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140 lines
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.721.21529" ID-GBIF-Dataset="8e4ab8e9-98d9-417c-9188-ea0b1161c40b" ID-PMC="PMC5740412" ID-Pensoft-Pub="1313-2970-721-45" ID-PubMed="29308024" ID-ZBK="1300B5278A5041E38D5E1C745846BBDF" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-721-45" ModsDocOrigin="ZooKeys 721" ModsDocTitle="The geographical patterns of distribution of the genus Teuthraustes Simon, 1878 in Ecuador and description of three new species (Scorpiones, Chactidae)" checkinTime="1513120619192" checkinUser="pensoft" docAuthor="Ythier, Eric & Lourenco, Wilson R." docDate="2017" docId="1DE0D60B066161B122E1123BBE986EFC" docLanguage="en" docName="ZooKeys 721: 45-63" docOrigin="ZooKeys 721" docSource="http://dx.doi.org/10.3897/zookeys.721.21529" docTitle="Teuthraustes kuryi Ythier & Lourenco, 2017, sp. n." docType="treatment" docUuid="DB39E8D8-51EB-4784-AECF-43FC3BE4A923" docUuidSource="ZooBank" docVersion="5" lastPageNumber="59" masterDocId="2E37FFB92571FFC5FFA7FFC34F5FFFB3" masterDocTitle="The geographical patterns of distribution of the genus Teuthraustes Simon, 1878 in Ecuador and description of three new species (Scorpiones, Chactidae)" masterLastPageNumber="63" masterPageNumber="45" pageNumber="56" updateTime="1668165195725" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The geographical patterns of distribution of the genus Teuthraustes Simon, 1878 in Ecuador and description of three new species (Scorpiones, Chactidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ythier, Eric</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lourenco, Wilson R.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>721</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>45</mods:start>
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<mods:end>63</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.721.21529</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.721.21529</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-721-45</mods:identifier>
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<mods:identifier type="ZBK">1300B5278A5041E38D5E1C745846BBDF</mods:identifier>
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<mods:identifier type="ZooBank">1300B5278A5041E38D5E1C745846BBDF</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="137679866" LSID="urn:lsid:zoobank.org:act:DB39E8D8-51EB-4784-AECF-43FC3BE4A923" httpUri="http://treatment.plazi.org/id/1DE0D60B066161B122E1123BBE986EFC" lastPageId="14" lastPageNumber="59" pageId="11" pageNumber="56">
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<subSubSection pageId="11" pageNumber="56" type="nomenclature">
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<paragraph pageId="11" pageNumber="56">
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<taxonomicName LSID="http://zoobank.org/DB39E8D8-51EB-4784-AECF-43FC3BE4A923" class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teuthraustes kuryi" order="Scorpiones" pageId="11" pageNumber="56" phylum="Arthropoda" rank="species" species="kuryi">Teuthraustes kuryi</taxonomicName>
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<taxonomicNameLabel pageId="11" pageNumber="56">sp. n.</taxonomicNameLabel>
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Fig. 7, 8, Table 1
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</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="56" type="type material">
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<paragraph pageId="11" pageNumber="56">Type material.</paragraph>
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<paragraph pageId="11" pageNumber="56">
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Ecuador, Esmeraldas Province (N00.88883, W078.53732), 873 m, in zone where rainforest was recently destroyed, under log, 17/III/2011 (A. Chagas, A. Giupponi, A. Kury, M. Vega). Female holotype and female paratype. Holotype deposited in the Museo de Zoologia, Pontificia Universidad
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<normalizedToken originalValue="Católica">Catolica</normalizedToken>
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de Quito, Ecuador; paratype deposited in the Museu Nacional, Rio de Janeiro, Brazil.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="56" type="etymology">
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<paragraph pageId="11" pageNumber="56">Etymology.</paragraph>
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<paragraph pageId="11" pageNumber="56">Specific name honours Prof. Adriano B. Kury, Museu Nacional, Universidade Federal do Rio de Janeiro, Brazil, for his important contribution to the study of arachnids.</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="56" type="diagnosis">
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<paragraph pageId="11" pageNumber="56">Diagnosis.</paragraph>
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<paragraph pageId="11" pageNumber="56">
