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<document id="1E4D9AD4361975B8610FFFA17D1D0EFF" ID-CLB-Dataset="51232" ID-DOI="10.5281/zenodo.157034" ID-GBIF-Dataset="3034e634-02b3-4e21-b89f-a785b2f545dd" ID-ISSN="1175­5326" ID-Zenodo-Dep="157034" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459640220932" checkinUser="plazi" docAuthor="Ahrens, Dirk" docDate="2003" docId="345087C7FFF3FFC9FEB9FDA6FB28352C" docLanguage="en" docName="zt00392.pdf" docOrigin="Zootaxa 392" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Maladera infuscata Moser 1915" docType="treatment" docVersion="9" lastPageNumber="7" masterDocId="C869FFBFFFF2FFCFFFB1FFD8FFAE375A" masterDocTitle="The identity of Maladera infuscata (Moser, 1915), with description of a new species from Taiwan (Coleoptera: Scarabaeidae: Sericini)" masterLastPageNumber="8" masterPageNumber="1" pageNumber="2" updateTime="1698206224494" updateUser="ExternalLinkService">
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<mods:title id="3662A3ECE3B93CCDFBA5400FBE9A331E">The identity of Maladera infuscata (Moser, 1915), with description of a new species from Taiwan (Coleoptera: Scarabaeidae: Sericini)</mods:title>
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<mods:date id="2186F0C0839D0E2024BC27763C3FFE2F">2003</mods:date>
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<treatment id="345087C7FFF3FFC9FEB9FDA6FB28352C" ID-DOI="http://doi.org/10.5281/zenodo.6275287" ID-GBIF-Taxon="119332683" ID-Zenodo-Dep="6275287" LSID="urn:lsid:plazi:treatment:345087C7FFF3FFC9FEB9FDA6FB28352C" httpUri="http://treatment.plazi.org/id/345087C7FFF3FFC9FEB9FDA6FB28352C" lastPageId="6" lastPageNumber="7" pageId="1" pageNumber="2">
<subSubSection id="F4E3655AFFF3FFCEFEB9FDA6FC8D35C2" box="[264,803,638,664]" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="BC4636D1FFF3FFCEFEB9FDA6FC8D35C2" blockId="1.[264,803,638,664]" box="[264,803,638,664]" pageId="1" pageNumber="2">
<heading id="E70E81BDFFF3FFCEFEB9FDA6FC8D35C2" bold="true" box="[264,803,638,664]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
<emphasis id="8E8DEAC3FFF3FFCEFEB9FDA6FC8D35C2" bold="true" box="[264,803,638,664]" pageId="1" pageNumber="2">
<taxonomicName id="7BF94D52FFF3FFCEFEB9FDA6FD0B35C2" authority="Moser, 1915" authorityName="Moser" authorityYear="1915" box="[264,677,638,664]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="infuscata">
<emphasis id="8E8DEAC3FFF3FFCEFEB9FDA6FE5A35C2" bold="true" box="[264,500,638,664]" italics="true" pageId="1" pageNumber="2">Maladera infuscata</emphasis>
(
<bibRefCitation id="D8684B20FFF3FFCEFDB5FDA6FD3335C2" author="Moser" box="[516,669,638,664]" pageId="1" pageNumber="7" refString="Moser, J. (1915) Neue Serica-Arten. Deutsche Entomologische Zeitschrift, 1915, 337 - 393." type="journal article" year="1915">Moser, 1915</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="24C22A54FFF3FFCEFD04FDA6FCB435C2" box="[693,794,638,664]" captionStart="FIGURES 1 – 7" captionStartId="3.[264,370,1792,1813]" captionTargetBox="[322,1265,344,1754]" captionTargetPageId="3" captionText="FIGURES 1 – 7: 1 – 3 Maladera infuscata (Moser, 1915) (Lectotype: “ Korea ”); 4 – 7: Maladera howdeni sp. n. (Holotype: Taiwan, Wushe); 1, 3, 4, 7: aedeagus, lateral view; 2: aedeagus, dorsal view; 5: parameres, dorsal view; 6: parameres, ventral view. Scale: 0.5 mm." httpUri="https://zenodo.org/record/168876/files/figure.png" pageId="1" pageNumber="2">Figs 1­3</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F4E3655AFFF3FFCEFEB9FD12FBE7331B" pageId="1" pageNumber="2" type="reference_group">
<paragraph id="BC4636D1FFF3FFCEFEB9FD12FC5635BA" blockId="1.[264,1322,713,1089]" box="[264,1016,713,737]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="9CE911FFFFF3FFCEFEB9FD12FC5635BA" box="[264,1016,713,737]" pageId="1" pageNumber="2">
<taxonomicName id="7BF94D52FFF3FFCEFEB9FD12FD0735BA" authority="Moser, 1915: 340" authorityName="Moser" authorityPageNumber="340" authorityYear="1915" box="[264,681,713,737]" class="Insecta" family="Melolonthidae" genus="Autoserica" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="infuscata">
<emphasis id="8E8DEAC3FFF3FFCEFEB9FD12FE4A35BB" box="[264,484,714,737]" italics="true" pageId="1" pageNumber="2">Autoserica infuscata</emphasis>
<treatmentCitation id="3D5810C0FFF3FFCEFE5AFD11FD0735BA" author="Moser" box="[491,681,713,736]" page="340" pageId="1" pageNumber="2" year="1915">
<bibRefCitation id="D8684B20FFF3FFCEFE5AFD11FDDA35BA" author="Moser" box="[491,628,713,736]" pageId="1" pageNumber="7" refString="Moser, J. (1915) Neue Serica-Arten. Deutsche Entomologische Zeitschrift, 1915, 337 - 393." type="journal article" year="1915">Moser, 1915</bibRefCitation>
: 340
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="3D5810C0FFF3FFCEFD05FD11FC5A35BA" author="Nijima" box="[692,1012,713,736]" page="6" pageId="1" pageNumber="2" year="1927">
<bibRefCitation id="D8684B20FFF3FFCEFD05FD11FC7735BA" author="Nijima" box="[692,985,713,736]" pageId="1" pageNumber="7" refString="Niijima, Y. &amp; Kinoshita, E. (1927): Revision of Japanese Melolonthids and allied families. III. Research Bulletin of the College Expermental Forestry Hokkaido Imperial University, 4, 1 - 97." type="journal article" year="1927">Nijima and Kinoshita 1927</bibRefCitation>
: 6
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="BC4636D1FFF3FFCEFEB9FD32FCFB345A" blockId="1.[264,1322,713,1089]" box="[264,853,745,769]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="9CE911FFFFF3FFCEFEB9FD32FCFB345A" box="[264,853,745,769]" pageId="1" pageNumber="2">
<taxonomicName id="7BF94D52FFF3FFCEFEB9FD32FD06345A" authority="Murayama 1938: 13" authorityName="Murayama" authorityPageNumber="13" authorityYear="1938" box="[264,680,745,769]" class="Insecta" family="Scarabaeidae" genus="Aserica" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="infuscata">
<emphasis id="8E8DEAC3FFF3FFCEFEB9FD32FE6C345B" box="[264,450,746,769]" italics="true" pageId="1" pageNumber="2">Aserica infuscata</emphasis>
:
<treatmentCitation id="3D5810C0FFF3FFCEFE61FD31FD06345A" author="Murayama" box="[464,680,745,768]" page="13" pageId="1" pageNumber="2" year="1938">
<bibRefCitation id="D8684B20FFF3FFCEFE61FD31FD2D345A" author="Murayama" box="[464,643,745,768]" pageId="1" pageNumber="7" refString="Murayama, J. (1938) Revision des Sericines (Col., Scar.) de la Coree. Annotiones Zoologicae Japonenses, 17 (1), 7 - 21." type="journal article" year="1938">Murayama 1938</bibRefCitation>
: 13
</treatmentCitation>
</taxonomicName>
[misidentified].
</treatmentCitationGroup>
</paragraph>
<paragraph id="BC4636D1FFF3FFCEFEB9FCD2FCE9347A" blockId="1.[264,1322,713,1089]" box="[264,839,777,801]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="9CE911FFFFF3FFCEFEB9FCD2FCE9347A" box="[264,839,777,801]" pageId="1" pageNumber="2">
<taxonomicName id="7BF94D52FFF3FFCEFEB9FCD2FD35347A" authority="Murayama 1954: 41" authorityName="Murayama" authorityPageNumber="41" authorityYear="1954" box="[264,667,777,801]" class="Insecta" family="Melolonthidae" genus="Serica" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="infuscata">
<emphasis id="8E8DEAC3FFF3FFCEFEB9FCD2FE1B347B" box="[264,437,778,801]" italics="true" pageId="1" pageNumber="2">Serica infuscata</emphasis>
:
<treatmentCitation id="3D5810C0FFF3FFCEFE72FCD1FD35347A" author="Murayama" box="[451,667,777,800]" page="41" pageId="1" pageNumber="2" year="1954">
<bibRefCitation id="D8684B20FFF3FFCEFE72FCD1FDD8347A" author="Murayama" box="[451,630,777,800]" pageId="1" pageNumber="7" refString="Murayama, J. (1954) Icones of the Scarabeid-Beetles from Manchuria and Korea. I. Nihon Gakujutu, Shinko-kai, 163 pp." type="book" year="1954">Murayama 1954</bibRefCitation>
: 41
</treatmentCitation>
</taxonomicName>
[misidentified].
