treatments-xml/data/09/A5/26/09A5260E31A15CBAA44DDF3A92883500.xml

<document id="F008027D6BC98D0AD3A4992A245563FD" ID-DOI="10.3897/herpetozoa.37.e137809" ID-ZooBank="F99D228B-B5CF-4BCE-A021-295D57FE2D9F" ID-publisher-id="137809" URI-arpha="F2A8F56B-9777-5DF7-BDED-6FFF919B5B1F" URI-zoobank="http://zoobank.org/F99D228B-B5CF-4BCE-A021-295D57FE2D9F" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" article-type="research-article" checkinTime="1733325286099" checkinUser="pensoft" docAuthor="Poyarkov, Nikolay A., Bragin, Andrey M. &amp; Nguyen, Tan Van" docDate="2024" docId="09A5260E31A15CBAA44DDF3A92883500" docLanguage="en" docName="Herpetozoa 37: 379-390" docOrigin="Herpetozoa 37" docSource="https://herpetozoa.pensoft.net/article/137809/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Colubroelaps adleri Poyarkov, Bragin &amp; Nguyen, 2024, sp. nov." docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="390" masterDocId="F2A8F56B97775DF7BDED6FFF919B5B1F" masterDocTitle="A new endemic insular species of the genus Colubroelaps (Squamata, Serpentes, Colubroidea) from Khanh Hoa Province, Vietnam" masterLastPageNumber="390" masterPageNumber="379" pageNumber="379" updateTime="1733334610492" updateUser="admin">
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<mods:title id="30C69C1E3100F9D1AF24F52B9A42C374">A new endemic insular species of the genus Colubroelaps (Squamata, Serpentes, Colubroidea) from Khanh Hoa Province, Vietnam</mods:title>
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<mods:name id="F2D8702B35A1F334760BE31979F330BF" type="personal">
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<mods:namePart id="26CB05AAD28B49A2EA7A1ECCE7A3B5A9">Poyarkov, Nikolay A.</mods:namePart>
<mods:nameIdentifier id="CECAC96B34F60786B544FAB27829109F" type="ORCID">0000-0002-7576-2283</mods:nameIdentifier>
<mods:affiliation id="FE51612D519A52AFC499D5406604FFBD">Department of Vertebrate Zoology, Lomonosov Moscow State University, Leninskiye Gory, GSP- 1, Moscow 119991, Russia &amp; Joint Vietnam - Russia Tropical Science and Technology Research Center, 63 Nguyen Van Huyen Road, Nghia Do, Cau Giay, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="EFC351BB3DD8F128CE018B3704F0650D">Bragin, Andrey M.</mods:namePart>
<mods:nameIdentifier id="C93C1E14EE939DA17B3F53D125D5775F" type="ORCID">0000-0002-3621-9763</mods:nameIdentifier>
<mods:affiliation id="875D178103F5330A2177C2BDE682953B">Department of Vertebrate Zoology, Lomonosov Moscow State University, Leninskiye Gory, GSP- 1, Moscow 119991, Russia</mods:affiliation>
</mods:name>
<mods:name id="C4F9E75A678D1F33C520519E6029D8D2" type="personal">
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<mods:namePart id="79CF076B3A38833174665AC31D1F5BA4">Nguyen, Tan Van</mods:namePart>
<mods:nameIdentifier id="B16B2D7B8D76949B7F6A20E345FA7F39" type="ORCID">0000-0001-5413-968X</mods:nameIdentifier>
<mods:affiliation id="8B4D51D664141249FE8D2E919D754C10">Institute for Research and Training in Medicine, Biology and Pharmacy, Duy Tan University, Da Nang, 550000, Vietnam</mods:affiliation>
</mods:name>
<mods:typeOfResource id="2327643DF56C15AA22611A68817D09C2">text</mods:typeOfResource>
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<mods:title id="065C147B74969BEEF755E40C650CC761">Herpetozoa</mods:title>
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<mods:part id="36FFB748B0DEF45A375821346A25FACE">
<mods:date id="86305ECFC7EB4D43DE8481C984531ED2">2024</mods:date>
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<mods:number id="098F698DF8F4DF1AB40980EE5DE70FE8">2024-12-04</mods:number>
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<mods:number id="CB56A303F375B76518B05AF8DB8CEAA6">37</mods:number>
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<mods:classification id="558B52B18A3CD69C90F552DCEF2DFF76">journal article</mods:classification>
