFDF6FF5B" bold="true" box="[199,563,222,249]" pageId="6" pageNumber="522">
Redescription. Male 
<typeStatus id="907288226043FFC96AD1FF2EFDEBFF5B" box="[453,558,223,249]" pageId="6" pageNumber="522" type="holotype">holotype</typeStatus>
.
</emphasis>
Carapace with moderate width (CW/FW 3.0), length (CL/ FW 2.0), height (
<collectingCountry id="37DE76106043FFC96B8BFEF2FF03FEBF" box="[159,198,259,285]" name="Switzerland" pageId="6" pageNumber="522">CH</collectingCountry>
/FW 1.0); surface completely smooth with no deep grooves, granules, or carinae; semicircular, urogastric, cardiac, branchial grooves all faint; epigastric crests low, separated by clear, short, forked mid-groove; postfrontal crest completely crossing carapace, low in middle, sharp laterally, meeting anterolateral margins (
<figureCitation id="D7F22A056043FFC96FD4FEBAFAD8FEC7" box="[1216,1309,331,357]" captionStart="FIGURE 3" captionStartId="9.[151,250,1957,1981]" captionTargetBox="[191,1396,181,1932]" captionTargetId="figure-21@9.[191,1396,181,1932]" captionTargetPageId="9" captionText="FIGURE 3. Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Whole animal, dorsal view (A), whole animal, ventral view (B). Scale bars: A = 12.5 mm, B = 5.38 mm." figureDoi="http://doi.org/10.5281/zenodo.5497105" httpUri="https://zenodo.org/record/5497105/files/figure.png" pageId="6" pageNumber="522">Fig. 3A</figureCitation>
). Exorbital tooth small, low, blunt, epibranchial tooth reduced to granule; anterolateral margin between exorbital, epibranchial teeth smooth, straight, lacking intermediate tooth; anterolateral margin posterior to epibranchial tooth smooth, posterior end continuous with posterolateral margin (
<figureCitation id="D7F22A056043FFC969D1FE46FCE5FE73" box="[709,800,439,465]" captionStart="FIGURE 3" captionStartId="9.[151,250,1957,1981]" captionTargetBox="[191,1396,181,1932]" captionTargetId="figure-21@9.[191,1396,181,1932]" captionTargetPageId="9" captionText="FIGURE 3. Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Whole animal, dorsal view (A), whole animal, ventral view (B). Scale bars: A = 12.5 mm, B = 5.38 mm." figureDoi="http://doi.org/10.5281/zenodo.5497105" httpUri="https://zenodo.org/record/5497105/files/figure.png" pageId="6" pageNumber="522">Fig. 3A</figureCitation>
). Branchiostegite with faintly granulated vertical sulcus; subhepatic, suborbital, pterygostomial regions all smooth (
<figureCitation id="D7F22A056043FFC9683CFE2AFC4EFE57" box="[808,907,475,501]" captionStart="FIGURE 3" captionStartId="9.[151,250,1957,1981]" captionTargetBox="[191,1396,181,1932]" captionTargetId="figure-21@9.[191,1396,181,1932]" captionTargetPageId="9" captionText="FIGURE 3. Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Whole animal, dorsal view (A), whole animal, ventral view (B). Scale bars: A = 12.5 mm, B = 5.38 mm." figureDoi="http://doi.org/10.5281/zenodo.5497105" httpUri="https://zenodo.org/record/5497105/files/figure.png" pageId="6" pageNumber="522">Figs. 3B</figureCitation>
, 
<figureCitation id="D7F22A056043FFC96882FE2AFC7EFE57" box="[918,955,475,501]" captionStart="FIGURE 4" captionStartId="10.[151,250,1747,1771]" captionTargetBox="[152,1436,309,1723]" captionTargetId="figure-93@10.[152,1436,309,1723]" captionTargetPageId="10" captionText="FIGURE 4. Frontal view of carapace. (A) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (B) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (C) Potamonautes macrobrachii Bott, 1953, paratype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Photograph B by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A = 10.4 mm; B = 5.7 mm; C = 4.7 mm." figureDoi="http://doi.org/10.5281/zenodo.5497107" httpUri="https://zenodo.org/record/5497107/files/figure.png" pageId="6" pageNumber="522">4C</figureCitation>
). Third maxillipeds filling entire oral field, except for transversely oval efferent respiratory openings; exopod with long flagellum, ischium vertical sulcus faint (
<figureCitation id="D7F22A056043FFC96BB4FDD2FF3DFD9F" box="[160,248,547,573]" captionStart="FIGURE 3" captionStartId="9.[151,250,1957,1981]" captionTargetBox="[191,1396,181,1932]" captionTargetId="figure-21@9.[191,1396,181,1932]" captionTargetPageId="9" captionText="FIGURE 3. Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Whole animal, dorsal view (A), whole animal, ventral view (B). Scale bars: A = 12.5 mm, B = 5.38 mm." figureDoi="http://doi.org/10.5281/zenodo.5497105" httpUri="https://zenodo.org/record/5497105/files/figure.png" pageId="6" pageNumber="522">Fig. 3B</figureCitation>
). Mandibular palp 2-articled; terminal article lacking anterior lobe or ledge. Thoracic sternal sulcii S1/2, S2/3 deep, completely traversing sternum; S3/4 complete, deep, v-shaped, meeting anterior margin of sternopleonal cavity; outer margins of S4 thickened, raised; thoracic episternal sulci S4/E4, S5/E5, S6/ E6, S7/E7 each obscure.
</paragraph>
<paragraph id="4F7636806043FFC96BD3FD7EFD27FC23" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="522">
Chelipeds unequal, right (major) longer, higher than left (minor) (
<figureCitation id="D7F22A056043FFC968A1FD7EFBD5FD0B" box="[949,1040,655,681]" captionStart="FIGURE 3" captionStartId="9.[151,250,1957,1981]" captionTargetBox="[191,1396,181,1932]" captionTargetId="figure-21@9.[191,1396,181,1932]" captionTargetPageId="9" captionText="FIGURE 3. Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Whole animal, dorsal view (A), whole animal, ventral view (B). Scale bars: A = 12.5 mm, B = 5.38 mm." figureDoi="http://doi.org/10.5281/zenodo.5497105" httpUri="https://zenodo.org/record/5497105/files/figure.png" pageId="6" pageNumber="522">Fig. 3A</figureCitation>
). Dactylus of right chela slim, distinctly arched, enclosing broad oval interspace when closed; propodus palm swollen. Movable finger (dactylus) lacking teeth proximally, distal half with 5 small well-spaced small teeth; fixed finger (pollex of propodus) of right chela with 4 widely spaced small teeth (
<figureCitation id="D7F22A056043FFC9695BFD0AFD04FCB7" box="[591,705,762,789]" captionStart="FIGURE 5" captionStartId="11.[151,250,1588,1612]" captionTargetBox="[159,1428,181,1565]" captionTargetId="figure-21@11.[159,1428,181,1565]" captionTargetPageId="11" captionText="FIGURE 5. (A–F) Frontal view of right and left chela; (G–L) cheliped merus and carpus. (A, B) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (C, D) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (E, F) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892); (G, H) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (G) ventral view of right chela merus; (H) dorsal view of right chela carpus and merus; (I, J) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province,Angola (AMG GEN 1887A) (I) ventral view of right chela merus; (J) dorsal view of right chela merus and carpus; (K, L) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892): (K) ventral view of right chela merus; (L) dorsal view of right chela merus and carpus;. Photographs C, D, I, J by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A = 18.4 mm; B = 17.9 mm; C, D = 8.9 mm; E = 8.33 mm; F = 5.81 mm; G = 17.9 mm; H = 7.3 mm; I = 8.9 mm; J = 5.9 mm; K = 5.81 mm; L = 5.6 mm." figureDoi="http://doi.org/10.5281/zenodo.5497109" httpUri="https://zenodo.org/record/5497109/files/figure.png" pageId="6" pageNumber="522">Fig. 5E, F</figureCitation>
). Cheliped carpus inner margin distal, proximal teeth both heavily reduced, proximal tooth granule-sized (
<figureCitation id="D7F22A056043FFC9695AFCEEFD61FC9B" box="[590,676,798,825]" captionStart="FIGURE 5" captionStartId="11.[151,250,1588,1612]" captionTargetBox="[159,1428,181,1565]" captionTargetId="figure-21@11.[159,1428,181,1565]" captionTargetPageId="11" captionText="FIGURE 5. (A–F) Frontal view of right and left chela; (G–L) cheliped merus and carpus. (A, B) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (C, D) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (E, F) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892); (G, H) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (G) ventral view of right chela merus; (H) dorsal view of right chela carpus and merus; (I, J) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province,Angola (AMG GEN 1887A) (I) ventral view of right chela merus; (J) dorsal view of right chela merus and carpus; (K, L) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892): (K) ventral view of right chela merus; (L) dorsal view of right chela merus and carpus;. Photographs C, D, I, J by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A = 18.4 mm; B = 17.9 mm; C, D = 8.9 mm; E = 8.33 mm; F = 5.81 mm; G = 17.9 mm; H = 7.3 mm; I = 8.9 mm; J = 5.9 mm; K = 5.81 mm; L = 5.6 mm." figureDoi="http://doi.org/10.5281/zenodo.5497109" httpUri="https://zenodo.org/record/5497109/files/figure.png" pageId="6" pageNumber="522">Fig. 5L</figureCitation>
). Cheliped merus lower margins both lined by granules, distal meral tooth low, blunt (
<figureCitation id="D7F22A056043FFC96A40FCB2FE6BFCFF" box="[340,430,834,861]" captionStart="FIGURE 5" captionStartId="11.[151,250,1588,1612]" captionTargetBox="[159,1428,181,1565]" captionTargetId="figure-21@11.[159,1428,181,1565]" captionTargetPageId="11" captionText="FIGURE 5. (A–F) Frontal view of right and left chela; (G–L) cheliped merus and carpus. (A, B) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (C, D) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (E, F) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892); (G, H) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (G) ventral view of right chela merus; (H) dorsal view of right chela carpus and merus; (I, J) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province,Angola (AMG GEN 1887A) (I) ventral view of right chela merus; (J) dorsal view of right chela merus and carpus; (K, L) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892): (K) ventral view of right chela merus; (L) dorsal view of right chela merus and carpus;. Photographs C, D, I, J by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A = 18.4 mm; B = 17.9 mm; C, D = 8.9 mm; E = 8.33 mm; F = 5.81 mm; G = 17.9 mm; H = 7.3 mm; I = 8.9 mm; J = 5.9 mm; K = 5.81 mm; L = 5.6 mm." figureDoi="http://doi.org/10.5281/zenodo.5497109" httpUri="https://zenodo.org/record/5497109/files/figure.png" pageId="6" pageNumber="522">Fig. 5K</figureCitation>
). Ambulatory legs (P2–5) slender, P5 shortest; dactyli P2–5 tapering to point, each bearing rows of downward-pointing sharp bristles (
<figureCitation id="D7F22A056043FFC9696FFC96FD10FC23" box="[635,725,871,897]" captionStart="FIGURE 3" captionStartId="9.[151,250,1957,1981]" captionTargetBox="[191,1396,181,1932]" captionTargetId="figure-21@9.[191,1396,181,1932]" captionTargetPageId="9" captionText="FIGURE 3. Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Whole animal, dorsal view (A), whole animal, ventral view (B). Scale bars: A = 12.5 mm, B = 5.38 mm." figureDoi="http://doi.org/10.5281/zenodo.5497105" httpUri="https://zenodo.org/record/5497105/files/figure.png" pageId="6" pageNumber="522">Fig. 3A</figureCitation>
).