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Moderate to large scorpions with 60-61 mm in total length. Colouration dark brown to blackish. Body and appendages strongly granulated. Pectines with 7-7 teeth in females. Pedipalps bulk with short fingers; fixed and movable fingers of chela with 5-6 rows of granules. Metasoma long and strong; ventral carinae strongly marked on metasomal segments I to IV. Trichobothrial pattern of type C neobothriotaxic
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<normalizedToken originalValue="‘majorante’">'majorante'</normalizedToken>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="56" type="description">
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<paragraph pageId="11" pageNumber="56">Description.</paragraph>
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<paragraph pageId="11" pageNumber="56">Based on female holotype and female paratype.</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="56" type="colouration">
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<paragraph pageId="11" pageNumber="56">Colouration.</paragraph>
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<paragraph pageId="11" pageNumber="56">General colouration basically dark brown to blackish. Prosoma: carapace dark brown; eyes blackish. Mesosoma: tergites reddish brown to dark brown, slightly paler than carapace; venter and sternites reddish brown to dark brown; pectines and genital operculum yellow. Metasoma: segments blackish brown, with darker zones over carinae; vesicle reddish brown with the base of the aculeus reddish yellow. Chelicerae: reddish yellow, with some diffuse variegated reddish spots at the base of the fingers; fingers dark reddish. Pedipalps dark brown to blackish; carinae blackish. Legs dark brown to blackish.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="12" lastPageNumber="57" pageId="11" pageNumber="56" type="morphology">
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<paragraph pageId="11" pageNumber="56">Morphology.</paragraph>
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<paragraph lastPageId="12" lastPageNumber="57" pageId="11" pageNumber="56">
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Prosoma: carapace almost acarinate, but with dense strongly marked granulations on the entire surface, except on the zone of furrows; furrows deep; sternum pentagonal, wider than long. Mesosoma: tergites intensely granulated but less marked than carapace; sternites, smooth and shiny; spiracles oval-shaped and con
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<pageBreakToken pageId="12" pageNumber="57" start="start">spicuous</pageBreakToken>
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; VII acarinate with granulations; pectinal tooth count 7-7 in both females, holotype and paratype, fulcra vestigial. Metasoma: segments I and II wider than long; metasomal tegument on segments I to IV strongly granulated including dorsal aspect; segment V with some spinoid granulations ventrally; carinae on segments
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<normalizedToken originalValue="I–V">I-V</normalizedToken>
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strongly developed; ventral present on all segments; telson strongly granulated; aculeus shorter than vesicle. Chelicerae: dentition typical of the family
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<taxonomicName family="Chactidae" lsidName="" pageId="12" pageNumber="57" rank="family">Chactidae</taxonomicName>
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(
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<bibRefCitation author="Vachon, M" journalOrPublisher="Bulletin du Museum d'histoire naturelle" pageId="17" pageNumber="62" pagination="161 - 166" title="De l'utilite, en systematique, d'une nomenclature des dents des cheliceres chez les Scorpions." volume="35" year="1963">Vachon 1963</bibRefCitation>
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), and with dense setation ventrally and internally. Pedipalps: femur with dorsal internal, dorsal external and ventral internal carinae strongly marked; ventral external carina vestigial; dorsal, ventral and internal aspects granulated; patella granulated with well-marked carinae; chela strongly granulated and with most carinae well-marked; dentate margins on movable and fixed fingers with 5-6 rows of granules; trichobothriotaxy of type C; neobothriotaxic
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<normalizedToken originalValue="‘majorante’">'majorante'</normalizedToken>
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(
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<bibRefCitation author="Vachon, M" journalOrPublisher="Bulletin du Museum d'histoire naturelle" pageId="17" pageNumber="62" pagination="857 - 958" title="Etude des caracteres utilises pour classer les familles et les genres de Scorpions (Arachnides). 1. La trichobothriotaxie en arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les Scorpions." volume="140" year="1974">Vachon 1974</bibRefCitation>
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). Legs: tarsi with short setae disposed in a single line.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="13" lastPageNumber="58" pageId="12" pageNumber="57" type="morphometric values">
