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<mods:title id="07049C5FD4F76663133A49291811C024">The genus Dicrania LePeletier &amp; Audinet-Serville, 1828 (Coleoptera: Scarabaeidae: Melolonthinae: Macrodactylini) from Colombia with the description of a new species</mods:title>
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Clavijo-Bustos, Neita Moreno, &amp; Fuhrmann
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<figureCitation id="E69507A3FFB50102FEF0A6F0FE0071A5" box="[347,389,703,729]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="4.[151,250,1618,1642]" captionStartId-1="5.[151,250,1400,1424]" captionTargetBox-0="[156,1436,202,1592]" captionTargetBox-1="[151,1437,181,1376]" captionTargetId-0="figure-21@4.[151,1436,181,1594]" captionTargetId-1="figure-24@5.[151,1437,181,1376]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 2. Dicrania chimu, new species, male holotype. A, Dorsal view; B, lateral view; C, ventral view; D, labels; E, head and pronotum in dorsal view; F, aedeagus in lateral view; G, aedeagus in distal view; H, pygidium in caudal view; I, antenna in dorsal view." captionText-1="FIGURE 3. Dicrania chimu, new species, male paratype. A, Protarsal claw; B, mesotarsal claw; C, metatarsal claw; D, apex of protibial; E, apex of mesotibia; F, apex of metatibia; G, mesotarsomere 1 in ventral view; H, mesotarsomere 1 in lateral internal view." figureDoi-0="http://doi.org/10.5281/zenodo.11548017" figureDoi-1="http://doi.org/10.5281/zenodo.11548019" httpUri-0="https://zenodo.org/record/11548017/files/figure.png" httpUri-1="https://zenodo.org/record/11548019/files/figure.png" pageId="2" pageNumber="574">2–3</figureCitation>
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/ Clavijo-Bustos, Neita Moreno &amp; Fuhrmann, / 2024” [red printed label] // “ICN 034823” [white printed label with barcode].
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ICN 035725 and 
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. 
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<elevation id="F583FC15FFB50102FDC2A7F5FD3170A9" box="[617,692,954,981]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="574" unit="m" value="450.0">450 m</elevation>
</quantity>
, 
<date id="0A103DE6FFB50102FD69A7F5FCB770A9" box="[706,818,954,981]" pageId="2" pageNumber="574" value="2005-05-27">
<collectingDate id="1A54C40EFFB50102FD69A7F5FCB770A9" box="[706,818,954,981]" pageId="2" pageNumber="574" value="2005-05-27">27.v.2005</collectingDate>
</date>
, 
<collectorName id="D35B7EF0FFB50102FCEBA7F4FC2170A9" box="[832,932,955,981]" pageId="2" pageNumber="574">J. Castro</collectorName>
(
<specimenCount id="68A8D0AFFFB50102FC1CA7F4FC5F70A9" box="[951,986,955,981]" count="1" pageId="2" pageNumber="574" type="male">1♂</specimenCount>
UnAB 
<quantity id="B956B6C3FFB50102FB9FA7F5FB0E70A8" box="[1076,1163,954,980]" metricMagnitude="2" metricUnit="m" metricValue="1.251204" pageId="2" pageNumber="574" unit="in" value="4926.0">4926 in</quantity>
<collectionCode id="18BF83E3FFB50102FB3FA7F4FB6D70A9" box="[1172,1256,955,981]" country="Colombia" httpUri="http://grbio.org/cool/hx5r-ym26" name="Universidad Nacional, Facultad de Agronomia" pageId="2" pageNumber="574">UNAB</collectionCode>
);
</materialsCitation>
<materialsCitation id="CEC6117BFFB50102FB56A7F4FEFA7761" ID-GBIF-Occurrence="4880078307" collectingDate="2011-04-29" collectingDateMax="2011-04-29" collectingDateMin="1996-05-10" collectionCode="MPUJ" collectorName="D. Bohorquez &amp; C. Chaparro &amp; S. Herrera" country="Colombia" elevation="240" latitude="4.100278" location="Remolinos" longLatPrecision="21" longitude="-72.950554" municipality="Puerto Lopez" pageId="2" pageNumber="574" specimenCode="ICN 034824, 035726" specimenCount="1" specimenCount-male="1" stateProvince="Meta" typeStatus="paratype">