</paragraph>
<paragraph id="4F7636806043FFC96BD3FC7AFB76FB4B" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="522">
Male pleon, telson together forming slim triangle, pleon edges slightly indented; telson triangular, apex round- ed, base broadest, sides outwardly sloping; pleomeres PL1–6 rectangular, wider than long, PL 6 longest, more than 1/2 as long as wide (
<figureCitation id="D7F22A056043FFC96A94FC22FE1CFC4F" box="[384,473,979,1005]" captionStart="FIGURE 3" captionStartId="9.[151,250,1957,1981]" captionTargetBox="[191,1396,181,1932]" captionTargetId="figure-21@9.[191,1396,181,1932]" captionTargetPageId="9" captionText="FIGURE 3. Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Whole animal, dorsal view (A), whole animal, ventral view (B). Scale bars: A = 12.5 mm, B = 5.38 mm." figureDoi="http://doi.org/10.5281/zenodo.5497105" httpUri="https://zenodo.org/record/5497105/files/figure.png" pageId="6" pageNumber="522">Fig. 3B</figureCitation>
); remaining pleomeres short, less than 1/3 as long as wide. G1 TA short, about 1/3 as long as SA (G1 TA/SA 0.33), angled outward at midpoint at 50° to longitudinal axis of G1 SA; G1 TA midsection distinctly widened by raised rounded mesial fold, lateral fold low throughout length (
<figureCitation id="D7F22A056043FFC96F45FBEAFB02FB97" box="[1105,1223,1051,1077]" captionStart="FIGURE 6" captionStartId="12.[151,250,1828,1852]" captionTargetBox="[197,1391,181,1805]" captionTargetId="figure-17@12.[197,1391,181,1805]" captionTargetPageId="12" captionText="FIGURE 6. Right G1, ventral view (A–C) and dorsal view (D, F). (A, D) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (B, E) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (C, F) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Photo- graphs B and E by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A= 6.7 mm; B = 2.3 mm; C = 1.13 mm; D = 2.7 mm; E = 2.3 mm; F = 1.2." figureDoi="http://doi.org/10.5281/zenodo.5497111" httpUri="https://zenodo.org/record/5497111/files/figure.png" pageId="6" pageNumber="522">Fig. 6C, F</figureCitation>
); distal third of G1 TA slim, tapering to pointed up-curved tip; proximal two-thirds of G1 TA with longitudinal groove between mesial, lateral folds. G1 SA widest at base, narrowest at G1 TA/SA junction; G1 SA mesial, lateral margins lined by conspicuous long setae; G1 SA ventral side with broad setae-lined fold that partly covers SA basally, completely covers SA at TA/SA junction. G2 TA long, flagellum-like (G2 TA/SA 0.33); G2 SA widest at base, last two-thirds tapering sharply inward forming long, thin, tapering, upright process supporting long G2 TA (
<figureCitation id="D7F22A056043FFC96F5AFB3EFB63FB4B" box="[1102,1190,1231,1257]" captionStart="FIGURE 7" captionStartId="13.[151,250,1855,1879]" captionTargetBox="[264,1323,181,1831]" captionTargetId="figure-21@13.[264,1323,181,1831]" captionTargetPageId="13" captionText="FIGURE 7. Right G2 ventral view (A–C); (A) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (B) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (C) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Photograph C by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A = 5.6 mm; B = 2.3 mm; C = 1.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5497113" httpUri="https://zenodo.org/record/5497113/files/figure.png" pageId="6" pageNumber="522">Fig. 7C</figureCitation>
).
</paragraph>
<paragraph id="4F7636806043FFC96BD3FB03FD06FAAF" blockId="6.[151,1437,151,2013]" box="[199,707,1266,1293]" pageId="6" pageNumber="522">
<emphasis id="7DBDEA926043FFC96BD3FB03FF32FAAE" bold="true" box="[199,247,1266,1292]" pageId="6" pageNumber="522">Size</emphasis>
. Medium species, adult at CW 32.0 mm.
</paragraph>
<paragraph id="4F7636806043FFC96BD3FAE7FA98FA93" blockId="6.[151,1437,151,2013]" box="[199,1373,1302,1329]" pageId="6" pageNumber="522">
<emphasis id="7DBDEA926043FFC96BD3FAE7FEE5FA92" bold="true" box="[199,288,1302,1328]" pageId="6" pageNumber="522">Colour.</emphasis>
The colour of living specimens is unknown, but specimens preserved in ethanol are light brown.
</paragraph>
</subSubSection>
<subSubSection id="07D3650B6043FFC96BD3FACAFEA3FA63" pageId="6" pageNumber="522" type="materials_examined">
<paragraph id="4F7636806043FFC96BD3FACAFEA3FA63" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="522">
<emphasis id="7DBDEA926043FFC96BD3FACAFEAEFAF6" bold="true" box="[199,363,1338,1365]" pageId="6" pageNumber="522">Type locality.</emphasis>
<materialsCitation id="FFA13CDD6043FFC96A62FACAFF68FA63" collectorName="Kuketa &amp; This &amp; de Moco" country="Angola" county="Benguala" elevation="2200" latitude="-12.466667" location="Catumbela River" longLatPrecision="1" longitude="15.166667" municipality="Serra do Moco" pageId="6" pageNumber="522" specimenCount="1" stateProvince="Huambo">
<collectingCountry id="37DE76106043FFC96A62FACAFE0EFAF7" box="[374,459,1339,1365]" name="Angola" pageId="6" pageNumber="522">Angola</collectingCountry>
, 
<collectingCounty id="A6174E0C6043FFC96AC8FACAFD8CFAF7" box="[476,585,1339,1365]" pageId="6" pageNumber="522">Benguala</collectingCounty>
, 
<collectingRegion id="8D0DF8626043FFC9694DFACAFCE9FAF7" box="[601,812,1339,1365]" country="Angola" name="Huambo" pageId="6" pageNumber="522">Huambo Province</collectingRegion>
, 
<collectingMunicipality id="AF12ACFA6043FFC96829FACAFC2BFAF7" box="[829,1006,1339,1365]" pageId="6" pageNumber="522">Serra do Moco</collectingMunicipality>
(escarpment of Moco) (
<geoCoordinate id="2AFD50476043FFC96E07FACAFA5DFAF7" box="[1299,1432,1339,1365]" degrees="12.466667" direction="south" orientation="latitude" pageId="6" pageNumber="522" precision="1" value="-12.466667">-12.466667</geoCoordinate>
, 
<geoCoordinate id="2AFD50476043FFC96B83FAAFFED5FADB" box="[151,272,1374,1401]" degrees="15.166667" direction="east" orientation="longitude" pageId="6" pageNumber="522" precision="1" value="15.166667">15.166667</geoCoordinate>
) 
<quantity id="88319B656043FFC96A0AFAAEFEBFFADB" box="[286,378,1375,1401]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="6" pageNumber="522" unit="m" value="2200.0">
<elevation id="C4E4D1B36043FFC96A0AFAAEFEBFFADB" box="[286,378,1375,1401]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="6" pageNumber="522" unit="m" value="2200.0">2,200 m</elevation>
</quantity>
ASL, 
<collectorName id="E23C53566043FFC96AD5FAAEFDD7FADB" box="[449,530,1375,1401]" pageId="6" pageNumber="522">Kuketa</collectorName>
stream, a tributary of the 
<location id="4A16605B6043FFC96825FAAEFC2AFADB" LSID="urn:lsid:plazi:treatment:C76087966040FFC16B83FA72FE1EF8A3:4A16605B6043FFC96825FAAEFC2AFADB" box="[817,1007,1375,1401]" country="Angola" county="Benguala" latitude="-12.466667" longLatPrecision="1" longitude="15.166667" municipality="Serra do Moco" name="Catumbela River" pageId="6" pageNumber="522" stateProvince="Huambo">Catumbela River</location>
draining the 
<location id="4A16605B6043FFC96F96FAAEFF17FA3F" LSID="urn:lsid:plazi:treatment:C76087966040FFC16B83FA72FE1EF8A3:4A16605B6043FFC96F96FAAEFF17FA3F" country="Angola" county="Benguala" latitude="-12.466667" longLatPrecision="1" longitude="15.166667" municipality="Serra do Moco" name="Mount Moco Special Reserve" pageId="6" pageNumber="522" stateProvince="Huambo">Mount Moco Special Reserve</location>
(
<quantity id="88319B656043FFC96BF6FA72FEFDFA3F" box="[226,312,1411,1437]" metricMagnitude="3" metricUnit="m" metricValue="1.898" pageId="6" pageNumber="522" unit="m" value="1898.0">
<elevation id="C4E4D1B36043FFC96BF6FA72FEFDFA3F" box="[226,312,1411,1437]" metricMagnitude="3" metricUnit="m" metricValue="1.898" pageId="6" pageNumber="522" unit="m" value="1898.0">1898 m</elevation>
</quantity>
ASL). 
<collectorName id="E23C53566043FFC96A9EFA72FE79FA3F" box="[394,444,1411,1437]" pageId="6" pageNumber="522">This</collectorName>
high-altitude locality is close to 
<location id="4A16605B6043FFC96839FA72FC18FA3E" LSID="urn:lsid:plazi:treatment:C76087966040FFC16B83FA72FE1EF8A3:4A16605B6043FFC96839FA72FC18FA3E" box="[813,989,1411,1437]" country="Angola" county="Benguala" latitude="-12.466667" longLatPrecision="1" longitude="15.166667" municipality="Serra do Moco" name="Morro de Moco" pageId="6" pageNumber="522" stateProvince="Huambo">
Morro 
<collectorName id="E23C53566043FFC96868FA72FC18FA3E" box="[892,989,1411,1437]" pageId="6" pageNumber="522">
de 
<emphasis id="7DBDEA926043FFC96889FA72FC18FA3E" box="[925,989,1411,1436]" italics="true" pageId="6" pageNumber="522">Moco</emphasis>
</collectorName>
</location>
(
<quantity id="88319B656043FFC968F8FA72FB8FFA3F" box="[1004,1098,1411,1437]" metricMagnitude="3" metricUnit="m" metricValue="2.62" pageId="6" pageNumber="522" unit="m" value="2620.0">
<elevation id="C4E4D1B36043FFC968F8FA72FB8FFA3F" box="[1004,1098,1411,1437]" metricMagnitude="3" metricUnit="m" metricValue="2.62" pageId="6" pageNumber="522" unit="m" value="2620.0">2,620 m</elevation>
</quantity>
ASL) which the highest point in
</materialsCitation>
<collectingCountry id="37DE76106043FFC96BA6FA56FEC1FA63" box="[178,260,1447,1473]" name="Angola" pageId="6" pageNumber="522">Angola</collectingCountry>
(
<figureCitation id="D7F22A056043FFC96A00FA56FE9CFA63" box="[276,345,1447,1473]" captionStart="FIGURE 8" captionStartId="14.[151,250,1336,1360]" captionTargetBox="[159,1428,189,1302]" captionTargetId="figure-17@14.[151,1436,181,1312]" captionTargetPageId="14" captionText="FIGURE 8. Map of the Angola showing the known localities of Potamonautes adeleae (black square, not shown to scale), P. kensleyi (black circles, not shown to scale), and P. macrobrachii (black diamond, not shown to scale). Scale bar = 520 km. See text for exact locality coordinates." figureDoi="http://doi.org/10.5281/zenodo.5497115" httpUri="https://zenodo.org/record/5497115/files/figure.png" pageId="6" pageNumber="522">Fig. 8</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="07D3650B6043FFC96BD3FA3BFB20F8EF" pageId="6" pageNumber="522" type="distribution">
<paragraph id="4F7636806043FFC96BD3FA3BFAE1F98F" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="522">
<emphasis id="7DBDEA926043FFC96BD3FA3BFE9CFA46" bold="true" box="[199,345,1482,1508]" pageId="6" pageNumber="522">Distribution</emphasis>
. 