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</paragraph>
<paragraph id="BC4636D1FFF3FFCEFEB9FCF2FBAB343A" blockId="1.[264,1322,713,1089]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="9CE911FFFFF3FFCEFEB9FCF2FBAB343A" pageId="1" pageNumber="2">
<taxonomicName id="7BF94D52FFF3FFCEFEB9FCF2FD07341A" authority="Nomura1959: 40" authorityName="Nomura" authorityPageNumber="40" authorityYear="1959" box="[264,681,809,833]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="infuscata">
<emphasis id="8E8DEAC3FFF3FFCEFEB9FCF2FE73341B" box="[264,477,810,833]" italics="true" pageId="1" pageNumber="2">Maladera infuscata</emphasis>
:
<treatmentCitation id="3D5810C0FFF3FFCEFE5CFCF1FD07341A" author="Nomura" box="[493,681,809,832]" page="40" pageId="1" pageNumber="2" year="1959">
<bibRefCitation id="D8684B20FFF3FFCEFE5CFCF1FDD1341A" author="Nomura" box="[493,639,809,832]" pageId="1" pageNumber="7" refString="Nomura, S. (1959) Notes on the Japanese Scarabaeidae with two subspecies from Formosa. Toho- Gakuho, 9, 39 - 54." type="journal article" year="1959">Nomura1959</bibRefCitation>
: 40
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="3D5810C0FFF3FFCEFD06FCF1FC26341A" author="Nomura" box="[695,904,809,832]" page="104" pageId="1" pageNumber="2" year="1974">
<bibRefCitation id="D8684B20FFF3FFCEFD06FCF1FCFF341A" author="Nomura" box="[695,849,809,832]" pageId="1" pageNumber="8" refString="Nomura, S. (1974) On the Sericini of Taiwan. Toho-Gakuho, 24, 81 - 115." type="journal article" year="1974">Nomura 1974</bibRefCitation>
: 104
</treatmentCitation>
, 109, fig. 48 [misidentified];
<treatmentCitation id="3D5810C0FFF3FFCEFB61FCF1FE5F343A" author="Sun" page="324" pageId="1" pageNumber="2" year="1982">
<bibRefCitation id="D8684B20FFF3FFCEFB61FCF1FE12343A" author="Sun" pageId="1" pageNumber="8" refString="Sun, Y. &amp; Zhang, Z. (1982) A study on three sibling white grubs. Entomotaxonomia, 4 (4), 323 - 324." type="journal article" year="1982">Sun and Zhang 1982</bibRefCitation>
: 324
</treatmentCitation>
;
<treatmentCitation id="3D5810C0FFF3FFCEFE4DFC91FD52343A" author="Kim" box="[508,764,841,864]" page="122" pageId="1" pageNumber="2" year="1997">
<bibRefCitation id="D8684B20FFF3FFCEFE4DFC91FD69343A" author="Kim" box="[508,711,841,864]" pageId="1" pageNumber="7" refString="Kim, J. I. &amp; Lee, O. J. (1997) Taxonomic study of Korean Sericinae (Melolonthidae, Coleoptera) II, Genus Maladera. Insecta Koreana, 14, 119 - 135." type="journal article" year="1997">Kim and Lee 1997</bibRefCitation>
: 122
</treatmentCitation>
;
<treatmentCitation id="3D5810C0FFF3FFCEFCB6FC91FBAF343A" author="Kim" box="[775,1025,841,864]" page="89" pageId="1" pageNumber="2" year="2003">
<bibRefCitation id="D8684B20FFF3FFCEFCB6FC91FC77343A" author="Kim" box="[775,985,841,864]" pageId="1" pageNumber="7" refString="Kim, J. I. &amp; Kim, A-Y. (2003) Taxonomic Review of Korean Sericinae (Coleoptera, Melolonthidae) II: Genus Maladera Mulsant. Insecta Koreana, 20 (1), 81 - 94." type="journal article" year="2003">Kim and Kim 2003</bibRefCitation>
: 89
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="BC4636D1FFF3FFCEFEB9FCB2FD0C34DA" blockId="1.[264,1322,713,1089]" box="[264,674,873,897]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="9CE911FFFFF3FFCEFEB9FCB2FD0C34DA" box="[264,674,873,897]" pageId="1" pageNumber="2">
<taxonomicName id="7BF94D52FFF3FFCEFEB9FCB2FD3034DA" authority="Moser, 1919: 330" authorityName="Moser" authorityPageNumber="330" authorityYear="1919" box="[264,670,873,897]" class="Insecta" family="Melolonthidae" genus="Autoserica" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="koreana">
<emphasis id="8E8DEAC3FFF3FFCEFEB9FCB2FE7634DB" box="[264,472,874,897]" italics="true" pageId="1" pageNumber="2">Autoserica koreana</emphasis>
<treatmentCitation id="3D5810C0FFF3FFCEFE6EFCB1FD3034DA" author="Moser" box="[479,670,873,896]" page="330" pageId="1" pageNumber="2" year="1919">
<bibRefCitation id="D8684B20FFF3FFCEFE6EFCB1FDC734DA" author="Moser" box="[479,617,873,896]" pageId="1" pageNumber="7" refString="Moser, J. (1919) Beitrag zur Kenntnis der Melolonthiden. Stettiner Entomologische Zeitung, 80: 330 - 364." type="journal article" year="1919">Moser, 1919</bibRefCitation>
: 330
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="BC4636D1FFF3FFCEFEB9FC52FCE734FA" blockId="1.[264,1322,713,1089]" box="[264,841,905,929]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="9CE911FFFFF3FFCEFEB9FC52FCE734FA" box="[264,841,905,929]" pageId="1" pageNumber="2">
<taxonomicName id="7BF94D52FFF3FFCEFEB9FC52FD3334FA" authority="Murayama 1938: 15" authorityName="Murayama" authorityPageNumber="15" authorityYear="1938" box="[264,669,905,929]" class="Insecta" family="Scarabaeidae" genus="Aserica" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="koreana">
<emphasis id="8E8DEAC3FFF3FFCEFEB9FC52FE1834FB" box="[264,438,906,929]" italics="true" pageId="1" pageNumber="2">Aserica koreana</emphasis>
:
<treatmentCitation id="3D5810C0FFF3FFCEFE74FC51FD3334FA" author="Murayama" box="[453,669,905,928]" page="15" pageId="1" pageNumber="2" year="1938">
<bibRefCitation id="D8684B20FFF3FFCEFE74FC51FDD634FA" author="Murayama" box="[453,632,905,928]" pageId="1" pageNumber="7" refString="Murayama, J. (1938) Revision des Sericines (Col., Scar.) de la Coree. Annotiones Zoologicae Japonenses, 17 (1), 7 - 21." type="journal article" year="1938">Murayama 1938</bibRefCitation>
: 15
</treatmentCitation>
</taxonomicName>
[misidentified].
</treatmentCitationGroup>
</paragraph>
<paragraph id="BC4636D1FFF3FFCEFEB9FC72FC92349A" blockId="1.[264,1322,713,1089]" box="[264,828,937,961]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="9CE911FFFFF3FFCEFEB9FC72FC92349A" box="[264,828,937,961]" pageId="1" pageNumber="2">
<taxonomicName id="7BF94D52FFF3FFCEFEB9FC72FD3E349A" authority="Murayama 1954: 65" authorityName="Murayama" authorityPageNumber="65" authorityYear="1954" box="[264,656,937,961]" class="Insecta" family="Melolonthidae" genus="Serica" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="koreana">
<emphasis id="8E8DEAC3FFF3FFCEFEB9FC72FE07349B" box="[264,425,938,961]" italics="true" pageId="1" pageNumber="2">Serica koreana</emphasis>
:
<treatmentCitation id="3D5810C0FFF3FFCEFE06FC71FD3E349A" author="Murayama" box="[439,656,937,960]" page="65" pageId="1" pageNumber="2" year="1954">
<bibRefCitation id="D8684B20FFF3FFCEFE06FC71FDC5349A" author="Murayama" box="[439,619,937,960]" pageId="1" pageNumber="7" refString="Murayama, J. (1954) Icones of the Scarabeid-Beetles from Manchuria and Korea. I. Nihon Gakujutu, Shinko-kai, 163 pp." type="book" year="1954">Murayama 1954</bibRefCitation>
: 65
</treatmentCitation>
</taxonomicName>
[misidentified].