<collectingMunicipality id="9E75815CFFB50102FB56A7F4FA1D70A9" box="[1277,1432,955,981]" pageId="2" pageNumber="574">Puerto López</collectingMunicipality>
, 
<location id="7B714DFDFFB50102FF3CA791FE967084" LSID="urn:lsid:plazi:treatment:F607AA30FFB50103FF3CA6D5FF557410:7B714DFDFFB50102FF3CA791FE967084" box="[151,275,990,1016]" country="Colombia" latitude="4.100278" longLatPrecision="21" longitude="-72.950554" municipality="Puerto Lopez" name="Remolinos" pageId="2" pageNumber="574" stateProvince="Meta">Remolinos</location>
, 
<location id="7B714DFDFFB50102FE8AA790FE407084" LSID="urn:lsid:plazi:treatment:F607AA30FFB50103FF3CA6D5FF557410:7B714DFDFFB50102FE8AA790FE407084" box="[289,453,990,1017]" country="Colombia" latitude="4.100278" longLatPrecision="21" longitude="-72.950554" municipality="Puerto Lopez" name="Centro Cafam" pageId="2" pageNumber="574" stateProvince="Meta">Centro Cafam</location>
, 
<location id="7B714DFDFFB50102FE79A791FD077084" LSID="urn:lsid:plazi:treatment:F607AA30FFB50103FF3CA6D5FF557410:7B714DFDFFB50102FE79A791FD077084" box="[466,642,990,1016]" country="Colombia" latitude="4.100278" longLatPrecision="21" longitude="-72.950554" municipality="Puerto Lopez" name="Piedra Candela" pageId="2" pageNumber="574" stateProvince="Meta">Piedra Candela</location>
, 
<quantity id="B956B6C3FFB50102FD24A790FD5F7085" box="[655,730,991,1017]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="574" unit="m" value="240.0">
<elevation id="F583FC15FFB50102FD24A790FD5F7085" box="[655,730,991,1017]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="574" unit="m" value="240.0">240 m</elevation>
</quantity>
, 
<date id="0A103DE6FFB50102FD43A791FCE57084" box="[744,864,990,1016]" pageId="2" pageNumber="574" value="2011-04-29">
<collectingDate id="1A54C40EFFB50102FD43A791FCE57084" box="[744,864,990,1016]" pageId="2" pageNumber="574" value="2011-04-29">29.iv.2011</collectingDate>
</date>
, Ad hoc diurna, 
<collectorName id="D35B7EF0FFB50102FB89A790FB417084" box="[1058,1220,990,1017]" pageId="2" pageNumber="574">D. Bohórquez</collectorName>
, 
<collectorName id="D35B7EF0FFB50102FB79A790FAD87084" box="[1234,1373,990,1017]" pageId="2" pageNumber="574">C. Chaparro</collectorName>
&amp; 
<collectorName id="D35B7EF0FFB50102FA2DA790FF697761" pageId="2" pageNumber="574">S. Herrera</collectorName>
(
<specimenCount id="68A8D0AFFFB50102FF54A04CFEA77761" box="[255,290,1027,1053]" count="1" pageId="2" pageNumber="574" type="male">1♂</specimenCount>
<collectionCode id="18BF83E3FFB50102FE80A04CFEFD7761" box="[299,376,1027,1053]" pageId="2" pageNumber="574">MPUJ</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="CEC6117BFFB50102FE20A04CFDB5773D" ID-GBIF-Occurrence="4880078309" country="Colombia" pageId="2" pageNumber="574" specimenCount="1" specimenCount-male="1" stateProvince="Meta" typeStatus="paratype">
<typeStatus id="A115A584FFB50102FE20A04CFE7D7761" box="[395,504,1027,1053]" pageId="2" pageNumber="574" type="paratype">Paratypes</typeStatus>
with yellow printed label “ 
<typeStatus id="A115A584FFB50102FC99A04CFC267761" box="[818,931,1027,1053]" pageId="2" pageNumber="574" type="paratype">Paratypus</typeStatus>
<specimenCount id="68A8D0AFFFB50102FC07A04CFC447761" box="[940,961,1027,1053]" count="1" pageId="2" pageNumber="574" type="male">♂</specimenCount>