<mods:identifier id="3698115E2E6DF2D6510055175BAFD7C1" type="DOI">10.3897/herpetozoa.37.e137809</mods:identifier>
<mods:identifier id="6747497F0C3CA936D0E060CDD2031339" type="ZooBank">F99D228B-B5CF-4BCE-A021-295D57FE2D9F</mods:identifier>
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<treatment id="09A5260E31A15CBAA44DDF3A92883500" ID-arpha="09A5260E-31A1-5CBA-A44D-DF3A92883500" ID-zoobank="https://zoobank.org/F64EE8E0-B851-4923-8299-75793283310E" LSID="urn:lsid:plazi:treatment:09A5260E31A15CBAA44DDF3A92883500" LSID-ZBK="urn:lsid:zoobank.org:act:F64EE8E0-B851-4923-8299-75793283310E" httpUri="http://treatment.plazi.org/id/09A5260E31A15CBAA44DDF3A92883500" scope_class="Squamata" scope_superFamily="Colubroidea">
<subSubSection id="6389689651204F8EEF335B78ACB7C43D" type="nomenclature">
<paragraph id="35FDD8D1CFCD899A02E528DEFD7C4966">
<taxonomicName id="25BD9A0BFB0E5F2C89A5E02D5EBE20D0" authorityName="Poyarkov &amp; Bragin &amp; Nguyen" authorityYear="2024" class="Squamata" family="Colubridae" genus="Colubroelaps" kingdom="Animalia" phylum="Chordata" rank="species" species="adleri" status="sp. nov.">
<emphasis id="D38059C7F2E71DA5E39D1C6DBF4BE811" italics="true">Colubroelaps adleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C35458050D870AF6043305909EEAA83C" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="B28ACC5A73837A9F99D25A494F414F76" type="description">
<paragraph id="4C76DD85C2C241583616577484F242C0">
<tableCitation id="98F0257C57A8A26A0B4A06B5E8868A25" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological data on the type series of Colubroelaps adleri sp. nov. and C. nguyenvansangi; diagnostic differences from the new species are marked in bold. Notes: for abbreviations, see the Materials and methods section; “ * ”: data on the holotype of C. nguyenvansangi as reported by Orlov et al. (2009), when different from our observations." httpUri="http://table.plazi.org/id/8E5F0BED38B5CEE14412962DA7D57026" tableUuid="8E5F0BED38B5CEE14412962DA7D57026">Table 1</tableCitation>
, 
<figureCitation id="2007CBC50170CBA31887F3F557E774FA" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The holotype of Colubroelaps adleri sp. nov. in life (ZMMU Re- 18000, adult female) from Hon Tre Island, Khanh Hoa Province, Vietnam. A. Dorsal view of the head; B. Ventral view of the head; C. Lateral view of the right side of the head; D. Lateral view of the left side of the head; E. Ventral view of the tail; F. Ventral view of the body; G. Dorsal view of the body. Scale bars: 5 mm (A – D); 10 mm (E – G). Photographs by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure2" httpUri="https://binary.pensoft.net/fig/1190525">Figs 2</figureCitation>
, 
<figureCitation id="4E7FC8C1A3CDDFACB342656135812934" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Line drawings of head scalation of the Colubroelaps adleri sp. nov. holotype (ZMMU Re- 18000, adult female; A – C) and of the C. nguyenvansangi holotype (ZISP 25682, adult female; D – F). Line drawings show the head scales in lateral (A, D), dorsal (B, E), and ventral (C, F) aspects. Abbreviations: ACS = anterior chin shield; AT = anterior temporal; F = frontal; G = gular scale; IL = infralabial; IN = internasal; L = loreal; LPT = lower posterior temporal; M = mental; N = nasal; P = parietal; PCS = posterior chin shield; PF = prefrontal; PO = postocular; PrO = preocular; R = rostral; SL = supralabial; SO = supraocular; UPT = upper posterior temporal. Scale bar equals 5 mm. Line drawings by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure3" httpUri="https://binary.pensoft.net/fig/1190526">3 A – C</figureCitation>
, 
