treatments-xml/data/FA/11/77/FA1177A074C859B19F69711C4654B852.xml

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<mods:title id="D3252BA2D38AB5E97BE40CAA8FE07B25">Systematics of the Dendropsophus leucophyllatus species group (Anura, Hylidae) from the Chocó region of Ecuador, with description of a new species</mods:title>
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<mods:namePart id="3CE7BE39BA085877B178124B4078A1BF">Aguirre, P. Doménica</mods:namePart>
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<mods:affiliation id="8FA460559ED82CC3DC50FE87B173BEA6">Museo de Zoología, Escuela de Biología, Pontificia Universidad Católica del Ecuador, Av. 12 de Octubre y Roca, Aptdo. 17 - 01 - 2184, Quito, Ecuador &amp; Department of Biological Science, Florida State University, Tallahassee, FL 32306, USA</mods:affiliation>
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<mods:namePart id="9B54AF4B4088CB5EA6EDBD0FB9DEC234">Ron, Santiago R.</mods:namePart>
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<mods:affiliation id="201BC866D05841951B511F86459B738C">Museo de Zoología, Escuela de Biología, Pontificia Universidad Católica del Ecuador, Av. 12 de Octubre y Roca, Aptdo. 17 - 01 - 2184, Quito, Ecuador</mods:affiliation>
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<mods:date id="E13E347833914AB7FCCE525498223CB9">2025</mods:date>
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<taxonomicName id="CD2F5D708242EE4C68CAE6550AF93F72" authorityName="Aguirre &amp; Apunte &amp; Ron" authorityYear="2025" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="cannatellai" status="sp. nov.">
<emphasis id="80F99C415059BF70CF77DDE1D51C7C06" italics="true">Dendropsophus cannatellai</emphasis>
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<taxonomicNameLabel id="4530D72E68B8816E6F955A1575B7A252" rank="species">sp. nov.</taxonomicNameLabel>
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<emphasis id="BB24D32248D3573B07E7FCD9D734D5AC" italics="true">Dendropsophus ebraccatus</emphasis>
</taxonomicName>
: 
<bibRefCitation id="1998CDC8BBE96660A733E09C1BBE0586" author="Ortega-Andrade" etAl="et al." firstAuthor="Ortega-Andrade" refId="B31" refString="Ortega-Andrade HM, Meza-Ramos P, Cisneros-Heredia DF, Yánez-Munoz MH, Altamirano-Benavidez M (2010) Los anfibios y reptiles del Chocó Esmeraldeño. In: Serie herpetofauna del Ecuador: el Chocó Esmeraldeño. Monografía. Museo Ecuatoriano de Ciencias Naturales, Quito, Ecuador, 95–232." year="2010">Ortega-Andrade et al. 2010</bibRefCitation>
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.
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<emphasis id="195A4EB97A39D850C8969C7D5444A835" italics="true">Dendropsophus ebraccatus</emphasis>
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: 
<bibRefCitation id="1819BC5A2CB01980B468FFD6340A7481" author="Ron" editor="Heatwole H &amp; Barrio-Amoros C &amp; Wilkinson HW" etAl="et al." firstAuthor="Ron" journalOrPublisher="9, Part 2. Surrey Beatty &amp; Sons Pty. Ltd., Australia" pagination="129-170" refId="B41" refString="Ron SR, Guayasamin JM, Menéndez-Guerrero PA (2011) Biodiversity and Conservation Status of Ecuadorian Amphibians. In: Heatwole H, Barrio-Amoros C, Wilkinson HW (Eds) Amphibian Biology, Vol. 9, Part 2. Surrey Beatty &amp; Sons Pty. Ltd., Australia, 129–170." title="Biodiversity and Conservation Status of Ecuadorian Amphibians." volumeTitle="Amphibian Biology, Vol." year="2011">Ron et al. 2011</bibRefCitation>
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<taxonomicName id="8CA4127AAC7A8E2F82983BF04375A855" ID-CoL="34S56" baseAuthorityName="Cope" baseAuthorityYear="1874" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="ebraccatus">
<emphasis id="0DC23699E9DD1992BAE9DE5ACCC77F99" italics="true">Dendropsophus ebraccatus</emphasis>
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: 
<bibRefCitation id="FEDAF54C41373E58F08BE753E3FBEE79" author="Simpson" etAl="et al." firstAuthor="Simpson" refId="B44" refString="Simpson N, Reyes-Puig C, Tobar C, Yánez-Muñoz M (2013) Herpetofauna en áreas prioritarias para la conservación: el sistema de Reservas Jocotoco y Ecominga. MECN-Jocotoco-ECOMINGA 2013." year="2013">Simpson et al. 2013</bibRefCitation>
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<emphasis id="7CBB4C348E740C22AC9CCCAC1E1FE5AB" italics="true">Dendropsophus ebraccatus</emphasis>
</taxonomicName>
: 
<bibRefCitation id="2086BAE091A9501D0D2DB230567A98CC" DOI="10.1371/journal.pone.0171785" author="Caminer" etAl="et al." firstAuthor="Caminer" refId="B1" refString="Caminer MA, Mila B, Jansen M, Fouquet A, Venegas PJ, Chavez G, Lougheed SC, Ron SR (2017) Systematics of the Dendropsophus leucophyllatus species complex (Anura: Hylidae): Cryptic diversity and the description of two new species. PLoS ONE 12 (3): e 0171785. https://doi.org/10.1371/journal.pone.0171785" year="2017">Caminer et al. 2017</bibRefCitation>
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<taxonomicName id="26E6040A83B15C8583E830C1A173C530" ID-CoL="34S56" baseAuthorityName="Cope" baseAuthorityYear="1874" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="ebraccatus">
<emphasis id="7CAA2C674A5B2F63BE0BBA5FAB93E036" italics="true">Dendropsophus ebraccatus</emphasis>
</taxonomicName>
(in part): 
<bibRefCitation id="5064C99F9E5C6AB896AD0B1F81DE5E7B" DOI="10.1016/j.ympev.2020.106877" author="Pirani" etAl="et al." firstAuthor="Pirani" refId="B35" refString="Pirani RM, Peloso PL, Prado JR, Polo ÉM, Knowles LL, Ron SR, Rodrigues MT, Sturaro MJ, Werneck FP (2020) Diversification history of clown tree frogs in Neotropical rainforests (Anura, Hylidae, Dendropsophus leucophyllatus group). Molecular Phylogenetics and Evolution 150: 106877. https://doi.org/10.1016/j.ympev.2020.106877" year="2020">Pirani et al. 2020</bibRefCitation>
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<taxonomicName id="C42DDF12244976D36A06E3A692FCF7EA" ID-CoL="34S56" baseAuthorityName="Cope" baseAuthorityYear="1874" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="ebraccatus">
<emphasis id="72D3D068E16756E3F2205F7FAB9350D3" italics="true">Dendropsophus ebraccatus</emphasis>
</taxonomicName>
(in part): 
<bibRefCitation id="138F85B275D8A99D8108DE3DABC24E3B" DOI="10.1111/cla.12429" author="Orrico" etAl="et al." firstAuthor="Orrico" issue="1" journalOrPublisher="Cladistics" pagination="73-105" refId="B30" refString="Orrico VG, Grant T, Faivovich J, Rivera-Correa M, Rada MA, Lyra ML, Cassini CS, Valdujo PH, Schargel WE, Machado DJ (2021) The phylogeny of Dendropsophini (Anura: Hylidae: Hylinae). Cladistics 37 (1): 73–105. https://doi.org/10.1111/cla.12429" title="The phylogeny of Dendropsophini (Anura: Hylidae: Hylinae)." volume="37" year="2021">Orrico et al. 2021</bibRefCitation>
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<taxonomicName id="4DB2D1D60B20183459A1936EDEAE7FA9" ID-CoL="34S56" baseAuthorityName="Cope" baseAuthorityYear="1874" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="ebraccatus">
<emphasis id="EDA6DD2B919CD3DA60C4EB2D21E7B734" italics="true">Dendropsophus ebraccatus</emphasis>
</taxonomicName>
: 
<bibRefCitation id="87457ECBD8DCB8ADFE6E860096FED10B" DOI="10.1111/ecog.05906" author="Pintanel" etAl="et al." firstAuthor="Pintanel" refId="B34" refString="Pintanel P, Tejedo M, Merino-Viteri A, Almeida-Reinoso F, Salinas-Ivanenko S, López-Rosero AC, Llorente GA, Gutiérrez-Pesquera LM (2022) Elevational and local climate variability predicts thermal breadth of mountain tropical tadpoles. Ecography 2022: e 05906. https://doi.org/10.1111/ecog.05906" year="2022">Pintanel et al. 2022</bibRefCitation>
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<taxonomicName id="CD28BCC5DF8060448410E4382E419A2C" ID-CoL="34S56" baseAuthorityName="Cope" baseAuthorityYear="1874" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="ebraccatus">
<emphasis id="542AC4DFFC1C66C88A2668DB9120991E" italics="true">Dendropsophus ebraccatus</emphasis>
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(in part): 
<bibRefCitation id="BAE39B013A469F6673154786E280FF80" DOI="10.1073/pnas.2312371121" author="Touchon" etAl="et al." firstAuthor="Touchon" refId="B47" refString="Touchon JC, McMillan WO, Ibáñez R, Lessios HA (2024) Flexible oviposition behavior enabled the evolution of terrestrial reproduction. Proceedings of the National Academy of Sciences 121: e 2312371121. https://doi.org/10.1073/pnas.2312371121" year="2024">Touchon et al. 2024</bibRefCitation>
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<subSubSection id="SECID0E1SAI" type="Holotype">
<paragraph id="31B7C6318DFF574E3E97206B372DBACD">
<heading id="6EEE5E693EB9B7BFAF0A2C48C3A4ACD7" reason="title">
<typeStatus id="8CCDBAB3CA30DC6AD6F4614CBD9E8159" type="holotype">Holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="1ECA083F90B445ADAB1DA85191273ED1">
<materialsCitation id="8CB58621C9DFB53C46F0E7B462575A67" collectingDate="2007-08-06" collectionCode="QCAZ" collectorName="Luis A. Coloma &amp; Santiago R. Ron &amp; Mark Weiner" country="Ecuador" elevation="64" latitude="1.0873" location="Playon San Francisco" longLatPrecision="7" longitude="-78.6905" municipality="Canton San Lorenzo" specimenCode="QCAZ 40847" specimenCount="1" specimenCount-male="1" stateProvince="Esmeraldas Province" typeStatus="holotype">
• 
<specimenCode id="2FC329301B2810C4CF690553842EB2B9" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" type="Museum">
<named-content id="DE271E98803EF593FE3F8A8B1510CC33" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
40847
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(field no. SC-PUCE 19751), 
<specimenCount id="0453E38BBED6229C11C203E72EF707E1" count="1" type="male">adult male</specimenCount>
from 
<collectingCountry id="DD2F9E6DBED97C4EC88928982E3ABE35" name="Ecuador">Ecuador</collectingCountry>
, 
<collectingRegion id="08BC89CF2FAF772B8631AD34C210AD1F" country="Ecuador" name="Esmeraldas">Esmeraldas Province</collectingRegion>
, 
<collectingMunicipality id="196A4B31156635938C208D9DD25B7ED9">Cantón San Lorenzo</collectingMunicipality>
, 
<location id="946DDA835A24A5BB49FD98B00C08E408" LSID="urn:lsid:plazi:treatment:FA1177A074C859B19F69711C4654B852:946DDA835A24A5BB49FD98B00C08E408" country="Ecuador" latitude="1.0873" longLatPrecision="7" longitude="-78.6905" municipality="Canton San Lorenzo" name="Playon San Francisco" stateProvince="Esmeraldas Province">Playón San Francisco</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EHTAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-78.690500,1.087300]}">
<geoCoordinate id="9EAFB3855E2BE45C5D92822D1126EE9C" degrees="1.0873" direction="north" orientation="latitude" precision="5" value="1.0873">1.0873 ° N</geoCoordinate>
, 
<geoCoordinate id="FFCAB55FCEE89AB77E02D0822C8067D3" degrees="78.6905" direction="west" orientation="longitude" precision="5" value="-78.6905">78.6905 ° W</geoCoordinate>
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), 
<quantity id="B8DD44797A487EBB3016958373536739" metricMagnitude="1" metricUnit="m" metricValue="6.4" unit="m" value="64.0">
<elevation id="114931E2C109A6483E011810CC82211D" metricMagnitude="1" metricUnit="m" metricValue="6.4" unit="m" value="64.0">64 m</elevation>
</quantity>
, collected by 
<collectorName id="16D2D1CB4708F33E1F500B2DC27DA78A">Luis A. Coloma</collectorName>
, 
<collectorName id="579F48FF99E0203CE010F44662A452C9">Santiago R. Ron</collectorName>
, and 
<collectorName id="2311944BA9520FF2DF7E9CA1EA8904C3">Mark Weiner</collectorName>
on 
<date id="05B5629D16C439D9D584625509BE5F40" value="2007-08-06">
<collectingDate id="C2B181A9B1D7868D86CC5B9D17580D80" value="2007-08-06">06 August 2007</collectingDate>
</date>
.
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</paragraph>
</subSubSection>
<subSubSection id="SECID0EMTAI" type="Paratypes">
<paragraph id="17B14C14B0F56B4D227D2E0D5F7F44A3">
<heading id="CCC37E65C59B3E44E0C659A6D01FB6DE" reason="title">
<typeStatus id="296F6C4153D1BA97ACE50D00C93FDD47" type="paratype">Paratypes</typeStatus>
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<paragraph id="48CDEC3766CD4942734A3D736D8919BB">
<emphasis id="3EB51E22E4EC50D13BCA316CC60E95D0" bold="true">(130)</emphasis>
: All specimens were collected in 
<collectingCountry id="791FFEDA855A70861B7B49F68AEA1A15" name="Ecuador">Ecuador</collectingCountry>
. 