/ 
<taxonomicName id="B9AE60A5FFB50102FC70A04CFB087760" authority="Clavijo-Bustos &amp; Moreno &amp; Fuhrmann, 2024" authorityName="Clavijo-Bustos &amp; Moreno &amp; Fuhrmann" authorityYear="2024" box="[987,1165,1026,1052]" class="Insecta" family="Scarabaeidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="574" phylum="Arthropoda" rank="species" species="chimu" status="sp. nov.">
<emphasis id="4CDAC734FFB50102FC70A04CFB087760" box="[987,1165,1026,1052]" italics="true" pageId="2" pageNumber="574">Dicrania chimu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="57E97A4FFFB50102FB3DA04CFB517761" box="[1174,1236,1027,1053]" pageId="2" pageNumber="574" rank="species">n. sp.</taxonomicNameLabel>
/ Clavijo-Bustos, Neita Moreno &amp; Fuhrmann, / 2024”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="36B448ADFFB50102FF6CA005FAC577AD" pageId="2" pageNumber="574" type="diagnosis">
<paragraph id="7E111B26FFB50102FF6CA005FAC577AD" blockId="2.[151,1437,774,1917]" pageId="2" pageNumber="574">
<emphasis id="4CDAC734FFB50102FF6CA005FEC77718" bold="true" box="[199,322,1098,1124]" pageId="2" pageNumber="574">Diagnosis.</emphasis>
Antenna with nine antennomeres; eyes in dorsal view large and head about 1.8 times wider than the narrowest interocular distance; pronotum with lateral areas without punctures or setae; elytra with puncture indistinct, disc glabrous; protarsomere 1 with a truncate inner tooth; protarsomere 2 longer than wide; mesotarsomere 1 with an inner spatulate tooth; metatarsomere 2 with sparse setae, medial setae shorter than the length of tarsomere.
</paragraph>
</subSubSection>
<subSubSection id="36B448ADFFB50103FF6CA095FA977530" lastPageId="3" lastPageNumber="575" pageId="2" pageNumber="574" type="description">
<paragraph id="7E111B26FFB50102FF6CA095FEE57400" blockId="2.[151,1437,774,1917]" pageId="2" pageNumber="574">
<emphasis id="4CDAC734FFB50102FF6CA095FEDC7788" bold="true" box="[199,345,1242,1268]" pageId="2" pageNumber="574">Description.</emphasis>
Male 
<typeStatus id="A115A584FFB50102FE34A095FD847788" box="[415,513,1242,1268]" pageId="2" pageNumber="574" type="holotype">holotype</typeStatus>
(
<figureCitation id="E69507A3FFB50102FDBBA095FDD07789" box="[528,597,1242,1269]" captionStart="FIGURE 2" captionStartId="4.[151,250,1618,1642]" captionTargetBox="[156,1436,202,1592]" captionTargetId="figure-21@4.[151,1436,181,1594]" captionTargetPageId="4" captionText="FIGURE 2. Dicrania chimu, new species, male holotype. A, Dorsal view; B, lateral view; C, ventral view; D, labels; E, head and pronotum in dorsal view; F, aedeagus in lateral view; G, aedeagus in distal view; H, pygidium in caudal view; I, antenna in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11548017" httpUri="https://zenodo.org/record/11548017/files/figure.png" pageId="2" pageNumber="574">Fig. 2</figureCitation>
). Body length: 
<quantity id="B956B6C3FFB50102FCAAA094FCE27789" box="[769,871,1243,1269]" metricMagnitude="-2" metricUnit="m" metricValue="1.49" pageId="2" pageNumber="574" unit="mm" value="14.9">14.9 mm</quantity>
; Body maximum width (at half of elytra): 
<quantity id="B956B6C3FFB50102FAEBA095FA1C7789" box="[1344,1433,1242,1269]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="2" pageNumber="574" unit="mm" value="7.6">7.6 mm</quantity>
. 
<emphasis id="4CDAC734FFB50102FF3CA0B1FF5D7664" box="[151,216,1278,1304]" italics="true" pageId="2" pageNumber="574">Color</emphasis>
: Entirely black. Setae yellowish brown, disc of pronotum with setae dark brown. 