</treatmentCitationGroup>
</paragraph>
<paragraph id="BC4636D1FFF3FFCEFEB9FC12FC7B34BB" blockId="1.[264,1322,713,1089]" box="[264,981,969,993]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="9CE911FFFFF3FFCEFEB9FC12FC7B34BB" box="[264,981,969,993]" pageId="1" pageNumber="2">
<taxonomicName id="7BF94D52FFF3FFCEFEB9FC12FD7434BA" authority="Kim and Lee 1997: 129" authorityName="Kim and Lee" authorityPageNumber="129" authorityYear="1997" box="[264,730,969,993]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="koreana">
<emphasis id="8E8DEAC3FFF3FFCEFEB9FC12FE6034BB" box="[264,462,970,993]" italics="true" pageId="1" pageNumber="2">Maladera koreana</emphasis>
:
<treatmentCitation id="3D5810C0FFF3FFCEFE6DFC11FD7434BA" author="Kim" box="[476,730,969,992]" page="129" pageId="1" pageNumber="2" year="1997">
<bibRefCitation id="D8684B20FFF3FFCEFE6DFC11FD0934BA" author="Kim" box="[476,679,969,992]" pageId="1" pageNumber="7" refString="Kim, J. I. &amp; Lee, O. J. (1997) Taxonomic study of Korean Sericinae (Melolonthidae, Coleoptera) II, Genus Maladera. Insecta Koreana, 14, 119 - 135." type="journal article" year="1997">Kim and Lee 1997</bibRefCitation>
: 129
</treatmentCitation>
</taxonomicName>
[misidentified],
<emphasis id="8E8DEAC3FFF3FFCEFC3DFC11FC7B34BB" bold="true" box="[908,981,969,993]" pageId="1" pageNumber="2">syn. n.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="BC4636D1FFF3FFCEFEB9FC32FD1E335A" blockId="1.[264,1322,713,1089]" box="[264,688,1001,1025]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="9CE911FFFFF3FFCEFEB9FC32FD1E335A" box="[264,688,1001,1025]" pageId="1" pageNumber="2">
<taxonomicName id="7BF94D52FFF3FFCEFEB9FC32FD05335A" ID-CoL="7RXYH" authority="Murayama, 1938: 18" authorityName="Murayama" authorityPageNumber="18" authorityYear="1938" box="[264,683,1001,1025]" class="Insecta" family="Scarabaeidae" genus="Aserica" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="okamotoi">
<emphasis id="8E8DEAC3FFF3FFCEFEB9FC32FE6D335B" box="[264,451,1002,1025]" italics="true" pageId="1" pageNumber="2">Aserica okamotoi</emphasis>
<treatmentCitation id="3D5810C0FFF3FFCEFE7BFC31FD05335A" author="Murayama" box="[458,683,1001,1024]" page="18" pageId="1" pageNumber="2" year="1938">
<bibRefCitation id="D8684B20FFF3FFCEFE7BFC31FD2A335A" author="Murayama" box="[458,644,1001,1024]" pageId="1" pageNumber="7" refString="Murayama, J. (1938) Revision des Sericines (Col., Scar.) de la Coree. Annotiones Zoologicae Japonenses, 17 (1), 7 - 21." type="journal article" year="1938">Murayama, 1938</bibRefCitation>
: 18
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="BC4636D1FFF3FFCEFEB9FBD2FD0A337A" blockId="1.[264,1322,713,1089]" box="[264,676,1033,1057]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="9CE911FFFFF3FFCEFEB9FBD2FD0A337A" box="[264,676,1033,1057]" pageId="1" pageNumber="2">
<taxonomicName id="7BF94D52FFF3FFCEFEB9FBD2FD0E337A" authority="Murayama 1954: 35" authorityName="Murayama" authorityPageNumber="35" authorityYear="1954" box="[264,672,1033,1057]" class="Insecta" family="Melolonthidae" genus="Serica" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="okamotoi">
<emphasis id="8E8DEAC3FFF3FFCEFEB9FBD2FE18337B" box="[264,438,1034,1057]" italics="true" pageId="1" pageNumber="2">Serica okamotoi</emphasis>
:
<treatmentCitation id="3D5810C0FFF3FFCEFE75FBD1FD0E337A" author="Murayama" box="[452,672,1033,1056]" page="35" pageId="1" pageNumber="2" year="1954">
<bibRefCitation id="D8684B20FFF3FFCEFE75FBD1FDD6337A" author="Murayama" box="[452,632,1033,1056]" pageId="1" pageNumber="7" refString="Murayama, J. (1954) Icones of the Scarabeid-Beetles from Manchuria and Korea. I. Nihon Gakujutu, Shinko-kai, 163 pp." type="book" year="1954">Murayama 1954</bibRefCitation>
: 35
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="BC4636D1FFF3FFCEFEB9FBF2FBE7331B" blockId="1.[264,1322,713,1089]" box="[264,1097,1065,1089]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="9CE911FFFFF3FFCEFEB9FBF2FBE7331B" box="[264,1097,1065,1089]" pageId="1" pageNumber="2">
<taxonomicName id="7BF94D52FFF3FFCEFEB9FBF2FD5F331A" authority="Kim and Lee, 1997: 126" authorityName="Kim and Lee" authorityPageNumber="126" authorityYear="1997" box="[264,753,1065,1089]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="okamotoi">
<emphasis id="8E8DEAC3FFF3FFCEFEB9FBF2FE75331B" box="[264,475,1066,1089]" italics="true" pageId="1" pageNumber="2">Maladera okamotoi</emphasis>
:
<treatmentCitation id="3D5810C0FFF3FFCEFE58FBF1FD5F331A" author="Kim" box="[489,753,1065,1088]" page="126" pageId="1" pageNumber="2" year="1997">
<bibRefCitation id="D8684B20FFF3FFCEFE58FBF1FD15331A" author="Kim" box="[489,699,1065,1088]" pageId="1" pageNumber="7" refString="Kim, J. I. &amp; Lee, O. J. (1997) Taxonomic study of Korean Sericinae (Melolonthidae, Coleoptera) II, Genus Maladera. Insecta Koreana, 14, 119 - 135." type="journal article" year="1997">Kim and Lee, 1997</bibRefCitation>
: 126
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="3D5810C0FFF3FFCEFD4DFBF1FC5C331A" author="Kim" box="[764,1010,1065,1088]" page="88" pageId="1" pageNumber="2" year="2003">
<bibRefCitation id="D8684B20FFF3FFCEFD4DFBF1FC61331A" author="Kim" box="[764,975,1065,1088]" pageId="1" pageNumber="7" refString="Kim, J. I. &amp; Kim, A-Y. (2003) Taxonomic Review of Korean Sericinae (Coleoptera, Melolonthidae) II: Genus Maladera Mulsant. Insecta Koreana, 20 (1), 81 - 94." type="journal article" year="2003">Kim and Kim 2003</bibRefCitation>
: 88
</treatmentCitation>
,
<emphasis id="8E8DEAC3FFF3FFCEFC4EFBF1FBE7331B" bold="true" box="[1023,1097,1065,1089]" pageId="1" pageNumber="2">syn. n.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="F4E3655AFFF3FFCDFEB9FBA9FD86307C" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="BC4636D1FFF3FFCEFEB9FBA9FD053271" blockId="1.[264,1323,1137,1883]" pageId="1" pageNumber="2">
<emphasis id="8E8DEAC3FFF3FFCEFEB9FBA9FDEE33D1" bold="true" box="[264,576,1137,1163]" pageId="1" pageNumber="2">
<typeStatus id="63428873FFF3FFCEFEB9FBA9FEEB33D1" box="[264,325,1137,1163]" pageId="1" pageNumber="2">Type</typeStatus>
material examined:
</emphasis>
<typeStatus id="63428873FFF3FFCEFDF6FBA9FD1233D1" box="[583,700,1137,1163]" pageId="1" pageNumber="2" type="lectotype">Lectotype</typeStatus>
(
<taxonomicName id="7BF94D52FFF3FFCEFD7CFBA9FC9633D0" box="[717,824,1137,1162]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="infuscata">
<emphasis id="8E8DEAC3FFF3FFCEFD7CFBA9FC9633D0" box="[717,824,1137,1162]" italics="true" pageId="1" pageNumber="2">infuscata</emphasis>
</taxonomicName>
, here designated): 1 ď “
<collectingCountry id="C4EE7641FFF3FFCEFBE3FBA9FB3233D1" box="[1106,1180,1137,1163]" name="South Korea" pageId="1" pageNumber="2">Korea</collectingCountry>
/
<taxonomicName id="7BF94D52FFF3FFCEFB1DFBA9FEDD33E8" class="Insecta" family="Melolonthidae" genus="Autoserica" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="infuscata">
<emphasis id="8E8DEAC3FFF3FFCEFB1DFBA9FEDD33E8" italics="true" pageId="1" pageNumber="2">Autoserica infuscata</emphasis>
</taxonomicName>
<typeStatus id="63428873FFF3FFCEFECAFB41FE1B33E9" box="[379,437,1177,1203]" pageId="1" pageNumber="2">Type</typeStatus>
ď Mos.” (
<collectionCode id="DAE8AE14FFF3FFCEFD80FB41FD2233E9" box="[561,652,1177,1203]" pageId="1" pageNumber="2">ZMHB</collectionCode>
).