<figureCitation id="74E767B6B4C0CA884374F3B0A3D2BDE7" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Two species of the genus Colubroelaps in life. A. Colubroelaps adleri sp. nov., holotype ZMMU Re- 18000, adult female from Dam Bay Research Station, Hon Tre Island, Khanh Hoa Province, Vietnam; B. Colubroelaps nguyenvansangi, ZMMU Re- 15485, adult male from Cat Tien NP., Tan Phu District, Dong Nai Province, Vietnam. Photographs by: A. M. Bragin (A); E. A. Galoyan (B)." figureDoi="10.3897/herpetozoa.37.e137809.figure4" httpUri="https://binary.pensoft.net/fig/1190527">4 A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E34AE" type="Holotype">
<paragraph id="AB1AC3382624CD404EB3AC629479499B">
<heading id="009FD1F0C08549993CF4E81E25F05FD3" reason="title">
<typeStatus id="1AC145039741CF1C0032F7EE66A12E30" type="holotype">Holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="4A7FCB5A35C532B65D2D083F05005481">
<materialsCitation id="177F6A5EB5895C869233F7A82CD8187A" collectingDate="2023-06-17" collectionCode="ZMMU" collectorName="Nikolay A. Poyarkov &amp; Andrey M. Bragin" country="Vietnam" county="Nha Trang City" elevation="30" latitude="12.198" location="Dam Bay Research Station" longLatPrecision="77" longitude="109.289" municipality="Hon Tre Island" specimenCode="ZMMU Re-18000" specimenCount="1" specimenCount-female="1" stateProvince="Khanh Hoa Province" typeStatus="holotype">
• 
<specimenCode id="BF27D3CBAAADF6A7AEBDA925233627B0">ZMMU Re-18000</specimenCode>
(field tag NAP-15227), 
<specimenCount id="A4B3A73BEC426747F124A4B9142CA853" count="1" type="female">adult female</specimenCount>
, collected by 
<collectorName id="1A916C206D817A2734C8F19C1697C679">Nikolay A. Poyarkov</collectorName>
and 
<collectorName id="22AE39E45ECD0B394484904ADC3DC9DF">Andrey M. Bragin</collectorName>
on 
<date id="57F3E462F7969C32B6E782DF204D5CFF" value="2023-06-17">
<collectingDate id="D4D58FDDF5A4B13F437903572F7E758F" value="2023-06-17">June 17, 2023</collectingDate>
</date>
, from 
<location id="E2956D1CE65B355962A77E7DF21CE756" LSID="urn:lsid:plazi:treatment:09A5260E31A15CBAA44DDF3A92883500:E2956D1CE65B355962A77E7DF21CE756" country="Vietnam" county="Nha Trang City" latitude="12.198" longLatPrecision="77" longitude="109.289" municipality="Hon Tre Island" name="Dam Bay Research Station" stateProvince="Khanh Hoa Province">Dam Bay Research Station</location>
, 
<collectingMunicipality id="6D3D9046E628ABCD0F5C992A876E5A3C">Hon Tre Island</collectingMunicipality>
, within 
<collectingMunicipality id="28009E2982BAA04D847F2788B84CD1B3">Nha Trang Bay in Vinh Nguyen Ward</collectingMunicipality>
, 
<collectingCounty id="BDA804EEB9C81FD6F1E283CDE77874FB">Nha Trang City</collectingCounty>
, 
<collectingRegion id="15B560511ED8794899015FEB2387D5BC" country="Vietnam" name="Khanh Hoa">Khanh Hoa Province</collectingRegion>
, 
<collectingCountry id="4615101F52BE71DB9C828DF22ABF9022" name="Vietnam">Vietnam</collectingCountry>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EF5AE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[109.289000,12.198000]}">
<geoCoordinate id="FF3AC19575316ADF011E795A670D3662" degrees="12.198" direction="north" orientation="latitude" precision="55" value="12.198">12.198 ° N</geoCoordinate>
, 
<geoCoordinate id="4069AACE40D772E2B0C46A4673F108DE" degrees="109.289" direction="east" orientation="longitude" precision="55" value="109.289">109.289 ° E</geoCoordinate>
</named-content>
; at the elevation of 
<elevation id="9AE2F083C2D5FB08693074515A44FE50" metricMagnitude="1" metricUnit="m" metricValue="3.0" unit="m" value="30.0">
<quantity id="9E5A2D0E674D383C7B78DE45D0CA9C26" metricMagnitude="1" metricUnit="m" metricValue="3.0" unit="m" value="30.0">30 m</quantity>
<abbrev id="ABBRID0EK5AE" xlink_title="above sea level">asl</abbrev>
</elevation>
.).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EO5AE" type="diagnosis">
<paragraph id="9102E4CC1F97BD06F7FCA91E02E4B426">
<heading id="BDD38CB0EB962C122710F57F3A58C01E" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="31A7FEFB10DEF212C335339A21E778C3">