<taxonomicName id="88C94D036043FFC96A7CFA3AFD63FA47" authorityName=", Cumberlidge &amp; Tavares" authorityYear="2006" box="[360,678,1483,1509]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="522" phylum="Arthropoda" rank="species" species="macrobrachii">
<emphasis id="7DBDEA926043FFC96A7CFA3AFD63FA47" box="[360,678,1483,1509]" italics="true" pageId="6" pageNumber="522">Potamonautes macrobrachii</emphasis>
</taxonomicName>
is known only from just one locality on the escarpment of Serra do Moco (
<geoCoordinate id="2AFD50476043FFC96BF2FA1EFEAEF9AB" box="[230,363,1519,1545]" degrees="12.466667" direction="south" orientation="latitude" pageId="6" pageNumber="522" precision="1" value="-12.466667">-12.466667</geoCoordinate>
, 
<geoCoordinate id="2AFD50476043FFC96A61FA1FFE2BF9AB" box="[373,494,1518,1545]" degrees="15.166667" direction="east" orientation="longitude" pageId="6" pageNumber="522" precision="1" value="15.166667">15.166667</geoCoordinate>
) 
<quantity id="88319B656043FFC96AE8FA1EFD92F9AB" box="[508,599,1519,1545]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="6" pageNumber="522" unit="m" value="2200.0">2,200 m</quantity>
ASL, in Benguala 
<collectingRegion id="8D0DF8626043FFC9683DFA1EFC33F9AB" box="[809,1014,1519,1545]" country="Angola" name="Huambo" pageId="6" pageNumber="522">Huambo Province</collectingRegion>
. The specimen was collected from the Kuketa stream, a tributary of the Catumbela River that drains the Mount Moco Special Reserve (
<figureCitation id="D7F22A056043FFC96FC6F9E2FAD2F98F" box="[1234,1303,1555,1581]" captionStart="FIGURE 8" captionStartId="14.[151,250,1336,1360]" captionTargetBox="[159,1428,189,1302]" captionTargetId="figure-17@14.[151,1436,181,1312]" captionTargetPageId="14" captionText="FIGURE 8. Map of the Angola showing the known localities of Potamonautes adeleae (black square, not shown to scale), P. kensleyi (black circles, not shown to scale), and P. macrobrachii (black diamond, not shown to scale). Scale bar = 520 km. See text for exact locality coordinates." figureDoi="http://doi.org/10.5281/zenodo.5497115" httpUri="https://zenodo.org/record/5497115/files/figure.png" pageId="6" pageNumber="522">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="4F7636806043FFC96BD3F9C7FCC1F93B" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="522">
<emphasis id="7DBDEA926043FFC96BD3F9C7FEEFF9F2" bold="true" box="[199,298,1590,1616]" pageId="6" pageNumber="522">Habitat.</emphasis>
<materialsCitation id="FFA13CDD6043FFC96A26F9C6FCC5F93B" country="Angola" county="Afromontane forest" elevation="2620" location="Mount Moco" municipality="Mount Moco" pageId="6" pageNumber="522" specimenCount="1" stateProvince="Huambo">
<location id="4A16605B6043FFC96A26F9C6FE00F9F3" LSID="urn:lsid:plazi:treatment:C76087966040FFC16B83FA72FE1EF8A3:4A16605B6043FFC96A26F9C6FE00F9F3" box="[306,453,1591,1617]" country="Angola" county="Afromontane forest" municipality="Mount Moco" name="Mount Moco" pageId="6" pageNumber="522" stateProvince="Huambo">Mount Moco</location>
is located in the mountainous province of 
<collectingRegion id="8D0DF8626043FFC968BEF9C6FBC8F9F3" box="[938,1037,1591,1617]" country="Angola" name="Huambo" pageId="6" pageNumber="522">Huambo</collectingRegion>
, in central-west 
<collectingCountry id="37DE76106043FFC96FD2F9C6FADEF9F3" box="[1222,1307,1591,1617]" name="Angola" pageId="6" pageNumber="522">Angola</collectingCountry>
, and is that country’s highest peak at 
<quantity id="88319B656043FFC96AA3F9AAFDD3F9D7" box="[439,534,1627,1653]" metricMagnitude="3" metricUnit="m" metricValue="2.62" pageId="6" pageNumber="522" unit="m" value="2620.0">
<elevation id="C4E4D1B36043FFC96AA3F9AAFDD3F9D7" box="[439,534,1627,1653]" metricMagnitude="3" metricUnit="m" metricValue="2.62" pageId="6" pageNumber="522" unit="m" value="2620.0">2,620 m</elevation>
</quantity>
. The slopes of 
<collectingMunicipality id="AF12ACFA6043FFC969ABF9AAFC97F9D7" box="[703,850,1627,1653]" pageId="6" pageNumber="522">Mount Moco</collectingMunicipality>
include steep valleys that support 
<collectingCounty id="A6174E0C6043FFC96FC2F9AAFF7CF93B" pageId="6" pageNumber="522">Afromontane forest</collectingCounty>
, while at lower altitudes there is miombo woodland
</materialsCitation>
.
</paragraph>
<paragraph id="4F7636806043FFC96BD3F953FB20F8EF" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="522">
<emphasis id="7DBDEA926043FFC96BD3F953FE70F91F" bold="true" box="[199,437,1698,1725]" pageId="6" pageNumber="522">Conservation status</emphasis>
. 
<taxonomicName id="88C94D036043FFC96AD1F952FCC1F91F" authorityName=", Cumberlidge &amp; Tavares" authorityYear="2006" box="[453,772,1699,1725]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="522" phylum="Arthropoda" rank="species" species="macrobrachii">
<emphasis id="7DBDEA926043FFC96AD1F952FCC1F91F" box="[453,772,1699,1725]" italics="true" pageId="6" pageNumber="522">Potamonautes macrobrachii</emphasis>
</taxonomicName>
was listed in the IUCN Red List of Threatened Species as Data Deficient (
<bibRefCitation id="2B584B716043FFC96A59F936FDF4F943" author="Cumberlidge, N." box="[333,561,1735,1761]" pageId="6" pageNumber="522" pagination="64386" refId="ref9561" refString="Cumberlidge, N. (2008 c) Potamonautes macrobrachii. The IUCN Red List of Threatened Species, 2008, e. T 64386 A 12766440. https: // doi. org / 10.2305 / IUCN. UK. 2008. RLTS. T 64386 A 12766440. en" type="journal article" year="2008">Cumberlidge 2008c</bibRefCitation>
) because there is very little information available on its distributional range, population size and trends, or habitat. Although Mount Moco Special Reserve is a protected area it currently receives little protection from unsustainable and unregulated wood collection for logging and firewood, and uncontrolled bush fires. All of this represents a potential threat to the habitat of this little-known species.
</paragraph>
</subSubSection>
<subSubSection id="07D3650B6043FFC16BD3F8A6FE1EF8A3" lastPageId="14" lastPageNumber="530" pageId="6" pageNumber="522" type="discussion">
<paragraph id="4F7636806043FFC56BD3F8A6FA50FEBF" blockId="6.[151,1437,151,2013]" lastBlockId="10.[151,1436,151,285]" lastPageId="10" lastPageNumber="526" pageId="6" pageNumber="522">
<emphasis id="7DBDEA926043FFC96BD3F8A6FEFEF8D3" bold="true" box="[199,315,1879,1905]" pageId="6" pageNumber="522">Remarks.</emphasis>
The affinities of 
<taxonomicName id="88C94D036043FFC96AE9F8A6FD75F8D3" authorityName=", Cumberlidge &amp; Tavares" authorityYear="2006" box="[509,688,1879,1905]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="522" phylum="Arthropoda" rank="species" species="macrobrachii">
<emphasis id="7DBDEA926043FFC96AE9F8A6FD75F8D3" box="[509,688,1879,1905]" italics="true" pageId="6" pageNumber="522">P. macrobrachii</emphasis>
</taxonomicName>
(based on similarities of characters of the carapace, chelipeds, and gonopods) place it close to southern African species such as 
<taxonomicName id="88C94D036043FFC96857F88AFAC9F837" authority="Daniels &amp; Bayliss, 2012" authorityName="Daniels &amp; Bayliss" authorityYear="2012" box="[835,1292,1915,1941]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="522" phylum="Arthropoda" rank="species" species="mulanjeensis">
<emphasis id="7DBDEA926043FFC96857F88AFC34F837" box="[835,1009,1915,1941]" italics="true" pageId="6" pageNumber="522">P. mulanjeensis</emphasis>
<bibRefCitation id="2B584B716043FFC968EDF88AFAC9F837" author="Daniels, S. R. &amp; Bayliss, J." box="[1017,1292,1915,1941]" pageId="6" pageNumber="522" pagination="498 - 509" refId="ref9826" refString="Daniels, S. R. &amp; Bayliss, J. (2012) Neglected refugia of biodiversity: Mountainous regions in Mozambique and Malawi yield three novel freshwater crab species (Potamonautidae: Potamonautes). Zoological Journal of the Linnean Society, 164, 498 - 509. https: // doi. org / 10.1111 / j. 1096 - 3642.2011.00773. x" type="journal article" year="2012">Daniels &amp; Bayliss, 2012</bibRefCitation>
</taxonomicName>
from Mount 
<collectingRegion id="8D0DF8626043FFC96B83F86EFF31F81B" box="[151,244,1951,1977]" country="Malawi" name="Mulanje" pageId="6" pageNumber="522">Mulanje</collectingRegion>
in 
<collectingCountry id="37DE76106043FFC96A0CF86EFEABF81B" box="[280,366,1951,1977]" name="Malawi" pageId="6" pageNumber="522">Malawi</collectingCountry>
and 
<taxonomicName id="88C94D036043FFC96AB1F86EFC80F81B" authority="Phiri &amp; Daniels, 2013" authorityName="Phiri &amp; Daniels" authorityYear="2013" box="[421,837,1951,1977]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="522" phylum="Arthropoda" rank="species" species="mutareensis">
<emphasis id="7DBDEA926043FFC96AB1F86EFD83F81A" box="[421,582,1951,1976]" italics="true" pageId="6" pageNumber="522">P. mutareensis</emphasis>
<bibRefCitation id="2B584B716043FFC96959F86EFC80F81B" author="Phiri, E. E. &amp; Daniels, S. R." box="[589,837,1951,1977]" pageId="6" pageNumber="522" pagination="530 - 539" refId="ref10406" refString="Phiri, E. E. &amp; Daniels, S. R. (2013) Hidden in the highlands: the description and phylogenetic positioning of a novel endemic freshwater crab species (Potamonautidae: Potamonautes) from Zimbabwe. Invertebrate Systematics, 27, 530 - 539. https: // doi. org / 10.1071 / is 13012" type="journal article" year="2013">
Phiri &amp; 
<emphasis id="7DBDEA926043FFC969BCF86EFD3AF81B" box="[680,767,1951,1977]" italics="true" pageId="6" pageNumber="522">Daniels</emphasis>
, 2013
</bibRefCitation>
</taxonomicName>
from the eastern Highlands of 
<collectingCountry id="37DE76106043FFC96FB6F86EFADFF81B" box="[1186,1306,1951,1977]" name="Zimbabwe" pageId="6" pageNumber="522">Zimbabwe</collectingCountry>
(
<bibRefCitation id="2B584B716043FFC96E3CF86EFEE8F87F" author="Daniels, S. R. &amp; Bayliss, J." pageId="6" pageNumber="522" pagination="498 - 509" refId="ref9826" refString="Daniels, S. R. &amp; Bayliss, J. (2012) Neglected refugia of biodiversity: Mountainous regions in Mozambique and Malawi yield three novel freshwater crab species (Potamonautidae: Potamonautes). Zoological Journal of the Linnean Society, 164, 498 - 509. https: // doi. org / 10.1111 / j. 1096 - 3642.2011.00773. x" type="journal article" year="2012">Daniels &amp; Bayliss 2012</bibRefCitation>
; 