<materialsCitation id="49D5CB2FC6D39961074C9BD8989E14FA" collectingDate="1993-03-13" collectionCode="QCAZ" collectorName="Stephan Lotters &amp; Luis A. Coloma" country="Ecuador" elevation="745" latitude="0.0832" location="Maldonado" longLatPrecision="7" longitude="-79.0394" municipality="Canton Pedro Vicente Maldonado" specimenCode="QCAZ 3768" specimenCount="1" stateProvince="Pichincha" typeStatus="paratype">
<collectingRegion id="39F76E0900B3E61A4A6801991D50929C" country="Ecuador" name="Pichincha">Pichincha Province</collectingRegion>
: • 
<specimenCode id="C18878FF7D637C16D65B162C06275B05">
<named-content id="89C43C12DC535BAF801D0091363FE7E7" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
3768
</specimenCode>
–3774, 4840–4843 
<collectingMunicipality id="32CAB7D0CC813B0FE8768AC55E62B89E">Cantón Pedro Vicente Maldonado</collectingMunicipality>
, 
<locationDeviation id="6F8E1E9E2E5722D2FA384266DEBD1735">
<quantity id="08DC92312CAA2DB2E6B1DDBBFB230522" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="km" value="1.0">1 km</quantity>
E from
</locationDeviation>
<location id="6336E7876FFB63FE298100260CEC4195" LSID="urn:lsid:plazi:treatment:FA1177A074C859B19F69711C4654B852:6336E7876FFB63FE298100260CEC4195" country="Ecuador" latitude="0.0832" longLatPrecision="7" longitude="-79.0394" municipality="Canton Pedro Vicente Maldonado" name="Maldonado" stateProvince="Pichincha">Maldonado</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E2TAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-79.039400,0.083200]}">
<geoCoordinate id="1C48AB0A8D58A3A6C877A94A8F27E4C9" degrees="0.0832" direction="north" orientation="latitude" precision="5" value="0.0832">0.0832 ° N</geoCoordinate>
, 
<geoCoordinate id="BBA7A18A95705D819B958D77C8D16D6C" degrees="79.0394" direction="west" orientation="longitude" precision="5" value="-79.0394">79.0394 ° W</geoCoordinate>
</named-content>
), 
<quantity id="4D3E58D8691ECE2B34364DEB9884C1CC" metricMagnitude="2" metricUnit="m" metricValue="7.45" unit="m" value="745.0">
<elevation id="683558107DCE1727EED6B9ECAA33E5DC" metricMagnitude="2" metricUnit="m" metricValue="7.45" unit="m" value="745.0">745 m</elevation>
</quantity>
, collected by 
<collectorName id="C6348D89EB401977DFB332CE9BD155F5">Stephan Lötters</collectorName>
and 
<collectorName id="383E67058161FC4AA8907211459670EE">Luis A. Coloma</collectorName>
on 
<date id="2B0A6DBAF7A3F8CB95346FFF8A96F243" value="1993-03-13">
<collectingDate id="AF2E2D6E5B4C4F9A25895138B2880B81" value="1993-03-13">13 March, 1993</collectingDate>
</date>
</materialsCitation>
; • 
<materialsCitation id="2A8A0C4301DC528C81CBDBAC5B127BC0" collectingDate="1995-08-01" collectingDateMax="1996-01-01" collectingDateMin="1995-08-01" collectionCode="QCAZ" collectorName="Giovanna Rodriguez" country="Ecuador" elevation="280" latitude="0.1192" location="Rio Pitzara" longLatPrecision="7" longitude="-79.2344" municipality="Canton Pedro Vicente Maldonado" specimenCode="QCAZ 9457" specimenCount="1" stateProvince="Pichincha" typeStatus="paratype">
<specimenCode id="D8F1FDF8C1288FD5C4CCB3BC3D50BDC2" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" type="Museum">
<named-content id="413F18C922C719FC6039E6C01C266C9B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
9457
</specimenCode>
, 9489, 
<collectingMunicipality id="1743A466BE22B8675A92263FA5B96770">Cantón Pedro Vicente Maldonado</collectingMunicipality>
, 
<location id="1BB604392D8E01C935C116C0511E5DB4" LSID="urn:lsid:plazi:treatment:FA1177A074C859B19F69711C4654B852:1BB604392D8E01C935C116C0511E5DB4" country="Ecuador" latitude="0.1192" longLatPrecision="7" longitude="-79.2344" municipality="Canton Pedro Vicente Maldonado" name="Rio Pitzara" stateProvince="Pichincha">Río Pitzará</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EHUAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-79.234400,0.119200]}">
<geoCoordinate id="B0815F9538029907538D4CF3D8B926A3" degrees="0.1192" direction="north" orientation="latitude" precision="5" value="0.1192">0.1192 ° N</geoCoordinate>
, 
<geoCoordinate id="74F282078C7183566187C571F3F351F3" degrees="79.2344" direction="west" orientation="longitude" precision="5" value="-79.2344">79.2344 ° W</geoCoordinate>
</named-content>
), 
<quantity id="A41BCF266A91A70A7D1A32E34CA58FD3" metricMagnitude="2" metricUnit="m" metricValue="2.8" unit="m" value="280.0">
<elevation id="F7B581A1532F41D11F44C11E1AB1B7DC" metricMagnitude="2" metricUnit="m" metricValue="2.8" unit="m" value="280.0">280 m</elevation>
</quantity>
, collected by 
<collectorName id="A035475324B4A904C659775BB36FFC93">Giovanna Rodríguez</collectorName>
on 
<date id="968869F2CDAAF4A7D3F7C3B06EF5C00B" value="1996-01-01">
<collectingDate id="BEA874D9C3CAF5F31031AE26A5B7AFEB" value="1996-01-01">01 January, 1996</collectingDate>
</date>
and 
<date id="20F689FA1FC2CB7CDCD98F9489D90692" value="1995-08-01">
<collectingDate id="9570A4F3362E551B46E69971C260D420" value="1995-08-01">01 August, 1995</collectingDate>
</date>
</materialsCitation>
; • 
<materialsCitation id="ED1EE7C6415E681151354EC602911923" collectingDate="2004-04-08" collectionCode="QCAZ" collectorName="David Cannatella" country="Ecuador" elevation="149" latitude="0.1192" location="Aldea Salamandra" longLatPrecision="7" longitude="-79.2344" municipality="Canton Puerto Quito" specimenCode="QCAZ 10434" specimenCount="1" stateProvince="Pichincha" typeStatus="paratype">
<specimenCode id="B42EBE8DAAB081CAC5E9779195D93307" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" type="Museum">
<named-content id="B474E9EF0FDEF13AB7DBFE1716C90948" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
10434
</specimenCode>
, 27207–27209 
<collectingMunicipality id="F9F486A254EE9C00922590D33F71F0D4">Cantón Puerto Quito</collectingMunicipality>
, 
<location id="99612170DFF50BB066FC6D5437095D4A" LSID="urn:lsid:plazi:treatment:FA1177A074C859B19F69711C4654B852:99612170DFF50BB066FC6D5437095D4A" country="Ecuador" latitude="0.1192" longLatPrecision="7" longitude="-79.2344" municipality="Canton Puerto Quito" name="Aldea Salamandra" stateProvince="Pichincha">Aldea Salamandra</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ETUAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-79.234400,0.119200]}">
<geoCoordinate id="B8DC3A2F2EE186F5E283E41E532F1A2E" degrees="0.1192" direction="north" orientation="latitude" precision="5" value="0.1192">0.1192 ° N</geoCoordinate>
, 
<geoCoordinate id="A43F89F7C88A65CC7A29DFAC99CEEEE5" degrees="79.2344" direction="west" orientation="longitude" precision="5" value="-79.2344">79.2344 ° W</geoCoordinate>
</named-content>
), 
<quantity id="762D01C06113B0648C54D9E8A86B347B" metricMagnitude="2" metricUnit="m" metricValue="1.49" unit="m" value="149.0">
<elevation id="8E4E66EC2D6A1DC817250122E57FB9A3" metricMagnitude="2" metricUnit="m" metricValue="1.49" unit="m" value="149.0">149 m</elevation>
</quantity>
, collected by 
<collectorName id="E19A8A06813EDB684DDA971AF15091F1">David Cannatella</collectorName>
on 
<date id="01A62795AA2A91C21959524A16FB0F58" value="2004-04-08">
<collectingDate id="B31AB27AA5FA67DB141D4D9EE86A6C51" value="2004-04-08">08 April, 2004</collectingDate>
</date>
</materialsCitation>
; • 
<materialsCitation id="6E905DA444F7C4730A74956B2944C9FE" collectingDate="1990-03-30" collectionCode="QCAZ" collectorName="John J. Wiens" country="Ecuador" elevation="670" latitude="0.0829" location="Maldonado" longLatPrecision="7" longitude="-79.0383" municipality="Canton Pedro Vicente Maldonado" specimenCode="QCAZ 27623" specimenCount="1" stateProvince="Pichincha" typeStatus="paratype">
<specimenCode id="DE439EDBA287415DEE120E16FF93A5A7">
<named-content id="BF5994374A2E251BBCC98BE0CB22A262" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
27623
</specimenCode>
–27624 
<collectingMunicipality id="D0C0B57D631B7204489E821F0D83B3E6">Cantón Pedro Vicente Maldonado</collectingMunicipality>
, 
<locationDeviation id="90FD42FCA6D9115C77A829FC179A3923">
<quantity id="975D48B72B4FBCA41FEFFC6F12F2D2AF" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="km" value="1.0">1 km</quantity>
E from
</locationDeviation>
<location id="0D6F5ADB55EF903F98F082991E075BE9" LSID="urn:lsid:plazi:treatment:FA1177A074C859B19F69711C4654B852:0D6F5ADB55EF903F98F082991E075BE9" country="Ecuador" latitude="0.0829" longLatPrecision="7" longitude="-79.0383" municipality="Canton Pedro Vicente Maldonado" name="Maldonado" stateProvince="Pichincha">Maldonado</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E6UAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-79.038300,0.082900]}">
<geoCoordinate id="F6A12FD9A0BF9C6668377E1176CE650C" degrees="0.0829" direction="north" orientation="latitude" precision="5" value="0.0829">0.0829 ° N</geoCoordinate>
, 
<geoCoordinate id="32E3CE0B776C3717780AFAB8872A0936" degrees="79.0383" direction="west" orientation="longitude" precision="5" value="-79.0383">79.0383 ° W</geoCoordinate>
</named-content>
), 
<quantity id="624D0EF3814DE0E3E3A8793D127EAACB" metricMagnitude="2" metricUnit="m" metricValue="6.7" unit="m" value="670.0">
<elevation id="F5FBB08E4B8822C1DB4789721DABF4F3" metricMagnitude="2" metricUnit="m" metricValue="6.7" unit="m" value="670.0">670 m</elevation>
</quantity>
, collected by 
<collectorName id="6B1F077F7582BE1A5506BD41BA304C29">John J. Wiens</collectorName>
on 
<date id="51E64719742BFF4788DB1013B18102FA" value="1990-03-30">
<collectingDate id="716CBFBAF6B766FFA89B673CC60B33EF" value="1990-03-30">30 March, 1990</collectingDate>
</date>
</materialsCitation>
; • 
<materialsCitation id="D0A5945A40C6654EBB20D1A0CAF20B43" collectingDate="2023-01-19" collectionCode="QCAZ" collectorName="Gabriela B. Bittencourt" country="Ecuador" elevation="369" latitude="0.1278" location="Rio Silanche" longLatPrecision="7" longitude="-79.1414" municipality="Canton Pedro Vicente Maldonado" specimenCode="QCAZ 78254" specimenCount="1" stateProvince="Pichincha" typeStatus="paratype">
<specimenCode id="2D48CFF0D4CA2E267BA6E241996017E3">
<named-content id="D5A5F38A5CD1469B5013CD77DCE76366" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
78254
</specimenCode>
–78259, 78309, 78576 
<collectingMunicipality id="133121135CBC70638C2D5C2BF610A325">Cantón Pedro Vicente Maldonado</collectingMunicipality>
, 
<location id="8F2DE19D61400DCEE33F0533DBFE8A51" LSID="urn:lsid:plazi:treatment:FA1177A074C859B19F69711C4654B852:8F2DE19D61400DCEE33F0533DBFE8A51" country="Ecuador" latitude="0.1278" longLatPrecision="7" longitude="-79.1414" municipality="Canton Pedro Vicente Maldonado" name="Rio Silanche" stateProvince="Pichincha">Río Silanche</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ELVAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-79.141400,0.127800]}">
<geoCoordinate id="1BAB31B9045642B67832F79FA5C4FEB6" degrees="0.1278" direction="north" orientation="latitude" precision="5" value="0.1278">0.1278 ° N</geoCoordinate>
, 
<geoCoordinate id="98CBE398675A0C1DA3F0E81753815939" degrees="79.1414" direction="west" orientation="longitude" precision="5" value="-79.1414">79.1414 ° W</geoCoordinate>
</named-content>
), 
<quantity id="71D2E27D607ADD1290B5AFCBB089F864" metricMagnitude="2" metricUnit="m" metricValue="3.69" unit="m" value="369.0">
<elevation id="46CAEBF6B1119A6F8EF8AD41A89A292C" metricMagnitude="2" metricUnit="m" metricValue="3.69" unit="m" value="369.0">369 m</elevation>
</quantity>
, collected by 
<collectorName id="8B9A250E47543A20133BD8F306A7610D">Gabriela B. Bittencourt</collectorName>
on 
<date id="8487796675F9B687038867F0BD3AE52E" value="2023-01-19">
<collectingDate id="ADEBE2585E3D67B0001572D128CBE91B" value="2023-01-19">19 January, 2023</collectingDate>
</date>
</materialsCitation>
; • 
<materialsCitation id="FADC7BA0B3BC5541FEB68D3E8F6C8783" collectingDate="2023-07-17" collectionCode="QCAZ" collectorName="Santiago" country="Ecuador" county="Canton Pedro Vicente Maldonado" elevation="369" latitude="0.8003" location="Rio Silanche" longLatPrecision="7" longitude="-79.0709" municipality="Rio Silanche" specimenCode="QCAZ 78809" specimenCount="1" stateProvince="Pichincha" typeStatus="paratype">
<specimenCode id="262C89E50F9DB8684B853C4B18D519BD">
<named-content id="10BF07E2CC9AE3FD37F604890D205317" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
78809
</specimenCode>
–78814 
<collectingCounty id="E9AEF42C17DA818B9E30E693E3B5FA6D">Cantón Pedro Vicente Maldonado</collectingCounty>
, 
<location id="89E9CC583D748EEED1B74035AB985DB5" LSID="urn:lsid:plazi:treatment:FA1177A074C859B19F69711C4654B852:89E9CC583D748EEED1B74035AB985DB5" country="Ecuador" county="Canton Pedro Vicente Maldonado" latitude="0.8003" longLatPrecision="7" longitude="-79.0709" municipality="Rio Silanche" name="Rio Silanche" stateProvince="Esmeraldas">Río Silanche</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EXVAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-79.141400,0.127800]}">
<geoCoordinate id="C65AD64FBA354BE06AA89D1A05C4D5A5" degrees="0.1278" direction="north" orientation="latitude" precision="5" value="0.1278">0.1278 ° N</geoCoordinate>
, 
<geoCoordinate id="7DA2D605D0AE3C18A1A34F7A501E465D" degrees="79.1414" direction="west" orientation="longitude" precision="5" value="-79.1414">79.1414 ° W</geoCoordinate>
</named-content>
), 
<quantity id="A624EAD4C7B322CD40090B6DDBF6C2E7" metricMagnitude="2" metricUnit="m" metricValue="3.69" unit="m" value="369.0">
<elevation id="0A92F86FB05A7021FCC6CB53DCEEC6BD" metricMagnitude="2" metricUnit="m" metricValue="3.69" unit="m" value="369.0">369 m</elevation>
</quantity>
, collected by 
<collectorName id="8CA6C74912018ACBBB363E43ADE5A3DF">Santiago</collectorName>
R. Ron on 
<date id="4853B10899D51391EECAD0767F99F3F2" value="2023-07-17">
<collectingDate id="19877D4D196B2037E5354447B8E0EAAD" value="2023-07-17">17 July, 2023</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="22FBA712FA1A5AB833F8A6A9905F9636" collectingDate="1997-01-18" collectorName="Nestor Acosta" country="Ecuador" elevation="110" latitude="0.8003" location="Santo Domingo de Onzole" longLatPrecision="7" longitude="-79.0709" specimenCount="1" stateProvince="Esmeraldas" typeStatus="paratype">
<collectingRegion id="8EB1C9F7F7751D00C31D747407267C0C" country="Ecuador" name="Esmeraldas">Esmeraldas Province</collectingRegion>
: 
<location id="B362E640CF863A1880E3672B3EF70400" LSID="urn:lsid:plazi:treatment:FA1177A074C859B19F69711C4654B852:B362E640CF863A1880E3672B3EF70400" latitude="0.8003" longLatPrecision="7" longitude="-79.0709" name="Santo Domingo de Onzole" stateProvince="Esmeraldas Province">Santo Domingo de Onzole</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E6VAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-79.070900,0.800300]}">
<geoCoordinate id="E02C6DE75225060735EEE79F9D49AAFC" degrees="0.8003" direction="north" orientation="latitude" precision="5" value="0.8003">0.8003 ° N</geoCoordinate>
, 
<geoCoordinate id="FD74D74164818391E4E6E2A058DEE1ED" degrees="79.0709" direction="west" orientation="longitude" precision="5" value="-79.0709">79.0709 ° W</geoCoordinate>