<typeStatus id="63428873FFF3FFCEFD2EFB41FCE433E9" box="[671,842,1177,1203]" pageId="1" pageNumber="2" type="paralectotype">Paralectotypes</typeStatus>
(
<taxonomicName id="7BF94D52FFF3FFCEFCEAFB41FC6833E8" box="[859,966,1177,1202]" class="Insecta" family="Melolonthidae" genus="Autoserica" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="infuscata">
<emphasis id="8E8DEAC3FFF3FFCEFCEAFB41FC6833E8" box="[859,966,1177,1202]" italics="true" pageId="1" pageNumber="2">infuscata</emphasis>
</taxonomicName>
, here designated): 1 Ψ “
<collectingCountry id="C4EE7641FFF3FFCEFB50FB41FA8433E9" box="[1249,1322,1177,1203]" name="China" pageId="1" pageNumber="2">China</collectingCountry>
Kiantschou/
<taxonomicName id="7BF94D52FFF3FFCEFE14FB19FD0E3380" box="[421,672,1217,1242]" class="Insecta" family="Melolonthidae" genus="Autoserica" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="infuscata">
<emphasis id="8E8DEAC3FFF3FFCEFE14FB19FD0E3380" box="[421,672,1217,1242]" italics="true" pageId="1" pageNumber="2">Autoserica infuscata</emphasis>
</taxonomicName>
<typeStatus id="63428873FFF3FFCEFD1EFB19FD473381" box="[687,745,1217,1243]" pageId="1" pageNumber="2">Type</typeStatus>
Ψ Mos.” (
<collectionCode id="DAE8AE14FFF3FFCEFCCDFB19FC793381" box="[892,983,1217,1243]" pageId="1" pageNumber="2">ZMHB</collectionCode>
), 1 ď „
<collectingCountry id="C4EE7641FFF3FFCEFB8EFB19FB263381" box="[1087,1160,1217,1243]" name="China" pageId="1" pageNumber="2">China</collectingCountry>
Kiantschou“ (
<collectionCode id="DAE8AE14FFF3FFCEFEA1FB31FEC53259" box="[272,363,1257,1283]" pageId="1" pageNumber="2">ZMHB</collectionCode>
).
<typeStatus id="63428873FFF3FFCEFE31FB31FE5B3259" box="[384,501,1257,1283]" pageId="1" pageNumber="2" type="lectotype">Lectotype</typeStatus>
(
<taxonomicName id="7BF94D52FFF3FFCEFDB9FB31FDC93258" box="[520,615,1257,1282]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="koreana">
<emphasis id="8E8DEAC3FFF3FFCEFDB9FB31FDC93258" box="[520,615,1257,1282]" italics="true" pageId="1" pageNumber="2">koreana</emphasis>
</taxonomicName>
, here designated): 1 ď “
<collectingCountry id="C4EE7641FFF3FFCEFC39FB31FC7D3259" box="[904,979,1257,1283]" name="South Korea" pageId="1" pageNumber="2">Korea</collectingCountry>
Chemulpo/
<taxonomicName id="7BF94D52FFF3FFCEFBD7FB31FE9A3270" class="Insecta" family="Melolonthidae" genus="Autoserica" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="koreana">
<emphasis id="8E8DEAC3FFF3FFCEFBD7FB31FE9A3270" italics="true" pageId="1" pageNumber="2">Autoserica koreana</emphasis>
</taxonomicName>
<typeStatus id="63428873FFF3FFCEFE8DFAC9FED83271" box="[316,374,1297,1323]" pageId="1" pageNumber="2">Type</typeStatus>
Mos./
<taxonomicName id="7BF94D52FFF3FFCEFE79FAC9FD893270" box="[456,551,1297,1322]" class="Insecta" family="Melolonthidae" genus="Autoserica" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="koreana">
<emphasis id="8E8DEAC3FFF3FFCEFE79FAC9FD893270" box="[456,551,1297,1322]" italics="true" pageId="1" pageNumber="2">koreana</emphasis>
</taxonomicName>
” (
<collectionCode id="DAE8AE14FFF3FFCEFDF5FAC9FD313271" box="[580,671,1297,1323]" pageId="1" pageNumber="2">ZMHB</collectionCode>
).
</paragraph>
<paragraph id="BC4636D1FFF3FFCEFE89FAE1FD4D32F9" blockId="1.[264,1323,1137,1883]" pageId="1" pageNumber="2">
<emphasis id="8E8DEAC3FFF3FFCEFE89FAE1FD623209" bold="true" box="[312,716,1337,1363]" pageId="1" pageNumber="2">Additional material examined:</emphasis>
1 ď „Tsingtau Lazarettgarten Prof. Hoffmann/
<date id="C8471011FFF3FFCEFEB9FAB9FEFA3221" box="[264,340,1377,1403]" pageId="1" pageNumber="2">22.VI.</date>
“ (
<collectionCode id="DAE8AE14FFF3FFCEFEC0FAB9FE653221" box="[369,459,1377,1403]" pageId="1" pageNumber="2">ZMHB</collectionCode>
), 1 ď, 1 Ψ “Tsingtau Prof. Hoffmann/ Februar” (
<collectionCode id="DAE8AE14FFF3FFCEFBA2FAB9FBC33221" box="[1043,1133,1377,1403]" pageId="1" pageNumber="2">ZMHB</collectionCode>
), 1 ex. “Seishin
<collectingCountry id="C4EE7641FFF3FFCEFEB9FA51FEFC32F9" box="[264,338,1417,1443]" name="South Korea" pageId="1" pageNumber="2">Korea</collectingCountry>
” (CF), 3 ex. “Ompo
<collectingCountry id="C4EE7641FFF3FFCEFDF9FA51FD3D32F9" box="[584,659,1417,1443]" name="South Korea" pageId="1" pageNumber="2">Korea</collectingCountry>
” (CF).
</paragraph>
<paragraph id="BC4636D1FFF3FFCEFE89FA69FBA03141" blockId="1.[264,1323,1137,1883]" pageId="1" pageNumber="2">
<emphasis id="8E8DEAC3FFF3FFCEFE89FA69FDDD3291" bold="true" box="[312,627,1457,1483]" pageId="1" pageNumber="2">Lectotype­Redescription.</emphasis>
Length:
<quantity id="7B019B34FFF3FFCEFD6FFA69FC953291" box="[734,827,1457,1483]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="1" pageNumber="2" unit="mm" value="9.1">9.1 mm</quantity>
, length of elytra:
<quantity id="7B019B34FFF3FFCEFBBDFA69FBC43291" box="[1036,1130,1457,1483]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="1" pageNumber="2" unit="mm" value="6.5">6.5 mm</quantity>
, width:
<quantity id="7B019B34FFF3FFCEFB78FA69FA893291" box="[1225,1319,1457,1483]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="1" pageNumber="2" unit="mm" value="5.2">5.2 mm</quantity>
. Body oval, reddish brown, antenna yellow, labroclypeus shiny, remainder of dorsal surface dull, glabrous, except for a few small setae on the head and elytra.
</paragraph>
<paragraph id="BC4636D1FFF3FFCDFE89F9F1FE0636F4" blockId="1.[264,1323,1137,1883]" lastBlockId="2.[264,1324,284,1830]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, straight and strongly convergent to moderately rounded anterior angles, lateral border and ocular canthus producing a distinct blunt angle, margins weakly reflexed, anteriorly very shallowly sinuate medially; surface weakly convex medially, shiny, finely punctate, distance between punctures irregular sometimes less than their diameter, with a few short, erect setae anteriorly; frontoclypeal suture feebly impressed and weakly curved medially; smooth area in front of eye approximately twice as wide as long; ocular canthus short and moderately broad, finely punctate, with a single short terminal seta. Frons with fine, moderately dense punctures, glabrous except for a few setae beside eyes. Eyes small, ratio of diameter/ interocular width: 0.57. Antenna yellow, with 10 antennomeres; club with three antennomeres, as long as remaining antennomeres together. Mentum anteriorly elevated and flattened.</paragraph>
<paragraph id="BC4636D1FFF0FFCDFE89FE64FD6135C4" blockId="2.[264,1324,284,1830]" pageId="2" pageNumber="3">Pronotum widest at base, lateral margins weakly and evenly convex and convergent anteriorly, anterior angles distinctly produced and sharp, anterior marginal line widely interrupted medially, margin weakly and convexly produced medially; surface moderately densely and finely punctate, with very minute setae in punctures only; anterior and lateral borders setose. Scutellum broad, triangular, with fine and moderately dense punctures, each bearing a single very minute seta.</paragraph>
<paragraph id="BC4636D1FFF0FFCDFE89FD74FBC6343C" blockId="2.[264,1324,284,1830]" pageId="2" pageNumber="3">Elytra oblong, widest at middle, striae distinctly impressed, finely and densely punctate, intervals moderately convex, with fine, moderately dense punctures, penultimate lateral interval with a few fine, erect, short setae, some punctures with very minute setae; epipleural edge robust, ending at the weakly convex external apical angle of elytra, epipleura densely setose, apical border chitinous, with short microtrichomes.</paragraph>
<paragraph id="BC4636D1FFF0FFCDFE89FCACFCC23394" blockId="2.[264,1324,284,1830]" pageId="2" pageNumber="3">Ventral surface dull, thorax and metacoxa with large dense punctures, sparsely setose, metacoxa glabrous except for a few long setae laterally; each abdominal sternite, in addition to generally distributed fine and moderately dense punctures, with a distinct transverse row of coarse punctures each bearing a short and robust seta, some punctures with microscopic setae, penultimate sternite apically with a shiny smooth chitinous border, which is one third as long as sternite. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/ metacoxa: 1/ 1.7. Pygidium strongly convex, finely and densely punctate, without smooth midline, punctures with microscopic setae and with a few long setae along apical margin.</paragraph>
<paragraph id="BC4636D1FFF0FFCDFE89FB04FB6F31A4" blockId="2.[264,1324,284,1830]" pageId="2" pageNumber="3">Legs moderately broad; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur dull, anterior edge acute, lacking an adjacent serrated line, posterior ventral margin medially straight, weakly widened in apical half and not serrate, dorsally not serrated, finely setose. Metatibia moderately broad and long, widest at apex, ratio width/ length: 1/ 3.4, dorsally sharply edged, with two groups of spines, basal one at middle, apical one at three fourth of metatibial length, basally with a few punctures, each bearing short single spines; lateral face longitudinally convex, with sparse and fine punctures in basal half, glabrous; ventral edge serrate, with three strong spines equidistant from each other, medial face not punctate, apex interiorly near tarsal articulation shallowly concave. Tarsomeres very sparsely and finely punctate dorsally, ventrally with sparse, short setae; metatarsomeres ventrally with a strongly serrated ridge, beside which is a strong longitudinal carina, first metatarsomere a little shorter than the two following segments combined and a little longer than the upper tibial spur. Protibia short, bidentate. All claws symmetrical, feebly curved and long, with normally developed basal tooth.</paragraph>
<paragraph id="BC4636D1FFF0FFCDFE89F8D4FD86307C" blockId="2.[264,1324,284,1830]" box="[312,552,1804,1830]" pageId="2" pageNumber="3">
Aedeagus:
<figureCitation id="24C22A54FFF0FFCDFE0AF8D4FD8A307C" box="[443,548,1804,1830]" captionStart="FIGURES 1 – 7" captionStartId="3.[264,370,1792,1813]" captionTargetBox="[322,1265,344,1754]" captionTargetPageId="3" captionText="FIGURES 1 – 7: 1 – 3 Maladera infuscata (Moser, 1915) (Lectotype: “ Korea ”); 4 – 7: Maladera howdeni sp. n. (Holotype: Taiwan, Wushe); 1, 3, 4, 7: aedeagus, lateral view; 2: aedeagus, dorsal view; 5: parameres, dorsal view; 6: parameres, ventral view. Scale: 0.5 mm." httpUri="https://zenodo.org/record/168876/files/figure.png" pageId="2" pageNumber="3">Figs 1–3</figureCitation>
.