<bibRefCitation id="2B584B716043FFC96A2CF832FDEAF87F" author="Phiri, E. E. &amp; Daniels, S. R." box="[312,559,1987,2013]" pageId="6" pageNumber="522" pagination="530 - 539" refId="ref10406" refString="Phiri, E. E. &amp; Daniels, S. R. (2013) Hidden in the highlands: the description and phylogenetic positioning of a novel endemic freshwater crab species (Potamonautidae: Potamonautes) from Zimbabwe. Invertebrate Systematics, 27, 530 - 539. https: // doi. org / 10.1071 / is 13012" type="journal article" year="2013">
Phiri &amp; 
<emphasis id="7DBDEA926043FFC96A87F832FE2FF87F" box="[403,490,1987,2013]" italics="true" pageId="6" pageNumber="522">Daniels</emphasis>
2013
</bibRefCitation>
). In 
<collectingCountry id="37DE76106043FFC9694BF832FD71F87F" box="[607,692,1987,2013]" name="Angola" pageId="6" pageNumber="522">Angola</collectingCountry>
, 
<taxonomicName id="88C94D036043FFC969D4F832FCB7F87F" authorityName=", Cumberlidge &amp; Tavares" authorityYear="2006" box="[704,882,1987,2013]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="522" phylum="Arthropoda" rank="species" species="macrobrachii">
<emphasis id="7DBDEA926043FFC969D4F832FCB7F87F" box="[704,882,1987,2013]" italics="true" pageId="6" pageNumber="522">P. macrobrachii</emphasis>
</taxonomicName>
is found at the same location as 
<taxonomicName id="88C94D036043FFC96FF0F832FAA8F87F" baseAuthorityName="de Brito Capello" baseAuthorityYear="1870" box="[1252,1389,1987,2013]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="522" phylum="Arthropoda" rank="species" species="anchietae">
<emphasis id="7DBDEA926043FFC96FF0F832FAA8F87F" box="[1252,1389,1987,2013]" italics="true" pageId="6" pageNumber="522">P. anchietae</emphasis>
</taxonomicName>
and both share a similar carapace height and frontal margin width (
<collectingCountry id="37DE7610604FFFC56870FF66FC4EFF13" box="[868,907,151,177]" name="Switzerland" pageId="10" pageNumber="526">CH</collectingCountry>
/FW 1.08, 
<taxonomicName id="88C94D03604FFFC56F12FF66FB7FFF13" authorityName=", Cumberlidge &amp; Tavares" authorityYear="2006" box="[1030,1210,151,177]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="526" phylum="Arthropoda" rank="species" species="macrobrachii">
<emphasis id="7DBDEA92604FFFC56F12FF66FB7FFF13" box="[1030,1210,151,177]" italics="true" pageId="10" pageNumber="526">P. macrobrachii</emphasis>
</taxonomicName>
, 1.07, 
<taxonomicName id="88C94D03604FFFC56E1FFF66FA50FF13" baseAuthorityName="de Brito Capello" baseAuthorityYear="1870" box="[1291,1429,151,177]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="526" phylum="Arthropoda" rank="species" species="anchietae">
<emphasis id="7DBDEA92604FFFC56E1FFF66FA50FF13" box="[1291,1429,151,177]" italics="true" pageId="10" pageNumber="526">P. anchietae</emphasis>
</taxonomicName>
; and FW/CW 0.34, 
<taxonomicName id="88C94D03604FFFC56A7DFF4AFDDEFF77" authorityName=", Cumberlidge &amp; Tavares" authorityYear="2006" box="[361,539,187,213]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="526" phylum="Arthropoda" rank="species" species="macrobrachii">
<emphasis id="7DBDEA92604FFFC56A7DFF4AFDDEFF77" box="[361,539,187,213]" italics="true" pageId="10" pageNumber="526">P. macrobrachii</emphasis>
</taxonomicName>
, 0.32, 
<taxonomicName id="88C94D03604FFFC56972FF4AFD2BFF77" baseAuthorityName="de Brito Capello" baseAuthorityYear="1870" box="[614,750,187,213]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="526" phylum="Arthropoda" rank="species" species="anchietae">
<emphasis id="7DBDEA92604FFFC56972FF4AFD2BFF77" box="[614,750,187,213]" italics="true" pageId="10" pageNumber="526">P. anchietae</emphasis>
</taxonomicName>
), and both have a prominent and complete postfrontal crest, a greatly reduced epibranchial tooth (either granular or missing), a v-shaped thoracic sternal sulcus S3/4 that is deep at the sides and shallow in the middle, and a cheliped merus whose inferior margins are either granular or smooth.
</paragraph>
<caption id="1BB666086042FFC86B83F85CFB3BF84A" ID-DOI="http://doi.org/10.5281/zenodo.5497101" ID-Zenodo-Dep="5497101" httpUri="https://zenodo.org/record/5497101/files/figure.png" pageId="7" pageNumber="523" startId="7.[151,250,1965,1989]" targetBox="[191,1397,179,1938]" targetPageId="7">
<paragraph id="4F7636806042FFC86B83F85CFB3BF84A" blockId="7.[151,1436,1965,2025]" pageId="7" pageNumber="523">
<emphasis id="7DBDEA926042FFC86B83F85CFED0F867" bold="true" box="[151,277,1965,1989]" pageId="7" pageNumber="523">FIGURE 1.</emphasis>
<taxonomicName id="88C94D036042FFC86A0AF85FFDB8F867" authority="Bott, 1968" authorityName="Bott" authorityYear="1968" box="[286,637,1965,1989]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="523" phylum="Arthropoda" rank="species" species="adeleae">
<emphasis id="7DBDEA926042FFC86A0AF85FFDC3F867" box="[286,518,1965,1989]" italics="true" pageId="7" pageNumber="523">Potamonautes adeleae</emphasis>
<bibRefCitation id="2B584B716042FFC8691AF85CFDB8F867" author="Bott, R." box="[526,637,1965,1989]" pageId="7" pageNumber="523" pagination="165 - 172" refId="ref9155" refString="Bott, R. (1968) Decapoden aus dem Museu do Dundo (Crust. Decap.). Publicacoes culturais da Companhia de Diamantes de Angola, Lisboa, 77, 165 - 172." type="journal article" year="1968">Bott, 1968</bibRefCitation>
</taxonomicName>
, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444). Whole animal, dorsal view 
<emphasis id="7DBDEA926042FFC86AE2F820FDE5F84B" bold="true" box="[502,544,2001,2025]" pageId="7" pageNumber="523">(A),</emphasis>
whole animal, ventral view 
<emphasis id="7DBDEA926042FFC86854F820FCA7F84B" bold="true" box="[832,866,2001,2025]" pageId="7" pageNumber="523">(B)</emphasis>
. Scale bars: 
<emphasis id="7DBDEA926042FFC868F7F820FC30F84B" bold="true" box="[995,1013,2001,2025]" pageId="7" pageNumber="523">A</emphasis>
= 10.17 mm, 
<emphasis id="7DBDEA926042FFC86F90F820FB50F84B" bold="true" box="[1156,1173,2001,2025]" pageId="7" pageNumber="523">B</emphasis>
= 10 mm.
</paragraph>
</caption>
<caption id="1BB66608604DFFC76B83F879FB1FF84A" ID-DOI="http://doi.org/10.5281/zenodo.5497103" ID-Zenodo-Dep="5497103" httpUri="https://zenodo.org/record/5497103/files/figure.png" pageId="8" pageNumber="524" startId="8.[151,250,1928,1952]" targetBox="[207,1380,181,1905]" targetPageId="8">
<paragraph id="4F763680604DFFC76B83F879FB1FF84A" blockId="8.[151,1437,1928,2025]" pageId="8" pageNumber="524">
<emphasis id="7DBDEA92604DFFC76B83F879FED6F802" bold="true" box="[151,275,1928,1952]" pageId="8" pageNumber="524">FIGURE 2.</emphasis>
<taxonomicName id="88C94D03604DFFC76A0DF878FCF9F803" authority="Cumberlidge &amp; Tavares, 2006" authorityName="Cumberlidge &amp; Tavares" authorityYear="2006" box="[281,828,1929,1953]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="524" phylum="Arthropoda" rank="species" species="kensleyi">
<emphasis id="7DBDEA92604DFFC76A0DF878FDC7F803" box="[281,514,1929,1953]" italics="true" pageId="8" pageNumber="524">Potamonautes kensleyi</emphasis>
<bibRefCitation id="2B584B71604DFFC7691CF878FCF9F803" author="Cumberlidge, N. &amp; Tavares, M." box="[520,828,1929,1953]" pageId="8" pageNumber="524" pagination="248 - 257" refId="ref9684" refString="Cumberlidge, N. &amp; Tavares, M. (2006) Remarks on the freshwater crabs of Angola, southwestern Africa, with the description of Potamonautes kensleyi, new species (Brachyura: Potamoidea: Potamonautidae). Journal of Crustacean Biology, 26 (2), 248 - 257. https: // doi. org / 10.1651 / s- 2653.1" type="journal article" year="2006">Cumberlidge &amp; Tavares, 2006</bibRefCitation>
</taxonomicName>
, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A). Whole animal, dorsal view 
<emphasis id="7DBDEA92604DFFC768E7F85DFBD8F866" bold="true" box="[1011,1053,1964,1988]" pageId="8" pageNumber="524">(A),</emphasis>
whole animal, ventral view 
<emphasis id="7DBDEA92604DFFC76E2AF85DFAA5F866" bold="true" box="[1342,1376,1964,1988]" pageId="8" pageNumber="524">(B)</emphasis>
. Photographs by T. Busschau, University of Stellenbosch, South Africa. Scale bars: 
<emphasis id="7DBDEA92604DFFC768ADF821FC0EF84A" bold="true" box="[953,971,2000,2024]" pageId="8" pageNumber="524">A</emphasis>
= 12.0 mm, 
<emphasis id="7DBDEA92604DFFC76F59F820FB9BF84B" bold="true" box="[1101,1118,2001,2025]" pageId="8" pageNumber="524">B</emphasis>
= 12.2 mm.
</paragraph>
</caption>
<caption id="1BB66608604CFFC66B83F854FAC1F842" ID-DOI="http://doi.org/10.5281/zenodo.5497105" ID-Zenodo-Dep="5497105" httpUri="https://zenodo.org/record/5497105/files/figure.png" pageId="9" pageNumber="525" startId="9.[151,250,1957,1981]" targetBox="[191,1396,181,1932]" targetPageId="9">
<paragraph id="4F763680604CFFC66B83F854FAC1F842" blockId="9.[151,1436,1957,2017]" pageId="9" pageNumber="525">
<emphasis id="7DBDEA92604CFFC66B83F854FED1F81F" bold="true" box="[151,276,1957,1981]" pageId="9" pageNumber="525">FIGURE 3.</emphasis>
<taxonomicName id="88C94D03604CFFC66A08F857FD76F81F" authority="Bott, 1953" authorityName="Bott" authorityYear="1953" box="[284,691,1957,1981]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="525" phylum="Arthropoda" rank="species" species="macrobrachii">
<emphasis id="7DBDEA92604CFFC66A08F857FDF8F81F" box="[284,573,1957,1981]" italics="true" pageId="9" pageNumber="525">Potamonautes macrobrachii</emphasis>
<bibRefCitation id="2B584B71604CFFC66951F854FD76F81F" author="Bott, R." box="[581,691,1957,1981]" pageId="9" pageNumber="525" pagination="133 - 148" refId="ref9019" refString="Bott, R. (1953) Potamoniden (Crust. Decap.) von Angola. Publicacoes culturais da Companhia de Diamantes de Angola, Lisboa, 16, 133 - 148." type="journal article" year="1953">Bott, 1953</bibRefCitation>
</taxonomicName>
, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Whole animal, dorsal view 
<emphasis id="7DBDEA92604CFFC66AE2F838FDE5F843" bold="true" box="[502,544,1993,2017]" pageId="9" pageNumber="525">(A),</emphasis>
whole animal, ventral view 
<emphasis id="7DBDEA92604CFFC66854F838FCA7F843" bold="true" box="[832,866,1993,2017]" pageId="9" pageNumber="525">(B)</emphasis>
. Scale bars: 
<emphasis id="7DBDEA92604CFFC668F7F838FC30F843" bold="true" box="[995,1013,1993,2017]" pageId="9" pageNumber="525">A</emphasis>
= 12.5 mm, 
<emphasis id="7DBDEA92604CFFC66F63F838FB4DF843" bold="true" box="[1143,1160,1993,2017]" pageId="9" pageNumber="525">B</emphasis>
= 5.38 mm.