</named-content>
), 
<quantity id="04DA71BE0059CC159AEBB8FDB33C2B65" metricMagnitude="2" metricUnit="m" metricValue="1.1" unit="m" value="110.0">
<elevation id="8DDF62EE5FC9485EB214184F41A78BF5" metricMagnitude="2" metricUnit="m" metricValue="1.1" unit="m" value="110.0">110 m</elevation>
</quantity>
, collected by 
<collectorName id="1E2E82879A3D564CAC72AAB0D8A81B7E">Néstor Acosta</collectorName>
on 
<date id="A99721557CAE810E0831D8939BD817D6" value="1997-01-18">
<collectingDate id="BCFE8424A341B901C78422DA1D8BC3ED" value="1997-01-18">18 January, 1997</collectingDate>
</date>
</materialsCitation>
; • 
<materialsCitation id="0451AD50151E432A8E8F64162165C430" collectingDate="1990-03-31" collectingDateMax="1993-03-13" collectingDateMin="1990-03-31" collectionCode="QCAZ" collectorName="Luis Coloma &amp; Sebastian Valdiviezo" country="Ecuador" elevation="670" latitude="0.0878" location="Canton Pedro Vicente Maldonado" longLatPrecision="7" longitude="-79.0491" specimenCode="QCAZ 10704" specimenCount="1" stateProvince="Esmeraldas" typeStatus="paratype">
<specimenCode id="7336C0F04AD4B9DE461406124ACE0D8C" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" type="Museum">
<named-content id="65681FBDE10FFB206C12C9F4456BE181" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
10704
</specimenCode>
, 22893, 
<location id="377A78C47536AAFFD73855096D5988E1" LSID="urn:lsid:plazi:treatment:FA1177A074C859B19F69711C4654B852:377A78C47536AAFFD73855096D5988E1" country="Ecuador" latitude="0.0878" longLatPrecision="7" longitude="-79.0491" name="Canton Pedro Vicente Maldonado" stateProvince="Esmeraldas">Cantón Pedro Vicente Maldonado</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ELWAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-79.049100,0.087800]}">
<geoCoordinate id="15059E2FCDF3871C127910C9DA4D5FB1" degrees="0.0878" direction="north" orientation="latitude" precision="5" value="0.0878">0.0878 ° N</geoCoordinate>
, 
<geoCoordinate id="772B238E188D274369BF9E33395702E4" degrees="79.0491" direction="west" orientation="longitude" precision="5" value="-79.0491">79.0491 ° W</geoCoordinate>
</named-content>
), 
<quantity id="3680BAC95BC48D5948BFE03F6EFC516A" metricMagnitude="2" metricUnit="m" metricValue="6.7" unit="m" value="670.0">
<elevation id="4646E644184E05189E3D0A4686AA667E" metricMagnitude="2" metricUnit="m" metricValue="6.7" unit="m" value="670.0">670 m</elevation>
</quantity>
, collected by 
<collectorName id="D01ACB4689842B3BA9FCDFA88E81C676">Luis Coloma</collectorName>
on 
<date id="511E96EE6F264FD10D1426E4AA0B253B" value="1993-03-13">
<collectingDate id="F730A7E3AE44E08DD444F938644E558E" value="1993-03-13">13 March, 1993</collectingDate>
</date>
and 
<collectorName id="35405C719865C84FB133EE8A269458BE">Sebastián Valdiviezo</collectorName>
on 
<date id="206CAB8D5DE41329CE2D083EE5394F8A" value="1990-03-31">
<collectingDate id="198D578E269ED9976498124A916A6F6E" value="1990-03-31">31 March, 1990</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="2D4A7E1EC61C91CCBEC46C2D9CBE9CFF" collectingDate="2001-06-15" collectingDateMax="2003-03-03" collectingDateMin="2001-06-15" collectionCode="QCAZ" collectorName="Italo Tapia &amp; David Cannatella" country="Ecuador" elevation="158" latitude="1.0643" location="Durango" longLatPrecision="7" longitude="-78.6443" municipality="Canton San Lorenzo" specimenCode="QCAZ 15519" specimenCount="1" stateProvince="Esmeraldas" typeStatus="paratype">
<collectingRegion id="FA4A670983E458133F2AF55069758B45" country="Ecuador" name="Esmeraldas">Esmeraldas Province</collectingRegion>
: • 
<specimenCode id="E851BAC78537779C66A7BD2FCD5896AD" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" type="Museum">
<named-content id="6C8D1E3D555F1AA086960108B5A82DE4" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
15519
</specimenCode>
, 23156, 23160, 23371, 
<collectingMunicipality id="178CC3A03475D7E31456307CD5AA0DEE">Cantón San Lorenzo</collectingMunicipality>
, 
<locationDeviation id="8BFF95FD2622E80F3D612A3A8BA7AEE9">
<quantity id="C451D0B10F9186742B67A6BBA8552E50" metricMagnitude="3" metricUnit="m" metricValue="3.0" unit="km" value="3.0">3 km</quantity>
west of
</locationDeviation>
<location id="A1D9116C4DE0120A0D5BCFCE462750E8" LSID="urn:lsid:plazi:treatment:FA1177A074C859B19F69711C4654B852:A1D9116C4DE0120A0D5BCFCE462750E8" country="Ecuador" latitude="1.0643" longLatPrecision="7" longitude="-78.6443" municipality="Canton San Lorenzo" name="Durango" stateProvince="Esmeraldas Province">Durango</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EXWAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-78.644300,1.064300]}">
<geoCoordinate id="742F7D57DB8066D36C176C8CE1A304CC" degrees="1.0643" direction="north" orientation="latitude" precision="5" value="1.0643">1.0643 ° N</geoCoordinate>
, 
<geoCoordinate id="A81A9D6607712D19AD35FC8C755EF2D1" degrees="78.6443" direction="west" orientation="longitude" precision="5" value="-78.6443">78.6443 ° W</geoCoordinate>
</named-content>
), 
<quantity id="06411B6607819F6E78360E47E1C306F2" metricMagnitude="2" metricUnit="m" metricValue="1.58" unit="m" value="158.0">
<elevation id="C1C4C3AC6CC4134E7DB5F4573D75BC79" metricMagnitude="2" metricUnit="m" metricValue="1.58" unit="m" value="158.0">158 m</elevation>
</quantity>
, collected by 
<collectorName id="A11DD84B5455189FF910CAE33E49D37B">Ítalo Tapia</collectorName>
on 
<date id="2E0F6AC0B0E837063B3C491FC5B71ADA" value="2001-06-15">
<collectingDate id="9BDCB47A58469F7538E2D677AD8CF943" value="2001-06-15">15 June, 2001</collectingDate>
</date>
and 
<collectorName id="1ABE4E581ABE81F0206ECFC388123D08">David Cannatella</collectorName>
on 
<date id="305A83FBB0C51644175C83A1647E0E7F" value="2003-03-03">
<collectingDate id="C59804F5DFB481AAA4CFAB7BB8761AF5" value="2003-03-03">03 March, 2003</collectingDate>
</date>
</materialsCitation>
; • 
<materialsCitation id="0701035350031EBA836313A7E2B5948D" collectingDate="2003-03-04" collectionCode="QCAZ" collectorName="David Cannatella" country="Ecuador" elevation="14" latitude="1.0127" location="Durango" longLatPrecision="7" longitude="-78.8954" municipality="Canton San Lorenzo" specimenCode="QCAZ 23162" specimenCount="1" stateProvince="Esmeraldas" typeStatus="paratype">
<specimenCode id="8E784FE034341F2F7A2378F5F9EAE18A" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" type="Museum">
<named-content id="F95A6A5BA1015C5E1ABB2350BFCAE5CE" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
23162
</specimenCode>
<collectingMunicipality id="CD7FAAADD5133C3079C8840FE55C742A">Cantón San Lorenzo</collectingMunicipality>
, 
<location id="B6F42DA8ADC2FFDD6F8DBAF06DF47FFB" LSID="urn:lsid:plazi:treatment:FA1177A074C859B19F69711C4654B852:B6F42DA8ADC2FFDD6F8DBAF06DF47FFB" country="Ecuador" latitude="1.0127" longLatPrecision="7" longitude="-78.8954" municipality="Canton San Lorenzo" name="Durango" stateProvince="Esmeraldas">Durango</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EDXAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-78.895400,1.012700]}">
<geoCoordinate id="E308216144CE3FD505689A931BBF6110" degrees="1.0127" direction="north" orientation="latitude" precision="5" value="1.0127">1.0127 ° N</geoCoordinate>
, 
<geoCoordinate id="BCFC44D5E7392082DE4C2CB701321A54" degrees="78.8954" direction="west" orientation="longitude" precision="5" value="-78.8954">78.8954 ° W</geoCoordinate>
</named-content>
), 
<quantity id="53488F3A84135BF53DDD33B81120461B" metricMagnitude="1" metricUnit="m" metricValue="1.4" unit="m" value="14.0">
<elevation id="A0E6C2664FDC230EA90236DFAF28A5C3" metricMagnitude="1" metricUnit="m" metricValue="1.4" unit="m" value="14.0">14 m</elevation>
</quantity>
, collected by 
<collectorName id="30D6DA26A8B450875263FA236092119A">David Cannatella</collectorName>
on 
<date id="C435B47B1284A25F0CFA7C63CE1066C6" value="2003-03-04">
<collectingDate id="044433D13C8A600F982CA600C89F65C5" value="2003-03-04">04 March, 2003</collectingDate>
</date>
</materialsCitation>
; • 
<materialsCitation id="72E79414E1468BED75A2D9C4942BF799" collectingDate="2003-06-08" collectionCode="QCAZ" collectorName="Nestor Acosta" country="Ecuador" elevation="74" latitude="1.0793" location="Durango - San Lorenzo" longLatPrecision="7" longitude="-78.6695" municipality="Canton San Lorenzo" specimenCode="QCAZ 26098" specimenCount="1" stateProvince="Esmeraldas" typeStatus="paratype">
<specimenCode id="BE2AE66165F609B746A24159A9FB22EB" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" type="Museum">
<named-content id="45A16530B21957F18C02C03C539E9B44" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
26098
</specimenCode>
, 26100 
<collectingMunicipality id="860150D7673A0A94B46F7970EB786247">Cantón San Lorenzo</collectingMunicipality>
, 
<location id="62DA73217D37A22BBB37BC1FD7340A7F" LSID="urn:lsid:plazi:treatment:FA1177A074C859B19F69711C4654B852:62DA73217D37A22BBB37BC1FD7340A7F" country="Ecuador" latitude="1.0793" longLatPrecision="7" longitude="-78.6695" municipality="Canton San Lorenzo" name="Durango - San Lorenzo" stateProvince="Esmeraldas">Durango – San Lorenzo</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EPXAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-78.669500,1.079300]}">
<geoCoordinate id="136F47179DCB94AFCC59F66D17168508" degrees="1.0793" direction="north" orientation="latitude" precision="5" value="1.0793">1.0793 ° N</geoCoordinate>
, 
<geoCoordinate id="A0E66ED2EC0D28F3D0D9A1EA35A97947" degrees="78.6695" direction="west" orientation="longitude" precision="5" value="-78.6695">78.6695 ° W</geoCoordinate>
</named-content>
), 
<quantity id="3A6613D78F516B38AE5E2BF0EBD05344" metricMagnitude="1" metricUnit="m" metricValue="7.4" unit="m" value="74.0">
<elevation id="774CC6445567125419338C7818214A8B" metricMagnitude="1" metricUnit="m" metricValue="7.4" unit="m" value="74.0">74 m</elevation>
</quantity>
, collected by 
<collectorName id="3B5613CDC627772DAF0B93E2BB3AEC52">Néstor Acosta</collectorName>
on 
<date id="98E50D441F0870B90F200A41310E6CAE" value="2003-06-08">
<collectingDate id="EE10AA3756F18CAAB40E2C3BE7A6AE98" value="2003-06-08">08 June, 2003</collectingDate>
</date>
</materialsCitation>
; • 
<materialsCitation id="D801EECF32B1AC429FB8A78D9C80C600" collectingDate="2004-12-19" collectingDateMax="2013-04-20" collectingDateMin="2004-12-19" collectionCode="QCAZ" collectorName="Italo Tapia &amp; Elicio Tapia &amp; Justin Touchon" country="Ecuador" elevation="85" latitude="1.0427" location="Durango" longLatPrecision="7" longitude="-78.6244" municipality="Canton San Lorenzo" specimenCode="QCAZ 29489" specimenCount="1" stateProvince="Esmeraldas" typeStatus="paratype">
<specimenCode id="17309E04A5CA6A497FB44E441D431C75">
<named-content id="EC0F766E4B829F937D4092E0F8D05697" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
29489
</specimenCode>
–29511, 30329–30330, 55529–55530 
<collectingMunicipality id="8EAA06DBAE0D0B259EE54D3147167E43">Cantón San Lorenzo</collectingMunicipality>
, 
<location id="0B91C7169A518EB3EAF5870F48A56FED" LSID="urn:lsid:plazi:treatment:FA1177A074C859B19F69711C4654B852:0B91C7169A518EB3EAF5870F48A56FED" country="Ecuador" latitude="1.0427" longLatPrecision="7" longitude="-78.6244" municipality="Canton San Lorenzo" name="Durango" stateProvince="Esmeraldas">Durango</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E2XAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-78.624400,1.042700]}">
<geoCoordinate id="EA2E2BB8267C86B5E82CB068C876F7BD" degrees="1.0427" direction="north" orientation="latitude" precision="5" value="1.0427">1.0427 ° N</geoCoordinate>
, 
<geoCoordinate id="612821A68C0585CD9CF38ED1FF230810" degrees="78.6244" direction="west" orientation="longitude" precision="5" value="-78.6244">78.6244 ° W</geoCoordinate>
</named-content>
), 
<quantity id="D229C9516266D6BE61B2F02ED9907300" metricMagnitude="1" metricUnit="m" metricValue="8.5" unit="m" value="85.0">
<elevation id="8F9E5B19B1C0C05ECF360A05D46195C5" metricMagnitude="1" metricUnit="m" metricValue="8.5" unit="m" value="85.0">85 m</elevation>
</quantity>
, collected by 
<collectorName id="CD69B3FD040BFAF7A1490D796BA441BB">Ítalo Tapia</collectorName>
on 
<date id="F34A4D3A0243A9359839FF6787E7699A" value="2005-02-02">
<collectingDate id="80267FF5999FDFBC4FEA977C1E5C0A05" value="2005-02-02">02 February, 2005</collectingDate>
</date>
, 
<collectorName id="8FEFF4A688E3DEECC2409F55623E4476">Elicio Tapia</collectorName>
on 
<date id="0C6292E0E7DF331523CDB779290321B4" value="2004-12-19">
<collectingDate id="7E6BF5E4BE0795DD84D2E4A6B886F794" value="2004-12-19">19 December, 2004</collectingDate>
</date>
and 
<collectorName id="BD641F9BC37AE2CB7CDCA65FB66CA86C">Justin Touchon</collectorName>
on 
<date id="49F234835E8F1A4A8845CEF4BCCA8663" value="2013-04-20">
<collectingDate id="F718219ABDDCD2BECCDA5DD661A98D82" value="2013-04-20">20 April, 2013</collectingDate>
</date>
</materialsCitation>
; • 
<materialsCitation id="5D59D512FF187A87FAF987C2C51A195D" collectingDate="2004-09-17" collectionCode="QCAZ" collectorName="Italo Tapia" country="Ecuador" latitude="1.0872" location="Durango - San Francisco" longLatPrecision="7" longitude="-78.6905" municipality="Canton San Lorenzo" specimenCode="QCAZ 29799" specimenCount="1" stateProvince="Esmeraldas" typeStatus="paratype">
<specimenCode id="7C5F03E476F138D193558ACA03915B9B">
<named-content id="5ABF1B402BF782AC55FE1EAD6E51356F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
29799
</specimenCode>
–29809, 30255 
<collectingMunicipality id="81B1BB90D3E4B503AEA3C7D792359EBC">Cantón San Lorenzo</collectingMunicipality>
, 
<location id="1CE93DFAC97E869F1E40468E8B99DB07" LSID="urn:lsid:plazi:treatment:FA1177A074C859B19F69711C4654B852:1CE93DFAC97E869F1E40468E8B99DB07" country="Ecuador" latitude="1.0872" longLatPrecision="7" longitude="-78.6905" municipality="Canton San Lorenzo" name="Durango - San Francisco" stateProvince="Esmeraldas">Durango – San Francisco</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EHYAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-78.690500,1.087200]}">
<geoCoordinate id="1A50D2B4867EBAB8692F16D850572179" degrees="1.0872" direction="north" orientation="latitude" precision="5" value="1.0872">1.0872 ° N</geoCoordinate>
, 
<geoCoordinate id="5032FD2D8183A84E26B5A0CDA8822235" degrees="78.6905" direction="west" orientation="longitude" precision="5" value="-78.6905">78.6905 ° W</geoCoordinate>
</named-content>
), collected by 
<collectorName id="37B74D3AA87D56BEE468AA3FCD5DEDB5">Ítalo Tapia</collectorName>
on 
<date id="BE6201582426CCAC233D196D33D185E5" value="2004-09-17">
<collectingDate id="8458CE3604E57962F9B44FEFB74EE241" value="2004-09-17">17 September, 2004</collectingDate>
</date>
</materialsCitation>
; • 
<materialsCitation id="BFCBA7AB471B50CE81E09824A46DE0D0" collectingDate="2006-05-26" collectionCode="QCAZ" collectorName="Diego Almeida" country="Ecuador" elevation="85" latitude="1.0888" location="San Francisco" longLatPrecision="5" longitude="-78.69563" specimenCode="QCAZ 32136" specimenCount="1" stateProvince="Esmeraldas" typeStatus="paratype">
