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<document id="6580C791642BE2D54610648C63A9B2D8" ID-CLB-Dataset="29036" ID-DOI="10.11646/zootaxa.4459.1.2" ID-GBIF-Dataset="acdf8f3d-d29f-463e-a431-191275b69a80" ID-ISSN="1175-5326" ID-Zenodo-Dep="1458336" ID-ZooBank="67E956AB-04B1-4EF7-8CC0-E152F95D0563" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539287820689" checkinUser="plazi" docAuthor="Liu, Weixin &amp; Golovatch, Sergei" docDate="2018" docId="9005EB48423A8E5348DAFAE705EDFCAD" docLanguage="en" docName="zootaxa.4459.1.2.pdf" docOrigin="Zootaxa 4459 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Epanerchodus latus Liu &amp; Golovatch 2018, new species" docType="treatment" docVersion="7" lastPageNumber="58" masterDocId="6C3C933042398E56484DFFE30629FF95" masterDocTitle="The millipede genus Epanerchodus Attems, 1901 in continental China, with descriptions of seven new cavernicolous species (Diplopoda, Polydesmida, Polydesmidae)" masterLastPageNumber="84" masterPageNumber="53" pageNumber="56" updateTime="1698577720608" updateUser="ExternalLinkService">
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<mods:title id="04216F7A31EFD5889F5EC91097C59536">The millipede genus Epanerchodus Attems, 1901 in continental China, with descriptions of seven new cavernicolous species (Diplopoda, Polydesmida, Polydesmidae)</mods:title>
</mods:titleInfo>
<mods:name id="92198398462B12F6786B6126EDFA7F62" type="personal">
<mods:role id="2C0532A23F28602B134D526A7DA7E4E7">
<mods:roleTerm id="55BF514011C6F646DF5E4E8A5659100A">Author</mods:roleTerm>
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<mods:namePart id="6BA29BEA882F0C549DE2B909F60EAFF4">Liu, Weixin</mods:namePart>
</mods:name>
<mods:name id="F514618B732926FE19E4B2963995DFB9" type="personal">
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<mods:roleTerm id="A145A9006D32CBFAB8919072157DE409">Author</mods:roleTerm>
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<mods:namePart id="74D874B1BCAB54FED2518560A04B3256">Golovatch, Sergei</mods:namePart>
</mods:name>
<mods:typeOfResource id="3CCFA86430ADFD4D51FD8BAEB17D2301">text</mods:typeOfResource>
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<mods:title id="65E3497F8C53637F1640227F6844D1BA">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A06E5E594FA2C7BEB604B2AE3145C150">
<mods:date id="418FEB074AD1816BB208DB12DA0DEAE6">2018</mods:date>
<mods:detail id="0E10F096D89D7247511932232EE6EB81" type="pubDate">
<mods:number id="4B3218F2C70D7E1C0F8DE1864D9FDA94">2018-08-14</mods:number>
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<mods:detail id="498F0240270BE8DC63592EC2B16B9A03" type="volume">
<mods:number id="C6890D6EA6511A471FB8CFC3C170A5E7">4459</mods:number>
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<mods:classification id="957C66BB9FEF98D1647083D7B9FEB96B">journal article</mods:classification>
<mods:identifier id="940AB1A39D28E2D65095778622856953" type="CLB-Dataset">29036</mods:identifier>
<mods:identifier id="8FE8274FD59672E259E032251E5467B8" type="DOI">10.11646/zootaxa.4459.1.2</mods:identifier>
<mods:identifier id="F17B8AA0EB5481360E1426201038C58D" type="GBIF-Dataset">acdf8f3d-d29f-463e-a431-191275b69a80</mods:identifier>
<mods:identifier id="C2AFDB29B3EF4761DA90D83987880876" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="91E07859DEF148ED563F6EC48A3B41E9" type="Zenodo-Dep">1458336</mods:identifier>
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<treatment id="9005EB48423A8E5348DAFAE705EDFCAD" ID-DOI="http://doi.org/10.5281/zenodo.5973764" ID-GBIF-Taxon="148692457" ID-Zenodo-Dep="5973764" LSID="urn:lsid:plazi:treatment:9005EB48423A8E5348DAFAE705EDFCAD" httpUri="http://treatment.plazi.org/id/9005EB48423A8E5348DAFAE705EDFCAD" lastPageId="5" lastPageNumber="58" pageId="3" pageNumber="56">
<subSubSection id="50B609D5423A8E5548DAFAE706D6FAAA" pageId="3" pageNumber="56" type="nomenclature">
<paragraph id="18135A5E423A8E5548DAFAE7040EFA8B" blockId="3.[151,551,1284,1343]" box="[151,551,1284,1310]" pageId="3" pageNumber="56">
<heading id="435BED32423A8E5548DAFAE7040EFA8B" bold="true" box="[151,551,1284,1310]" fontSize="11" level="1" pageId="3" pageNumber="56" reason="1">
<emphasis id="2AD8864C423A8E5548DAFAE7040EFA8B" bold="true" box="[151,551,1284,1310]" pageId="3" pageNumber="56">
<taxonomicName id="DFAC21DD423A8E5548DAFAE707A1FA8B" ID-CoL="6FMTL" authority="Liu &amp; Golovatch, 2018" authorityName="Liu &amp; Golovatch" authorityYear="2018" box="[151,392,1284,1310]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="latus" status="sp. nov.">
<emphasis id="2AD8864C423A8E5548DAFAE707A1FA8B" bold="true" box="[151,392,1284,1310]" italics="true" pageId="3" pageNumber="56">Epanerchodus latus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="31EB3B37423A8E5549DAFAE7040EFA8B" box="[407,551,1284,1310]" pageId="3" pageNumber="56" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="18135A5E423A8E5548DAFAC506D6FAAA" blockId="3.[151,551,1284,1343]" box="[151,255,1318,1343]" pageId="3" pageNumber="56">