</paragraph>
</caption>
<caption id="1BB66608604FFFC56B83F922FBC4F8D9" ID-DOI="http://doi.org/10.5281/zenodo.5497107" ID-Zenodo-Dep="5497107" httpUri="https://zenodo.org/record/5497107/files/figure.png" pageId="10" pageNumber="526" startId="10.[151,250,1747,1771]" targetBox="[152,1436,309,1723]" targetPageId="10">
<paragraph id="4F763680604FFFC56B83F922FBC4F8D9" blockId="10.[151,1436,1747,1916]" pageId="10" pageNumber="526">
<emphasis id="7DBDEA92604FFFC56B83F922FED0F949" bold="true" box="[151,277,1747,1771]" pageId="10" pageNumber="526">FIGURE 4.</emphasis>
Frontal view of carapace. (
<emphasis id="7DBDEA92604FFFC56923F922FD8CF949" bold="true" box="[567,585,1747,1771]" pageId="10" pageNumber="526">A</emphasis>
) 
<taxonomicName id="88C94D03604FFFC5694FF925FC7EF94E" authority="Bott, 1968" authorityName="Bott" authorityYear="1968" box="[603,955,1748,1772]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="526" phylum="Arthropoda" rank="species" species="adeleae">
<emphasis id="7DBDEA92604FFFC5694FF925FC86F94E" box="[603,835,1748,1772]" italics="true" pageId="10" pageNumber="526">Potamonautes adeleae</emphasis>
<bibRefCitation id="2B584B71604FFFC56858F925FC7EF94E" author="Bott, R." box="[844,955,1748,1772]" pageId="10" pageNumber="526" pagination="165 - 172" refId="ref9155" refString="Bott, R. (1968) Decapoden aus dem Museu do Dundo (Crust. Decap.). Publicacoes culturais da Companhia de Diamantes de Angola, Lisboa, 77, 165 - 172." type="journal article" year="1968">Bott, 1968</bibRefCitation>
</taxonomicName>
, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (
<emphasis id="7DBDEA92604FFFC56909F909FDEBF8B2" bold="true" box="[541,558,1784,1808]" pageId="10" pageNumber="526">B</emphasis>
) 
<taxonomicName id="88C94D03604FFFC56929F909FB9BF8B2" authority="Cumberlidge &amp; Tavares, 2006" authorityName="Cumberlidge &amp; Tavares" authorityYear="2006" box="[573,1118,1784,1808]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="526" phylum="Arthropoda" rank="species" species="kensleyi">
<emphasis id="7DBDEA92604FFFC56929F909FCE0F8B2" box="[573,805,1784,1808]" italics="true" pageId="10" pageNumber="526">Potamonautes kensleyi</emphasis>
<bibRefCitation id="2B584B71604FFFC5683FF909FB9BF8B2" author="Cumberlidge, N. &amp; Tavares, M." box="[811,1118,1784,1808]" pageId="10" pageNumber="526" pagination="248 - 257" refId="ref9684" refString="Cumberlidge, N. &amp; Tavares, M. (2006) Remarks on the freshwater crabs of Angola, southwestern Africa, with the description of Potamonautes kensleyi, new species (Brachyura: Potamoidea: Potamonautidae). Journal of Crustacean Biology, 26 (2), 248 - 257. https: // doi. org / 10.1651 / s- 2653.1" type="journal article" year="2006">Cumberlidge &amp; Tavares, 2006</bibRefCitation>
</taxonomicName>
, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (
<emphasis id="7DBDEA92604FFFC56F08F8EAFBEBF891" bold="true" box="[1052,1070,1819,1843]" pageId="10" pageNumber="526">C</emphasis>
) 
<taxonomicName id="88C94D03604FFFC56F2BF8EDFF09F8F5" authority="Bott, 1953" authorityName="Bott" authorityYear="1953" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="526" phylum="Arthropoda" rank="species" species="macrobrachii">
<emphasis id="7DBDEA92604FFFC56F2BF8EDFAA7F896" box="[1087,1378,1820,1844]" italics="true" pageId="10" pageNumber="526">Potamonautes macrobrachii</emphasis>
<bibRefCitation id="2B584B71604FFFC56E7EF8EDFF09F8F5" author="Bott, R." pageId="10" pageNumber="526" pagination="133 - 148" refId="ref9019" refString="Bott, R. (1953) Potamoniden (Crust. Decap.) von Angola. Publicacoes culturais da Companhia de Diamantes de Angola, Lisboa, 16, 133 - 148." type="journal article" year="1953">Bott, 1953</bibRefCitation>
</taxonomicName>
, paratype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Photograph 
<emphasis id="7DBDEA92604FFFC56F64F8B1FB44F8FA" bold="true" box="[1136,1153,1856,1880]" pageId="10" pageNumber="526">B</emphasis>
by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: 
<emphasis id="7DBDEA92604FFFC56970F892FDB3F8D9" bold="true" box="[612,630,1891,1915]" pageId="10" pageNumber="526">A</emphasis>
= 10.4 mm; 
<emphasis id="7DBDEA92604FFFC569EDF895FCCFF8DE" bold="true" box="[761,778,1892,1916]" pageId="10" pageNumber="526">B</emphasis>
= 5.7 mm; 
<emphasis id="7DBDEA92604FFFC5686BF892FC54F8D9" bold="true" box="[895,913,1891,1915]" pageId="10" pageNumber="526">C</emphasis>
= 4.7 mm.
</paragraph>
</caption>
<caption id="1BB66608604EFFC46B83F9C5FC0BF87A" ID-DOI="http://doi.org/10.5281/zenodo.5497109" ID-Zenodo-Dep="5497109" httpUri="https://zenodo.org/record/5497109/files/figure.png" pageId="11" pageNumber="527" startId="11.[151,250,1588,1612]" targetBox="[159,1428,181,1565]" targetPageId="11">
<paragraph id="4F763680604EFFC46B83F9C5FC0BF87A" blockId="11.[151,1437,1588,2009]" pageId="11" pageNumber="527">
<emphasis id="7DBDEA92604EFFC46B83F9C5FED3F9EF" bold="true" box="[151,278,1588,1613]" pageId="11" pageNumber="527">FIGURE 5.</emphasis>
(
<emphasis id="7DBDEA92604EFFC46A3DF9C5FE92F9EF" bold="true" box="[297,343,1588,1613]" pageId="11" pageNumber="527">A–F</emphasis>
) Frontal view of right and left chela; (
<emphasis id="7DBDEA92604EFFC469EDF9C5FCEFF9EF" bold="true" box="[761,810,1588,1613]" pageId="11" pageNumber="527">G–L</emphasis>
) cheliped merus and carpus. (
<emphasis id="7DBDEA92604EFFC46F79F9C5FB65F9EF" bold="true" box="[1133,1184,1588,1613]" pageId="11" pageNumber="527">A, B</emphasis>
) 
<taxonomicName id="88C94D03604EFFC46FA7F9C4FEC0F9D3" authority="Bott, 1968" authorityName="Bott" authorityYear="1968" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="527" phylum="Arthropoda" rank="species" species="adeleae">
<emphasis id="7DBDEA92604EFFC46FA7F9C4FA59F9EF" box="[1203,1436,1589,1613]" italics="true" pageId="11" pageNumber="527">Potamonautes adeleae</emphasis>
<bibRefCitation id="2B584B71604EFFC46B83F9A8FEC0F9D3" author="Bott, R." box="[151,261,1625,1649]" pageId="11" pageNumber="527" pagination="165 - 172" refId="ref9155" refString="Bott, R. (1968) Decapoden aus dem Museu do Dundo (Crust. Decap.). Publicacoes culturais da Companhia de Diamantes de Angola, Lisboa, 77, 165 - 172." type="journal article" year="1968">Bott, 1968</bibRefCitation>
</taxonomicName>
, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (
<emphasis id="7DBDEA92604EFFC46F64F9A9FB67F9D3" bold="true" box="[1136,1186,1624,1649]" pageId="11" pageNumber="527">C, D</emphasis>
) 
<taxonomicName id="88C94D03604EFFC46FA6F9A8FE08F937" authority="Cumberlidge &amp; Tavares, 2006" authorityName="Cumberlidge &amp; Tavares" authorityYear="2006" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="527" phylum="Arthropoda" rank="species" species="kensleyi">
<emphasis id="7DBDEA92604EFFC46FA6F9A8FA58F9D3" box="[1202,1437,1625,1649]" italics="true" pageId="11" pageNumber="527">Potamonautes kensleyi</emphasis>
<bibRefCitation id="2B584B71604EFFC46B83F98CFE08F937" author="Cumberlidge, N. &amp; Tavares, M." box="[151,461,1661,1685]" pageId="11" pageNumber="527" pagination="248 - 257" refId="ref9684" refString="Cumberlidge, N. &amp; Tavares, M. (2006) Remarks on the freshwater crabs of Angola, southwestern Africa, with the description of Potamonautes kensleyi, new species (Brachyura: Potamoidea: Potamonautidae). Journal of Crustacean Biology, 26 (2), 248 - 257. https: // doi. org / 10.1651 / s- 2653.1" type="journal article" year="2006">Cumberlidge &amp; Tavares, 2006</bibRefCitation>
</taxonomicName>
, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (
<emphasis id="7DBDEA92604EFFC46A6BF950FE69F91B" bold="true" box="[383,428,1697,1721]" pageId="11" pageNumber="527">E, F</emphasis>
) 
<taxonomicName id="88C94D03604EFFC46AAFF950FC8AF91B" authority="Bott, 1953" authorityName="Bott" authorityYear="1953" box="[443,847,1697,1721]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="527" phylum="Arthropoda" rank="species" species="macrobrachii">
<emphasis id="7DBDEA92604EFFC46AAFF950FD19F91B" box="[443,732,1697,1721]" italics="true" pageId="11" pageNumber="527">Potamonautes macrobrachii</emphasis>
<bibRefCitation id="2B584B71604EFFC469F6F950FC8AF91B" author="Bott, R." box="[738,847,1697,1721]" pageId="11" pageNumber="527" pagination="133 - 148" refId="ref9019" refString="Bott, R. (1953) Potamoniden (Crust. Decap.) von Angola. Publicacoes culturais da Companhia de Diamantes de Angola, Lisboa, 16, 133 - 148." type="journal article" year="1953">Bott, 1953</bibRefCitation>
</taxonomicName>
, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892); (
<emphasis id="7DBDEA92604EFFC46A69F935FE77F97F" bold="true" box="[381,434,1732,1757]" pageId="11" pageNumber="527">G, H</emphasis>
) 
<taxonomicName id="88C94D03604EFFC46AD6F934FCDBF97F" authority="Bott, 1968" authorityName="Bott" authorityYear="1968" box="[450,798,1733,1757]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="527" phylum="Arthropoda" rank="species" species="adeleae">
<emphasis id="7DBDEA92604EFFC46AD6F934FD6CF97F" box="[450,681,1733,1757]" italics="true" pageId="11" pageNumber="527">Potamonautes adeleae</emphasis>
<bibRefCitation id="2B584B71604EFFC469A4F934FCDBF97F" author="Bott, R." box="[688,798,1733,1757]" pageId="11" pageNumber="527" pagination="165 - 172" refId="ref9155" refString="Bott, R. (1968) Decapoden aus dem Museu do Dundo (Crust. Decap.). Publicacoes culturais da Companhia de Diamantes de Angola, Lisboa, 77, 165 - 172." type="journal article" year="1968">Bott, 1968</bibRefCitation>
</taxonomicName>
, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (
<emphasis id="7DBDEA92604EFFC46A6BF919FE56F8A2" bold="true" box="[383,403,1768,1792]" pageId="11" pageNumber="527">G</emphasis>
) ventral view of right chela merus; (
<emphasis id="7DBDEA92604EFFC46805F918FCE0F8A3" bold="true" box="[785,805,1769,1793]" pageId="11" pageNumber="527">H</emphasis>
) dorsal view of right chela carpus and merus; (
<emphasis id="7DBDEA92604EFFC46E04F918FAF0F8A3" bold="true" box="[1296,1333,1769,1793]" pageId="11" pageNumber="527">I, J</emphasis>
) 