<specimenCode id="8EF0C0887C0EBFA4E81EBB7894306FE0" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" type="Museum">
<named-content id="A5B2884EDC39FC6DF421F15756DCCDEF" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
32136
</specimenCode>
<location id="52017D2362F816EFC0E0FDF7CF13801D" LSID="urn:lsid:plazi:treatment:FA1177A074C859B19F69711C4654B852:52017D2362F816EFC0E0FDF7CF13801D" country="Ecuador" latitude="1.0888" longLatPrecision="5" longitude="-78.69563" name="San Francisco" stateProvince="Esmeraldas">San Francisco</location>
, 
<quantity id="48C7FD091124C8E98A0C6E6B4E0754C2" metricMagnitude="3" metricUnit="m" metricValue="1.3" unit="km" value="1.3">
<locationDeviation id="FE2B45546F3478465A6033080199839C" metricMagnitude="3" metricUnit="m" metricValue="1.3" unit="km" value="1.3">1.3 km</locationDeviation>
</quantity>
east (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ETYAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-78.695630,1.088800]}">
<geoCoordinate id="CEFFE643F1CF7F058A7142AB0AE40AB5" degrees="1.0888" direction="north" orientation="latitude" precision="5" value="1.0888">1.0888 ° N</geoCoordinate>
, 
<geoCoordinate id="FC1D67E85D8DE6EA54AC4667DEE1F095" degrees="78.69563" direction="west" orientation="longitude" precision="1" value="-78.69563">78.69563 ° W</geoCoordinate>
</named-content>
), 
<quantity id="D4932193E0F97B74F5D2B59FAB57B497" metricMagnitude="1" metricUnit="m" metricValue="8.5" unit="m" value="85.0">
<elevation id="7106C48B1EA1EB167EDA6FAF34064372" metricMagnitude="1" metricUnit="m" metricValue="8.5" unit="m" value="85.0">85 m</elevation>
</quantity>
, collected by 
<collectorName id="BAF9A1D0A6BBD3FC8737B9E1389BD909">Diego Almeida</collectorName>
on 
<date id="C390B735183646E7E75F521366E7FA51" value="2006-05-26">
<collectingDate id="D1756B12C91434C8D2B8251D0B663EAD" value="2006-05-26">26 May, 2006</collectingDate>
</date>
</materialsCitation>
; • 
<materialsCitation id="EEC8B3A262B2590C1983DBF36222FAB7" collectingDate="1998-11-27" collectionCode="QCAZ" collectorName="La Mayronga &amp; November" country="Ecuador" elevation="150" latitude="1.05" location="La Mayronga" longLatPrecision="785" longitude="-79.27" specimenCode="QCAZ 35709" specimenCount="1" stateProvince="Esmeraldas" typeStatus="paratype">
<specimenCode id="684897A8F6A6C5B88BF20066ADB97EB4">
<named-content id="94D54DAE30C9CC27D7B5736615E55AC9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
35709
</specimenCode>
–35710 
<location id="A0A7CB1E2EB6F54F5176FCE1AE4FAD13" LSID="urn:lsid:plazi:treatment:FA1177A074C859B19F69711C4654B852:A0A7CB1E2EB6F54F5176FCE1AE4FAD13" country="Ecuador" latitude="1.05" longLatPrecision="785" longitude="-79.27" name="La Mayronga" stateProvince="Esmeraldas">La Mayronga</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E6YAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-79.270000,1.050000]}">
<geoCoordinate id="733F4910C0CE226A70DD16DC5550C7FD" degrees="1.05" direction="north" orientation="latitude" precision="555" value="1.05">1.05 ° N</geoCoordinate>
, 
<geoCoordinate id="5889ADE5D5F567A576958BA8C7C8F730" degrees="79.27" direction="west" orientation="longitude" precision="555" value="-79.27">79.27 ° W</geoCoordinate>
</named-content>
), 
<quantity id="DB3B95C134776D71711E433532BB6F94" metricMagnitude="2" metricUnit="m" metricValue="1.5" unit="m" value="150.0">
<elevation id="9119307B58B9770A4B652EA29A7C663B" metricMagnitude="2" metricUnit="m" metricValue="1.5" unit="m" value="150.0">150 m</elevation>
</quantity>
, collected on 
<date id="FEFBCB3697F6E408621C347F85A340DF" value="1998-11-27">
<collectingDate id="4E5A57996D59DCC2A5614ED851A50598" value="1998-11-27">27 November, 1998</collectingDate>
</date>
</materialsCitation>
; • 
<materialsCitation id="7BC2BA9FE781FD2B4A252DFF985149EE" collectingDate="2008-01-03" collectionCode="QCAZ" collectorName="Italo Tapia" country="Ecuador" elevation="284" latitude="1.0248" location="Durango" longLatPrecision="7" longitude="-78.6175" municipality="Canton San Lorenzo" specimenCode="QCAZ 37741" specimenCount="1" stateProvince="Esmeraldas" typeStatus="paratype">
<specimenCode id="FF70B3FFC17412AC779ADCA931CE4D54" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" type="Museum">
<named-content id="6F06B2DAF1CDE8CAE47AC0D2A2742354" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
37741
</specimenCode>
, 37845 
<collectingMunicipality id="FADC24BAD16F48447AF9499D3481A2C1">Cantón San Lorenzo</collectingMunicipality>
, 
<locationDeviation id="6AD5ECF3148A0A6E5F1B2DAB5EC18387">
<quantity id="912D15B832284C39D96E7BFAA598A078" metricMagnitude="4" metricUnit="m" metricValue="2.1" unit="km" value="21.0">21 km</quantity>
of
</locationDeviation>
<location id="9964D26C963422BC76D2B0DEA6053EF8" LSID="urn:lsid:plazi:treatment:FA1177A074C859B19F69711C4654B852:9964D26C963422BC76D2B0DEA6053EF8" country="Ecuador" latitude="1.0248" longLatPrecision="7" longitude="-78.6175" municipality="Canton San Lorenzo" name="Durango" stateProvince="Esmeraldas">Durango</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ELZAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-78.617500,1.024800]}">
<geoCoordinate id="F0828109C83400F1D78B0C371F694685" degrees="1.0248" direction="north" orientation="latitude" precision="5" value="1.0248">1.0248 ° N</geoCoordinate>
, 
<geoCoordinate id="2A4C082A1BBD0F6810BA41FD939797A0" degrees="78.6175" direction="west" orientation="longitude" precision="5" value="-78.6175">78.6175 ° W</geoCoordinate>
</named-content>
), 
<quantity id="5F0BFDE929D061A6D22580941CA3FA1C" metricMagnitude="2" metricUnit="m" metricValue="2.84" unit="m" value="284.0">
<elevation id="BE26D9D2FFBA398A88196F24B3330FD0" metricMagnitude="2" metricUnit="m" metricValue="2.84" unit="m" value="284.0">284 m</elevation>
</quantity>
, collected by 
<collectorName id="0BDD14C129B19B032D8C23D76E984ADE">Ítalo Tapia</collectorName>
on 
<date id="0BFE961878F325A455779DD55BAB4F6F" value="2008-01-03">
<collectingDate id="5D4BA5FEAE1C8CC9E79D4ADEF0EDA3D7" value="2008-01-03">03 January, 2008</collectingDate>
</date>
</materialsCitation>
; • 
<materialsCitation id="E5DADCD2C62704FEC8577C2B4668DE52" collectingDate="2007-12-15" collectionCode="QCAZ" collectorName="Italo Tapia" country="Ecuador" elevation="104" latitude="1.047" location="Durango" longLatPrecision="7" longitude="-78.6182" municipality="Canton San Lorenzo" specimenCode="QCAZ 37749" specimenCount="1" stateProvince="Esmeraldas" typeStatus="paratype">
<specimenCode id="0BA455DFAAA642D10D6F4EAAA9998CC1" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" type="Museum">
<named-content id="ACF088282851C34307851F03FA623D60" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
37749
</specimenCode>
<collectingMunicipality id="E8E35116524DFF9641F668311A3BEBA3">Cantón San Lorenzo</collectingMunicipality>
, 
<locationDeviation id="68214B66D76925D050014A13234C36E2">
<quantity id="9A8B2BF3D5149C4D1D4DB46EB41F815E" metricMagnitude="4" metricUnit="m" metricValue="2.1" unit="km" value="21.0">21 km</quantity>
of
</locationDeviation>
<location id="62621052116DCB4D861D831389E8F8D8" LSID="urn:lsid:plazi:treatment:FA1177A074C859B19F69711C4654B852:62621052116DCB4D861D831389E8F8D8" country="Ecuador" latitude="1.047" longLatPrecision="7" longitude="-78.6182" municipality="Canton San Lorenzo" name="Durango" stateProvince="Esmeraldas">Durango</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EXZAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-78.618200,1.047000]}">
<geoCoordinate id="788D388602187FF1E0D8CCC82FA20050" degrees="1.0470" direction="north" orientation="latitude" precision="5" value="1.047">1.0470 ° N</geoCoordinate>
, 
<geoCoordinate id="88F578B1BB8E53CB7794FF680C26D4F9" degrees="78.6182" direction="west" orientation="longitude" precision="5" value="-78.6182">78.6182 ° W</geoCoordinate>
</named-content>
), 
<quantity id="7DD8D4831D93B48B1A70E5ECD2646047" metricMagnitude="2" metricUnit="m" metricValue="1.04" unit="m" value="104.0">
<elevation id="8FA74DA88415D4C09F2E0D041A929121" metricMagnitude="2" metricUnit="m" metricValue="1.04" unit="m" value="104.0">104 m</elevation>
</quantity>
, collected by 
<collectorName id="27F12467AA9157FCAAA2E2BFBDA78C3B">Ítalo Tapia</collectorName>
on 
<date id="37340B64701BE91A81289B6A9FCC8BCC" value="2007-12-15">
<collectingDate id="2957B3647CA0324857D1B7458B4421B2" value="2007-12-15">15 December, 2007</collectingDate>
</date>
</materialsCitation>
; • 
<materialsCitation id="DA84E8402AD9B477FE32CEC791610BCC" collectingDate="2009-03-13" collectionCode="QCAZ" collectorName="Italo Tapia" country="Ecuador" elevation="85" latitude="1.0247" location="Durango" longLatPrecision="7" longitude="-78.6174" municipality="Canton San Lorenzo" specimenCode="QCAZ 40276" specimenCount="1" stateProvince="Esmeraldas" typeStatus="paratype">
<specimenCode id="4177F4E354420BD4D3A97B34C491D00F" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" type="Museum">
<named-content id="5FEB13B16C22CA5E2164CD2C8969EA9A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
40276
</specimenCode>
, 40279 
<collectingMunicipality id="311AE4AEC5E8EE3783E392B3CD5C0ACF">Cantón San Lorenzo</collectingMunicipality>
, 
<locationDeviation id="B7DC8334C43022479B5AE014A8A2EEBD">
<quantity id="BF2F557BC882B7868AEDF2851282C0BD" metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="km" value="2.0">2 km</quantity>
east of
</locationDeviation>
<location id="84225D1B026E3D44592969FFF3054246" LSID="urn:lsid:plazi:treatment:FA1177A074C859B19F69711C4654B852:84225D1B026E3D44592969FFF3054246" country="Ecuador" latitude="1.0247" longLatPrecision="7" longitude="-78.6174" municipality="Canton San Lorenzo" name="Durango" stateProvince="Esmeraldas">Durango</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ED1AI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-78.617400,1.024700]}">
<geoCoordinate id="D344591625E66348E5BD5436AABB72C9" degrees="1.0247" direction="north" orientation="latitude" precision="5" value="1.0247">1.0247 ° N</geoCoordinate>
, 
<geoCoordinate id="C0911BB4F1BFCACDF75254D4E2FCC2A1" degrees="78.6174" direction="west" orientation="longitude" precision="5" value="-78.6174">78.6174 ° W</geoCoordinate>
</named-content>
), 
<quantity id="F0AD5599E083E30A03C42A911A21269F" metricMagnitude="1" metricUnit="m" metricValue="8.5" unit="m" value="85.0">
<elevation id="11DCFA0EA9ABB585042C7C888D96D84D" metricMagnitude="1" metricUnit="m" metricValue="8.5" unit="m" value="85.0">85 m</elevation>
</quantity>
, collected by 
<collectorName id="EA135D786E22F19AB38A92735B4FB933">Ítalo Tapia</collectorName>
on 
<date id="06D06E1F1ADCC92E29CED8CC5942D32D" value="2009-03-13">
<collectingDate id="AE3A036CBED8B47739985D756DA37315" value="2009-03-13">13 March, 2009</collectingDate>
</date>
</materialsCitation>
; • 
<materialsCitation id="80C46B1A3AC179119E045C4F3719A61F" collectingDate="2011-09-10" collectionCode="QCAZ" collectorName="Andres Merino" country="Ecuador" elevation="283" latitude="1.0379" location="Durango" longLatPrecision="7" longitude="-78.6222" municipality="Canton San Lorenzo" specimenCode="QCAZ 51854" specimenCount="1" stateProvince="Esmeraldas" typeStatus="paratype">
<specimenCode id="8C0D048E927B63F0FFB0FEB96A9BF002" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" type="Museum">
<named-content id="DA7E29AD0F07BE9CC5754CAFD3D82A53" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
51854
</specimenCode>
, 51863, 51877 
<collectingMunicipality id="AA7B3C84E625200CC64FDB62953F364D">Cantón San Lorenzo</collectingMunicipality>
, 
<location id="7534BD624F285DCB8D85BD91EFCAA8A9" LSID="urn:lsid:plazi:treatment:FA1177A074C859B19F69711C4654B852:7534BD624F285DCB8D85BD91EFCAA8A9" country="Ecuador" latitude="1.0379" longLatPrecision="7" longitude="-78.6222" municipality="Canton San Lorenzo" name="Durango" stateProvince="Esmeraldas">Durango</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EP1AI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-78.622200,1.037900]}">
<geoCoordinate id="C65F3278E8D3E9BAAF682611A0CCD6A8" degrees="1.0379" direction="north" orientation="latitude" precision="5" value="1.0379">1.0379 ° N</geoCoordinate>
, 
<geoCoordinate id="A11C0BD68ACF44686C687DF3E192C643" degrees="78.6222" direction="west" orientation="longitude" precision="5" value="-78.6222">78.6222 ° W</geoCoordinate>
</named-content>
), 
<quantity id="16D9D5DF4AAABAC6AE91B5FABF5CA064" metricMagnitude="2" metricUnit="m" metricValue="2.83" unit="m" value="283.0">
<elevation id="2B155C0040FE69347653F77C00ACA4C3" metricMagnitude="2" metricUnit="m" metricValue="2.83" unit="m" value="283.0">283 m</elevation>
</quantity>
, collected by 
<collectorName id="8170599813E00D00A0797A6177B80DF2">Andrés Merino</collectorName>
on 
<date id="F2623F34B39131F86591D31030191529" value="2011-09-10">
<collectingDate id="7A4CE246C2165B54EC4ACB7618EF2DBA" value="2011-09-10">10 September, 2011</collectingDate>
</date>
</materialsCitation>
; • 
<materialsCitation id="F28800C6F15AB84F99AC8BE3E6CA90B6" collectingDate="2013-04-26" collectionCode="QCAZ" collectorName="Justin Touchon" country="Ecuador" elevation="91" latitude="1.2083" location="Durango" longLatPrecision="7" longitude="-78.7413" municipality="Canton San Lorenzo" specimenCode="QCAZ 55531" specimenCount="1" stateProvince="Esmeraldas" typeStatus="paratype">
<specimenCode id="9AD4C084248606B58C6F63722972E8A5">
<named-content id="7FAC6C4A25DABC3979661F4FCC879837" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
55531
</specimenCode>
–540, 55545–55550, 55555–55558 
<collectingMunicipality id="906109B8EDF7BBA1E733D139FBED0023">Cantón San Lorenzo</collectingMunicipality>
, 
<locationDeviation id="C1154C167B56C1C2E7CE3381997554F6">
<quantity id="70D622A0B5344B17128FFAEC9DFEA8CA" metricMagnitude="3" metricUnit="m" metricValue="9.0" unit="km" value="9.0">9 km</quantity>
west of
</locationDeviation>
<location id="AE2F84A9D96244821895D1D1896320E6" LSID="urn:lsid:plazi:treatment:FA1177A074C859B19F69711C4654B852:AE2F84A9D96244821895D1D1896320E6" country="Ecuador" latitude="1.2083" longLatPrecision="7" longitude="-78.7413" municipality="Canton San Lorenzo" name="Durango" stateProvince="Esmeraldas">Durango</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E21AI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-78.741300,1.208300]}">
<geoCoordinate id="B721D8962B5FD1D808E6F694DDF98FB1" degrees="1.2083" direction="north" orientation="latitude" precision="5" value="1.2083">1.2083 ° N</geoCoordinate>
, 
<geoCoordinate id="853750782E32FC822AAFF1EB2E7E32A0" degrees="78.7413" direction="west" orientation="longitude" precision="5" value="-78.7413">78.7413 ° W</geoCoordinate>
</named-content>
), 
<quantity id="3069FD1A1721C7720BD03F7BA8749387" metricMagnitude="1" metricUnit="m" metricValue="9.1" unit="m" value="91.0">