<figureCitation id="809746DB423A8E5548DAFAC506D2FAAA" box="[151,251,1318,1343]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="4.[151,250,1782,1804]" captionStartId-1="5.[151,250,1837,1859]" captionTargetBox-0="[205,1388,198,1761]" captionTargetBox-1="[192,1395,905,1791]" captionTargetId-0="figure@4.[204,1394,195,1767]" captionTargetId-1="figure@5.[188,1398,877,1816]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 1. Epanerchodus latus n. sp., male paratype. A: anterior part of body, ventral view. B: anterior part of body, dorsal view. C: posterior part of body, dorsal view. D: gonopods in situ, ventral view. Scale bar: A–C, 0.2 mm; D, no scale." captionText-1="FIGURE 2. Epanerchodus latus n. sp., male paratype. A: right gonopod, lateral view. B: right gonopod, mesal view. Abbreviations: cl = clivus; en = endomere; l = lobe; p1 and p2 = processes 1 and 2; sg = seminal groove." httpUri-0="https://zenodo.org/record/1458338/files/figure.png" httpUri-1="https://zenodo.org/record/1458340/files/figure.png" pageId="3" pageNumber="56">Figs 1–2</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="50B609D5423A8E5548DAFA8D037EFA5A" pageId="3" pageNumber="56" type="materials_examined">
<paragraph id="18135A5E423A8E5548DAFA8D037EFA5A" blockId="3.[151,1436,1390,2027]" pageId="3" pageNumber="56">
<materialsCitation id="A8C45003423A8E5548DAFA8D042EFA5A" ID-GBIF-Occurrence="1917355134" collectingDate="2014-07-06" collectionCode="SCAU" country="China" county="Wushan County" location="Qinglong Village" municipality="Luoping Town" pageId="3" pageNumber="56" specimenCount="1" stateProvince="Chongqing" typeStatus="holotype">
<emphasis id="2AD8864C423A8E5548DAFA8D0756FA12" bold="true" box="[151,383,1390,1415]" pageId="3" pageNumber="56">Material examined</emphasis>
:
<typeStatus id="C717E4FC423A8E5549DEFA8D07D2FA12" box="[403,507,1390,1415]" pageId="3" pageNumber="56" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="7EBDC29B423A8E554A1FFA8C04B7FA12" box="[594,670,1391,1415]" pageId="3" pageNumber="56">SCAU</collectionCode>
eCQ2-1) from
<collectingCountry id="60BB1ACE423A8E554B11FA8D058BFA12" box="[860,930,1390,1415]" name="China" pageId="3" pageNumber="56">China</collectingCountry>
,
<collectingRegion id="DA6894BC423A8E554BFFFA8D0207FA12" box="[946,1070,1390,1415]" country="China" name="Chongqing" pageId="3" pageNumber="56">Chongqing</collectingRegion>
Municipality,
<collectingCounty id="F17222D2423A8E554C90FA8D03B1FA12" box="[1245,1432,1390,1415]" pageId="3" pageNumber="56">Wushan County</collectingCounty>
,
<collectingMunicipality id="F877C024423A8E5548DAFA710716FA3F" box="[151,319,1426,1451]" pageId="3" pageNumber="56">Luoping Town</collectingMunicipality>
,
<location id="1D730C85423A8E554907FA710426FA3E" LSID="urn:lsid:plazi:treatment:9005EB48423A8E5348DAFAE705EDFCAD:1D730C85423A8E554907FA710426FA3E" box="[330,527,1426,1451]" country="China" county="Wushan County" municipality="Luoping Town" name="Qinglong Village" pageId="3" pageNumber="56" stateProvince="Chongqing">Qinglong Village</location>
,
<location id="1D730C85423A8E554A57FA710525FA3E" LSID="urn:lsid:plazi:treatment:9005EB48423A8E5348DAFAE705EDFCAD:1D730C85423A8E554A57FA710525FA3E" box="[538,780,1426,1451]" country="China" county="Wushan County" municipality="Luoping Town" name="Qinglong Dong Cave" pageId="3" pageNumber="56" stateProvince="Chongqing">Qinglong Dong Cave</location>
,
<collectingDate id="7C568576423A8E554B55FA7105BFFA3E" box="[792,918,1426,1451]" pageId="3" pageNumber="56" value="2014-07-06">
<date id="6C127C9E423A8E554B55FA7105BFFA3E" box="[792,918,1426,1451]" pageId="3" pageNumber="56" value="2014-07-06">6.VII.2014</date>
</collectingDate>
, colls:
<location id="1D730C85423A8E554BA8FA71025DFA3E" LSID="urn:lsid:plazi:treatment:9005EB48423A8E5348DAFAE705EDFCAD:1D730C85423A8E554BA8FA71025DFA3E" box="[997,1140,1426,1451]" country="China" county="Wushan County" municipality="Luoping Town" name="Mingyi Tian" pageId="3" pageNumber="56" stateProvince="Chongqing">Mingyi Tian</location>
,
<location id="1D730C85423A8E554C32FA7102D6FA3E" LSID="urn:lsid:plazi:treatment:9005EB48423A8E5348DAFAE705EDFCAD:1D730C85423A8E554C32FA7102D6FA3E" box="[1151,1279,1426,1451]" country="China" county="Wushan County" municipality="Luoping Town" name="Weixin Liu" pageId="3" pageNumber="56" stateProvince="Chongqing">Weixin Liu</location>
,
<location id="1D730C85423A8E554D41FA7103B1FA3E" LSID="urn:lsid:plazi:treatment:9005EB48423A8E5348DAFAE705EDFCAD:1D730C85423A8E554D41FA7103B1FA3E" box="[1292,1432,1426,1451]" country="China" county="Wushan County" municipality="Luoping Town" name="Haomin Yin" pageId="3" pageNumber="56" stateProvince="Chongqing">Haomin Yin</location>
,
<location id="1D730C85423A8E5548DAFA550711FA5A" LSID="urn:lsid:plazi:treatment:9005EB48423A8E5348DAFAE705EDFCAD:1D730C85423A8E5548DAFA550711FA5A" box="[151,312,1462,1487]" country="China" county="Wushan County" municipality="Luoping Town" name="Sunbin Huang" pageId="3" pageNumber="56" stateProvince="Chongqing">Sunbin Huang</location>
and
<location id="1D730C85423A8E554923FA55042EFA5A" LSID="urn:lsid:plazi:treatment:9005EB48423A8E5348DAFAE705EDFCAD:1D730C85423A8E554923FA55042EFA5A" box="[366,519,1462,1487]" country="China" county="Wushan County" municipality="Luoping Town" name="Xinhui Wang" pageId="3" pageNumber="56" stateProvince="Chongqing">Xinhui Wang</location>
</materialsCitation>
.