<taxonomicName id="88C94D03604EFFC46E51F918FDB0F887" authority="Cumberlidge &amp; Tavares, 2006" authorityName="Cumberlidge &amp; Tavares" authorityYear="2006" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="527" phylum="Arthropoda" rank="species" species="kensleyi">
<emphasis id="7DBDEA92604EFFC46E51F918FEF6F887" italics="true" pageId="11" pageNumber="527">Potamonautes kensleyi</emphasis>
<bibRefCitation id="2B584B71604EFFC46A2FF8FCFDB0F887" author="Cumberlidge, N. &amp; Tavares, M." box="[315,629,1805,1829]" pageId="11" pageNumber="527" pagination="248 - 257" refId="ref9684" refString="Cumberlidge, N. &amp; Tavares, M. (2006) Remarks on the freshwater crabs of Angola, southwestern Africa, with the description of Potamonautes kensleyi, new species (Brachyura: Potamoidea: Potamonautidae). Journal of Crustacean Biology, 26 (2), 248 - 257. https: // doi. org / 10.1651 / s- 2653.1" type="journal article" year="2006">Cumberlidge &amp; Tavares, 2006</bibRefCitation>
</taxonomicName>
, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A) (
<emphasis id="7DBDEA92604EFFC4693DF8C0FDF6F8EB" bold="true" box="[553,563,1841,1865]" pageId="11" pageNumber="527">I</emphasis>
) ventral view of right chela merus; (
<emphasis id="7DBDEA92604EFFC468B4F8C0FC68F8EB" bold="true" box="[928,941,1841,1865]" pageId="11" pageNumber="527">J</emphasis>
) dorsal view of right chela merus and carpus; (
<emphasis id="7DBDEA92604EFFC46E96F8C0FF6DF8CF" bold="true" pageId="11" pageNumber="527">K, L</emphasis>
) 
<taxonomicName id="88C94D03604EFFC46BA3F8A4FD8EF8CF" authority="Bott, 1953" authorityName="Bott" authorityYear="1953" box="[183,587,1877,1901]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="527" phylum="Arthropoda" rank="species" species="macrobrachii">
<emphasis id="7DBDEA92604EFFC46BA3F8A4FE12F8CF" box="[183,471,1877,1901]" italics="true" pageId="11" pageNumber="527">Potamonautes macrobrachii</emphasis>
<bibRefCitation id="2B584B71604EFFC46ACAF8A4FD8EF8CF" author="Bott, R." box="[478,587,1877,1901]" pageId="11" pageNumber="527" pagination="133 - 148" refId="ref9019" refString="Bott, R. (1953) Potamoniden (Crust. Decap.) von Angola. Publicacoes culturais da Companhia de Diamantes de Angola, Lisboa, 16, 133 - 148." type="journal article" year="1953">Bott, 1953</bibRefCitation>
</taxonomicName>
, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892): (
<emphasis id="7DBDEA92604EFFC46E94F8A4FA51F8CF" bold="true" box="[1408,1428,1877,1901]" pageId="11" pageNumber="527">K</emphasis>
) ventral view of right chela merus; (
<emphasis id="7DBDEA92604EFFC46AEAF888FDCAF833" bold="true" box="[510,527,1913,1937]" pageId="11" pageNumber="527">L</emphasis>
) dorsal view of right chela merus and carpus;. Photographs 
<emphasis id="7DBDEA92604EFFC46F67F889FB40F832" bold="true" box="[1139,1157,1912,1936]" pageId="11" pageNumber="527">C</emphasis>
, 
<emphasis id="7DBDEA92604EFFC46F86F888FB10F833" bold="true" box="[1170,1237,1913,1937]" pageId="11" pageNumber="527">D, I, J</emphasis>
by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: 
<emphasis id="7DBDEA92604EFFC469B8F86DFD78F816" bold="true" box="[684,701,1948,1972]" pageId="11" pageNumber="527">A</emphasis>
= 18.4 mm; 
<emphasis id="7DBDEA92604EFFC46854F86CFC94F817" bold="true" box="[832,849,1949,1973]" pageId="11" pageNumber="527">B</emphasis>
= 17.9 mm; 
<emphasis id="7DBDEA92604EFFC468C0F86DFBC0F817" bold="true" box="[980,1029,1948,1973]" pageId="11" pageNumber="527">C, D</emphasis>
= 8.9 mm; 
<emphasis id="7DBDEA92604EFFC46F68F86CFB48F817" bold="true" box="[1148,1165,1949,1973]" pageId="11" pageNumber="527">E</emphasis>
= 8.33 mm; 
<emphasis id="7DBDEA92604EFFC46E04F86CFADAF817" bold="true" box="[1296,1311,1949,1973]" pageId="11" pageNumber="527">F</emphasis>
= 5.81 mm; 
<emphasis id="7DBDEA92604EFFC46B83F831FF6EF87A" bold="true" box="[151,171,1984,2008]" pageId="11" pageNumber="527">G</emphasis>
= 17.9 mm; 
<emphasis id="7DBDEA92604EFFC46A39F830FE84F87B" bold="true" box="[301,321,1985,2009]" pageId="11" pageNumber="527">H</emphasis>
= 7.3 mm; 
<emphasis id="7DBDEA92604EFFC46AA3F830FE04F87B" bold="true" box="[439,449,1985,2009]" pageId="11" pageNumber="527">I</emphasis>
= 8.9 mm; 
<emphasis id="7DBDEA92604EFFC46922F830FD86F87B" bold="true" box="[566,579,1985,2009]" pageId="11" pageNumber="527">J</emphasis>
= 5.9 mm; 
<emphasis id="7DBDEA92604EFFC469ADF830FD08F87B" bold="true" box="[697,717,1985,2009]" pageId="11" pageNumber="527">K</emphasis>
= 5.81 mm; 
<emphasis id="7DBDEA92604EFFC4685BF830FC9AF87B" bold="true" box="[847,863,1985,2009]" pageId="11" pageNumber="527">L</emphasis>
= 5.6 mm.
</paragraph>
</caption>
<caption id="1BB666086049FFC36B83F8D5FE32F853" ID-DOI="http://doi.org/10.5281/zenodo.5497111" ID-Zenodo-Dep="5497111" httpUri="https://zenodo.org/record/5497111/files/figure.png" pageId="12" pageNumber="528" startId="12.[151,250,1828,1852]" targetBox="[197,1391,181,1805]" targetPageId="12">
<paragraph id="4F7636806049FFC36B83F8D5FE32F853" blockId="12.[151,1436,1828,2033]" pageId="12" pageNumber="528">
<emphasis id="7DBDEA926049FFC36B83F8D5FED6F89E" bold="true" box="[151,275,1828,1852]" pageId="12" pageNumber="528">FIGURE 6.</emphasis>
Right G1, ventral view (
<emphasis id="7DBDEA926049FFC3691BF8D5FD85F89E" bold="true" box="[527,576,1828,1852]" pageId="12" pageNumber="528">A–C</emphasis>
) and dorsal view (
<emphasis id="7DBDEA926049FFC369EAF8D4FCE9F89F" bold="true" box="[766,812,1829,1853]" pageId="12" pageNumber="528">D, F</emphasis>
). (
<emphasis id="7DBDEA926049FFC3685EF8D5FCBFF89F" bold="true" box="[842,890,1828,1853]" pageId="12" pageNumber="528">A, D</emphasis>
) 
<taxonomicName id="88C94D036049FFC3689DF8D4FB27F89F" authority="Bott, 1968" authorityName="Bott" authorityYear="1968" box="[905,1250,1829,1853]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="528" phylum="Arthropoda" rank="species" species="adeleae">
<emphasis id="7DBDEA926049FFC3689DF8D4FBAAF89F" box="[905,1135,1829,1853]" italics="true" pageId="12" pageNumber="528">Potamonautes adeleae</emphasis>
<bibRefCitation id="2B584B716049FFC36F61F8D4FB27F89F" author="Bott, R." box="[1141,1250,1829,1853]" pageId="12" pageNumber="528" pagination="165 - 172" refId="ref9155" refString="Bott, R. (1968) Decapoden aus dem Museu do Dundo (Crust. Decap.). Publicacoes culturais da Companhia de Diamantes de Angola, Lisboa, 77, 165 - 172." type="journal article" year="1968">Bott, 1968</bibRefCitation>
</taxonomicName>
, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (
<emphasis id="7DBDEA926049FFC36822F8B8FCA8F8C2" bold="true" box="[822,877,1864,1889]" pageId="12" pageNumber="528">B, E)</emphasis>
<taxonomicName id="88C94D036049FFC36860F8B8FA5DF8C3" authority="Cumberlidge &amp; Tavares, 2006" authorityName="Cumberlidge &amp; Tavares" authorityYear="2006" box="[884,1432,1865,1889]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="528" phylum="Arthropoda" rank="species" species="kensleyi">
<emphasis id="7DBDEA926049FFC36860F8B8FB98F8C3" box="[884,1117,1865,1889]" italics="true" pageId="12" pageNumber="528">Potamonautes kensleyi</emphasis>
<bibRefCitation id="2B584B716049FFC36F77F8B8FA5DF8C3" author="Cumberlidge, N. &amp; Tavares, M." box="[1123,1432,1865,1889]" pageId="12" pageNumber="528" pagination="248 - 257" refId="ref9684" refString="Cumberlidge, N. &amp; Tavares, M. (2006) Remarks on the freshwater crabs of Angola, southwestern Africa, with the description of Potamonautes kensleyi, new species (Brachyura: Potamoidea: Potamonautidae). Journal of Crustacean Biology, 26 (2), 248 - 257. https: // doi. org / 10.1651 / s- 2653.1" type="journal article" year="2006">Cumberlidge &amp; Tavares, 2006</bibRefCitation>
</taxonomicName>
, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (
<emphasis id="7DBDEA926049FFC36E77F89DFAB0F826" bold="true" box="[1379,1397,1900,1924]" pageId="12" pageNumber="528">C</emphasis>
, 
<emphasis id="7DBDEA926049FFC36E90F89CFA56F827" bold="true" box="[1412,1427,1901,1925]" pageId="12" pageNumber="528">F</emphasis>
) 
<taxonomicName id="88C94D036049FFC36B83F860FDE9F80A" authority="Bott, 1953" authorityName="Bott" authorityYear="1953" box="[151,556,1936,1961]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="528" phylum="Arthropoda" rank="species" species="macrobrachii">
<emphasis id="7DBDEA926049FFC36B83F860FE7DF80B" box="[151,440,1937,1961]" italics="true" pageId="12" pageNumber="528">Potamonautes macrobrachii</emphasis>
<bibRefCitation id="2B584B716049FFC36AAAF860FDE9F80A" author="Bott, R." box="[446,556,1936,1960]" pageId="12" pageNumber="528" pagination="133 - 148" refId="ref9019" refString="Bott, R. (1953) Potamoniden (Crust. Decap.) von Angola. Publicacoes culturais da Companhia de Diamantes de Angola, Lisboa, 16, 133 - 148." type="journal article" year="1953">Bott, 1953</bibRefCitation>
</taxonomicName>
, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Photo- graphs 
<emphasis id="7DBDEA926049FFC36BF4F844FF34F86F" bold="true" box="[224,241,1973,1997]" pageId="12" pageNumber="528">B</emphasis>
and 
<emphasis id="7DBDEA926049FFC36A35F844FEF7F86F" bold="true" box="[289,306,1973,1997]" pageId="12" pageNumber="528">E</emphasis>
by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: 
<emphasis id="7DBDEA926049FFC36F18F845FBDBF86E" bold="true" box="[1036,1054,1972,1996]" pageId="12" pageNumber="528">A</emphasis>
= 6.7 mm; 
<emphasis id="7DBDEA926049FFC36F98F844FB58F86F" bold="true" box="[1164,1181,1973,1997]" pageId="12" pageNumber="528">B</emphasis>
= 2.3 mm; 
<emphasis id="7DBDEA926049FFC36E04F845FAE7F86E" bold="true" box="[1296,1314,1972,1996]" pageId="12" pageNumber="528">C</emphasis>
= 1.13 mm; 
<emphasis id="7DBDEA926049FFC36B83F828FF6CF853" bold="true" box="[151,169,2009,2033]" pageId="12" pageNumber="528">D</emphasis>
= 2.7 mm; 
<emphasis id="7DBDEA926049FFC36A0BF828FEF5F853" bold="true" box="[287,304,2009,2033]" pageId="12" pageNumber="528">E</emphasis>
= 2.3 mm; 
<emphasis id="7DBDEA926049FFC36AB2F828FE70F853" bold="true" box="[422,437,2009,2033]" pageId="12" pageNumber="528">F</emphasis>
= 1.2.