<taxonomicName id="505E3212602E9F811E7A4597A435E8D1" authorityName="Poyarkov &amp; Bragin &amp; Nguyen" authorityYear="2024" class="Squamata" family="Colubridae" genus="Colubroelaps" kingdom="Animalia" phylum="Chordata" rank="species" species="adleri" status="sp. nov.">
<emphasis id="450D61B3D6E876EE7381149DF902B5AD" italics="true">Colubroelaps adleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CD34CFC99531436EA269921B79D2D655" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from 
<taxonomicName id="D58CE34358545A09BB6DDDAB787ECD16" authorityName="Orlov, Kharin, Ananjeva, Nguyen &amp; Nguyen" authorityYear="2009" class="Squamata" family="Colubridae" genus="Colubroelaps" kingdom="Animalia" phylum="Chordata" rank="species" species="nguyenvansangi">
<emphasis id="A6CA3A111605E613C71ECF6D1CDE5D74" italics="true">C. nguyenvansangi</emphasis>
</taxonomicName>
by the following combination of morphological characters: body size small (
<abbrev id="ABBRID0EK6AE" xlink_title="total length">TL</abbrev>
<quantity id="D0E118F589566FB257AE9046BBB7F354" metricMagnitude="-1" metricUnit="m" metricValue="4.02" unit="mm" value="402.0">402 mm</quantity>
); tail relatively short (ratio 
<abbrev id="ABBRID0EO6AE" xlink_title="tail length">TaL</abbrev>
/ 
<abbrev id="ABBRID0ES6AE" xlink_title="total length">TL</abbrev>
0.10); dorsal scales in 14–14 – 14 rows; supralabials six, third and fourth entering orbit; infralabials seven; loreal present; ventral scales 234; subcaudals 30, all divided; cloacal plate divided; dorsal coloration pale brown with narrow and interrupted dark stripe along spine; body flanks dark gray lacking bluish iridescence; ventrally uniform off-white; head black with rostral, nasals, prefrontals, preoculars, loreal, and the two anterior supralabials, as well as the anterior parts of supraoculars and frontal shields dirty yellowish-brown with dark brown spots.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EW6AE" type="description">
<paragraph id="3941C156E27E6B92E87E2F4D8DE2F0F7">
<heading id="7B3ADF882BEE6DF07D8832547196F580" reason="title">
Description of the 
<typeStatus id="F0728A774A0AE196DD15CD318C4D8F79" type="holotype">holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="AFAC4E53180CB463747D9C43342145F5">
Adult female specimen in a good state of preservation (Fig. 
<figureCitation id="EF40E4C1D02C373FF5C64EBDB3EA0764" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The holotype of Colubroelaps adleri sp. nov. in life (ZMMU Re- 18000, adult female) from Hon Tre Island, Khanh Hoa Province, Vietnam. A. Dorsal view of the head; B. Ventral view of the head; C. Lateral view of the right side of the head; D. Lateral view of the left side of the head; E. Ventral view of the tail; F. Ventral view of the body; G. Dorsal view of the body. Scale bars: 5 mm (A – D); 10 mm (E – G). Photographs by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure2" httpUri="https://binary.pensoft.net/fig/1190525">2</figureCitation>
). Body strongly elongated, very thin, vermiform, round in cross-section. Tail obtusely rounded, comparatively short, the tip of tail blunt (
<abbrev id="ABBRID0EBAAG" xlink_title="snout-vent length">SVL</abbrev>
<quantity id="46FBC4EF1222CC015D439D2E80EE6E27" metricMagnitude="-1" metricUnit="m" metricValue="3.62" unit="mm" value="362.0">362 mm</quantity>
; 
<abbrev id="ABBRID0EFAAG" xlink_title="tail length">TaL</abbrev>
<quantity id="29E0AEAE39B8239D2AD9B0DED6E9B99F" metricMagnitude="-2" metricUnit="m" metricValue="4.02" unit="mm" value="40.2">40.2 mm</quantity>
; 
<abbrev id="ABBRID0EJAAG" xlink_title="total length">TL</abbrev>
<quantity id="FFDF9E4BD52DED1F4B6C2866F780FDF0" metricMagnitude="-1" metricUnit="m" metricValue="4.022" unit="mm" value="402.2">402.2 mm</quantity>
; ratio 
<abbrev id="ABBRID0ENAAG" xlink_title="tail length">TaL</abbrev>
/ 
<abbrev id="ABBRID0ERAAG" xlink_title="total length">TL</abbrev>
0.10) (Fig. 