<emphasis id="4CDAC734FFB50102FBCAA0B1FB1B7664" box="[1121,1182,1278,1304]" italics="true" pageId="2" pageNumber="574">Head</emphasis>
(
<figureCitation id="E69507A3FFB50102FB06A0B1FA9D7665" box="[1197,1304,1278,1305]" captionStart="FIGURE 2" captionStartId="4.[151,250,1618,1642]" captionTargetBox="[156,1436,202,1592]" captionTargetId="figure-21@4.[151,1436,181,1594]" captionTargetPageId="4" captionText="FIGURE 2. Dicrania chimu, new species, male holotype. A, Dorsal view; B, lateral view; C, ventral view; D, labels; E, head and pronotum in dorsal view; F, aedeagus in lateral view; G, aedeagus in distal view; H, pygidium in caudal view; I, antenna in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11548017" httpUri="https://zenodo.org/record/11548017/files/figure.png" pageId="2" pageNumber="574">Fig. 2E, I</figureCitation>
): Large and pronotum about 1.8 times wider than head, eyes large and head about 1.8 times wider than the narrowest interocular distance, frontoclypeal suture slightly sinuous. Clypeus trapezoid, sparsely punctate than frons; anterior angles obtuse and deflected. Antenna with nine antennomeres, antennal club about 4.0 times longer than wide and with three lamellae. 
<emphasis id="4CDAC734FFB50102FEEEA1C0FE3176D4" box="[325,436,1423,1448]" italics="true" pageId="2" pageNumber="574">Pronotum</emphasis>
: anterior, medial posterior areas and margins punctate and setose, setae on anterior area thin, short, setae on medial and posterior areas long; lateral areas without punctures or setae. 
<taxonomicName id="B9AE60A5FFB50102FB34A1FDFA8876B0" box="[1183,1293,1458,1484]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="2" pageNumber="574" phylum="Arthropoda" rank="genus">
<emphasis id="4CDAC734FFB50102FB34A1FDFA8876B0" box="[1183,1293,1458,1484]" italics="true" pageId="2" pageNumber="574">Scutellum</emphasis>
</taxonomicName>
: Setose with a medial longitudinal impunctate area. 
<emphasis id="4CDAC734FFB50102FDE2A199FD0A768C" box="[585,655,1494,1520]" italics="true" pageId="2" pageNumber="574">Elytra</emphasis>
: Glossy, striae indistinct, humerus rounded. Disc smooth, impunctate, and glabrous. Distal margin with thin setae, length of setae about two times longer than the distance between related punctures. 
<emphasis id="4CDAC734FFB50102FEB9A251FEF87544" box="[274,381,1566,1592]" italics="true" pageId="2" pageNumber="574">Pygidium</emphasis>
(
<figureCitation id="E69507A3FFB50102FE26A251FE627545" box="[397,487,1566,1593]" captionStart="FIGURE 2" captionStartId="4.[151,250,1618,1642]" captionTargetBox="[156,1436,202,1592]" captionTargetId="figure-21@4.[151,1436,181,1594]" captionTargetPageId="4" captionText="FIGURE 2. Dicrania chimu, new species, male holotype. A, Dorsal view; B, lateral view; C, ventral view; D, labels; E, head and pronotum in dorsal view; F, aedeagus in lateral view; G, aedeagus in distal view; H, pygidium in caudal view; I, antenna in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11548017" httpUri="https://zenodo.org/record/11548017/files/figure.png" pageId="2" pageNumber="574">Fig. 2H</figureCitation>
): As wide as long or almost so, surface punctate and setose; setae thin, short or long; punctures barely defined and thin. 
<emphasis id="4CDAC734FFB50102FD88A20CFD007520" box="[547,645,1603,1628]" italics="true" pageId="2" pageNumber="574">Ventrites</emphasis>
: Ventrite V shorter than ventrite VI. 
<emphasis id="4CDAC734FFB50102FB81A20CFB217520" box="[1066,1188,1603,1628]" italics="true" pageId="2" pageNumber="574">Parameres</emphasis>
(
<figureCitation id="E69507A3FFB50102FB1EA20DFAB57521" box="[1205,1328,1602,1629]" captionStart="FIGURE 2" captionStartId="4.[151,250,1618,1642]" captionTargetBox="[156,1436,202,1592]" captionTargetId="figure-21@4.[151,1436,181,1594]" captionTargetPageId="4" captionText="FIGURE 2. Dicrania chimu, new species, male holotype. A, Dorsal view; B, lateral view; C, ventral view; D, labels; E, head and pronotum in dorsal view; F, aedeagus in lateral view; G, aedeagus in distal view; H, pygidium in caudal view; I, antenna in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11548017" httpUri="https://zenodo.org/record/11548017/files/figure.png" pageId="2" pageNumber="574">Fig. 2F–G</figureCitation>
): Slightly constricted medially in dorsal and lateral views. 