</paragraph>
</subSubSection>
<caption id="E8866659FFF1FFCCFEB9F8D8FB9F303A" httpUri="https://zenodo.org/record/168876/files/figure.png" pageId="3" pageNumber="4" targetBox="[322,1265,344,1754]" targetPageId="3">
<paragraph id="BC4636D1FFF1FFCCFEB9F8D8FB9F303A" blockId="3.[264,1323,1790,1888]" pageId="3" pageNumber="4">
<emphasis id="8E8DEAC3FFF1FFCCFEB9F8D8FE0D304F" bold="true" box="[264,419,1792,1813]" pageId="3" pageNumber="4">FIGURES 1–7</emphasis>
:
<emphasis id="8E8DEAC3FFF1FFCCFE09F8D8FE72304F" bold="true" box="[440,476,1792,1813]" pageId="3" pageNumber="4">1–3</emphasis>
<taxonomicName id="7BF94D52FFF1FFCCFE5AF927FCDD304C" authority="(Moser, 1915)" authorityName="Moser" authorityYear="1915" baseAuthorityName="Moser" baseAuthorityYear="1915" box="[491,883,1791,1814]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="infuscata">
<emphasis id="8E8DEAC3FFF1FFCCFE5AF927FD6B304C" box="[491,709,1791,1814]" italics="true" pageId="3" pageNumber="4">Maladera infuscata</emphasis>
(Moser, 1915)
</taxonomicName>
(Lectotype: “Korea”);
<emphasis id="8E8DEAC3FFF1FFCCFB32F926FB05304C" bold="true" box="[1155,1195,1790,1814]" pageId="3" pageNumber="4">4–7</emphasis>
:
<taxonomicName id="7BF94D52FFF1FFCCFB70F927FECC3061" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="howdeni" status="sp. nov.">
<emphasis id="8E8DEAC3FFF1FFCCFB70F927FECC3061" italics="true" pageId="3" pageNumber="4">Maladera howdeni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95BE57B8FFF1FFCCFEDAF8FCFE083061" box="[363,422,1828,1851]" pageId="3" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
(Holotype: Taiwan, Wushe); 1, 3, 4, 7: aedeagus, lateral view; 2: aedeagus, dorsal view; 5: parameres, dorsal view; 6: parameres, ventral view. Scale: 0.5 mm.
</paragraph>
</caption>
<subSubSection id="F4E3655AFFF6FFCBFE89FEC3FD0035A2" pageId="4" pageNumber="5" type="discussion">
<paragraph id="BC4636D1FFF6FFCBFE89FEC3FC65357C" blockId="4.[264,1323,283,630]" pageId="4" pageNumber="5">
<emphasis id="8E8DEAC3FFF6FFCBFE89FEC3FE1F366F" bold="true" box="[312,433,283,309]" pageId="4" pageNumber="5">Remarks.</emphasis>
Since the original descriptions of
<taxonomicName id="7BF94D52FFF6FFCBFCEDFEC4FC5D366F" box="[860,1011,284,309]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="infuscata">
<emphasis id="8E8DEAC3FFF6FFCBFCEDFEC4FC5D366F" box="[860,1011,284,309]" italics="true" pageId="4" pageNumber="5">M. infuscata</emphasis>
</taxonomicName>
and
<taxonomicName id="7BF94D52FFF6FFCBFB85FEC4FB11366F" box="[1076,1215,284,309]" class="Insecta" family="Melolonthidae" genus="Autoserica" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="koreana">
<emphasis id="8E8DEAC3FFF6FFCBFB85FEC4FB11366F" box="[1076,1215,284,309]" italics="true" pageId="4" pageNumber="5">M. koreana</emphasis>
</taxonomicName>
gave no information about the number of specimens on which the descriptions were based, it is necessary to fix the taxonomic status of both taxa by designating
<typeStatus id="63428873FFF6FFCBFBA9FEB4FB3D36DC" box="[1048,1171,364,390]" pageId="4" pageNumber="5" type="lectotype">lectotypes</typeStatus>
. The respective
<typeStatus id="63428873FFF6FFCBFE8CFE4CFE1A36F4" box="[317,436,404,430]" pageId="4" pageNumber="5" type="lectotype">lectotypes</typeStatus>
selected here represent the only remaining
<typeStatus id="63428873FFF6FFCBFC72FE4CFB8436F4" box="[963,1066,404,430]" pageId="4" pageNumber="5" type="syntype">syntypes</typeStatus>
of each taxon occurring in the Moser collection. The
<typeStatus id="63428873FFF6FFCBFD29FE64FCA1368C" box="[664,783,444,470]" pageId="4" pageNumber="5" type="lectotype">lectotypes</typeStatus>
of the two taxa and the figure of aedeagus in original description of
<taxonomicName id="7BF94D52FFF6FFCBFDAAFE3CFC3C36A4" authority="Murayama 1938" authorityName="Murayama" authorityYear="1938" box="[539,914,484,510]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="okamotoi">
<emphasis id="8E8DEAC3FFF6FFCBFDAAFE3CFD1D36A7" box="[539,691,484,509]" italics="true" pageId="4" pageNumber="5">M. okamotoi</emphasis>
(
<bibRefCitation id="D8684B20FFF6FFCBFD72FE3CFC2436A4" author="Murayama" box="[707,906,484,510]" pageId="4" pageNumber="7" refString="Murayama, J. (1938) Revision des Sericines (Col., Scar.) de la Coree. Annotiones Zoologicae Japonenses, 17 (1), 7 - 21." type="journal article" year="1938">Murayama 1938</bibRefCitation>
)
</taxonomicName>
are virtually identical in shape of parameres, consequently I consider them to be synonymous.
</paragraph>
<paragraph id="BC4636D1FFF6FFCBFEB9FDECFBE2352C" blockId="4.[264,1323,283,630]" pageId="4" pageNumber="5">
Since previous authors have not examined the
<typeStatus id="63428873FFF6FFCBFC9EFDECFCCF3514" box="[815,865,564,590]" pageId="4" pageNumber="5">type</typeStatus>
specimens of Moser,
<taxonomicName id="7BF94D52FFF6FFCBFBD9FDECFB523517" box="[1128,1276,564,589]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="infuscata">
<emphasis id="8E8DEAC3FFF6FFCBFBD9FDECFB523517" box="[1128,1276,564,589]" italics="true" pageId="4" pageNumber="5">M. infuscata</emphasis>
</taxonomicName>
nor
<emphasis id="8E8DEAC3FFF6FFCBFEB9FD84FE5A352F" box="[264,500,604,629]" italics="true" pageId="4" pageNumber="5">
<taxonomicName id="7BF94D52FFF6FFCBFEB9FD84FE5E352F" box="[264,496,604,629]" class="Insecta" family="Melolonthidae" genus="Autoserica" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="koreana">Autoserica koreana</taxonomicName>
,
</emphasis>
these species have been misidentified consistently.