<materialsCitation id="A8C45003423A8E554A5FFA54037AFA5A" ID-GBIF-Occurrence="1917355152" box="[530,1363,1462,1487]" collectingDate="2014-07-06" collectionCode="SCAU" country="China" county="Wushan County" location="Qinglong Village" municipality="Luoping Town" pageId="3" pageNumber="56" specimenCount="12" specimenCount-female="9" specimenCount-male="3" stateProvince="Chongqing" typeStatus="paratype">
<typeStatus id="C717E4FC423A8E554A5FFA5404AAFA5A" box="[530,643,1463,1487]" pageId="3" pageNumber="56" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="0EAA91D7423A8E554AC0FA5404CFFA5A" box="[653,742,1462,1487]" pageId="3" pageNumber="56" type="male">3 males</specimenCount>
,
<specimenCount id="0EAA91D7423A8E554ABFFA540577FA5A" box="[754,862,1462,1487]" pageId="3" pageNumber="56" type="female">9 females</specimenCount>
(
<collectionCode id="7EBDC29B423A8E554B21FA540590FA5A" box="[876,953,1463,1487]" pageId="3" pageNumber="56">SCAU</collectionCode>
eCQ2-2), same data as the
<typeStatus id="C717E4FC423A8E554CA0FA55037AFA5A" box="[1261,1363,1462,1487]" pageId="3" pageNumber="56" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="50B609D5423A8E55488AFA3906EAF97A" pageId="3" pageNumber="56" type="diagnosis">
<paragraph id="18135A5E423A8E55488AFA39029EF9CA" blockId="3.[151,1436,1390,2027]" pageId="3" pageNumber="56">
<emphasis id="2AD8864C423A8E55488AFA390713FA66" bold="true" box="[199,314,1498,1523]" pageId="3" pageNumber="56">Diagnosis</emphasis>
: Adult males of
<taxonomicName id="DFAC21DD423A8E5549BAFA3F0464FA66" authority="Liu &amp; Golovatch, 2018" authorityName="Liu &amp; Golovatch" authorityYear="2018" box="[503,589,1498,1523]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="latus" status="sp. nov.">
<emphasis id="2AD8864C423A8E5549BAFA3F0464FA66" box="[503,589,1498,1523]" italics="true" pageId="3" pageNumber="56">E. latus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31EB3B37423A8E554A19FA3904BAFA66" box="[596,659,1498,1523]" pageId="3" pageNumber="56" rank="species">
<emphasis id="2AD8864C423A8E554A19FA3904BAFA66" bold="true" box="[596,659,1498,1523]" pageId="3" pageNumber="56">n. sp.</emphasis>
</taxonomicNameLabel>
are distinct from other
<taxonomicName id="DFAC21DD423A8E554BD1FA390214FA66" authorityName="Attems" authorityYear="1901" box="[924,1085,1498,1523]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="2AD8864C423A8E554BD1FA390214FA66" box="[924,1085,1498,1523]" italics="true" pageId="3" pageNumber="56">Epanerchodus</emphasis>
</taxonomicName>
species based on the following combination of characters: (1) broad paraterga (widths of pro- and metazonae 1.5–1.6 and
<quantity id="DF54F7BB423A8E554CD9FA1D0334F982" box="[1172,1309,1534,1559]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="3.6" metricValueMin="3.5" pageId="3" pageNumber="56" unit="mm" value="3.55" valueMax="3.6" valueMin="3.5">3.5–3.6 mm</quantity>
) (
<figureCitation id="809746DB423A8E554D79FA1D0684F9AE" captionStart="FIGURE 1" captionStartId="4.[151,250,1782,1804]" captionTargetBox="[205,1388,198,1761]" captionTargetId="figure@4.[204,1394,195,1767]" captionTargetPageId="4" captionText="FIGURE 1. Epanerchodus latus n. sp., male paratype. A: anterior part of body, ventral view. B: anterior part of body, dorsal view. C: posterior part of body, dorsal view. D: gonopods in situ, ventral view. Scale bar: A–C, 0.2 mm; D, no scale." httpUri="https://zenodo.org/record/1458338/files/figure.png" pageId="3" pageNumber="56">Figs 1B– C</figureCitation>
); (2) gonopodal femorite with a finger-shaped process distally and a semi-circular process medially; (3) endomere tip unequally bifid; (4) endomere carrying a strongly curved subapical lobe (
<figureCitation id="809746DB423A8E554C28F9A50283F9CA" box="[1125,1194,1606,1631]" captionStart="FIGURE 2" captionStartId="5.[151,250,1837,1859]" captionTargetBox="[192,1395,905,1791]" captionTargetId="figure@5.[188,1398,877,1816]" captionTargetPageId="5" captionText="FIGURE 2. Epanerchodus latus n. sp., male paratype. A: right gonopod, lateral view. B: right gonopod, mesal view. Abbreviations: cl = clivus; en = endomere; l = lobe; p1 and p2 = processes 1 and 2; sg = seminal groove." httpUri="https://zenodo.org/record/1458340/files/figure.png" pageId="3" pageNumber="56">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="18135A5E423A8E55488AF98906EAF97A" blockId="3.[151,1436,1390,2027]" pageId="3" pageNumber="56">