<elevation id="1E853DB765A8946B946E7F97D0023FC5" metricMagnitude="1" metricUnit="m" metricValue="9.1" unit="m" value="91.0">91 m</elevation>
</quantity>
, collected by 
<collectorName id="3C4F9BB27B12C5341B00FC8E0B9E6BD4">Justin Touchon</collectorName>
on 
<date id="459FC03E706CE3473EAEEB6E24388DBF" value="2013-04-26">
<collectingDate id="B5E8AF5DA109ED499B8F66B3EE733BB9" value="2013-04-26">26 April, 2013</collectingDate>
</date>
</materialsCitation>
; • 
<materialsCitation id="743432713FC7DF08A47B6AE45A4BA9F4" collectingDate="2013-04-22" collectionCode="QCAZ" collectorName="Justin Touchon" country="Ecuador" elevation="139" latitude="1.2044" location="Durango" longLatPrecision="7" longitude="-78.8447" municipality="Canton San Lorenzo" specimenCode="QCAZ 55541" specimenCount="1" stateProvince="Esmeraldas" typeStatus="paratype">
<specimenCode id="6ADD260D94C2FCF0AEAA220D25D8EB93">
<named-content id="465900C8BC601CE62229FFBFC8030DC1" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
55541
</specimenCode>
–55544, 55551–55554 
<collectingMunicipality id="148C746CF6893A2CBFA90D1316F1A562">Cantón San Lorenzo</collectingMunicipality>
, 
<locationDeviation id="0846219B48AB6D99CDA315A41AA87E1A">
<quantity id="3A588A62DDABCF4F66AA212B282A6C96" metricMagnitude="3" metricUnit="m" metricValue="7.0" unit="km" value="7.0">7 km</quantity>
west of
</locationDeviation>
<location id="FBDFE62A86652ECCFD5C445ABC9AEC9C" LSID="urn:lsid:plazi:treatment:FA1177A074C859B19F69711C4654B852:FBDFE62A86652ECCFD5C445ABC9AEC9C" country="Ecuador" latitude="1.2044" longLatPrecision="7" longitude="-78.8447" municipality="Canton San Lorenzo" name="Durango" stateProvince="Esmeraldas">Durango</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EH2AI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-78.844700,1.204400]}">
<geoCoordinate id="4EE820004D18F94A1EF30087A9BE2560" degrees="1.2044" direction="north" orientation="latitude" precision="5" value="1.2044">1.2044 ° N</geoCoordinate>
, 
<geoCoordinate id="9FDF57E6E9E3347C0E6751B142AC5BA7" degrees="78.8447" direction="west" orientation="longitude" precision="5" value="-78.8447">78.8447 ° W</geoCoordinate>
</named-content>
), 
<quantity id="032B0E570DDECA6FE6F0B58AAA2962E6" metricMagnitude="2" metricUnit="m" metricValue="1.39" unit="m" value="139.0">
<elevation id="3DBBA2DFA0F8E49713657048A2B7A695" metricMagnitude="2" metricUnit="m" metricValue="1.39" unit="m" value="139.0">139 m</elevation>
</quantity>
, collected by 
<collectorName id="9ECFD2A54021BBD3E05863BFC9B5F866">Justin Touchon</collectorName>
on 
<date id="55C351E55BD24D8821A50DC12DCCF6DD" value="2013-04-22">
<collectingDate id="EA0907C2AE903FC39C4FFFF06661AFE8" value="2013-04-22">22 April, 2013</collectingDate>
</date>
</materialsCitation>
; • 
<materialsCitation id="A3086ABB01E18FCD23828539F073FAF6" collectingDate="2017-02-10" collectionCode="QCAZ" collectorName="Diego Almeida" country="Ecuador" elevation="265" latitude="1.0874" location="Durango" longLatPrecision="7" longitude="-78.6221" municipality="Canton San Lorenzo" specimenCode="QCAZ 66629" specimenCount="1" stateProvince="Esmeraldas" typeStatus="paratype">
<specimenCode id="6583477B9CA8B3FA995C30909E442C74">
<named-content id="9D18C69D054ABD6690451063544F1510" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
66629
</specimenCode>
–66635 
<collectingMunicipality id="1987372385B46455254FC5514426D37A">Cantón San Lorenzo</collectingMunicipality>
, 
<location id="0F2C5D53B91FB1800D1BDA3AE5765493" LSID="urn:lsid:plazi:treatment:FA1177A074C859B19F69711C4654B852:0F2C5D53B91FB1800D1BDA3AE5765493" country="Ecuador" latitude="1.0874" longLatPrecision="7" longitude="-78.6221" municipality="Canton San Lorenzo" name="Durango" stateProvince="Esmeraldas">Durango</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ET2AI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-78.622100,1.087400]}">
<geoCoordinate id="E2BB5E50A7D7138C1DFAC988A57A8C02" degrees="1.0874" direction="north" orientation="latitude" precision="5" value="1.0874">1.0874 ° N</geoCoordinate>
, 
<geoCoordinate id="C4E22B4F490B38867D7AF9E2C7996EB7" degrees="78.6221" direction="west" orientation="longitude" precision="5" value="-78.6221">78.6221 ° W</geoCoordinate>
</named-content>
), 
<quantity id="77E344E1ACB7C5BFD7F5EF9854EB599D" metricMagnitude="2" metricUnit="m" metricValue="2.65" unit="m" value="265.0">
<elevation id="2CE0C4B21EC5443E793F61ADCD485ACE" metricMagnitude="2" metricUnit="m" metricValue="2.65" unit="m" value="265.0">265 m</elevation>
</quantity>
, collected by 
<collectorName id="307B665EFE3CA98612F6D3A3892D802B">Diego Almeida</collectorName>
on 
<date id="9B560098B375B9571A5ABDC9C9B2B42E" value="2017-02-10">
<collectingDate id="4E0D0B07A10ED0AE43E92B10880CB7ED" value="2017-02-10">10 February, 2017</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EY2AI" type="Common names">
<paragraph id="1F3EFD60A6BCB0B1E3943CBF4427FB28">
<heading id="6438BE41932DEEB2B1BF5FFB4A7599AA" reason="title">Common names.</heading>
</paragraph>
<paragraph id="2C6BFAC4E5C92B5FC4859FBE1AB98D37">English: Cannatella’s treefrog. Spanish: Ranita de Cannatella.</paragraph>
</subSubSection>
<subSubSection id="SECID0E42AI" type="Definition">
<paragraph id="0EEA28897E36F7C3590EA4DF18700E56">
<heading id="8306FA38A9E883A60E2FAAA1C903C58A" reason="title">Definition</heading>
</paragraph>
<paragraph id="F9EA0F60AADAEEE576D536C577418E52">
<emphasis id="130168173473B72B58D5A0E88ED6E28C" bold="true">
(Figs 
<figureCitation id="CA32B6852CAA047FB34FA1BEF97D47F2" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Life coloration of Dendropsophus cannatellai sp. nov. First and third rows, dorsolateral views; second and fourth, ventral views. From left to right, first and second rows: QCAZ 51877 (SVL = 22.26 mm, male); QCAZ 78809 (SVL = 33.61 mm, female); QCAZ 55558 (SVL = 21.96 mm, male); third and fourth rows: QCAZ 40847 (holotype; SVL = 22.32 mm, male); QCAZ 78812 (SVL = 25.86 mm, male); QCAZ 78813 (SVL = 24.94 mm, male). See type series for locality data. All specimens are shown at the same scale." httpUri="https://binary.pensoft.net/fig/1257792">8</figureCitation>
, 
<figureCitation id="68A6E29A3FB930339297EB1343AE6AB9" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Adult preserved specimens of Dendropsophus cannatellai sp. nov. First and third rows, dorsolateral views; second and fourth, ventral views. From left to right, first and second rows: QCAZ 9457 (SVL = 28.65 mm, female); QCAZ 23162 (22.83 mm, male); QCAZ 26098 (30.73 mm, female); third and fourth rows: QCAZ 35710 (24.74 mm, male); QCAZ 78254 (24.71 mm, male); QCAZ 78257 (31.94 mm, female). See type-series for locality data. All specimens are shown at the same scale." httpUri="https://binary.pensoft.net/fig/1257793">9</figureCitation>
).
</emphasis>
In this section, we describe coloration in life. We assign the new species to the genus 
<taxonomicName id="9BF4FC84F3BC99B2DF088B0917B9EE8B" authorityName="Fitzinger" authorityYear="1843" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="genus">
<emphasis id="FE8BACD83416C0B632D5FDAAE1005682" italics="true">Dendropsophus</emphasis>
</taxonomicName>
based on the phylogeny (Fig. 
<figureCitation id="305B8640409C90939D446961B134462E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Maximum likelihood tree of the Dendropsophus leucophyllatus group based on concatenated DNA sequences of mitochondrial and nuclear genes. Blue numbers on branches show SH-aLRT values (before slash) and ultrafast bootstrap support values (after slash). Support values not shown for intraspecific relationships, except for deepest nodes within D. ebraccatus." httpUri="https://binary.pensoft.net/fig/1257785">1</figureCitation>
). The new species is characterized by: (1) mean SVL 
<quantity id="2702DA35E3E12CBB254997CBD9367CD5" metricMagnitude="-2" metricUnit="m" metricValue="2.34" unit="mm" value="23.4">23.4 mm</quantity>
in males (range 21–26; n = 115); 
<quantity id="32ECD55048AD2274071254F297A7ADB1" metricMagnitude="-2" metricUnit="m" metricValue="3.07" unit="mm" value="30.7">30.7 mm</quantity>
in females (range 28–33; n = 14); (2) axillary membrane reaching arm halfway to elbow; (3) reduced webbing on fingers; (4) webbing on feet; (5) palmar tubercle single; (6) pectoral glands present; (7) dorsal background coloration brown with white to bright yellow dorsolateral bands, extending to the eyes and top of head; white or bright yellow well defined sacral mark, fused with dorsolateral bands in some individuals; (8) absence of a light labial stripe in contact with the orbit; (9) dorsal surfaces of the limbs brown, with white to bright yellow ovoid marks, with the forearm typically bearing one to two marks, and the shank one to three; (10) webbing and ventral surfaces vary from orange to yellow; (11) iris dull bronze to coppery bronze, and (12) advertisement call pulsed with a mean dominant frequency = 3274.7 Hz (range 3169.0–3498.7) and mean pulse rate of the primary note = 95 pulses / s (range 87–100).
</paragraph>
<caption id="7B0E2830643E2FCE2A9D3DE7B97E8492" ID-arpha="70E1649A-266E-51DA-AB7F-877A160A1569" httpUri="https://binary.pensoft.net/fig/1257792" startId="F8">
<paragraph id="2683B44A9079E2D37331308C90A6357D">
<label id="3CFD6360DAB0DC80B1CE35B745C86BC8">Figure 8.</label>
</paragraph>
<paragraph id="1F0B532E477585BBCD1DDA98C8F3CBB7">
Life coloration of 
<taxonomicName id="15536504EE860EBE1FFB274E9F98B9D7" authorityName="Aguirre &amp; Apunte &amp; Ron" authorityYear="2025" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="cannatellai" status="sp. nov.">
<emphasis id="434C0C596275930836FA09D861450D30" italics="true">Dendropsophus cannatellai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07B4623247D814A18B981D623F79CDFB" rank="species">sp. nov.</taxonomicNameLabel>
First and third rows, dorsolateral views; second and fourth, ventral views. From left to right, first and second rows: 
<named-content id="29BD91D833D9921C8023DF500BC98B56" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
51877 (SVL = 22.26 mm, male); 
<named-content id="D99BF4C0B5D94DFF52419A3B88286D4D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
78809 (SVL = 33.61 mm, female); 
<named-content id="BBF667E94617AEEB04622F43D1637EA0" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
55558 (SVL = 21.96 mm, male); third and fourth rows: 
<named-content id="0C8C292D79555B5EDFCF5B4C19D2F5D6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
40847 (holotype; SVL = 22.32 mm, male); 
<named-content id="8CDE2248B6367FF427BF83F5EAC74985" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
78812 (SVL = 25.86 mm, male); 
<named-content id="0AAA25FC931AA5A3BB33F4319DDFC0B7" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
78813 (SVL = 24.94 mm, male). See type series for locality data. All specimens are shown at the same scale.
</paragraph>
</caption>
<caption id="1E1A8AD699FA5E903B91BA0D09F363EC" ID-arpha="8D8FE406-C185-5DB3-A2FE-336995165B09" httpUri="https://binary.pensoft.net/fig/1257793" startId="F9">
<paragraph id="E83D30177B1919277F23918B9F5C9D4E">
<label id="13D0268255EB0265C46C4BA57ED3D93B">Figure 9.</label>
</paragraph>
<paragraph id="28FE5E5B9F12BF1A6B606A174106731F">
Adult preserved specimens of 
<taxonomicName id="97929F17AEB628EC10C31481CD452BE9" authorityName="Aguirre &amp; Apunte &amp; Ron" authorityYear="2025" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="cannatellai" status="sp. nov.">
<emphasis id="96364F21F96A72F8AAE40580531B825B" italics="true">Dendropsophus cannatellai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="986A38C27AD01A1B36F67128DBE7D0B5" rank="species">sp. nov.</taxonomicNameLabel>
First and third rows, dorsolateral views; second and fourth, ventral views. From left to right, first and second rows: 
<named-content id="389EB54145F2DFD59B508DFE06655627" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
9457 (SVL = 28.65 mm, female); 
<named-content id="0F2309416B4D1C5459C5C24485F11BF7" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
23162 (22.83 mm, male); 
<named-content id="00720D5A529BB0C39CC3E06E9FF02677" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
26098 (30.73 mm, female); third and fourth rows: 
<named-content id="E8C5B385433688B2FB330415436E736F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
35710 (24.74 mm, male); 
<named-content id="0174B17B4FE78423A64784045F4FF402" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
78254 (24.71 mm, male); 
<named-content id="078E2C9F2951CD47E5552C7E0533B426" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
78257 (31.94 mm, female). See type-series for locality data. All specimens are shown at the same scale.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EZ6AI" type="diagnosis">
<paragraph id="BFAD38FFC9B4C9F0A3F73A0B8FF0BDDB">
<heading id="5CE8850074E1D5B96010D3B51A6309E8" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="506541CDE1826EE7088F8CE9C966D1F9">
In this section, coloration refers to live adult individuals, unless otherwise noted. Adult individuals of 
<taxonomicName id="C2FB3FEF3BBC82D10D30D35E59F00492" authorityName="Aguirre &amp; Apunte &amp; Ron" authorityYear="2025" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="cannatellai" status="sp. nov.">
<emphasis id="5BEFB0B08A62E3ED2C8A9C45F8062EDC" italics="true">Dendropsophus cannatellai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="56F9D7E74A754ADC0A9E39CAB86F879B" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from other species of the genus 
<taxonomicName id="679C948DB3EBB29B55C4113A85E951A2" authorityName="Fitzinger" authorityYear="1843" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="genus">
<emphasis id="F68E6BA571F714A6824517A9AA38671A" italics="true">Dendropsophus</emphasis>
</taxonomicName>
that do not belong to the 
<taxonomicName id="8DE2D88281236EE9AB7085BDF0042BC0" baseAuthorityName="Beireis" baseAuthorityYear="1783" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="leucophyllatus">
<emphasis id="FC1E07EFA8CA92CE6DA87B1C4B0F8A34" italics="true">D. leucophyllatus</emphasis>
</taxonomicName>
group by the presence of two pectoral glands. Within the 
<taxonomicName id="F25F735906C98DCDDEA15F139E7B41D0" baseAuthorityName="Beireis" baseAuthorityYear="1783" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="leucophyllatus">
<emphasis id="9751094D081C3D365CFEDD04FBE65ECA" italics="true">D. leucophyllatus</emphasis>
</taxonomicName>
group, we focus our comparisons with species from Central America and the 
<collectingRegion id="7AD85192A8B24F7B14792B83E2A7C99F" country="Panama" name="Embera-Wounaan">Chocó region</collectingRegion>
, also the most closely related. Other species of the group have a cis-Andean distribution and, therefore, cannot be confused with the new species. Nevertheless, we include diagnosis for species from the western Amazon basin, which are geographically closest to the new species range.