<emphasis id="4CDAC734FFB50102FD14A228FD7175FC" box="[703,756,1639,1664]" italics="true" pageId="2" pageNumber="574">Legs</emphasis>
: protarsomere 1 with a truncate inner tooth, protarsomere 2 longer than wide, and protarsomeres 2–4 slightly laterally compressed; each protarsal claw gradually deflected, outer tooth about 1.5 times longer than and 0.9 as wide as inner tooth. Mesotibia about 2.6 times longer than the width at the apex; mesotarsomere 1 with a spatulate inner tooth. Apex of metatibia with inner area slightly projected, projection blunt and about 0.3 times as long as the length of remainder of apex; metatarsomere 2 with sparse setae, medial setae shorter than the length of tarsomere; metatarsomere 4 with distal inner setae as wide as other tarsomere setae or almost so; each metatarsal claw gradually deflected, outer tooth about 1.8 times longer than and 0.7 as wide as the inner tooth.
</paragraph>
<caption id="2AD14BAEFFB40103FF3CA178FDD376C7" ID-DOI="http://doi.org/10.5281/zenodo.11548013" ID-Zenodo-Dep="11548013" httpUri="https://zenodo.org/record/11548013/files/figure.png" pageId="3" pageNumber="575" startId="3.[151,250,1335,1359]" targetBox="[151,1436,183,1311]" targetPageId="3" targetType="figure">
<paragraph id="7E111B26FFB40103FF3CA178FDD376C7" blockId="3.[151,1437,1335,1467]" pageId="3" pageNumber="575">
<emphasis id="4CDAC734FFB40103FF3CA178FE937633" bold="true" box="[151,278,1335,1359]" pageId="3" pageNumber="575">FIGURE 1.</emphasis>
<taxonomicName id="B9AE60A5FFB40103FEB5A177FEFC7633" authorityName="LePeletier &amp; Audinet-Serville" authorityYear="1828" box="[286,377,1336,1359]" class="Insecta" family="Scarabaeidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis id="4CDAC734FFB40103FEB5A177FEFC7633" box="[286,377,1336,1359]" italics="true" pageId="3" pageNumber="575">Dicrania</emphasis>
</taxonomicName>
diagnostic morphological characters. 
<emphasis id="4CDAC734FFB40103FCADA178FC9A7633" bold="true" box="[774,799,1335,1359]" pageId="3" pageNumber="575">A,</emphasis>
<taxonomicName id="B9AE60A5FFB40103FC83A177FB0F7633" authority="Moser, 1921" authorityName="Moser" authorityYear="1921" box="[808,1162,1335,1359]" class="Insecta" family="Scarabaeidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="575" phylum="Arthropoda" rank="species" species="laevipennis">
<emphasis id="4CDAC734FFB40103FC83A177FC7A7633" box="[808,1023,1335,1359]" italics="true" pageId="3" pageNumber="575">Dicrania laevipennis</emphasis>
<bibRefCitation id="1A3F66D7FFB40103FBA2A177FB0F7633" author="Moser, J." box="[1033,1162,1335,1359]" pageId="3" pageNumber="575" pagination="133 - 182" refId="ref5317" refString="Moser, J. (1921) Neue Melolonthiden von Mittel- und Sud-Amerika. Stettiner Entomologische Zeitung, 82, 133 - 182." type="journal article" year="1921">Moser, 1921</bibRefCitation>
</taxonomicName>
male, head and pronotum in dorsal view; 
<emphasis id="4CDAC734FFB40103FE96A114FED1760F" bold="true" box="[317,340,1371,1395]" pageId="3" pageNumber="575">B,</emphasis>
<taxonomicName id="B9AE60A5FFB40103FEF6A113FE38760F" authority="Clavijo-Bustos &amp; Moreno &amp; Fuhrmann, 2024" authorityName="Clavijo-Bustos &amp; Moreno &amp; Fuhrmann" authorityYear="2024" box="[349,445,1371,1395]" class="Insecta" family="Scarabaeidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="575" phylum="Arthropoda" rank="species" species="chimu" status="new species">
<emphasis id="4CDAC734FFB40103FEF6A113FEEA760F" box="[349,367,1372,1395]" italics="true" pageId="3" pageNumber="575">D</emphasis>
. 