</paragraph>
</caption>
<caption id="1BB666086048FFC26B83F8CEFC36F844" ID-DOI="http://doi.org/10.5281/zenodo.5497113" ID-Zenodo-Dep="5497113" httpUri="https://zenodo.org/record/5497113/files/figure.png" pageId="13" pageNumber="529" startId="13.[151,250,1855,1879]" targetBox="[264,1323,181,1831]" targetPageId="13">
<paragraph id="4F7636806048FFC26B83F8CEFC36F844" blockId="13.[151,1437,1855,2023]" pageId="13" pageNumber="529">
<emphasis id="7DBDEA926048FFC26B83F8CEFED6F8F5" bold="true" box="[151,275,1855,1879]" pageId="13" pageNumber="529">FIGURE 7.</emphasis>
Right G2 ventral view (
<emphasis id="7DBDEA926048FFC2691CF8CEFDFCF8F5" bold="true" box="[520,569,1855,1879]" pageId="13" pageNumber="529">A–C</emphasis>
); (
<emphasis id="7DBDEA926048FFC26943F8CEFDACF8F5" bold="true" box="[599,617,1855,1879]" pageId="13" pageNumber="529">A</emphasis>
) 
<taxonomicName id="88C94D036048FFC2696CF8B1FC14F8F5" authority="Bott, 1968" authorityName="Bott" authorityYear="1968" box="[632,977,1855,1879]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="529" phylum="Arthropoda" rank="species" species="adeleae">
<emphasis id="7DBDEA926048FFC2696CF8B1FC9BF8F5" box="[632,862,1855,1879]" italics="true" pageId="13" pageNumber="529">Potamonautes adeleae</emphasis>
<bibRefCitation id="2B584B716048FFC26870F8CEFC14F8F5" author="Bott, R." box="[868,977,1855,1879]" pageId="13" pageNumber="529" pagination="165 - 172" refId="ref9155" refString="Bott, R. (1968) Decapoden aus dem Museu do Dundo (Crust. Decap.). Publicacoes culturais da Companhia de Diamantes de Angola, Lisboa, 77, 165 - 172." type="journal article" year="1968">Bott, 1968</bibRefCitation>
</taxonomicName>
, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (
<emphasis id="7DBDEA926048FFC26909F892FDEBF8D9" bold="true" box="[541,558,1891,1915]" pageId="13" pageNumber="529">B</emphasis>
) 
<taxonomicName id="88C94D036048FFC26929F895FB9BF8D9" authority="Cumberlidge &amp; Tavares, 2006" authorityName="Cumberlidge &amp; Tavares" authorityYear="2006" box="[573,1118,1891,1915]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="529" phylum="Arthropoda" rank="species" species="kensleyi">
<emphasis id="7DBDEA926048FFC26929F895FCE0F8D9" box="[573,805,1891,1915]" italics="true" pageId="13" pageNumber="529">Potamonautes kensleyi</emphasis>
<bibRefCitation id="2B584B716048FFC2683FF892FB9BF8D9" author="Cumberlidge, N. &amp; Tavares, M." box="[811,1118,1891,1915]" pageId="13" pageNumber="529" pagination="248 - 257" refId="ref9684" refString="Cumberlidge, N. &amp; Tavares, M. (2006) Remarks on the freshwater crabs of Angola, southwestern Africa, with the description of Potamonautes kensleyi, new species (Brachyura: Potamoidea: Potamonautidae). Journal of Crustacean Biology, 26 (2), 248 - 257. https: // doi. org / 10.1651 / s- 2653.1" type="journal article" year="2006">Cumberlidge &amp; Tavares, 2006</bibRefCitation>
</taxonomicName>
, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (
<emphasis id="7DBDEA926048FFC26F08F876FBEBF83D" bold="true" box="[1052,1070,1927,1951]" pageId="13" pageNumber="529">C</emphasis>
) 
<taxonomicName id="88C94D036048FFC26F2BF879FF09F861" authority="Bott, 1953" authorityName="Bott" authorityYear="1953" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="529" phylum="Arthropoda" rank="species" species="macrobrachii">
<emphasis id="7DBDEA926048FFC26F2BF879FAA7F83D" box="[1087,1378,1927,1951]" italics="true" pageId="13" pageNumber="529">Potamonautes macrobrachii</emphasis>
<bibRefCitation id="2B584B716048FFC26E7EF876FF09F861" author="Bott, R." pageId="13" pageNumber="529" pagination="133 - 148" refId="ref9019" refString="Bott, R. (1953) Potamoniden (Crust. Decap.) von Angola. Publicacoes culturais da Companhia de Diamantes de Angola, Lisboa, 16, 133 - 148." type="journal article" year="1953">Bott, 1953</bibRefCitation>
</taxonomicName>
, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Photograph 
<emphasis id="7DBDEA926048FFC26F7BF85AFB44F861" bold="true" box="[1135,1153,1963,1987]" pageId="13" pageNumber="529">C</emphasis>
by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: 
<emphasis id="7DBDEA926048FFC26970F83EFDB0F845" bold="true" box="[612,629,1999,2023]" pageId="13" pageNumber="529">A</emphasis>
= 5.6 mm; 
<emphasis id="7DBDEA926048FFC269FFF83EFD39F845" bold="true" box="[747,764,1999,2023]" pageId="13" pageNumber="529">B</emphasis>
= 2.3 mm; 
<emphasis id="7DBDEA926048FFC26865F83EFC46F845" bold="true" box="[881,899,1999,2023]" pageId="13" pageNumber="529">C</emphasis>
= 1.2 mm.
</paragraph>
</caption>
<caption id="1BB66608604BFFC16B83FAC9FE37FA3A" ID-DOI="http://doi.org/10.5281/zenodo.5497115" ID-Zenodo-Dep="5497115" httpUri="https://zenodo.org/record/5497115/files/figure.png" pageId="14" pageNumber="530" startId="14.[151,250,1336,1360]" targetBox="[159,1428,189,1302]" targetPageId="14">
<paragraph id="4F763680604BFFC16B83FAC9FE37FA3A" blockId="14.[151,1437,1336,1432]" pageId="14" pageNumber="530">
<emphasis id="7DBDEA92604BFFC16B83FAC9FED1FAF2" bold="true" box="[151,276,1336,1360]" pageId="14" pageNumber="530">FIGURE 8.</emphasis>
Map of the Angola showing the known localities of 
<taxonomicName id="88C94D03604BFFC16825FAC8FBDDFAF2" authorityName="Bott" authorityYear="1968" box="[817,1048,1336,1360]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="530" phylum="Arthropoda" rank="species" species="adeleae">
<emphasis id="7DBDEA92604BFFC16825FAC8FBDDFAF2" box="[817,1048,1336,1360]" italics="true" pageId="14" pageNumber="530">Potamonautes adeleae</emphasis>
</taxonomicName>
(black square, not shown to scale), 
<taxonomicName id="88C94D03604BFFC16E9EFAC8FF2CFAD6" authorityName="Cumberlidge &amp; Tavares" authorityYear="2006" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="530" phylum="Arthropoda" rank="species" species="kensleyi">
<emphasis id="7DBDEA92604BFFC16E9EFAC8FF2CFAD6" italics="true" pageId="14" pageNumber="530">P. kensleyi</emphasis>
</taxonomicName>
(black circles, not shown to scale), and 
<taxonomicName id="88C94D03604BFFC16994FAACFCE7FAD6" authorityName="Bott" authorityYear="1953" box="[640,802,1372,1396]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="530" phylum="Arthropoda" rank="species" species="macrobrachii">
<emphasis id="7DBDEA92604BFFC16994FAACFCE7FAD6" box="[640,802,1372,1396]" italics="true" pageId="14" pageNumber="530">P. macrobrachii</emphasis>
</taxonomicName>
(black diamond, not shown to scale). Scale bar = 520 km. See text for exact locality coordinates.