</paragraph>
<paragraph id="BC4636D1FFF6FFCBFEB9FD06FD0035A2" blockId="4.[264,686,734,760]" box="[264,686,734,760]" pageId="4" pageNumber="5">
<emphasis id="8E8DEAC3FFF6FFCBFEB9FD06FDE835A2" bold="true" box="[264,582,734,760]" pageId="4" pageNumber="5">
<taxonomicName id="7BF94D52FFF6FFCBFEB9FD06FE4535A2" box="[264,491,734,760]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="howdeni" status="sp. nov.">
<emphasis id="8E8DEAC3FFF6FFCBFEB9FD06FE4535A2" bold="true" box="[264,491,734,760]" italics="true" pageId="4" pageNumber="5">Maladera howdeni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95BE57B8FFF6FFCBFE42FD06FD9B35A2" box="[499,565,734,760]" pageId="4" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
(
</emphasis>
<emphasis id="8E8DEAC3FFF6FFCBFDF4FD38FD0035A2" bold="true" box="[581,686,734,760]" pageId="4" pageNumber="5">
<figureCitation id="24C22A54FFF6FFCBFDF4FD38FD0D35A2" box="[581,675,736,760]" captionStart="FIGURES 1 – 7" captionStartId="3.[264,370,1792,1813]" captionTargetBox="[322,1265,344,1754]" captionTargetPageId="3" captionText="FIGURES 1 – 7: 1 – 3 Maladera infuscata (Moser, 1915) (Lectotype: “ Korea ”); 4 – 7: Maladera howdeni sp. n. (Holotype: Taiwan, Wushe); 1, 3, 4, 7: aedeagus, lateral view; 2: aedeagus, dorsal view; 5: parameres, dorsal view; 6: parameres, ventral view. Scale: 0.5 mm." httpUri="https://zenodo.org/record/168876/files/figure.png" pageId="4" pageNumber="5">Figs 4–7</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="F4E3655AFFF6FFCBFEB9FCF4FD113419" box="[264,703,812,835]" pageId="4" pageNumber="5" type="reference_group">
<paragraph id="BC4636D1FFF6FFCBFEB9FCF4FD113419" blockId="4.[264,703,812,835]" box="[264,703,812,835]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="9CE911FFFFF6FFCBFEB9FCF4FD113419" box="[264,703,812,835]" pageId="4" pageNumber="5">
<taxonomicName id="7BF94D52FFF6FFCBFEB9FCF4FD153419" authority="Nomura, 1974: 109" authorityName="Nomura" authorityPageNumber="109" authorityYear="1974" box="[264,699,812,835]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="infuscata">
<emphasis id="8E8DEAC3FFF6FFCBFEB9FCF4FE743419" box="[264,474,812,835]" italics="true" pageId="4" pageNumber="5">Maladera infuscata</emphasis>
:
<treatmentCitation id="3D5810C0FFF6FFCBFE59FCF4FD153419" author="Nomura" box="[488,699,812,835]" page="109" pageId="4" pageNumber="5" year="1974">
<bibRefCitation id="D8684B20FFF6FFCBFE59FCF4FD283419" author="Nomura" box="[488,646,812,835]" pageId="4" pageNumber="8" refString="Nomura, S. (1974) On the Sericini of Taiwan. Toho-Gakuho, 24, 81 - 115." type="journal article" year="1974">Nomura, 1974</bibRefCitation>
: 109
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="F4E3655AFFF6FFCBFEB9FCA1FD793211" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="BC4636D1FFF6FFCBFEB9FCA1FABB33A1" blockId="4.[264,1324,889,1875]" pageId="4" pageNumber="5">
<emphasis id="8E8DEAC3FFF6FFCBFEB9FCA1FDEC34C9" bold="true" box="[264,578,889,915]" pageId="4" pageNumber="5">
<typeStatus id="63428873FFF6FFCBFEB9FCA1FEEB34C9" box="[264,325,889,915]" pageId="4" pageNumber="5">Type</typeStatus>
material examined:
</emphasis>
<typeStatus id="63428873FFF6FFCBFDFDFCA1FD1034C9" box="[588,702,889,915]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
: ɗ “
<collectingCountry id="C4EE7641FFF6FFCBFD41FCA1FCE234C9" box="[752,844,889,915]" name="Taiwan" pageId="4" pageNumber="5">Taiwan</collectingCountry>
, Wushe
<date id="C8471011FFF6FFCBFC03FCA1FB9D34C9" box="[946,1075,889,915]" pageId="4" pageNumber="5" value="1983-04-29">29.IV.1983</date>
H. Townes
<quantity id="7B019B34FFF6FFCBFB75FCA1FA8E34C9" box="[1220,1312,889,915]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="4" pageNumber="5" unit="m" value="1150.0">1150 m</quantity>
” (
<collectionCode id="DAE8AE14FFF6FFCBFEA1FC79FEC434E1" box="[272,362,929,955]" pageId="4" pageNumber="5">HAHC</collectionCode>
).
<typeStatus id="63428873FFF6FFCBFE31FC79FE5934E1" box="[384,503,929,955]" pageId="4" pageNumber="5" type="paratype">Paratypes</typeStatus>
: 1 ɗ ­ same data as
<typeStatus id="63428873FFF6FFCBFD45FC79FCF434E1" box="[756,858,929,955]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
(
<collectionCode id="DAE8AE14FFF6FFCBFCDCFC79FC6834E1" box="[877,966,929,955]" pageId="4" pageNumber="5">HAHC</collectionCode>
), 3 ɗ, 3 ΨΨ “
<collectingCountry id="C4EE7641FFF6FFCBFBC1FC79FB6234E1" box="[1136,1228,929,955]" name="Taiwan" pageId="4" pageNumber="5">Taiwan</collectingCountry>
, Wushe
<date id="C8471011FFF6FFCBFEB9FC11FEDF34B9" box="[264,369,969,995]" pageId="4" pageNumber="5" value="1983-05-03">3.V.1983</date>
H. Townes
<quantity id="7B019B34FFF6FFCBFDB8FC11FDC834B9" box="[521,614,969,995]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="4" pageNumber="5" unit="m" value="1150.0">1150 m</quantity>
” (
<collectionCode id="DAE8AE14FFF6FFCBFD35FC11FD7234B9" box="[644,732,969,995]" pageId="4" pageNumber="5">HAHC</collectionCode>
, CA), 1 ɗ, 3 ΨΨ “
<collectingCountry id="C4EE7641FFF6FFCBFC74FC11FB8F34B9" box="[965,1057,969,995]" name="Taiwan" pageId="4" pageNumber="5">Taiwan</collectingCountry>
, Wushe
<date id="C8471011FFF6FFCBFB3AFC11FAAA34B9" box="[1163,1284,969,995]" pageId="4" pageNumber="5" value="1983-05-15">15.V.1983</date>
H. Townes
<quantity id="7B019B34FFF6FFCBFEDFFC29FE623351" box="[366,460,1009,1035]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="4" pageNumber="5" unit="m" value="1150.0">1150 m</quantity>
” (
<collectionCode id="DAE8AE14FFF6FFCBFE5BFC29FDEC3351" box="[490,578,1009,1035]" pageId="4" pageNumber="5">HAHC</collectionCode>
, CA), 3 ɗɗ “
<collectingCountry id="C4EE7641FFF6FFCBFD5FFC29FCE43351" box="[750,842,1009,1035]" name="Taiwan" pageId="4" pageNumber="5">Taiwan</collectingCountry>
, Wushe
<date id="C8471011FFF6FFCBFC04FC29FB863351" box="[949,1064,1009,1035]" pageId="4" pageNumber="5" value="1983-04-07">7.IV.1983</date>
H. Townes
<quantity id="7B019B34FFF6FFCBFB70FC29FAB13351" box="[1217,1311,1009,1035]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="4" pageNumber="5" unit="m" value="1150.0">1150 m</quantity>
” (
<collectionCode id="DAE8AE14FFF6FFCBFEA1FBC1FEC93369" box="[272,359,1049,1075]" pageId="4" pageNumber="5">HAHC</collectionCode>
, CA), 1 ɗ “Wushe,
<collectingCountry id="C4EE7641FFF6FFCBFDD9FBC1FD103369" box="[616,702,1049,1075]" name="Taiwan" pageId="4" pageNumber="5">Taiwan</collectingCountry>