The new species is similar to
<taxonomicName id="DFAC21DD423A8E554A61F98F04FDF916" authorityName="Chen &amp; Zhang" authorityYear="1990" box="[556,724,1642,1667]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="stylotarseus">
<emphasis id="2AD8864C423A8E554A61F98F04FDF916" box="[556,724,1642,1667]" italics="true" pageId="3" pageNumber="56">E. stylotarseus</emphasis>
</taxonomicName>
(from caves in
<collectingRegion id="DA6894BC423A8E554BD8F98905D1F916" box="[917,1016,1642,1667]" country="China" name="Guizhou" pageId="3" pageNumber="56">Guizhou</collectingRegion>
,
<collectingCountry id="60BB1ACE423A8E554C45F9890266F916" box="[1032,1103,1642,1667]" name="China" pageId="3" pageNumber="56">China</collectingCountry>
), but is distinguished by (1) caudolateral corners of paraterga being faintly acute-angled (
<figureCitation id="809746DB423A8E554B72F96D05BAF932" box="[831,915,1678,1703]" captionStart="FIGURE 1" captionStartId="4.[151,250,1782,1804]" captionTargetBox="[205,1388,198,1761]" captionTargetId="figure@4.[204,1394,195,1767]" captionTargetPageId="4" captionText="FIGURE 1. Epanerchodus latus n. sp., male paratype. A: anterior part of body, ventral view. B: anterior part of body, dorsal view. C: posterior part of body, dorsal view. D: gonopods in situ, ventral view. Scale bar: A–C, 0.2 mm; D, no scale." httpUri="https://zenodo.org/record/1458338/files/figure.png" pageId="3" pageNumber="56">Fig. 1B</figureCitation>
)
<emphasis id="2AD8864C423A8E554BEEF96C05EBF933" box="[931,962,1679,1702]" italics="true" pageId="3" pageNumber="56">vs.</emphasis>
strongly triangular in
<taxonomicName id="DFAC21DD423A8E554CF6F96C0349F932" authorityName="Chen &amp; Zhang" authorityYear="1990" box="[1211,1376,1678,1703]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="stylotarseus">
<emphasis id="2AD8864C423A8E554CF6F96C0349F932" box="[1211,1376,1678,1703]" italics="true" pageId="3" pageNumber="56">E. stylotarseus</emphasis>
</taxonomicName>
(
<figureCitation id="809746DB423A8E554D23F96D069EF95E" captionStart="FIGURE 22" captionStartId="24.[151,250,1956,1978]" captionTargetBox="[337,1255,1139,1944]" captionTargetId="figure@24.[331,1262,1138,1947]" captionTargetPageId="24" captionText="FIGURE 22. Epanerchodus stylotarseus Chen and Zhang, 1990, male from Biyun Dong Cave. A: habitus, subdorsal view. B: habitus, sublateral view. Scale bar: 0.5 mm." httpUri="https://zenodo.org/record/1458380/files/figure.png" pageId="3" pageNumber="56">Fig. 22</figureCitation>
); (2) endomere ribbon-shaped, strongly curved (
<figureCitation id="809746DB423A8E554AAFF9510501F95E" box="[738,808,1714,1739]" captionStart="FIGURE 2" captionStartId="5.[151,250,1837,1859]" captionTargetBox="[192,1395,905,1791]" captionTargetId="figure@5.[188,1398,877,1816]" captionTargetPageId="5" captionText="FIGURE 2. Epanerchodus latus n. sp., male paratype. A: right gonopod, lateral view. B: right gonopod, mesal view. Abbreviations: cl = clivus; en = endomere; l = lobe; p1 and p2 = processes 1 and 2; sg = seminal groove." httpUri="https://zenodo.org/record/1458340/files/figure.png" pageId="3" pageNumber="56">Fig. 2</figureCitation>
)
<emphasis id="2AD8864C423A8E554B77F9570571F95E" box="[826,856,1716,1739]" italics="true" pageId="3" pageNumber="56">vs.</emphasis>
subspiniform and suberect in
<taxonomicName id="DFAC21DD423A8E554CFBF9570374F95E" authorityName="Chen &amp; Zhang" authorityYear="1990" box="[1206,1373,1714,1739]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="stylotarseus">
<emphasis id="2AD8864C423A8E554CFBF9570374F95E" box="[1206,1373,1714,1739]" italics="true" pageId="3" pageNumber="56">E. stylotarseus</emphasis>
</taxonomicName>
(
<figureCitation id="809746DB423A8E554D23F951069EF97A" captionStart="FIGURE 23" captionStartId="25.[151,250,1572,1594]" captionTargetBox="[282,1318,709,1553]" captionTargetId="figure@25.[272,1324,694,1561]" captionTargetPageId="25" captionText="FIGURE 23. Epanerchodus stylotarseus Chen and Zhang, 1990, male from Biyun Dong Cave. A: left gonopod, mesal view. B: left gonopod, lateral view. Abbreviation: en = endomere." httpUri="https://zenodo.org/record/1458382/files/figure.png" pageId="3" pageNumber="56">Fig. 23</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="50B609D5423A8E53488AF9190722FC80" lastPageId="5" lastPageNumber="58" pageId="3" pageNumber="56" type="description">