<emphasis id="4CDAC734FFB40103FED4A114FE38760F" box="[383,445,1371,1395]" italics="true" pageId="3" pageNumber="575">chimu</emphasis>
</taxonomicName>
<emphasis id="4CDAC734FFB40103FE6DA114FDC3760F" bold="true" box="[454,582,1371,1395]" pageId="3" pageNumber="575">
<taxonomicNameLabel id="57E97A4FFFB40103FE6DA114FDC3760F" box="[454,582,1371,1395]" pageId="3" pageNumber="575" rank="species">new species</taxonomicNameLabel>
</emphasis>
male paratype, scutellum in dorsal view; 
<emphasis id="4CDAC734FFB40103FBA9A114FB9E760F" bold="true" box="[1026,1051,1371,1395]" pageId="3" pageNumber="575">C,</emphasis>
<taxonomicName id="B9AE60A5FFB40103FB88A113FB3D760F" authorityName="Moser" authorityYear="1921" box="[1059,1208,1371,1395]" class="Insecta" family="Scarabaeidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="575" phylum="Arthropoda" rank="species" species="laevipennis">
<emphasis id="4CDAC734FFB40103FB88A113FBB0760F" box="[1059,1077,1372,1395]" italics="true" pageId="3" pageNumber="575">D</emphasis>
. 
<emphasis id="4CDAC734FFB40103FBEEA114FB3D760F" box="[1093,1208,1371,1395]" italics="true" pageId="3" pageNumber="575">laevipennis</emphasis>
</taxonomicName>
male, protibia; 
<emphasis id="4CDAC734FFB40103FAC9A114FAFF760F" bold="true" box="[1378,1402,1371,1395]" pageId="3" pageNumber="575">D,</emphasis>
<taxonomicName id="B9AE60A5FFB40103FA2FA113FE8F76EB" authorityName="Moser" authorityYear="1921" class="Insecta" family="Scarabaeidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="575" phylum="Arthropoda" rank="species" species="laevipennis">
<emphasis id="4CDAC734FFB40103FA2FA113FA13760F" box="[1412,1430,1372,1395]" italics="true" pageId="3" pageNumber="575">D</emphasis>
. 
<emphasis id="4CDAC734FFB40103FF3CA130FE8F76EB" box="[151,266,1407,1431]" italics="true" pageId="3" pageNumber="575">laevipennis</emphasis>
</taxonomicName>
male, metacoxa in dorsal view; 
<emphasis id="4CDAC734FFB40103FDFBA130FDE276EB" bold="true" box="[592,615,1407,1431]" pageId="3" pageNumber="575">E,</emphasis>
<taxonomicName id="B9AE60A5FFB40103FDC6A1CFFD4C76EB" authorityName="Clavijo-Bustos &amp; Moreno &amp; Fuhrmann" authorityYear="2024" box="[621,713,1407,1431]" class="Insecta" family="Scarabaeidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="575" phylum="Arthropoda" rank="species" species="chimu">
<emphasis id="4CDAC734FFB40103FDC6A1CFFDFA76EB" box="[621,639,1408,1431]" italics="true" pageId="3" pageNumber="575">D</emphasis>
. 
<emphasis id="4CDAC734FFB40103FD20A130FD4C76EB" box="[651,713,1407,1431]" italics="true" pageId="3" pageNumber="575">chimu</emphasis>
</taxonomicName>
male paratype, metafemora in ventral view; 
<emphasis id="4CDAC734FFB40103FB27A130FB2576EB" bold="true" box="[1164,1184,1407,1431]" pageId="3" pageNumber="575">F,</emphasis>
<taxonomicName id="B9AE60A5FFB40103FB0EA1CFFA8776EB" authorityName="Clavijo-Bustos &amp; Moreno &amp; Fuhrmann" authorityYear="2024" box="[1189,1282,1407,1431]" class="Insecta" family="Scarabaeidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="575" phylum="Arthropoda" rank="species" species="chimu">
<emphasis id="4CDAC734FFB40103FB0EA1CFFB3276EB" box="[1189,1207,1408,1431]" italics="true" pageId="3" pageNumber="575">D</emphasis>
. 
<emphasis id="4CDAC734FFB40103FB6FA130FA8776EB" box="[1220,1282,1407,1431]" italics="true" pageId="3" pageNumber="575">chimu</emphasis>
</taxonomicName>
male paratype, lateral internal view of metatarsomeres 4–5.