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<paragraph pageId="12" pageNumber="57">Morphometric values.</paragraph>
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<paragraph lastPageId="13" lastPageNumber="58" pageId="12" pageNumber="57">
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Female holotype of
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<taxonomicName lsidName="T. kuryi" pageId="12" pageNumber="57" rank="species" species="kuryi">T. kuryi</taxonomicName>
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sp. n. Total length (in mm) including the telson, 60.4. Carapace: length, 8.0; anterior width, 4.8; posterior width, 8.6. Mesosoma length, 16.2. Metasomal segments. I: length, 3.6; width, 4.9; II: length, 4.1; width, 4.6; III: length, 4.6; width, 4.5; IV: length, 5.6; width, 4.3; V: length, 9.4;
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<pageBreakToken pageId="13" pageNumber="58" start="start">width</pageBreakToken>
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, 4.2; depth, 3.6. Telson length, 8.9; vesicle: width, 4.3; depth, 3.6. Pedipalp: femur length, 5.7, width, 2.4; patella length, 6.1, width, 2.8; chela length, 11.1, width, 5.1, depth, 7.2; movable finger length, 5.9.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="14" lastPageNumber="59" pageId="13" pageNumber="58" type="relationships">
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<paragraph pageId="13" pageNumber="58">Relationships.</paragraph>
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<paragraph lastPageId="14" lastPageNumber="59" pageId="13" pageNumber="58">
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The new species can be distinguished from the others congeners in particular from
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<taxonomicName class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teuthraustes atramentarius" order="Scorpiones" pageId="13" pageNumber="58" phylum="Arthropoda" rank="species" species="atramentarius">Teuthraustes atramentarius</taxonomicName>
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Simon, which is also distributed in Ecuador,
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<pageBreakToken pageId="14" pageNumber="59" start="start">but</pageBreakToken>
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exclusively in the high central Andes, by the following features: (i) carapace, tergites, pedipalps, metasoma, and telson strongly granulated, (ii) metasomal segments I to V long and strong with well-marked ventral carinae, (iii) pedipalps bulk with short fingers on chela, (iv) ventral aspect with a darker pigmentation, reddish brown to dark brown.
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</paragraph>
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<caption pageId="14" pageNumber="59">
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<paragraph pageId="14" pageNumber="59">
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Figure 7.
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<taxonomicName class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teuthraustes kuryi" order="Scorpiones" pageId="14" pageNumber="59" phylum="Arthropoda" rank="species" species="kuryi">Teuthraustes kuryi</taxonomicName>
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sp. n. Female holotype.
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<normalizedToken originalValue="A–B">A-B</normalizedToken>
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Habitus, dorsal A and ventral B aspects.
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</paragraph>
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</caption>
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<caption pageId="14" pageNumber="59">
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<paragraph pageId="14" pageNumber="59">
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Figure 8.
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<taxonomicName class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Teuthraustes kuryi" order="Scorpiones" pageId="14" pageNumber="59" phylum="Arthropoda" rank="species" species="kuryi">Teuthraustes kuryi</taxonomicName>
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sp. n. Female holotype. A Chelicera, dorsal aspect B Disposition of the granulation over the dentate margins of pedipalp-chela movable finger
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<normalizedToken originalValue="C–H">C-H</normalizedToken>
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Trichobothrial pattern
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<normalizedToken originalValue="C–D">C-D</normalizedToken>
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Chela, dorso-external and ventral aspects E Femur, dorsal aspect
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<normalizedToken originalValue="F–H">F-H</normalizedToken>
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Patella, dorsal F external G and ventral H aspects. Scale bars 1 mm.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document>
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