</paragraph>
<caption id="6EAB6EC2E23A2388FC7D55668C494135" ID-arpha="C1495D83-5807-524E-B67A-530259A08E0A" httpUri="https://binary.pensoft.net/fig/1257794" startId="F10">
<paragraph id="3EF8D5B9C6A0329DE7D5A7BCE8A95508">
<label id="010BEF2B7659F6060794124A07F55CC5">Figure 10.</label>
</paragraph>
<paragraph id="88B7D307C15FA88F8B2C5148DB8DDB86">
Tadpole (
<named-content id="B1625068477780631D11AF0A6C461B7F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
78576) of 
<taxonomicName id="45B9E7B8D1DD21E2D85C6E58D84D8475" authorityName="Aguirre &amp; Apunte &amp; Ron" authorityYear="2025" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="cannatellai" status="sp. nov.">
<emphasis id="F5AB790576BA971462EA16496020887F" italics="true">Dendropsophus cannatellai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="730C0EA4911A1C43D34E9B629FF09124" rank="species">sp. nov</taxonomicNameLabel>
, stage 25 (
<bibRefCitation id="0CC999770715BC3ACA659BDE4D971044" author="Gosner" firstAuthor="Gosner" issue="3" journalOrPublisher="Herpetologica" pagination="183-190" refId="B14" refString="Gosner KL (1960) A simplified table for staging anuran embryos and larvae with notes on identification. Herpetologica 16 (3): 183–190. https://www.jstor.org/stable/3890061" title="A simplified table for staging anuran embryos and larvae with notes on identification." url="https://www.jstor.org/stable/3890061" volume="16" year="1960">Gosner 1960</bibRefCitation>
). Dorsal, ventral, and lateral views. Body length = 8.14 mm; tail length = 16.24 mm; total length = 24.38 mm.
</paragraph>
</caption>
<paragraph id="E5B406F9F83A1C8BBD62505FD2F15343">
<taxonomicName id="A6FD08B6FB39C88233CA6F27FA3DC58A" baseAuthorityName="Cope" baseAuthorityYear="1874" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="ebraccatus">
<emphasis id="BE581B63269B163F128A5964E78F31F4" italics="true">Dendropsophus ebraccatus</emphasis>
</taxonomicName>
differs from the new species (characters of the later in parenthesis) in having a light labial stripe reaching the orbit (area below the eye has dark background color, sometimes with one or two small light spots). In addition, 
<taxonomicName id="82C7F7E93BDACD7E6F424DBAAFE3B845" baseAuthorityName="Cope" baseAuthorityYear="1874" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="ebraccatus">
<emphasis id="3D0EDBCE8F41920386D7E9C94BDF4552" italics="true">D. ebraccatus</emphasis>
</taxonomicName>
either lacks dorsolateral yellow bands or, if present, they are wide and sinuous (if present, yellow dorsolateral bands straight to moderately curved; variation in coloration of 
<taxonomicName id="825A0396607ABA24B8146F937CDCCC70" baseAuthorityName="Cope" baseAuthorityYear="1874" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="ebraccatus">
<emphasis id="A7080D0E8BFEAA6FD439AFCE8E1982BA" italics="true">D. ebraccatus</emphasis>
</taxonomicName>
from 
<bibRefCitation id="80DE609B3F0F3F078E9110EF6F8B12AD" DOI="10.5962/bhl.title.2835" author="Duellman" firstAuthor="Duellman" refId="B6" refString="Duellman WE (1970) The hylid frogs of Middle America. Natural History Museum, University of Kansas. https://doi.org/10.5962/bhl.title.2835" year="1970">Duellman 1970</bibRefCitation>
and 
<bibRefCitation id="FB7E00182550E4D8A0680FDD00CA9D97" DOI="10.1111/j.1095-8312.2009.01210.x" author="Ohmer" etAl="et al." firstAuthor="Ohmer" journalOrPublisher="Biological Journal of the Linnean Society" pagination="298-313" refId="B29" refString="Ohmer ME, Robertson JM, Zamudio KR (2009) Discordance in body size, colour pattern, and advertisement call across genetically distinct populations in a Neotropical anuran (Dendropsophus ebraccatus). Biological Journal of the Linnean Society 97: 298–313. https://doi.org/10.1111/j.1095-8312.2009.01210.x" title="Discordance in body size, colour pattern, and advertisement call across genetically distinct populations in a Neotropical anuran (Dendropsophus ebraccatus)." volume="97" year="2009">Ohmer et al. 2009</bibRefCitation>
).
</paragraph>
<paragraph id="83F8D6E5229B18AD58462509C24949A7">
<taxonomicName id="FF966A00BB7F70FB2474FC575C156A0F" authorityName="Aguirre &amp; Apunte &amp; Ron" authorityYear="2025" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="cannatellai" status="sp. nov.">
<emphasis id="32ECD0AC289EEC50FBC67BF6B20DB31D" italics="true">Dendropsophus cannatellai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1A48C50EA2BC21792C382E42DAB6AF78" rank="species">sp. nov.</taxonomicNameLabel>
clearly differs from its sister species, 
<taxonomicName id="93491FB8D6D44CB27B25FAA3E6F97189" baseAuthorityName="Duellman" baseAuthorityYear="1973" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="gryllatus">
<emphasis id="7AF537D054323EDDC2BF8FEABF892083" italics="true">D. gryllatus</emphasis>
</taxonomicName>
, in dorsal coloration and by being larger (although SVL ranges overlap: male average SVL = 21.95, range 
<quantity id="071BDDF99741C47B02777F870C68D9A9" metricMagnitude="-2" metricUnit="m" metricValue="2.17" metricValueMax="2.469" metricValueMin="1.871" unit="mm" value="21.7" valueMax="24.69" valueMin="18.71">18.71–24.69 mm</quantity>
vs. SVL = 23.43, range 
<quantity id="FD950F599DC41DA1A98EE5CC8176B99A" metricMagnitude="-2" metricUnit="m" metricValue="2.3815" metricValueMax="2.658" metricValueMin="2.105" unit="mm" value="23.814999999999998" valueMax="26.58" valueMin="21.05">21.05–26.58 mm</quantity>
in 
<taxonomicName id="42039E8F288BE8EA3B94253C7C7042D9" authorityName="Aguirre &amp; Apunte &amp; Ron" authorityYear="2025" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="cannatellai" status="sp. nov.">
<emphasis id="66D731CCE487A2CDE6044506E3380003" italics="true">D. cannatellai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D043DAA0C283F7CF4C2354EA66C86E10" rank="species">sp. nov.</taxonomicNameLabel>
) The dorsal pattern of 
<taxonomicName id="5D494237571755E723758563F3B47FDD" baseAuthorityName="Duellman" baseAuthorityYear="1973" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="gryllatus">
<emphasis id="F093C9B52639275940657BE51434FFE2" italics="true">D. gryllatus</emphasis>
</taxonomicName>
consists of two longitudinal dark brown lines against a pale-yellow background (Fig. 
<figureCitation id="F51E13F9CCB225BBABF259F62517E3AE" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Life coloration of Dendropsophus gryllatus. First and third rows, dorsolateral views; second and fourth, ventral views. From left to right, first and second rows: QCAZ 75534 (SVL = 20.80 mm); QCAZ 75535 (SVL = 21.19 mm); QCAZ 75536 (SVL = 22.24 mm); third and fourth rows: QCAZ 75538 (SVL = 18.71 mm); QCAZ 75539 (SVL = 21.89 mm); QCAZ 75540 (SVL = 20.26 mm); all adult males. See Suppl. material 1, table S 1 for locality data. All specimens are shown at the same scale." httpUri="https://binary.pensoft.net/fig/1257798">14</figureCitation>
; brown background with clear yellow dorsolateral bands in the new species). The new species also differs from 
<taxonomicName id="987938445D95F8310ED82A08D3264BB4" baseAuthorityName="Duellman" baseAuthorityYear="1973" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="gryllatus">
<emphasis id="0987FC07EFB978121D131CF61575D0B8" italics="true">D. gryllatus</emphasis>
</taxonomicName>
by having bright orange flash coloration (pale yellow in 
<taxonomicName id="6697156F8A642D81DFDD0348B1806064" baseAuthorityName="Duellman" baseAuthorityYear="1973" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="gryllatus">
<emphasis id="97B9C92717FCDEF87082C523E3A527BE" italics="true">D. gryllatus</emphasis>
</taxonomicName>
). 
<taxonomicName id="B5762C5657511306B5D2C37AB847FB4C" baseAuthorityName="Duellman" baseAuthorityYear="1973" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="gryllatus">
<emphasis id="6C82A65D6FDD336FA5982C014B5385D0" italics="true">Dendropsophus gryllatus</emphasis>
</taxonomicName>
exhibits a sacral spot that extends forward, making it more elongated and continuous with other dorsal markings (the sacral spot of 
<taxonomicName id="B69E9D99F28DEC7BE8539A495A96123B" baseAuthorityName="Cope" baseAuthorityYear="1874" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="ebraccatus">
<emphasis id="E10161E13677F0B6F52EA89C79240C2C" italics="true">D. ebraccatus</emphasis>
</taxonomicName>
is distinct, circular or slightly oval, and symmetrically positioned near the lower back, with well-defined edges that contrast against its darker dorsal pattern; sacral spot is absent in some individuals).
</paragraph>
<paragraph id="FFFC647E5E02616B37F4C88FCB4C2F24">
The Amazonian species 
<taxonomicName id="A31FBD265D8D9B97F53563A2B0FE7FF8" baseAuthorityName="Shreve" baseAuthorityYear="1935" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="sarayacuensis">
<emphasis id="CFB12FBC019D60C56B3C87FD8BB07B03" italics="true">D. sarayacuensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="A5B7C4F35CF2B88E1AF536FD0D33F049" baseAuthorityName="Andersson" baseAuthorityYear="1945" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="bifurcus">
<emphasis id="BBA3D49970ED3F83C9D6ED80D2B7CF39" italics="true">D. bifurcus</emphasis>
</taxonomicName>
also has a brown background color with yellow bands and marks. However, 
<taxonomicName id="0D65ACBA7170D6442F50D676D2A4385D" baseAuthorityName="Shreve" baseAuthorityYear="1935" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="sarayacuensis">
<emphasis id="C3D49A11063C2F46AF317155D7667F81" italics="true">D. sarayacuensis</emphasis>
</taxonomicName>
differs in that the yellow areas in the shanks have irregular borders (borders are more linear in 
<taxonomicName id="04ED43D0D0856BA8F532D5E59034918E" authorityName="Aguirre &amp; Apunte &amp; Ron" authorityYear="2025" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="cannatellai" status="sp. nov.">
<emphasis id="CCEFA17BC152C075FFC943CD8E6A87CA" italics="true">D. cannatellai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7E7B3D366ECB10A40BB0A06370FC016E" rank="species">sp. nov.</taxonomicNameLabel>
) 
<taxonomicName id="42655741683E6FA823E0B197230A4A86" baseAuthorityName="Andersson" baseAuthorityYear="1945" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="bifurcus">
<emphasis id="F98DB68CDC8F3CC4C541D6F33434320E" italics="true">Dendropsophus bifurcus</emphasis>
</taxonomicName>
differs by having less extensive yellow bands and marks, especially in the shanks where, in most individuals, they are absent, except for the heel (extensive yellow marks along the shank in 
<taxonomicName id="56EE3987133F579B92800336082075F8" authorityName="Aguirre &amp; Apunte &amp; Ron" authorityYear="2025" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="cannatellai">
<emphasis id="B1F32B3B55E2E0248ED348C21D83A940" italics="true">D. cannatellai</emphasis>
</taxonomicName>
). 
<taxonomicName id="725E9843362B2AF59857011F3DFA134B" baseAuthorityName="Jimenez de la Espada" baseAuthorityYear="1870" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="reticulatus">
<emphasis id="4B5C6B659680F5A73052A6737304900F" italics="true">Dendropsophus reticulatus</emphasis>
</taxonomicName>
differs by having a uniform clear dorsal color pattern, sometimes with one or several dark brown round marks (
<bibRefCitation id="FE2725A57343D9CCDB25966E67DE3408" DOI="10.1371/journal.pone.0171785" author="Caminer" etAl="et al." firstAuthor="Caminer" refId="B1" refString="Caminer MA, Mila B, Jansen M, Fouquet A, Venegas PJ, Chavez G, Lougheed SC, Ron SR (2017) Systematics of the Dendropsophus leucophyllatus species complex (Anura: Hylidae): Cryptic diversity and the description of two new species. PLoS ONE 12 (3): e 0171785. https://doi.org/10.1371/journal.pone.0171785" year="2017">Caminer et al. 2017</bibRefCitation>
) (clear dorsal pattern restricted to head, dorsolateral bands, and rump in 
<taxonomicName id="9F99C014F9EB08C709BFB60C7691D5BD" authorityName="Aguirre &amp; Apunte &amp; Ron" authorityYear="2025" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="cannatellai">
<emphasis id="AC8E87092D9168CE5891107C6CE43DF7" italics="true">D. cannatellai</emphasis>
</taxonomicName>
). 
<taxonomicName id="1291CED4E70F58C88D8F9C1534508B34" baseAuthorityName="Gunther" baseAuthorityYear="1869" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="triangulum">
<emphasis id="434C35243BAC31080B8EF9FFC6AC317C" italics="true">Dendropsophus triangulum</emphasis>
</taxonomicName>
and 
<taxonomicName id="F8D570B148437423CE5782D0AB0BFA65" authorityName="Caminer, Mila, Jansen, Fouquet, Venegas, Chavez, Lougheed &amp; Ron" authorityYear="2017" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="arndti">
<emphasis id="4289D4E5B0C1935E4DFBE30EC3F2C0CC" italics="true">D. arndti</emphasis>
</taxonomicName>
differ by being larger, SVL&gt; 28.0 mm in males and&gt; 33.0 mm in females (
<bibRefCitation id="AE6A2CAF30B2A2CDD756B56BC92F423E" DOI="10.1371/journal.pone.0171785" author="Caminer" etAl="et al." firstAuthor="Caminer" refId="B1" refString="Caminer MA, Mila B, Jansen M, Fouquet A, Venegas PJ, Chavez G, Lougheed SC, Ron SR (2017) Systematics of the Dendropsophus leucophyllatus species complex (Anura: Hylidae): Cryptic diversity and the description of two new species. PLoS ONE 12 (3): e 0171785. https://doi.org/10.1371/journal.pone.0171785" year="2017">Caminer et al. 2017</bibRefCitation>
) (
<taxonomicName id="077299C09C6BEC945C823D17104C50B1" authorityName="Aguirre &amp; Apunte &amp; Ron" authorityYear="2025" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="cannatellai">
<emphasis id="F6CE0BC4FF47CC6D3C71B3EDE9BA5DB8" italics="true">D. cannatellai</emphasis>
</taxonomicName>
males SVL range 
<quantity id="61B125CE0DB8E89CA5EC7D8E0319620E" metricMagnitude="-2" metricUnit="m" metricValue="2.3815" metricValueMax="2.658" metricValueMin="2.105" unit="mm" value="23.814999999999998" valueMax="26.58" valueMin="21.05">21.05–26.58 mm</quantity>
, females = 
<quantity id="FB5AA7A3C0DD6986F190D123D9213A70" metricMagnitude="-2" metricUnit="m" metricValue="3.0095" metricValueMax="3.194" metricValueMin="2.825" unit="mm" value="30.095" valueMax="31.94" valueMin="28.25">28.25–31.94 mm</quantity>
).