<quantity id="7B019B34FFF6FFCBFD7BFBC1FC883369" box="[714,806,1049,1075]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="4" pageNumber="5" unit="m" value="1150.0">1150 m</quantity>
<date id="C8471011FFF6FFCBFC83FBC1FC1B3369" box="[818,949,1049,1075]" pageId="4" pageNumber="5" value="1983-03-16">16.III.1983</date>
H. &amp; M. Townes” (CA), 1 ɗ “Wushe,
<collectingCountry id="C4EE7641FFF6FFCBFEC6FB99FE633301" box="[375,461,1089,1115]" name="Taiwan" pageId="4" pageNumber="5">Taiwan</collectingCountry>
<quantity id="7B019B34FFF6FFCBFE69FB99FD9D3301" box="[472,563,1089,1115]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="4" pageNumber="5" unit="m" value="1150.0">1150 m</quantity>
<date id="C8471011FFF6FFCBFD8FFB99FD6F3301" box="[574,705,1089,1115]" pageId="4" pageNumber="5" value="1983-03-23">23.III.1983</date>
H. &amp; M. Townes” (
<collectionCode id="DAE8AE14FFF6FFCBFC0CFB99FBB83301" box="[957,1046,1089,1115]" pageId="4" pageNumber="5">HAHC</collectionCode>
), 1 Ψ “
<collectingCountry id="C4EE7641FFF6FFCBFBDEFB99FB643301" box="[1135,1226,1089,1115]" name="Taiwan" pageId="4" pageNumber="5">Taiwan</collectingCountry>
, Wushe
<date id="C8471011FFF6FFCBFEB9FBB1FE2E33D9" box="[264,384,1129,1155]" pageId="4" pageNumber="5" value="1983-05-10">10.V.1983</date>
H. Townes
<quantity id="7B019B34FFF6FFCBFDA6FBB1FDDA33D9" box="[535,628,1129,1155]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="4" pageNumber="5" unit="m" value="1150.0">1150 m</quantity>
” (
<collectionCode id="DAE8AE14FFF6FFCBFD20FBB1FD4433D9" box="[657,746,1129,1155]" pageId="4" pageNumber="5">HAHC</collectionCode>
), 2 ΨΨ “
<collectingCountry id="C4EE7641FFF6FFCBFCE6FBB1FC1D33D9" box="[855,947,1129,1155]" name="Taiwan" pageId="4" pageNumber="5">Taiwan</collectingCountry>
, Wushe
<date id="C8471011FFF6FFCBFBACFBB1FB3033D9" box="[1053,1182,1129,1155]" pageId="4" pageNumber="5" value="1983-04-13">13.IV.1983</date>
H. Townes
<quantity id="7B019B34FFF6FFCBFEB9FB49FEC833F1" box="[264,358,1169,1195]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="4" pageNumber="5" unit="m" value="1150.0">1150 m</quantity>
” (
<collectionCode id="DAE8AE14FFF6FFCBFE35FB49FE7333F1" box="[388,477,1169,1195]" pageId="4" pageNumber="5">HAHC</collectionCode>
), 1 ɗ “
<collectingCountry id="C4EE7641FFF6FFCBFD89FB49FD3A33F1" box="[568,660,1169,1195]" name="Taiwan" pageId="4" pageNumber="5">Taiwan</collectingCountry>
, Wushe
<date id="C8471011FFF6FFCBFD4EFB49FCD833F1" box="[767,886,1169,1195]" pageId="4" pageNumber="5" value="1983-05-22">22.V.1983</date>
H. Townes
<quantity id="7B019B34FFF6FFCBFBA1FB49FBC033F1" box="[1040,1134,1169,1195]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="4" pageNumber="5" unit="m" value="1150.0">1150 m</quantity>
” (
<collectionCode id="DAE8AE14FFF6FFCBFB3DFB49FB4B33F1" box="[1164,1253,1169,1195]" pageId="4" pageNumber="5">HAHC</collectionCode>
), 1 Ψ “Wushe,
<collectingCountry id="C4EE7641FFF6FFCBFEC5FB61FE643389" box="[372,458,1209,1235]" name="Taiwan" pageId="4" pageNumber="5">Taiwan</collectingCountry>
<quantity id="7B019B34FFF6FFCBFE63FB61FD843389" box="[466,554,1209,1235]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="4" pageNumber="5" unit="m" value="1150.0">1150 m</quantity>
<date id="C8471011FFF6FFCBFD83FB61FD0B3389" box="[562,677,1209,1235]" pageId="4" pageNumber="5" value="1983-04-02">2.IV.1983</date>
Henry Townes” (
<collectionCode id="DAE8AE14FFF6FFCBFCC7FB61FC613389" box="[886,975,1209,1235]" pageId="4" pageNumber="5">HAHC</collectionCode>
), 1 ɗ “
<collectingCountry id="C4EE7641FFF6FFCBFB91FB61FBD33389" box="[1056,1149,1209,1235]" name="Taiwan" pageId="4" pageNumber="5">Taiwan</collectingCountry>
: Hualien Prov. Taroko N. P. betw.
<quantity id="7B019B34FFF6FFCBFE59FB39FDD733A1" box="[488,633,1249,1275]" metricMagnitude="3" metricUnit="m" metricValue="1.335" metricValueMax="2.4" metricValueMin="0.27" pageId="4" pageNumber="5" unit="m" value="1335.0" valueMax="2400.0" valueMin="270.0">270­2400 m</quantity>
, at light
<date id="C8471011FFF6FFCBFD6FFB39FCC133A1" box="[734,879,1249,1275]" pageId="4" pageNumber="5" value="1997-04-02" valueMax="1997-04-05" valueMin="1997-04-02">2–5.IV.1997</date>
G. Csorba &amp; L. Ronkay” (
<collectionCode id="DAE8AE14FFF6FFCBFB19FB39FAA733A1" LSID="urn:lsid:biocol.org:col:34991" box="[1192,1289,1249,1275]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="4" pageNumber="5">HNHM</collectionCode>
).
</paragraph>
<paragraph id="BC4636D1FFF6FFCBFE89FAD1FD793211" blockId="4.[264,1324,889,1875]" pageId="4" pageNumber="5">
<emphasis id="8E8DEAC3FFF6FFCBFE89FAD1FD903279" bold="true" box="[312,574,1289,1315]" pageId="4" pageNumber="5">Additional Material:</emphasis>
1 ex. “
<collectingCountry id="C4EE7641FFF6FFCBFD26FAD1FD5A3279" box="[663,756,1289,1315]" name="Taiwan" pageId="4" pageNumber="5">Taiwan</collectingCountry>
: Prov. Nan­Tou Tayuling,
<quantity id="7B019B34FFF6FFCBFB85FAD1FB3D3279" box="[1076,1171,1289,1315]" metricMagnitude="3" metricUnit="m" metricValue="2.55" pageId="4" pageNumber="5" unit="m" value="2550.0">2550 m</quantity>
, 07–08.1996 leg. Gy. Fábián &amp; F. Nemes” (
<collectionCode id="DAE8AE14FFF6FFCBFDDBFAE9FD643211" LSID="urn:lsid:biocol.org:col:34991" box="[618,714,1329,1355]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="4" pageNumber="5">HNHM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="F4E3655AFFF6FFC9FE89FA81FCA636DC" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5" type="description">
<paragraph id="BC4636D1FFF6FFCBFE89FA81FC7C3299" blockId="4.[264,1324,889,1875]" pageId="4" pageNumber="5">
<emphasis id="8E8DEAC3FFF6FFCBFE89FA81FE7E3229" bold="true" box="[312,464,1369,1395]" pageId="4" pageNumber="5">Description.</emphasis>
Length:
<quantity id="7B019B34FFF6FFCBFD8CFA81FD043229" box="[573,682,1369,1395]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="4" pageNumber="5" unit="mm" value="10.5">10.5 mm</quantity>
, length of elytra:
<quantity id="7B019B34FFF6FFCBFCCFFA81FC733229" box="[894,989,1369,1395]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="4" pageNumber="5" unit="mm" value="7.4">7.4 mm</quantity>
, width:
<quantity id="7B019B34FFF6FFCBFB8FFA81FB323229" box="[1086,1180,1369,1395]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="4" pageNumber="5" unit="mm" value="5.9">5.9 mm</quantity>
. Body oval, dark reddish brown, antenna yellow, labroclypeus shiny, remainder of dorsal surface dull, glabrous, except for a few small setae on the head and elytra.