</paragraph>
</caption>
<paragraph id="4F763680604BFFC16BD3FA37FE1EF8A3" blockId="14.[151,1437,1478,1793]" pageId="14" pageNumber="530">
Characters that distinguish 
<taxonomicName id="88C94D03604BFFC16AE8FA36FD75FA42" authorityName="Bott" authorityYear="1953" box="[508,688,1478,1504]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="530" phylum="Arthropoda" rank="species" species="macrobrachii">
<emphasis id="7DBDEA92604BFFC16AE8FA36FD75FA42" box="[508,688,1478,1504]" italics="true" pageId="14" pageNumber="530">P. macrobrachii</emphasis>
</taxonomicName>
from 
<taxonomicName id="88C94D03604BFFC169E3FA36FC44FA42" baseAuthorityName="de Brito Capello" baseAuthorityYear="1870" box="[759,897,1478,1504]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="530" phylum="Arthropoda" rank="species" species="anchietae">
<emphasis id="7DBDEA92604BFFC169E3FA36FC44FA42" box="[759,897,1478,1504]" italics="true" pageId="14" pageNumber="530">P. anchietae</emphasis>
</taxonomicName>
include the following: the cheliped merus of 
<taxonomicName id="88C94D03604BFFC16E9CFA36FEEBF9A6" authorityName="Bott" authorityYear="1953" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="530" phylum="Arthropoda" rank="species" species="macrobrachii">
<emphasis id="7DBDEA92604BFFC16E9CFA36FEEBF9A6" italics="true" pageId="14" pageNumber="530">P. macrobrachii</emphasis>
</taxonomicName>
is greatly elongated (
<figureCitation id="D7F22A05604BFFC16935FA1BFD41F9A6" box="[545,644,1514,1540]" captionStart="FIGURE 3" captionStartId="9.[151,250,1957,1981]" captionTargetBox="[191,1396,181,1932]" captionTargetId="figure-21@9.[191,1396,181,1932]" captionTargetPageId="9" captionText="FIGURE 3. Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Whole animal, dorsal view (A), whole animal, ventral view (B). Scale bars: A = 12.5 mm, B = 5.38 mm." figureDoi="http://doi.org/10.5281/zenodo.5497105" httpUri="https://zenodo.org/record/5497105/files/figure.png" pageId="14" pageNumber="530">Figs. 3A</figureCitation>
, 
<figureCitation id="D7F22A05604BFFC16984FA1BFD16F9A7" box="[656,723,1514,1541]" captionStart="FIGURE 5" captionStartId="11.[151,250,1588,1612]" captionTargetBox="[159,1428,181,1565]" captionTargetId="figure-21@11.[159,1428,181,1565]" captionTargetPageId="11" captionText="FIGURE 5. (A–F) Frontal view of right and left chela; (G–L) cheliped merus and carpus. (A, B) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (C, D) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (E, F) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892); (G, H) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (G) ventral view of right chela merus; (H) dorsal view of right chela carpus and merus; (I, J) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province,Angola (AMG GEN 1887A) (I) ventral view of right chela merus; (J) dorsal view of right chela merus and carpus; (K, L) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892): (K) ventral view of right chela merus; (L) dorsal view of right chela merus and carpus;. Photographs C, D, I, J by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A = 18.4 mm; B = 17.9 mm; C, D = 8.9 mm; E = 8.33 mm; F = 5.81 mm; G = 17.9 mm; H = 7.3 mm; I = 8.9 mm; J = 5.9 mm; K = 5.81 mm; L = 5.6 mm." figureDoi="http://doi.org/10.5281/zenodo.5497109" httpUri="https://zenodo.org/record/5497109/files/figure.png" pageId="14" pageNumber="530">5K, L</figureCitation>
) (vs a cheliped merus of normal length in 
<taxonomicName id="88C94D03604BFFC16FBBFA1AFAFDF9A6" baseAuthorityName="de Brito Capello" baseAuthorityYear="1870" box="[1199,1336,1514,1540]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="530" phylum="Arthropoda" rank="species" species="anchietae">
<emphasis id="7DBDEA92604BFFC16FBBFA1AFAFDF9A6" box="[1199,1336,1514,1540]" italics="true" pageId="14" pageNumber="530">P. anchietae</emphasis>
</taxonomicName>
(cf. 
<bibRefCitation id="2B584B71604BFFC16E78FA1AFF16F98B" author="Bott, R." pageId="14" pageNumber="530" pagination="209 - 352" refId="ref9053" refString="Bott, R. (1955) Die Susswasserkrabben von Afrika (Crust. Decap.) und Ihre Stammesgeschichte. Annales du Musee Du Congo Belge, Tervuren, Belgique), C Zoologie, (3, 3), 3 (1), 209 - 352." type="journal article" year="1955">Bott 1955</bibRefCitation>
: fig. 1a, d); the distal and proximal teeth on the inner margin of the cheliped carpus of 
<taxonomicName id="88C94D03604BFFC16F9FF9FEFAF9F98A" authorityName="Bott" authorityYear="1953" box="[1163,1340,1550,1576]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="530" phylum="Arthropoda" rank="species" species="macrobrachii">
<emphasis id="7DBDEA92604BFFC16F9FF9FEFAF9F98A" box="[1163,1340,1550,1576]" italics="true" pageId="14" pageNumber="530">P. macrobrachii</emphasis>
</taxonomicName>
are both rounded and low (
<figureCitation id="D7F22A05604BFFC16A77F9C3FE20F9EF" box="[355,485,1586,1613]" captionStart="FIGURE 5" captionStartId="11.[151,250,1588,1612]" captionTargetBox="[159,1428,181,1565]" captionTargetId="figure-21@11.[159,1428,181,1565]" captionTargetPageId="11" captionText="FIGURE 5. (A–F) Frontal view of right and left chela; (G–L) cheliped merus and carpus. (A, B) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (C, D) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (E, F) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892); (G, H) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (G) ventral view of right chela merus; (H) dorsal view of right chela carpus and merus; (I, J) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province,Angola (AMG GEN 1887A) (I) ventral view of right chela merus; (J) dorsal view of right chela merus and carpus; (K, L) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892): (K) ventral view of right chela merus; (L) dorsal view of right chela merus and carpus;. Photographs C, D, I, J by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A = 18.4 mm; B = 17.9 mm; C, D = 8.9 mm; E = 8.33 mm; F = 5.81 mm; G = 17.9 mm; H = 7.3 mm; I = 8.9 mm; J = 5.9 mm; K = 5.81 mm; L = 5.6 mm." figureDoi="http://doi.org/10.5281/zenodo.5497109" httpUri="https://zenodo.org/record/5497109/files/figure.png" pageId="14" pageNumber="530">Figs. 5K, L</figureCitation>
) (vs two pointed teeth in 
<taxonomicName id="88C94D03604BFFC16811F9C2FC48F9EE" baseAuthorityName="de Brito Capello" baseAuthorityYear="1870" box="[773,909,1586,1612]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="530" phylum="Arthropoda" rank="species" species="anchietae">
<emphasis id="7DBDEA92604BFFC16811F9C2FC48F9EE" box="[773,909,1586,1612]" italics="true" pageId="14" pageNumber="530">P. anchietae</emphasis>
</taxonomicName>
(cf. 
<bibRefCitation id="2B584B71604BFFC168D6F9C2FBF0F9EF" author="Bott, R." box="[962,1077,1586,1613]" pageId="14" pageNumber="530" pagination="209 - 352" refId="ref9053" refString="Bott, R. (1955) Die Susswasserkrabben von Afrika (Crust. Decap.) und Ihre Stammesgeschichte. Annales du Musee Du Congo Belge, Tervuren, Belgique), C Zoologie, (3, 3), 3 (1), 209 - 352." type="journal article" year="1955">Bott 1955</bibRefCitation>
: fig. 1a)); the branchiostegite of 
<taxonomicName id="88C94D03604BFFC16B83F9A6FE8FF9D2" authorityName="Bott" authorityYear="1953" box="[151,330,1622,1648]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="530" phylum="Arthropoda" rank="species" species="macrobrachii">
<emphasis id="7DBDEA92604BFFC16B83F9A6FE8FF9D2" box="[151,330,1622,1648]" italics="true" pageId="14" pageNumber="530">P. macrobrachii</emphasis>
</taxonomicName>
is divided by two sutures into three parts (
<figureCitation id="D7F22A05604BFFC1683FF9A7FC48F9D2" box="[811,909,1622,1648]" captionStart="FIGURE 3" captionStartId="9.[151,250,1957,1981]" captionTargetBox="[191,1396,181,1932]" captionTargetId="figure-21@9.[191,1396,181,1932]" captionTargetPageId="9" captionText="FIGURE 3. Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Whole animal, dorsal view (A), whole animal, ventral view (B). Scale bars: A = 12.5 mm, B = 5.38 mm." figureDoi="http://doi.org/10.5281/zenodo.5497105" httpUri="https://zenodo.org/record/5497105/files/figure.png" pageId="14" pageNumber="530">Figs. 3B</figureCitation>
, 
<figureCitation id="D7F22A05604BFFC1688DF9A7FC7EF9D2" box="[921,955,1622,1648]" captionStart="FIGURE 4" captionStartId="10.[151,250,1747,1771]" captionTargetBox="[152,1436,309,1723]" captionTargetId="figure-93@10.[152,1436,309,1723]" captionTargetPageId="10" captionText="FIGURE 4. Frontal view of carapace. (A) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (B) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (C) Potamonautes macrobrachii Bott, 1953, paratype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Photograph B by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A = 10.4 mm; B = 5.7 mm; C = 4.7 mm." figureDoi="http://doi.org/10.5281/zenodo.5497107" httpUri="https://zenodo.org/record/5497107/files/figure.png" pageId="14" pageNumber="530">4C</figureCitation>
) (vs divided into four parts in 
<taxonomicName id="88C94D03604BFFC16E00F9A6FA59F9D2" baseAuthorityName="Brito-Capello" baseAuthorityYear="1871" box="[1300,1436,1622,1648]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="530" phylum="Arthropoda" rank="species" species="anchetiae">
<emphasis id="7DBDEA92604BFFC16E00F9A6FA59F9D2" box="[1300,1436,1622,1648]" italics="true" pageId="14" pageNumber="530">P. anchetiae</emphasis>
</taxonomicName>
(cf. 
<bibRefCitation id="2B584B71604BFFC16BD1F98BFDDDF937" author="Cumberlidge, N. &amp; Tavares, M." box="[197,536,1658,1685]" pageId="14" pageNumber="530" pagination="248 - 257" refId="ref9684" refString="Cumberlidge, N. &amp; Tavares, M. (2006) Remarks on the freshwater crabs of Angola, southwestern Africa, with the description of Potamonautes kensleyi, new species (Brachyura: Potamoidea: Potamonautidae). Journal of Crustacean Biology, 26 (2), 248 - 257. https: // doi. org / 10.1651 / s- 2653.1" type="journal article" year="2006">Cumberlidge &amp; Tavares 2006</bibRefCitation>
)); and the G1 TA of 
<taxonomicName id="88C94D03604BFFC16810F98AFC72F936" authorityName="Bott" authorityYear="1953" box="[772,951,1658,1684]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="530" phylum="Arthropoda" rank="species" species="macrobrachii">
<emphasis id="7DBDEA92604BFFC16810F98AFC72F936" box="[772,951,1658,1684]" italics="true" pageId="14" pageNumber="530">P. macrobrachii</emphasis>
</taxonomicName>
is strikingly widened in the midsection by a raised, rounded mesial fold that is significantly higher than the lateral fold (
<figureCitation id="D7F22A05604BFFC16F12F96FFBB9F91B" box="[1030,1148,1694,1721]" captionStart="FIGURE 6" captionStartId="12.[151,250,1828,1852]" captionTargetBox="[197,1391,181,1805]" captionTargetId="figure-17@12.[197,1391,181,1805]" captionTargetPageId="12" captionText="FIGURE 6. Right G1, ventral view (A–C) and dorsal view (D, F). (A, D) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (B, E) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (C, F) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Photo- graphs B and E by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A= 6.7 mm; B = 2.3 mm; C = 1.13 mm; D = 2.7 mm; E = 2.3 mm; F = 1.2." figureDoi="http://doi.org/10.5281/zenodo.5497111" httpUri="https://zenodo.org/record/5497111/files/figure.png" pageId="14" pageNumber="530">Fig. 6C, F</figureCitation>
) (vs a G1 TA that is only slightly widened in the midsection by a low mesial fold that is only slightly higher than the lateral fold in 
<taxonomicName id="88C94D03604BFFC16E2DF932FF03F8A2" baseAuthorityName="de Brito Capello" baseAuthorityYear="1870" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="530" phylum="Arthropoda" rank="species" species="anchietae">
<emphasis id="7DBDEA92604BFFC16E2DF932FF03F8A2" italics="true" pageId="14" pageNumber="530">P. anchietae</emphasis>
</taxonomicName>
(cf. 
<bibRefCitation id="2B584B71604BFFC16BEEF916FEA8F8A3" author="Bott, R." box="[250,365,1766,1793]" pageId="14" pageNumber="530" pagination="209 - 352" refId="ref9053" refString="Bott, R. (1955) Die Susswasserkrabben von Afrika (Crust. Decap.) und Ihre Stammesgeschichte. Annales du Musee Du Congo Belge, Tervuren, Belgique), C Zoologie, (3, 3), 3 (1), 209 - 352." type="journal article" year="1955">Bott 1955</bibRefCitation>
: fig. 25)).
</paragraph>
</subSubSection>
</treatment>
</document>