<paragraph id="18135A5E423A8E53488AF9190764FD58" blockId="3.[151,1436,1390,2027]" lastBlockId="5.[151,1437,151,824]" lastPageId="5" lastPageNumber="58" pageId="3" pageNumber="56">
<emphasis id="2AD8864C423A8E55488AF9190778F886" bold="true" box="[199,337,1786,1811]" pageId="3" pageNumber="56">Description</emphasis>
: Based on
<typeStatus id="C717E4FC423A8E55499FF918042BF886" box="[466,514,1787,1811]" pageId="3" pageNumber="56">type</typeStatus>
specimens. Lengths of body
<emphasis id="2AD8864C423A8E554B02F91F0543F886" box="[847,874,1788,1811]" italics="true" pageId="3" pageNumber="56">ca</emphasis>
<quantity id="DF54F7BB423A8E554B3FF91905C2F887" box="[882,1003,1786,1811]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="3" pageNumber="56" unit="mm" value="24.0" valueMax="25.0" valueMin="23.0">23–25 mm</quantity>
(males),
<quantity id="DF54F7BB423A8E554C1BF91802F9F887" box="[1110,1232,1787,1811]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="2.8" metricValueMin="2.7" pageId="3" pageNumber="56" unit="mm" value="27.5" valueMax="28.0" valueMin="27.0">27–28 mm</quantity>
(females); widths of pro- and metazonae 1.5–1.6 and
<quantity id="DF54F7BB423A8E554A6CF8FD0481F8A3" box="[545,680,1822,1847]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="3.6" metricValueMin="3.5" pageId="3" pageNumber="56" unit="mm" value="3.55" valueMax="3.6" valueMin="3.5">3.5–3.6 mm</quantity>
(males), 2.0–2.2 and 4.0–
<quantity id="DF54F7BB423A8E554B86F8FC0209F8A3" box="[971,1056,1823,1847]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="3" pageNumber="56" unit="mm" value="4.2">4.2 mm</quantity>
(females).
<emphasis id="2AD8864C423A8E554CD0F8FD0337F8A2" bold="true" box="[1181,1310,1822,1847]" pageId="3" pageNumber="56">Coloration</emphasis>
: in alcohol nearly pallid. Only mouthparts light grey-brown and gonopodal telopodites yellowish (
<figureCitation id="809746DB423A8E554C23F8A10299F8CE" box="[1134,1200,1858,1883]" captionStart="FIGURE 1" captionStartId="4.[151,250,1782,1804]" captionTargetBox="[205,1388,198,1761]" captionTargetId="figure@4.[204,1394,195,1767]" captionTargetPageId="4" captionText="FIGURE 1. Epanerchodus latus n. sp., male paratype. A: anterior part of body, ventral view. B: anterior part of body, dorsal view. C: posterior part of body, dorsal view. D: gonopods in situ, ventral view. Scale bar: A–C, 0.2 mm; D, no scale." httpUri="https://zenodo.org/record/1458338/files/figure.png" pageId="3" pageNumber="56">Fig. 1</figureCitation>
).
<emphasis id="2AD8864C423A8E554C84F8A1032EF8CE" bold="true" box="[1225,1287,1858,1883]" pageId="3" pageNumber="56">Body</emphasis>
: Adults with 20 rings. Width: head &lt;&lt; collum &lt;ring 2 &lt;3 &lt;4 &lt;5–13, thereafter (rings 14–19) body gradually tapering posteriorly towards telson.
<emphasis id="2AD8864C423A8E554986F8690423F836" bold="true" box="[459,522,1930,1955]" pageId="3" pageNumber="56">Head</emphasis>
: only clypeus densely pilose, vertex smooth, epicranial suture conspicuous (
<figureCitation id="809746DB423A8E554D22F8690692F852" captionStart="FIGURE 1" captionStartId="4.[151,250,1782,1804]" captionTargetBox="[205,1388,198,1761]" captionTargetId="figure@4.[204,1394,195,1767]" captionTargetPageId="4" captionText="FIGURE 1. Epanerchodus latus n. sp., male paratype. A: anterior part of body, ventral view. B: anterior part of body, dorsal view. C: posterior part of body, dorsal view. D: gonopods in situ, ventral view. Scale bar: A–C, 0.2 mm; D, no scale." httpUri="https://zenodo.org/record/1458338/files/figure.png" pageId="3" pageNumber="56">Fig. 1A</figureCitation>
). Antennae long, reaching past posterior margin of ring 3 when extended posteriorly, slightly clavate (
<figureCitation id="809746DB423A8E554D7EF84D0685F87E" captionStart="FIGURE 1" captionStartId="4.[151,250,1782,1804]" captionTargetBox="[205,1388,198,1761]" captionTargetId="figure@4.[204,1394,195,1767]" captionTargetPageId="4" captionText="FIGURE 1. Epanerchodus latus n. sp., male paratype. A: anterior part of body, ventral view. B: anterior part of body, dorsal view. C: posterior part of body, dorsal view. D: gonopods in situ, ventral view. Scale bar: A–C, 0.2 mm; D, no scale." httpUri="https://zenodo.org/record/1458338/files/figure.png" pageId="3" pageNumber="56">Figs 1A– B</figureCitation>
).