</paragraph>
</caption>
<paragraph id="7E111B26FFB40103FF6CA242FE27755B" blockId="3.[151,1437,1549,1900]" box="[199,418,1549,1575]" pageId="3" pageNumber="575">
<emphasis id="4CDAC734FFB40103FF6CA242FEA0755B" bold="true" box="[199,293,1549,1575]" pageId="3" pageNumber="575">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="7E111B26FFB40103FF6CA27EFA977530" blockId="3.[151,1437,1549,1900]" box="[199,1298,1585,1612]" pageId="3" pageNumber="575">
<emphasis id="4CDAC734FFB40103FF6CA27EFE5A7537" bold="true" box="[199,479,1585,1611]" pageId="3" pageNumber="575">Variation. Body length:</emphasis>
<quantity id="B956B6C3FFB40103FE4DA27EFD097530" box="[486,652,1585,1612]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" metricValueMax="1.41" metricValueMin="1.25" pageId="3" pageNumber="575" unit="mm" value="13.3" valueMax="14.1" valueMin="12.5">12.5–14.1 mm</quantity>
; 
<emphasis id="4CDAC734FFB40103FD3CA27EFBF97537" bold="true" box="[663,1148,1585,1611]" pageId="3" pageNumber="575">Body maximum width (at half of elytra):</emphasis>
<quantity id="B956B6C3FFB40103FB2FA27EFA8B7530" box="[1156,1294,1585,1612]" metricMagnitude="-3" metricUnit="m" metricValue="6.65" metricValueMax="7.4" metricValueMin="5.9" pageId="3" pageNumber="575" unit="mm" value="6.65" valueMax="7.4" valueMin="5.9">5.9–7.4 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="36B448ADFFB40103FF6CA21AFEF975E8" pageId="3" pageNumber="575" type="distribution">
<paragraph id="7E111B26FFB40103FF6CA21AFEF975E8" blockId="3.[151,1437,1549,1900]" pageId="3" pageNumber="575">
<emphasis id="4CDAC734FFB40103FF6CA21AFEE47513" bold="true" box="[199,353,1621,1647]" pageId="3" pageNumber="575">Distribution.</emphasis>
Known only from the 
<typeStatus id="A115A584FFB40103FDF2A219FD0C750C" box="[601,649,1622,1648]" pageId="3" pageNumber="575">type</typeStatus>
series collected in the municipality of Puerto López, 
<collectingRegion id="BC6AD5C4FFB40103FB79A219FA1D750C" box="[1234,1432,1622,1648]" country="Colombia" name="Meta" pageId="3" pageNumber="575">Meta Department</collectingRegion>
, 
<collectingCountry id="06B95BB6FFB40103FF3CA236FE8075EF" box="[151,261,1657,1683]" name="Colombia" pageId="3" pageNumber="575">Colombia</collectingCountry>
(
<figureCitation id="E69507A3FFB40103FEBEA236FEEA75E8" box="[277,367,1657,1684]" captionStart="FIGURE 5" captionStartId="7.[151,250,1980,2004]" captionTargetBox="[191,1396,1253,1956]" captionTargetId="figure-534@7.[183,1404,1245,1956]" captionTargetPageId="7" captionText="FIGURE 5. Distribution of the Dicrania species from Colombia. A, Colombia; B, known distribution of Dicrania laevipennis Moser, 1921." figureDoi="http://doi.org/10.5281/zenodo.11548027" httpUri="https://zenodo.org/record/11548027/files/figure.png" pageId="3" pageNumber="575">Fig. 5A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="36B448ADFFB40103FF6CA2D2FE06745F" pageId="3" pageNumber="575" type="etymology">
<paragraph id="7E111B26FFB40103FF6CA2D2FE06745F" blockId="3.[151,1437,1549,1900]" pageId="3" pageNumber="575">
<emphasis id="4CDAC734FFB40103FF6CA2D2FECB75CB" bold="true" box="[199,334,1693,1719]" pageId="3" pageNumber="575">Etymology.</emphasis>
<emphasis id="4CDAC734FFB40103FEF3A2D2FE2675CB" box="[344,419,1693,1719]" italics="true" pageId="3" pageNumber="575">Chimú</emphasis>
(or 
<emphasis id="4CDAC734FFB40103FE7DA2D2FDA475CB" box="[470,545,1693,1719]" italics="true" pageId="3" pageNumber="575">Chimó</emphasis>
) is a traditional pre-Colombian dark, shiny, and paste-like extract of tobacco (
<taxonomicName id="B9AE60A5FFB40103FF0AA28DFEF075A7" box="[161,373,1729,1755]" class="Magnoliopsida" family="Solanaceae" genus="Nicotiana" kingdom="Plantae" order="Solanales" pageId="3" pageNumber="575" phylum="Tracheophyta" rank="species" species="tabacum">
<emphasis id="4CDAC734FFB40103FF0AA28DFEF075A7" box="[161,373,1729,1755]" italics="true" pageId="3" pageNumber="575">Nicotiana tabacum</emphasis>
</taxonomicName>
L.; 
<taxonomicName id="B9AE60A5FFB40103FE0AA28EFDA475A7" box="[417,545,1729,1755]" class="Magnoliopsida" family="Solanaceae" kingdom="Plantae" order="Solanales" pageId="3" pageNumber="575" phylum="Tracheophyta" rank="family">Solanaceae</taxonomicName>
) made by aboriginals of the Llanos region of 
<collectingCountry id="06B95BB6FFB40103FBBEA28EFB0675A7" box="[1045,1155,1729,1755]" name="Colombia" pageId="3" pageNumber="575">Colombia</collectingCountry>
and 
<collectingCountry id="06B95BB6FFB40103FB1CA28EFAAB75A7" box="[1207,1326,1729,1755]" name="Venezuela" pageId="3" pageNumber="575">Venezuela</collectingCountry>
. 