</paragraph>
<paragraph id="5ADF6CD5440B1E2D89ED64A499450C47">
<taxonomicName id="92EA3A9DFC4F70198B7682ADB8B41E66" authorityName="Aguirre &amp; Apunte &amp; Ron" authorityYear="2025" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="cannatellai" status="sp. nov.">
<emphasis id="453E18DBF0380C81AC020D8B99A2C693" italics="true">Dendropsophus cannatellai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="962A3AAB52CD1C3F37B1D87A9D5219A8" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from 
<taxonomicName id="B0DDADC183404B7D9ABA8045D9A344FE" baseAuthorityName="Duellman" baseAuthorityYear="1969" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="carnifex">
<emphasis id="4014A16D178195DF9EA7117F1F26D5D4" italics="true">Dendropsophus carnifex</emphasis>
</taxonomicName>
(
<taxonomicName id="F3AD74367DE2D0F273F3EB64B4270AA2" baseAuthorityName="Boettger" baseAuthorityYear="1892" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="columbianus">
<emphasis id="9DF964163B6549BEBC1E3A57F8587104" italics="true">D. columbianus</emphasis>
</taxonomicName>
group), a species from the Andean 
<collectingRegion id="40CC3DB8EFCDB685F85570EAD4AA4459" country="Panama" name="Embera-Wounaan">Chocó</collectingRegion>
, by having a short axillary membrane that reaches halfway to the elbow (axillary membrane absent in 
<taxonomicName id="3750F7C1D902A89C4F9C8CD4A9D05AA0" baseAuthorityName="Duellman" baseAuthorityYear="1969" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="carnifex">
<emphasis id="6519EBF4543021FB29FC7065A240DF67" italics="true">D. carnifex</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBMBI" type="description">
<paragraph id="FDD02CD353881C070995A6BE022A40B5">
<heading id="584F6636C9FD388027E4FBC158F44426" reason="title">
Description of 
<typeStatus id="66E3F2BD5651A7FA27ACBD3120216433" type="holotype">holotype</typeStatus>
</heading>
</paragraph>
<paragraph id="D8DC20582321A646F7890523F6D15857">
<emphasis id="46722BCEF5669F70BC7BAE2BC5E4F453" bold="true">
(Figs 
<figureCitation id="9C186F787565A5EEDCA01A37ACD4AEFA" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Dorsal, ventral and lateral views of the holotype of Dendropsophus cannatellai sp. nov. (QCAZ 40847, SVL 22.32 mm)." httpUri="https://binary.pensoft.net/fig/1257796">12</figureCitation>
, 
<figureCitation id="D4922858CCA88ED7F266BFE0843663F9" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Ventral views of the holotype’s hand (left) and foot (right) of Dendropsophus cannatellai sp. nov. (QCAZ 40847)." httpUri="https://binary.pensoft.net/fig/1257797">13</figureCitation>
).
</emphasis>
Adult male, SVL 
<quantity id="EBC837A6900990D28F10E600DDC5FF93" metricMagnitude="-2" metricUnit="m" metricValue="2.232" unit="mm" value="22.32">22.32 mm</quantity>
, foot length 
<quantity id="9A661717A3322BFC509EE9D5280B70B1" metricMagnitude="-3" metricUnit="m" metricValue="3.04" unit="mm" value="3.04">3.04 mm</quantity>
, head length 
<quantity id="F876B0324255A7C9E1A0FA950495B197" metricMagnitude="-3" metricUnit="m" metricValue="7.42" unit="mm" value="7.42">7.42 mm</quantity>
, head width 
<quantity id="A8F59BE3309FAD54338BBE7EAFFD08ED" metricMagnitude="-3" metricUnit="m" metricValue="7.58" unit="mm" value="7.58">7.58 mm</quantity>
, eye diameter 
<quantity id="EDB9DCF4167B82826B4DD2D151C78AD1" metricMagnitude="-3" metricUnit="m" metricValue="3.34" unit="mm" value="3.34">3.34 mm</quantity>
, tympanum diameter 
<quantity id="4F61118FFAAA7B4272BD67C1D1C0A47D" metricMagnitude="-3" metricUnit="m" metricValue="1.25" unit="mm" value="1.25">1.25 mm</quantity>
, tibia length 
<quantity id="340195051736381FFCEDF44081A316AC" metricMagnitude="-2" metricUnit="m" metricValue="1.18" unit="mm" value="11.8">11.8 mm</quantity>
, femur length 
<quantity id="0E587E5B534581FF1AEA201A8D5D44EE" metricMagnitude="-2" metricUnit="m" metricValue="1.088" unit="mm" value="10.88">10.88 mm</quantity>
, arm length 
<quantity id="DB9C658891CDC18BC6E93ACE5628C9F4" metricMagnitude="-3" metricUnit="m" metricValue="4.55" unit="mm" value="4.55">4.55 mm</quantity>
, eye-nostril distance 
<quantity id="C5EA9CACA67DA43DF32F7C2399B88CFB" metricMagnitude="-3" metricUnit="m" metricValue="1.66" unit="mm" value="1.66">1.66 mm</quantity>
. Body as wide as head, head wider than long; snout short, truncated in dorsal view and profile; distance from nostril to eye shorter than eye diameter; canthus rostralis barely distinct, rounded; loreal region slightly concave; internarial area subtly depressed; nostrils slightly protruding, directed posterolaterally; interorbital region flat; eyes large, prominent; eye diameter 2.7 times tympanic annulus diameter; tympanum hidden beneath skin; tympanic membrane absent, tympanic annulus visible under skin, oval, longer dorsoventrally and concealed dorsally by supratympanic fold, separated from eye by about 57 % of its diameter; faint supratympanic fold, extending posteriorly from rear corner of eye to anterior edge of arm insertion. Arm sturdy and short, axillary membrane extends halfway to elbow; relative length of fingers I &lt;II &lt;IV &lt;III; fingers with large rounded discs, that of Finger III approximately the same diameter as tympanum; subarticular tubercles prominent, round, subconical, single; distal tubercle on Finger IV bifid; supernumerary tubercles present; palmar tubercle indistinct; prepollex sharp-pointed (based on x-ray image); nuptial excrescences absent; hand webbing formula I basal II 1 ½ — 2 ½ III 2 ⅔ — 2 IV. Hindlimbs large; toes with discs slightly wider than long, smaller than those on fingers; relative toe length I &lt;II &lt;V &lt;III &lt;IV; outer metatarsal tubercle indistinct, small, round; inner metatarsal tubercle large, elongated, elliptical; subarticular tubercles single, round, flat; supernumerary tubercles limited to soles, small; foot webbing formula I 1 + — 2 II 1 + — 2 – III 1 + — 2 IV 2 + — 1 + V (Fig. 
<figureCitation id="39E6B0C7E3380AA71702AD7A8C392C00" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Metamorph, juvenile and subadults of Dendropsophus cannatellai sp. nov. Dorsolateral views of individuals raised in captivity in Balsa de los Sapos. From left to right, first row: QCAZA l 78576 (total length = 23.27 mm); QCAZA n 79747 (SVL = 11.57 mm); second row: (18.46 mm); (19.59 mm); third row: (19.38 mm); (17.89 mm). All subadults approximately 1 year old since hatching." httpUri="https://binary.pensoft.net/fig/1257795">11</figureCitation>
). Two pectoral glands bellow the insertion of the arms, separated from each other by about half their individual width. Skin on dorsum, head, flanks, and dorsal surfaces of limbs smooth. Ventral skin coarsely granular, except for pectoral glands, which are finely granular. Ventral surfaces of body and thighs granular. Longitudinal wrinkles of vocal sac from throat to front of the insertion of the arms. Skin on arms and shanks smooth. Cloacal opening directed posteriorly at upper level of the thighs, covered by a long, broad cloacal sheath. Tongue broadly heart-shaped, free laterally and posteriorly, widely attached to mouth floor. Vomerine teeth between choanae, in two slightly angled rows, not contacting each other, smaller than ovoid choanae. In life dorsal background coloration is brown with yellow dorsolateral bands, extending to the eyes and top of head (Fig. 
<figureCitation id="D0E92864F5B896DCE7E550D327A4CA83" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Life coloration of Dendropsophus cannatellai sp. nov. First and third rows, dorsolateral views; second and fourth, ventral views. From left to right, first and second rows: QCAZ 51877 (SVL = 22.26 mm, male); QCAZ 78809 (SVL = 33.61 mm, female); QCAZ 55558 (SVL = 21.96 mm, male); third and fourth rows: QCAZ 40847 (holotype; SVL = 22.32 mm, male); QCAZ 78812 (SVL = 25.86 mm, male); QCAZ 78813 (SVL = 24.94 mm, male). See type series for locality data. All specimens are shown at the same scale." httpUri="https://binary.pensoft.net/fig/1257792">8</figureCitation>
); bright yellow well defined sacral mark; absence of a light labial stripe and large suborbital mark; the dorsal surfaces of the thighs is light brown; the forearm and shank have two yellow transversal marks each; coloration of webbing and ventral surfaces yellow; iris coppery bronze. Color of 
<typeStatus id="89814EC1F7B8E8461AC229583BD84046" type="holotype">holotype</typeStatus>
in preservative shown in Fig. 
<figureCitation id="1A57CF99252A24C8874C72913178B93B" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Dorsal, ventral and lateral views of the holotype of Dendropsophus cannatellai sp. nov. (QCAZ 40847, SVL 22.32 mm)." httpUri="https://binary.pensoft.net/fig/1257796">12</figureCitation>
.
</paragraph>
<caption id="A847FAD894422A392CBB7B75C100C7AA" ID-arpha="7F1907BD-9AD6-511A-A564-8FF0B2D719E1" httpUri="https://binary.pensoft.net/fig/1257795" startId="F11">
<paragraph id="9086BD9CA3455A2ADFD51A0AA8503F85">
<label id="07E997A76B6BAA05C43C474C248D5DBD">Figure 11.</label>
</paragraph>
<paragraph id="47E68C0CDC94DF486A0E8CB734539110">
Metamorph, juvenile and subadults of 
<taxonomicName id="5AF15C149E3CA3DC46D494DDE89E4BF7" authorityName="Aguirre &amp; Apunte &amp; Ron" authorityYear="2025" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="cannatellai" status="sp. nov.">
<emphasis id="8D38E392A33341DC33C0D4AF0A59064C" italics="true">Dendropsophus cannatellai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0D0C481FB4A2346A0BDC752A735E73D9" rank="species">sp. nov.</taxonomicNameLabel>
Dorsolateral views of individuals raised in captivity in Balsa de los Sapos. From left to right, first row: QCAZA l 78576 (total length = 23.27 mm); QCAZA n 79747 (SVL = 11.57 mm); second row: (18.46 mm); (19.59 mm); third row: (19.38 mm); (17.89 mm). All subadults approximately 1 year old since hatching.
</paragraph>
</caption>
<caption id="3516284DB7A31094CEFDD49882C1270C" ID-arpha="E87BA2D7-4349-5AB3-B450-8A307BBE06A9" httpUri="https://binary.pensoft.net/fig/1257796" startId="F12">
<paragraph id="D24CF69FB16FBD51D75005B46BC35EF6">
<label id="388D79523EDD828E5B159B1E61A95AEC">Figure 12.</label>
</paragraph>
<paragraph id="DA7CC3E0BA574F4F2F5EEF5C71AF71D0">
Dorsal, ventral and lateral views of the holotype of 
<taxonomicName id="CA466D71A7A4D204D63B42D11EC91969" authorityName="Aguirre &amp; Apunte &amp; Ron" authorityYear="2025" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="cannatellai" status="sp. nov.">
<emphasis id="6CCE5139593D5BA2345176EEAC24D66F" italics="true">Dendropsophus cannatellai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AEEB4B4A09B7292FFF0B84EDC10A5AE7" rank="species">sp. nov.</taxonomicNameLabel>
(
<named-content id="09A480C2B19ABB20DA3B7D8A465A1F71" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
40847, SVL 22.32 mm).
</paragraph>
</caption>
<caption id="24164134641522A9BE6D5D491D5AB7FF" ID-arpha="1C90E158-FD3D-59AA-A5F7-423D0EED3740" httpUri="https://binary.pensoft.net/fig/1257797" startId="F13">
<paragraph id="567ABAFCD04384FE573425D2EA1C2FDF">
<label id="6E892ACD9CA2FB958043D61DC5E3A3C5">Figure 13.</label>
</paragraph>
<paragraph id="06A902E8803FFC1A8058561D191895CE">
Ventral views of the holotype’s hand (left) and foot (right) of 
<taxonomicName id="65479A83637D259399F3309F6BDC4D05" authorityName="Aguirre &amp; Apunte &amp; Ron" authorityYear="2025" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="cannatellai" status="sp. nov.">
<emphasis id="E228CC01E9B472B3ED657F40B920382C" italics="true">Dendropsophus cannatellai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="664D2A112BA3ED72FFE74152735F327D" rank="species">sp. nov.</taxonomicNameLabel>
(
<named-content id="931A2E011F121AF7EF070AE27C15F26B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
40847).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EOPBI" type="variation">
<paragraph id="A5854C1D39DB49FB866F2B0A44640B1B">
<heading id="1969CB39307E49EDA63A7C22FB3A5529" reason="title">Variation.</heading>
</paragraph>
<paragraph id="3D0BB5796811B61B70F361BDF1CAE2CA">
Sexual dimorphism is observed in the size of adult individuals, where males are smaller (SVL males = 
<quantity id="616D3F6471BDD77C2D856CB12CE919BA" metricMagnitude="-2" metricUnit="m" metricValue="2.3815" metricValueMax="2.658" metricValueMin="2.105" unit="mm" value="23.814999999999998" valueMax="26.58" valueMin="21.05">21.05–26.58 mm</quantity>
; SD = 1.235. SVL females = 
<quantity id="0475795376C226CE87B9B647E51833DF" metricMagnitude="-2" metricUnit="m" metricValue="3.0095" metricValueMax="3.194" metricValueMin="2.825" unit="mm" value="30.095" valueMax="31.94" valueMin="28.25">28.25–31.94 mm</quantity>
; SD = 2.078).
</paragraph>
<paragraph id="8140242AC392754A4C2B1FA578AE0A3F">
Color variation in life and preservative is shown in Figs 
<figureCitation id="34B9B970AB750A079564A22081771888" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Life coloration of Dendropsophus cannatellai sp. nov. First and third rows, dorsolateral views; second and fourth, ventral views. From left to right, first and second rows: QCAZ 51877 (SVL = 22.26 mm, male); QCAZ 78809 (SVL = 33.61 mm, female); QCAZ 55558 (SVL = 21.96 mm, male); third and fourth rows: QCAZ 40847 (holotype; SVL = 22.32 mm, male); QCAZ 78812 (SVL = 25.86 mm, male); QCAZ 78813 (SVL = 24.94 mm, male). See type series for locality data. All specimens are shown at the same scale." httpUri="https://binary.pensoft.net/fig/1257792">8</figureCitation>
, 
<figureCitation id="C67BA71813A84231E348C51F25D3E888" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Adult preserved specimens of Dendropsophus cannatellai sp. nov. First and third rows, dorsolateral views; second and fourth, ventral views. From left to right, first and second rows: QCAZ 9457 (SVL = 28.65 mm, female); QCAZ 23162 (22.83 mm, male); QCAZ 26098 (30.73 mm, female); third and fourth rows: QCAZ 35710 (24.74 mm, male); QCAZ 78254 (24.71 mm, male); QCAZ 78257 (31.94 mm, female). See type-series for locality data. All specimens are shown at the same scale." httpUri="https://binary.pensoft.net/fig/1257793">9</figureCitation>
. In some individuals (e. g., 
<named-content id="B82ACD6E74A74FC7FC8B6201D66843F9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">
<collectionCode id="A6A14FB3C735A1A3880B73AE4DBD97EB" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" type="Museum">QCAZ</collectionCode>
</named-content>
55555) the dorsolateral bands are extensive and fused with the sacral mark. There is variation in the extent of membrane between the digits of the forelimbs, particularly between fingers II, III, and IV, where adult females have more extensive membrane. In dorsal view, snout shape varies from truncate to slightly rounded. In preserved male specimens, the vocal sac is evident as wrinkles in the gular region. However, in some individuals, these wrinkles are less noticeable.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBQBI" type="distribution">
<paragraph id="53F03530A1B52485265E9882460C53D4">
<heading id="3B216542F93914B4514847CBAE3CD3A3" reason="title">Distribution, natural history, and conservation status.</heading>
</paragraph>
<paragraph id="C2567D59AF59A6BF0C9C30979B24E340">
<taxonomicName id="30361F608CD3D14CC2E86D350A2D527A" authorityName="Aguirre &amp; Apunte &amp; Ron" authorityYear="2025" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="cannatellai" status="sp. nov.">
<emphasis id="CFC67B620415B888CCA7D45209B42AB1" italics="true">Dendropsophus cannatellai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CB53A6B1DD550F8BF120CAD08491223D" rank="species">sp. nov.</taxonomicNameLabel>
occurs in Western Foothill Forest and the Chocó Tropical Rainforest in NW 
<collectingCountry id="0127508767413EFB2F1E7A522C4342EF" name="Ecuador">Ecuador</collectingCountry>
and SW 
<collectingCountry id="F5C6624342E07C5A84BFBE9DE7DDA788" name="Colombia">Colombia</collectingCountry>
. Although the occurrence in 
<collectingCountry id="851276B10ADACF826CFD199804D2E522" name="Colombia">Colombia</collectingCountry>
needs to be corroborated, we found the following evidence of its presence: (1) localities in 
<collectingCountry id="3B19CA2172704677492D719F600296FC" name="Ecuador">Ecuador</collectingCountry>
near the border with 
<collectingCountry id="103D24C369DF520EBA334983434EF664" name="Colombia">Colombia</collectingCountry>
, and (2) records on iNaturalist (observation numbers 
<quantity id="BDF2946F5530E8E9BD3F6A2FD3B0B171" metricMagnitude="6" metricUnit="m" metricValue="3.7574482636000006" unit="in" value="1.47931034E8">147931034 in</quantity>
Tumaco, 
<collectingRegion id="8332042179CCB2C99132EB9539327227" country="Colombia" name="Narino">Nariño</collectingRegion>
, and 
<quantity id="9B9A4FDBDEEA4E786D6B7B93CA9D647A" metricMagnitude="5" metricUnit="m" metricValue="9.193954942000001" unit="in" value="3.6196673E7">36196673 in</quantity>
<collectingRegion id="AA2CE3973D0519535BDA9FB5C0ECE573" country="Colombia" name="Providencia y Santa Catalina">San Andrés</collectingRegion>
de Tumaco), morphologically indistinguishable from the new species.