<emphasis id="2AD8864C423A8E55488CF831077BF87E" bold="true" box="[193,338,2002,2027]" pageId="3" pageNumber="56">Exoskeleton</emphasis>
: Collum with subrectangular caudolateral corners and a faint lateral incision/denticle on each side. Paraterga broad (
<figureCitation id="809746DB423D8E5249D9F8830438F8ED" box="[404,529,1888,1913]" captionStart="FIGURE 1" captionStartId="4.[151,250,1782,1804]" captionTargetBox="[205,1388,198,1761]" captionTargetId="figure@4.[204,1394,195,1767]" captionTargetPageId="4" captionText="FIGURE 1. Epanerchodus latus n. sp., male paratype. A: anterior part of body, ventral view. B: anterior part of body, dorsal view. C: posterior part of body, dorsal view. D: gonopods in situ, ventral view. Scale bar: A–C, 0.2 mm; D, no scale." httpUri="https://zenodo.org/record/1458338/files/figure.png" pageId="4" pageNumber="57">Figs 1A–E</figureCitation>
), midbody paraterga extend metatergite to
<emphasis id="2AD8864C423D8E524BB5F882023AF8ED" box="[1016,1043,1889,1912]" italics="true" pageId="4" pageNumber="57">ca</emphasis>
2.3x width of prozonite. Paraterga 2–5 clearly upturned dorsally above a faintly convex dorsum, other paraterga flat (
<figureCitation id="809746DB423D8E524C66F86702A8F809" box="[1067,1153,1924,1949]" captionStart="FIGURE 1" captionStartId="4.[151,250,1782,1804]" captionTargetBox="[205,1388,198,1761]" captionTargetId="figure@4.[204,1394,195,1767]" captionTargetPageId="4" captionText="FIGURE 1. Epanerchodus latus n. sp., male paratype. A: anterior part of body, ventral view. B: anterior part of body, dorsal view. C: posterior part of body, dorsal view. D: gonopods in situ, ventral view. Scale bar: A–C, 0.2 mm; D, no scale." httpUri="https://zenodo.org/record/1458338/files/figure.png" pageId="4" pageNumber="57">Fig. 1B</figureCitation>
). Caudolateral corners of paraterga posterior to collum increasingly acute-angled and projecting posteriorly past tergal margin, especially acute on rings 17–19 (
<figureCitation id="809746DB423D8E5249D1F82F0435F871" box="[412,540,1996,2021]" captionStart="FIGURE 1" captionStartId="4.[151,250,1782,1804]" captionTargetBox="[205,1388,198,1761]" captionTargetId="figure@4.[204,1394,195,1767]" captionTargetPageId="4" captionText="FIGURE 1. Epanerchodus latus n. sp., male paratype. A: anterior part of body, ventral view. B: anterior part of body, dorsal view. C: posterior part of body, dorsal view. D: gonopods in situ, ventral view. Scale bar: A–C, 0.2 mm; D, no scale." httpUri="https://zenodo.org/record/1458338/files/figure.png" pageId="4" pageNumber="57">Figs 1B–C</figureCitation>
). Integument shining (
<figureCitation id="809746DB423D8E524B6CF82F054FF871" box="[801,870,1996,2021]" captionStart="FIGURE 1" captionStartId="4.[151,250,1782,1804]" captionTargetBox="[205,1388,198,1761]" captionTargetId="figure@4.[204,1394,195,1767]" captionTargetPageId="4" captionText="FIGURE 1. Epanerchodus latus n. sp., male paratype. A: anterior part of body, ventral view. B: anterior part of body, dorsal view. C: posterior part of body, dorsal view. D: gonopods in situ, ventral view. Scale bar: A–C, 0.2 mm; D, no scale." httpUri="https://zenodo.org/record/1458338/files/figure.png" pageId="4" pageNumber="57">Fig. 1</figureCitation>
), translucent, prozonae very delicately alveolate. Limbus regularly denticulate. Constriction between pro- and metazonae narrow, shallow and smooth (
<figureCitation id="809746DB423C8E534D79FF7403A7FF25" box="[1332,1422,151,176]" captionStart="FIGURE 1" captionStartId="4.[151,250,1782,1804]" captionTargetBox="[205,1388,198,1761]" captionTargetId="figure@4.[204,1394,195,1767]" captionTargetPageId="4" captionText="FIGURE 1. Epanerchodus latus n. sp., male paratype. A: anterior part of body, ventral view. B: anterior part of body, dorsal view. C: posterior part of body, dorsal view. D: gonopods in situ, ventral view. Scale bar: A–C, 0.2 mm; D, no scale." httpUri="https://zenodo.org/record/1458338/files/figure.png" pageId="5" pageNumber="58">Fig. 1B</figureCitation>
). Metatergal sculpture faint, with three irregular transverse rows of setigerous polygonal bosses. Sulcus between front and middle rows of setae a little deeper than that between middle and caudal rows (