</paragraph>
<paragraph id="BC4636D1FFF6FFCAFE89FA09FB063604" blockId="4.[264,1324,889,1875]" lastBlockId="5.[264,1325,284,1830]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins weakly convex and strongly convergent to strongly rounded anterior angles, lateral border and ocular canthus producing an indistinct blunt angle, margins strongly reflexed, anteriorly shallowly sinuate medially; surface weakly convex medially, shiny, basally narrowly dull, very coarse punctures mixed fine very fine ones, distance between punctures irregular sometimes less than their diameter, with a few long, erect setae anteriorly; frontoclypeal suture feebly impressed and weakly curved medially; smooth area in front of eye approximately twice as wide as long; ocular canthus short and moderately broad, finely and sparsely punctate, with one or two short setae. Frons dull, with fine, sparse punctures, glabrous except for a few setae beside eyes. Eyes small, ratio of diameter/ interocular width: 0.53. Antenna yellow, with 10 antennomeres; club with three antennomeres, as long as remaining antennomeres together. Mentum anteriorly elevated and flattened.</paragraph>
<paragraph id="BC4636D1FFF7FFCAFE89FEB4FE93352C" blockId="5.[264,1325,284,1830]" pageId="5" pageNumber="6">Pronotum widest at base, lateral margins straight, weakly narrowed to the middle, in anterior half strongly convex and convergent anteriorly, anterior angles distinctly produced and sharp, anterior marginal line widely interrupted medially, margin weakly and convexly produced medially; surface moderately densely and finely punctate, with very minute setae in the punctures only; anterior and lateral borders setose. Scutellum broad, triangular, with fine and moderately dense punctures, each bearing a single very minute seta.</paragraph>
<paragraph id="BC4636D1FFF7FFCAFE89FD5CFC163464" blockId="5.[264,1325,284,1830]" pageId="5" pageNumber="6">Elytra oblong, widest at middle, striae distinctly impressed, finely and densely punctate, intervals almost flat, with fine, moderately dense punctures, penultimate lateral interval with a few fine, erect, short setae, some punctures with very minute setae; epipleural edge robust, ending at the weakly convex external apical angle of elytra, epipleura densely setose, apical border chitinous, with short microtrichomes.</paragraph>
<paragraph id="BC4636D1FFF7FFCAFE89FC94FC0733FC" blockId="5.[264,1325,284,1830]" pageId="5" pageNumber="6">Ventral surface dull, thorax and metacoxa with large and dense punctures, sparsely setose, metacoxa glabrous except for numerous long setae laterally; each abdominal sternite, in addition to generally distributed fine and moderately dense punctures, with a distinct transverse row of coarse punctures each bearing a long and robust seta, some punctures with microscopic setae, penultimate sternite apically with a shiny smooth chitinous border, which is one third as long as sternite. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/ metacoxa: 1/ 1.67. Pygidium moderately convex, finely and moderately densely punctate, without smooth midline, punctures with microscopic setae and with numerous long setae apically.</paragraph>
<paragraph id="BC4636D1FFF7FFCAFE89FB6CFD2E31A4" blockId="5.[264,1325,284,1830]" pageId="5" pageNumber="6">Legs moderately broad; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur dull, anterior edge acute, lacking an adjacent serrated line, posterior ventral margin medially straight, weakly widened in apical half and not serrate, dorsally not serrated, finely setose. Metatibia moderately broad and long, widest at apex, ratio width/ length: 1/ 3.1, dorsally sharply edged, with two groups of spines, basal one at one third, apical one at three fourths of metatibial length, basally with a few punctures, each bearing short single spines; lateral face longitudinally convex, with moderately dense and fine punctures dorsally and in basal half, glabrous, apically almost smooth; ventral edge serrate, with four strong spines equidistant from each other, medial face not punctate, apex interiorly near tarsal articulation shallowly concave. Tarsomeres very sparsely and finely punctate dorsally, ventrally with sparse, short setae; metatarsomeres ventrally with a strongly serrated ridge, beside which is a strong longitudinal carina, first metatarsomere a little shorter than the following two combined and a little longer than the upper tibial spur. Protibia short, bidentate. All claws symmetrical, feebly curved and long, with normally developed basal tooth.</paragraph>
<paragraph id="BC4636D1FFF7FFCAFE89F8D4FD86307C" blockId="5.[264,1325,284,1830]" box="[312,552,1804,1830]" pageId="5" pageNumber="6">
Aedeagus:
<figureCitation id="24C22A54FFF7FFCAFE0BF8D4FD8A307C" box="[442,548,1804,1830]" captionStart="FIGURES 1 – 7" captionStartId="3.[264,370,1792,1813]" captionTargetBox="[322,1265,344,1754]" captionTargetPageId="3" captionText="FIGURES 1 – 7: 1 – 3 Maladera infuscata (Moser, 1915) (Lectotype: “ Korea ”); 4 – 7: Maladera howdeni sp. n. (Holotype: Taiwan, Wushe); 1, 3, 4, 7: aedeagus, lateral view; 2: aedeagus, dorsal view; 5: parameres, dorsal view; 6: parameres, ventral view. Scale: 0.5 mm." httpUri="https://zenodo.org/record/168876/files/figure.png" pageId="5" pageNumber="6">Figs 4–7</figureCitation>
.
</paragraph>
<paragraph id="BC4636D1FFF4FFC9FE89FEC3FCA636DC" blockId="6.[264,1324,283,630]" pageId="6" pageNumber="7">
<emphasis id="8E8DEAC3FFF4FFC9FE89FEC3FE6A366F" bold="true" box="[312,452,283,309]" pageId="6" pageNumber="7">Variability.</emphasis>
Length:
<quantity id="7B019B34FFF4FFC9FD81FEC4FD61366C" box="[560,719,284,310]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.13" metricValueMin="0.97" pageId="6" pageNumber="7" unit="mm" value="10.5" valueMax="11.3" valueMin="9.7">9.7–11.3 mm</quantity>
, length of elytra: 7.0–
<quantity id="7B019B34FFF4FFC9FC66FEC4FB9A366C" box="[983,1076,284,310]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="6" pageNumber="7" unit="mm" value="8.2">8.2 mm</quantity>
, width:
<quantity id="7B019B34FFF4FFC9FB24FEC4FA86366C" box="[1173,1320,284,310]" metricMagnitude="-3" metricUnit="m" metricValue="5.65" metricValueMax="6.2" metricValueMin="5.1" pageId="6" pageNumber="7" unit="mm" value="5.65" valueMax="6.2" valueMin="5.1">5.1–6.2 mm</quantity>
. Color: Dark brown to reddish brown. Female: pygidium strongly convex, antennal club shorter than the remaining joints combined.
</paragraph>
</subSubSection>
<subSubSection id="F4E3655AFFF4FFC9FE89FE4BFA8E357C" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="BC4636D1FFF4FFC9FE89FE4BFA8E357C" blockId="6.[264,1324,283,630]" pageId="6" pageNumber="7">
<emphasis id="8E8DEAC3FFF4FFC9FE89FE4BFE1936F7" bold="true" box="[312,439,403,429]" pageId="6" pageNumber="7">Diagnosis.</emphasis>
<taxonomicName id="7BF94D52FFF4FFC9FE73FE4CFD0D36F7" box="[450,675,404,429]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="howdeni" status="sp. nov.">
<emphasis id="8E8DEAC3FFF4FFC9FE73FE4CFD0D36F7" box="[450,675,404,429]" italics="true" pageId="6" pageNumber="7">Maladera howdeni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95BE57B8FFF4FFC9FD1CFE4CFD4136F4" box="[685,751,404,430]" pageId="6" pageNumber="7" rank="species">sp. n.</taxonomicNameLabel>
is very similar to
<taxonomicName id="7BF94D52FFF4FFC9FC63FE4CFB6936F4" authority="Frey, 1972" authorityName="Frey" authorityYear="1972" box="[978,1223,404,430]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="fusca">
<emphasis id="8E8DEAC3FFF4FFC9FC63FE4CFB9436F7" box="[978,1082,404,429]" italics="true" pageId="6" pageNumber="7">M. fusca</emphasis>
<bibRefCitation id="D8684B20FFF4FFC9FBF5FE4CFB6936F4" author="Frey" box="[1092,1223,404,430]" pageId="6" pageNumber="7" refString="Frey, G. (1972) Neue Sericinen der Klapperich-Ausbeute aus Fukien des Alexander Koenig Museum in Bonn (Col., Scarab., Melolonth.). Entomologische Arbeiten aus dem Museum Frey 23, 162 - 177." type="journal article" year="1972">Frey, 1972</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="7BF94D52FFF4FFC9FABAFE4CFE55368C" authority="Moser, 1918" authorityName="Moser" authorityYear="1918" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis id="8E8DEAC3FFF4FFC9FABAFE4CFEF7368F" italics="true" pageId="6" pageNumber="7">M. sauteri</emphasis>
<bibRefCitation id="D8684B20FFF4FFC9FED2FE64FE55368C" author="Moser" box="[355,507,444,470]" pageId="6" pageNumber="7" refString="Moser, J. (1918) Neue Melolonthiden aus der Sammlung des Deutschen Entomologischen Museums zu Berlin Dahlem. Stettiner Entomologische Zeitung, 79, 209 - 247." type="journal article" year="1918">Moser, 1918</bibRefCitation>
</taxonomicName>
, in habitus and shape of the male genitalia. It may be differentiated from both by the shorter phallobase and shorter parameres, as well as by the shape of the right paramere, which, in
<taxonomicName id="7BF94D52FFF4FFC9FD86FDD4FD6C357F" box="[567,706,524,549]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="howdeni" status="sp. nov.">
<emphasis id="8E8DEAC3FFF4FFC9FD86FDD4FD6C357F" box="[567,706,524,549]" italics="true" pageId="6" pageNumber="7">M. howdeni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95BE57B8FFF4FFC9FD79FDD4FCA6357C" box="[712,776,524,550]" pageId="6" pageNumber="7" rank="species">sp. n.</taxonomicNameLabel>
, is strongly flattened and not sinuate apically.
</paragraph>
</subSubSection>
<subSubSection id="F4E3655AFFF4FFC9FE89FDEBFB28352C" pageId="6" pageNumber="7" type="etymology">
<paragraph id="BC4636D1FFF4FFC9FE89FDEBFB28352C" blockId="6.[264,1324,283,630]" pageId="6" pageNumber="7">
<emphasis id="8E8DEAC3FFF4FFC9FE89FDEBFD8E3517" bold="true" box="[312,544,563,589]" pageId="6" pageNumber="7">Derivatio nominis.</emphasis>
The new species is dedicated to my dear colleague H.F. Howden (Ottawa) thanking him for made this interesting material available for study.
</paragraph>
</subSubSection>
</treatment>
</document>
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