</paragraph>
<caption id="9CFF935407C423FAE0E8CEFDFF377EC1" ID-arpha="C4B8C7B7-5AC2-5A67-BD7A-C6189345ACEE" httpUri="https://binary.pensoft.net/fig/1257798" startId="F14">
<paragraph id="F38024404A19BD13EC0BB68487CA21B1">
<label id="18112FAA50B699DF69AEEEAD34F6DE36">Figure 14.</label>
</paragraph>
<paragraph id="B2E8EB51B9169C6C12F6A6E23F42AEEA">
Life coloration of 
<taxonomicName id="7C3B31A40803337BEA1A8CAA650ED079" baseAuthorityName="Duellman" baseAuthorityYear="1973" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="gryllatus">
<emphasis id="93DA21C5B52475A675D89ECBE37D88B8" italics="true">Dendropsophus gryllatus</emphasis>
</taxonomicName>
. First and third rows, dorsolateral views; second and fourth, ventral views. From left to right, first and second rows: 
<named-content id="A31C3D3837434AE24DEA0F2ACACD807A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
75534 (SVL = 20.80 mm); 
<named-content id="C424C205AE1A5C080ECF93EF39F47719" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
75535 (SVL = 21.19 mm); 
<named-content id="05B44CD96DBE6618B783F51B35C654B6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
75536 (SVL = 22.24 mm); third and fourth rows: 
<named-content id="535D5069A80F09338086B443F5628AE0" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
75538 (SVL = 18.71 mm); 
<named-content id="A6C9C61903F8F3E56C01CA4B1F6C352B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
75539 (SVL = 21.89 mm); 
<named-content id="EDCA2E7D57AD3A97FE8E439854E68910" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
75540 (SVL = 20.26 mm); all adult males. See Suppl. material 
<xref id="C7766108367E95507416A39D5FFD8499" ref-type="supplementary-material" rid="S1">1</xref>
, table S 1 for locality data. All specimens are shown at the same scale.
</paragraph>
</caption>
<paragraph id="6B16CBD0983910893D16DCDFA05829BC">
Within its distribution range, climatic conditions are generally warm and humid, with an annual average temperature ranging from 22.5 to 25.4 ° C and an average annual precipitation with a range from 
<quantity id="183629768CC11F444C33DB9AA874DF0D" metricMagnitude="0" metricUnit="m" metricValue="2.1865" metricValueMax="3.348" metricValueMin="1.025" unit="mm" value="2186.5" valueMax="3348.0" valueMin="1025.0">1025 to 3348 mm</quantity>
. The physiognomy of these ecosystems consists of big trees that can reach almost 
<quantity id="5E45BF22B7ACD0ADBB770BC8DC5E506C" metricMagnitude="1" metricUnit="m" metricValue="3.0" unit="m" value="30.0">30 m</quantity>
tall, and a great abundance of palms. The Western Foothill Forest has an elevation range from 
<quantity id="9968467DE183C6DA2D577AADF2824D0E" metricMagnitude="2" metricUnit="m" metricValue="8.0" metricValueMax="13.0" metricValueMin="3.0" unit="m" value="800.0" valueMax="1300.0" valueMin="300.0">300 to 1300 m</quantity>
, with a dominance of plants from the families 
<taxonomicName id="C7EE161D8E1DAFC3C47CC68E0232E38B" class="Magnoliopsida" family="Mimosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="family">Mimosaceae</taxonomicName>
, 
<taxonomicName id="CBAD1EA7C48B155735C8DA1C5D8E7426" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
, and 
<taxonomicName id="A1A2EBA345A87B2FFA55C9C58F3AFFFF" class="Magnoliopsida" family="Burseraceae" kingdom="Plantae" order="Sapindales" phylum="Tracheophyta" rank="family">Burseraceae</taxonomicName>
with a high percentage of endemism. The presence of epiphytes is common. The 
<collectingRegion id="12C99E40C325C675D92C34203B6C93FA" country="Panama" name="Embera-Wounaan">Chocó</collectingRegion>
Tropical Rainforest has an elevation from 
<quantity id="E736DB21F5A9E5DC1531F8646E596218" metricMagnitude="2" metricUnit="m" metricValue="1.5" metricValueMax="3.0" metricValueMin="0.0" unit="m" value="150.0" valueMax="300.0" valueMin="0.0">0 to 300 m</quantity>
, with an abundance of ferns and a high diversity of trees (
<bibRefCitation id="6C678D13B746A32F9BBAB9910DA3C441" author="Ron" etAl="et al." firstAuthor="Ron" refId="B42" refString="Ron SR, Merino-Viteri A, Ortiz DA (2024) Anfibios del Ecuador. Version 2024.0. https://bioweb.bio/faunaweb/amphibiaweb/" url="https://bioweb.bio/faunaweb/amphibiaweb/" year="2024">Ron et al. 2024</bibRefCitation>
).
</paragraph>
<caption id="0D847DAB7611321352E7141CCBEC8663" ID-arpha="7D25B151-5A67-50FC-BB73-B2DCB9511073" httpUri="https://binary.pensoft.net/fig/1257799" startId="F15">
<paragraph id="A1119AE77A8C5D05AD22D731D90ECA66">
<label id="4B8781E67D360022EEE3917546361844">Figure 15.</label>
</paragraph>
<paragraph id="240DAC64F86FBDE43122FD5E94DBAD2F">
Adult preserved specimens of 
<taxonomicName id="BC26E7475243A19434DBCDEA7ACF24FF" baseAuthorityName="Duellman" baseAuthorityYear="1973" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="gryllatus">
<emphasis id="5CB9853765D65A07F6CF80DFF853AC4E" italics="true">Dendropsophus gryllatus</emphasis>
</taxonomicName>
showing variation in dorsal and ventral coloration. From left to right, first and second rows: 
<named-content id="7CBE76E7DD67713DBE68FEC1FC0A6CA9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
75532 (22.54 mm); 
<named-content id="4403F5252473841C50D3BE6D8491ED60" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
75533 (21.96 mm); 
<named-content id="BC47A6895D183B35D0486362B30AC0F2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
75535 (21.19 mm); third and fourth rows: 
<named-content id="82A2B113F5D8E6A47E9CAD9FE700921E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
75536 (22.24 mm); 
<named-content id="CC33EE6F607BFCBAFABAEB10FFCDE995" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
75537 (22.85 mm); 
<named-content id="20B052FE9EA6253C25646181C8592D8C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
75539 (21.89 mm); all adult males. See Suppl. material 
<xref id="E9B0FEDEC59BB4ECBE931608853D8DF4" ref-type="supplementary-material" rid="S1">1</xref>
, table S 1 for locality data. All specimens are shown at the same scale.
</paragraph>
</caption>
<paragraph id="1CDB019320A2A2D69A3DE82C7B27D7EE">
The new species is found in artificial open areas, near ponds and puddles where herbaceous vegetation is abundant. It is less frequent in secondary forest. Males call from leaves or grasses less than 
<quantity id="02313FBC32FE71EEF8351B72E8C3FFA8" metricMagnitude="0" metricUnit="m" metricValue="2.0" unit="m" value="2.0">2 m</quantity>
above water, between the 18: 00 and 0: 00 hours. A couple collected in Río Silanche, laid a clutch in captivity with 
<specimenCount id="343B9D17624B4DC81BF175CCAD38A839" count="190" type="egg">190 eggs</specimenCount>
. Experiments in captivity report average clutch sizes of 
<specimenCount id="3DE1BCA8C2FC3C711F0DF586E64CD17A" count="84" type="egg">84 eggs</specimenCount>
under shaded conditions and 98 under unshaded conditions (
<bibRefCitation id="CC46157FC3C421C761ACA8F4AF497A6C" DOI="10.1073/pnas.2312371121" author="Touchon" etAl="et al." firstAuthor="Touchon" refId="B47" refString="Touchon JC, McMillan WO, Ibáñez R, Lessios HA (2024) Flexible oviposition behavior enabled the evolution of terrestrial reproduction. Proceedings of the National Academy of Sciences 121: e 2312371121. https://doi.org/10.1073/pnas.2312371121" year="2024">Touchon et al. 2024</bibRefCitation>
). In the same experiments, most females laid terrestrial eggs, but 30 % of females laid a few eggs in water, under unshaded conditions.
</paragraph>
<caption id="2B0A3C9D7F9575A5829013F4920D3C88" ID-arpha="A256B96F-2A0F-5C79-B388-71773F7CD1CA" httpUri="https://binary.pensoft.net/fig/1257800" startId="F16">
<paragraph id="87F947B6709ADBA108C6EF8D4A8D63F3">
<label id="EAC53202A203F853A32CC6AA9272693C">Figure 16.</label>
</paragraph>
<paragraph id="987CE8478C2D2B9EB8E601F8F7280E9C">
Distribution of the species 
<taxonomicName id="DAEFEAF1A144BFBD8258B23DD956333B" authorityName="Aguirre &amp; Apunte &amp; Ron" authorityYear="2025" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="cannatellai" status="sp. nov.">
<emphasis id="75EB7A1AF337B8BE5E6846B68CD5B2CC" italics="true">Dendropsophus cannatellai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0C4BFDD833302A53ED47F2B3475E0D10" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4AC70046ED28A2F184A6FCAB4A52F7E4" baseAuthorityName="Duellman" baseAuthorityYear="1973" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="gryllatus">
<emphasis id="84EC9368EC29A745AC841B6D54947699" italics="true">D. gryllatus</emphasis>
</taxonomicName>
. Localities are based of specimens deposited at Pontificia Universidad Católica del Ecuador, 
<named-content id="23133DB60C7DB461B9294A0DE924121B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador">QCAZ</named-content>
Museum, 
<bibRefCitation id="5EDB8D1C3879004526E9D7B9A16C7289" author="Duellman WE" firstAuthor="Duellman" journalOrPublisher="Occasional Papers of the Museum of Natural History, University of Kansas" pagination="1-27" refId="B8" refString="Duellman WE (1974) A reassessment of the taxonomic status of some Neotropical hylid frogs. Occasional Papers of the Museum of Natural History, University of Kansas 27: 1–27." title="A reassessment of the taxonomic status of some Neotropical hylid frogs." volume="27" year="1974">Duellman (1974)</bibRefCitation>
, and 
<bibRefCitation id="3785409B8E09A949B9D3AA01F69961CD" author="Michelsohn MJ &amp; Barrow LN &amp; Moretta M &amp; Achig-Vega A &amp; Nusirwuia K &amp; Ron SR" etAl="et al." firstAuthor="Michelsohn" issue="2" journalOrPublisher="Herpetological Review" pagination="257-258" refId="B26" refString="Michelsohn MJ, Barrow LN, Moretta M, Achig-Vega A, Nusirwuia K, Ron SR (2022) Geographic distribution notes. Dendropsophus gryllatus (Pacific Lowland Treefrog). Herpetological Review 53 (2): 257–258." title="Geographic distribution notes. Dendropsophus gryllatus (Pacific Lowland Treefrog)." volume="53" year="2022">Michelsohn et al. (2022)</bibRefCitation>
. Shapes with white outlines represent type localities. All known localities for both species are shown.
</paragraph>
</caption>
<paragraph id="FF6D7510BFC618D94E8413AA9CF497C8">
The new species is assigned to the Red List category Near Threatened (
<abbrev id="ABBRID0EGWBI" xlink_title="Near Threatened">NTU</abbrev>
). Its extent of occurrence (
<abbrev id="ABBRID0EKWBI" xlink_title="extent of occurrence">EOO</abbrev>
) has &lt;
<quantity id="3838CBC1C73D6C6A17C4C24F47D50663" metricMagnitude="7" metricUnit="m" metricValue="2.0" unit="km" value="20000.0">20000 km</quantity>
<superScript id="14CF825E06C7219C0E9BF51506BBDA46">2</superScript>
(minimum convex polygon of the known localities of 
<taxonomicName id="87A977F3225873C62B25911F53480F03" authorityName="Aguirre &amp; Apunte &amp; Ron" authorityYear="2025" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="cannatellai" status="sp. nov.">
<emphasis id="4E17E18BD6F6FBA4B5EC7F1BF13DFC07" italics="true">D. cannatellai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0E1444B62C69721CF452BB52597CF09F" rank="species">sp. nov.</taxonomicNameLabel>
~ 
<quantity id="961F4D4BECA2702FB1C571CF933AD6F2" metricMagnitude="6" metricUnit="m" metricValue="6.1339999999999995" unit="km" value="6134.0">6134 km</quantity>
<superScript id="32CB01EBB296F5680BCD1B619F447C9E">2</superScript>
). However, populations are predominantly located on disturbed and deforested areas, near roads. The species appears to thrive in human modified habitats including pastures for cattle raising and ponds in artificial open areas with abundant herbaceous vegetation.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4WBI" type="etymology">
<paragraph id="11846F5B95303E9471D524EC949FB399">
<heading id="A4802A3BDF0EBEF9CAE9CA1AB3696E04" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="5CDE4D168BD34B225A2BF428F2551D66">
The specific name 
<taxonomicName id="0E58D023E447445A378F8C9683FFBED3" authorityName="Aguirre &amp; Apunte &amp; Ron" authorityYear="2025" class="Amphibia" family="Hylidae" genus="Dendropsophus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="cannatellai">
<emphasis id="727961394118F7D53F9F24AF289C2923" italics="true">cannatellai</emphasis>
</taxonomicName>
is a noun in the genitive case and is a patronym for David C. Cannatella, who collected 
<typeStatus id="73A2E000DED5516E936F9F87973B7AEA">type</typeStatus>
specimens of the new species. Moreover, during his career he has contributed extensively to the study of systematics and evolution of neotropical amphibians. He has been integral part of scientific societies, exemplified by his presidencies of the American Society of Ichthyologists and Herpetologists (2002–2003) and the Society of Systematic Biologists (2004–2005). Currently he is Dean of the Department of Integrative Biology at the University of 
<collectingRegion id="B951BA72A1728BF1D1091C041107CF1D" country="United States of America" name="Texas">Texas</collectingRegion>
at Austin, 
<collectingCountry id="CAD81C1FA1AA13DE8F32AEAF762082FB" name="United States of America">USA</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>