<figureCitation id="809746DB423C8E534C3EFF3C02E0FF6D" box="[1139,1225,223,248]" captionStart="FIGURE 1" captionStartId="4.[151,250,1782,1804]" captionTargetBox="[205,1388,198,1761]" captionTargetId="figure@4.[204,1394,195,1767]" captionTargetPageId="4" captionText="FIGURE 1. Epanerchodus latus n. sp., male paratype. A: anterior part of body, ventral view. B: anterior part of body, dorsal view. C: posterior part of body, dorsal view. D: gonopods in situ, ventral view. Scale bar: A–C, 0.2 mm; D, no scale." httpUri="https://zenodo.org/record/1458338/files/figure.png" pageId="5" pageNumber="58">Fig. 1B</figureCitation>
). Tergal setae very short, sometimes present only on collum and rings 18–19. Three or four faint setigerous incisions at lateral margins of poreless and pore-bearing rings, respectively (
<figureCitation id="809746DB423C8E534A83FEC40566FED5" box="[718,847,295,320]" captionStart="FIGURE 1" captionStartId="4.[151,250,1782,1804]" captionTargetBox="[205,1388,198,1761]" captionTargetId="figure@4.[204,1394,195,1767]" captionTargetPageId="4" captionText="FIGURE 1. Epanerchodus latus n. sp., male paratype. A: anterior part of body, ventral view. B: anterior part of body, dorsal view. C: posterior part of body, dorsal view. D: gonopods in situ, ventral view. Scale bar: A–C, 0.2 mm; D, no scale." httpUri="https://zenodo.org/record/1458338/files/figure.png" pageId="5" pageNumber="58">Figs 1B–C</figureCitation>
). Pore formula normal: 5, 7, 9, 10, 12, 13, 15–19, ozopores evident, dorsal, clearly set off from lateral margin and located between last and penultimate marginal incisions. Epiproct short, conical, pre-apical lateral papillae small. Hypoproct subtrapeziform, with two setigerous papillae. Pleurosternal carinae absent. Sterna sparsely setose, cross-shaped impressions (both axial and transverse) shallow (
<figureCitation id="809746DB423C8E53494CFE540776FE45" box="[257,351,439,464]" captionStart="FIGURE 1" captionStartId="4.[151,250,1782,1804]" captionTargetBox="[205,1388,198,1761]" captionTargetId="figure@4.[204,1394,195,1767]" captionTargetPageId="4" captionText="FIGURE 1. Epanerchodus latus n. sp., male paratype. A: anterior part of body, ventral view. B: anterior part of body, dorsal view. C: posterior part of body, dorsal view. D: gonopods in situ, ventral view. Scale bar: A–C, 0.2 mm; D, no scale." httpUri="https://zenodo.org/record/1458338/files/figure.png" pageId="5" pageNumber="58">Fig. 1A</figureCitation>
). Legs long and slender, about 2.0 times as long as body ring height in both sexes, without sphaerotrichomes or sternal cones, prefemora not bulging laterally (
<figureCitation id="809746DB423C8E534BD1FE3F05DEFE61" box="[924,1015,476,501]" captionStart="FIGURE 1" captionStartId="4.[151,250,1782,1804]" captionTargetBox="[205,1388,198,1761]" captionTargetId="figure@4.[204,1394,195,1767]" captionTargetPageId="4" captionText="FIGURE 1. Epanerchodus latus n. sp., male paratype. A: anterior part of body, ventral view. B: anterior part of body, dorsal view. C: posterior part of body, dorsal view. D: gonopods in situ, ventral view. Scale bar: A–C, 0.2 mm; D, no scale." httpUri="https://zenodo.org/record/1458338/files/figure.png" pageId="5" pageNumber="58">Fig. 1A</figureCitation>
).
<emphasis id="2AD8864C423C8E534C40FE3F02AFFE60" bold="true" box="[1037,1158,476,501]" pageId="5" pageNumber="58">Gonopods</emphasis>
: (
<figureCitation id="809746DB423C8E534CD2FE3F02D4FE61" box="[1183,1277,476,501]" captionStart="FIGURE 1" captionStartId="4.[151,250,1782,1804]" captionTargetBox="[205,1388,198,1761]" captionTargetId="figure@4.[204,1394,195,1767]" captionTargetPageId="4" captionText="FIGURE 1. Epanerchodus latus n. sp., male paratype. A: anterior part of body, ventral view. B: anterior part of body, dorsal view. C: posterior part of body, dorsal view. D: gonopods in situ, ventral view. Scale bar: A–C, 0.2 mm; D, no scale." httpUri="https://zenodo.org/record/1458338/files/figure.png" pageId="5" pageNumber="58">Figs 1D</figureCitation>
,
<figureCitation id="809746DB423C8E534D41FE3F0379FE61" box="[1292,1360,476,500]" captionStart="FIGURE 2" captionStartId="5.[151,250,1837,1859]" captionTargetBox="[192,1395,905,1791]" captionTargetId="figure@5.[188,1398,877,1816]" captionTargetPageId="5" captionText="FIGURE 2. Epanerchodus latus n. sp., male paratype. A: right gonopod, lateral view. B: right gonopod, mesal view. Abbreviations: cl = clivus; en = endomere; l = lobe; p1 and p2 = processes 1 and 2; sg = seminal groove." httpUri="https://zenodo.org/record/1458340/files/figure.png" pageId="5" pageNumber="58">2A–B</figureCitation>