<taxonomicName id="B9AE60A5FFB40103FA9CA28DFF5E7583" authorityName="Clavijo-Bustos &amp; Moreno &amp; Fuhrmann" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="575" phylum="Arthropoda" rank="species" species="chimu">
<emphasis id="4CDAC734FFB40103FA9CA28DFF5E7583" italics="true" pageId="3" pageNumber="575">Dicrania chimu</emphasis>
</taxonomicName>
is also from the Llanos Orientales region of 
<collectingCountry id="06B95BB6FFB40103FD64A2AAFCB87583" box="[719,829,1765,1791]" name="Colombia" pageId="3" pageNumber="575">Colombia</collectingCountry>
and is dark shiny black, just like 
<taxonomicName id="B9AE60A5FFB40103FB18A2AAFB727583" authorityName="Clavijo-Bustos &amp; Moreno &amp; Fuhrmann" authorityYear="2024" box="[1203,1271,1765,1791]" class="Insecta" family="Scarabaeidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="575" phylum="Arthropoda" rank="species" species="chimu">
<emphasis id="4CDAC734FFB40103FB18A2AAFB727583" box="[1203,1271,1765,1791]" italics="true" pageId="3" pageNumber="575">chimú</emphasis>
</taxonomicName>
. The epithet is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="36B448ADFFB40103FF6CA362FF557410" pageId="3" pageNumber="575" type="discussion">
<paragraph id="7E111B26FFB40103FF6CA362FF557410" blockId="3.[151,1437,1549,1900]" pageId="3" pageNumber="575">
<emphasis id="4CDAC734FFB40103FF6CA362FE45743B" bold="true" box="[199,448,1837,1863]" pageId="3" pageNumber="575">Taxonomic remarks.</emphasis>
<taxonomicName id="B9AE60A5FFB40103FE62A361FDF9743B" authorityName="Clavijo-Bustos &amp; Moreno &amp; Fuhrmann" authorityYear="2024" box="[457,636,1837,1863]" class="Insecta" family="Scarabaeidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="575" phylum="Arthropoda" rank="species" species="chimu">
<emphasis id="4CDAC734FFB40103FE62A361FDF9743B" box="[457,636,1837,1863]" italics="true" pageId="3" pageNumber="575">Dicrania chimu</emphasis>
</taxonomicName>
is the only species of the 
<taxonomicName id="B9AE60A5FFB40103FC1BA361FBFF743B" authorityName="Chevrolat" authorityYear="1844" box="[944,1146,1838,1864]" class="Insecta" family="Scarabaeidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="575" phylum="Arthropoda" rank="species" species="castaneipennis">
<emphasis id="4CDAC734FFB40103FC1BA361FC41743B" box="[944,964,1838,1863]" italics="true" pageId="3" pageNumber="575">D</emphasis>
. 
<emphasis id="4CDAC734FFB40103FC7FA361FBFF743B" box="[980,1146,1838,1863]" italics="true" pageId="3" pageNumber="575">castaneipennis</emphasis>
</taxonomicName>
species group with large eyes.
</paragraph>
</subSubSection>
</treatment>
</document>