) Coxa large and squarish, as usual. Prefemur densely setose and nearly half the length of telopodite. Clivus (
<emphasis id="2AD8864C423C8E534D19FE1C0341FD8D" bold="true" box="[1364,1384,511,536]" pageId="5" pageNumber="58">cl</emphasis>
), or distolateral ridge on femorite, prominent; femorite with a curved finger-shaped process (
<emphasis id="2AD8864C423C8E534C30FDC702B2FDA8" bold="true" box="[1149,1179,548,573]" pageId="5" pageNumber="58">p1</emphasis>
) distally and a smaller semi-circular process (
<emphasis id="2AD8864C423C8E5349D4FDA4079EFDF5" bold="true" box="[409,439,583,608]" pageId="5" pageNumber="58">p2</emphasis>
) medially. Endomere (
<emphasis id="2AD8864C423C8E534AF3FDA404F3FDF5" bold="true" box="[702,730,583,608]" pageId="5" pageNumber="58">en</emphasis>
) strongly curved, tip unequally bifid, with a small, subapical, curved lobe (
<emphasis id="2AD8864C423C8E534978FD8F0714FD10" bold="true" box="[309,317,620,645]" pageId="5" pageNumber="58">l</emphasis>
). Seminal groove (
<emphasis id="2AD8864C423C8E534A69FD8F0414FD10" bold="true" box="[548,573,620,645]" pageId="5" pageNumber="58">sg</emphasis>
) starting mesally, distally recurved laterad near base of
<emphasis id="2AD8864C423C8E534C93FD8F02D5FD10" bold="true" box="[1246,1276,620,645]" pageId="5" pageNumber="58">p1</emphasis>
, then turning laterobasad to run into an accessory seminal chamber, the latter opening on a hairy pulvillus on lateral face. An exomere absent.
</paragraph>
<caption id="4CD30AD6423D8E5248DAF9150318F8A5" httpUri="https://zenodo.org/record/1458338/files/figure.png" pageId="4" pageNumber="57" startId="4.[151,250,1782,1804]" targetBox="[205,1388,198,1761]" targetPageId="4">
<paragraph id="18135A5E423D8E5248DAF9150318F8A5" blockId="4.[151,1436,1782,1840]" pageId="4" pageNumber="57">
<emphasis id="2AD8864C423D8E5248DAF915073DF899" bold="true" box="[151,276,1782,1804]" pageId="4" pageNumber="57">FIGURE 1.</emphasis>
<taxonomicName id="DFAC21DD423D8E524956F91507CCF899" authority="Liu &amp; Golovatch, 2018" authorityName="Liu &amp; Golovatch" authorityYear="2018" box="[283,485,1782,1804]" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="latus" status="sp. nov.">
<emphasis id="2AD8864C423D8E524956F91507CCF899" box="[283,485,1782,1804]" italics="true" pageId="4" pageNumber="57">Epanerchodus latus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31EB3B37423D8E5249A1F915040EF899" box="[492,551,1782,1804]" pageId="4" pageNumber="57" rank="species">
<emphasis id="2AD8864C423D8E5249A1F915040EF899" bold="true" box="[492,551,1782,1804]" pageId="4" pageNumber="57">n. sp.</emphasis>
</taxonomicNameLabel>
, male paratype.
<emphasis id="2AD8864C423D8E524A9FF91504CDF899" bold="true" box="[722,740,1782,1804]" pageId="4" pageNumber="57">A</emphasis>
: anterior part of body, ventral view.
<emphasis id="2AD8864C423D8E524C13F9150246F899" bold="true" box="[1118,1135,1782,1804]" pageId="4" pageNumber="57">B</emphasis>
: anterior part of body, dorsal view.
<emphasis id="2AD8864C423D8E52489EF8F906CCF8A5" bold="true" box="[211,229,1818,1840]" pageId="4" pageNumber="57">C</emphasis>
: posterior part of body, dorsal view.
<emphasis id="2AD8864C423D8E524A17F8F90445F8A5" bold="true" box="[602,620,1818,1840]" pageId="4" pageNumber="57">D</emphasis>
: gonopods
<emphasis id="2AD8864C423D8E524AAEF8F80509F8A5" box="[739,800,1819,1840]" italics="true" pageId="4" pageNumber="57">in situ</emphasis>
, ventral view. Scale bar: A–C, 0.2 mm; D, no scale.
</paragraph>
</caption>
<paragraph id="18135A5E423C8E53488AFD340722FC80" blockId="5.[151,1437,151,824]" pageId="5" pageNumber="58">
<emphasis id="2AD8864C423C8E53488AFD340723FD65" bold="true" box="[199,266,727,752]" pageId="5" pageNumber="58">Notes</emphasis>
: Based on the long slender antennae and legs, and a depigmented cuticle, the species is most likely a troglobite.
</paragraph>
</subSubSection>
<subSubSection id="50B609D5423C8E53488AFCFC05EDFCAD" box="[199,964,799,824]" pageId="5" pageNumber="58" type="etymology">
<paragraph id="18135A5E423C8E53488AFCFC05EDFCAD" blockId="5.[151,1437,151,824]" box="[199,964,799,824]" pageId="5" pageNumber="58">
<emphasis id="2AD8864C423C8E53488AFCFC0761FCAD" bold="true" box="[199,328,799,824]" pageId="5" pageNumber="58">Etymology</emphasis>
: To emphasize the broad paraterga; masculine adjective.
</paragraph>
</subSubSection>
</treatment>
</document>
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