<figureCitation id="3D2DDEC510EA05603F4A8EEE84A8C9BB" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The holotype of Colubroelaps adleri sp. nov. in life (ZMMU Re- 18000, adult female) from Hon Tre Island, Khanh Hoa Province, Vietnam. A. Dorsal view of the head; B. Ventral view of the head; C. Lateral view of the right side of the head; D. Lateral view of the left side of the head; E. Ventral view of the tail; F. Ventral view of the body; G. Dorsal view of the body. Scale bars: 5 mm (A – D); 10 mm (E – G). Photographs by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure2" httpUri="https://binary.pensoft.net/fig/1190525">2 E – G</figureCitation>
). Head very small, rounded, slightly dorso-ventrally depressed, covered by large, regular, symmetric shields (Figs 
<figureCitation id="0C4F71B5F2134EE4D71BB6302960BDF0" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The holotype of Colubroelaps adleri sp. nov. in life (ZMMU Re- 18000, adult female) from Hon Tre Island, Khanh Hoa Province, Vietnam. A. Dorsal view of the head; B. Ventral view of the head; C. Lateral view of the right side of the head; D. Lateral view of the left side of the head; E. Ventral view of the tail; F. Ventral view of the body; G. Dorsal view of the body. Scale bars: 5 mm (A – D); 10 mm (E – G). Photographs by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure2" httpUri="https://binary.pensoft.net/fig/1190525">2 A – D</figureCitation>
, 
<figureCitation id="408C2437BBEA6BDF54B970F4C7477B55" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Line drawings of head scalation of the Colubroelaps adleri sp. nov. holotype (ZMMU Re- 18000, adult female; A – C) and of the C. nguyenvansangi holotype (ZISP 25682, adult female; D – F). Line drawings show the head scales in lateral (A, D), dorsal (B, E), and ventral (C, F) aspects. Abbreviations: ACS = anterior chin shield; AT = anterior temporal; F = frontal; G = gular scale; IL = infralabial; IN = internasal; L = loreal; LPT = lower posterior temporal; M = mental; N = nasal; P = parietal; PCS = posterior chin shield; PF = prefrontal; PO = postocular; PrO = preocular; R = rostral; SL = supralabial; SO = supraocular; UPT = upper posterior temporal. Scale bar equals 5 mm. Line drawings by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure3" httpUri="https://binary.pensoft.net/fig/1190526">3 A – C</figureCitation>
), not distinct from the neck (
<abbrev id="ABBRID0EBBAG" xlink_title="head length">HL</abbrev>
<quantity id="8F51469614161B651AE7112DE6A03DB3" metricMagnitude="-3" metricUnit="m" metricValue="7.99" unit="mm" value="7.99">7.99 mm</quantity>
; 
<abbrev id="ABBRID0EFBAG" xlink_title="maximum head width">HW</abbrev>
<quantity id="EDD65047DBF1FE3E592EE29500AA6893" metricMagnitude="-3" metricUnit="m" metricValue="5.16" unit="mm" value="5.16">5.16 mm</quantity>
; ratio 
<abbrev id="ABBRID0EJBAG" xlink_title="maximum head width">HW</abbrev>
/ 
<abbrev id="ABBRID0ENBAG" xlink_title="head length">HL</abbrev>
0.65). Snout wide, short, bluntly rounded in dorsal view (Fig. 
<figureCitation id="AD7E6BE8111FDCDD9339851203DCFA0B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The holotype of Colubroelaps adleri sp. nov. in life (ZMMU Re- 18000, adult female) from Hon Tre Island, Khanh Hoa Province, Vietnam. A. Dorsal view of the head; B. Ventral view of the head; C. Lateral view of the right side of the head; D. Lateral view of the left side of the head; E. Ventral view of the tail; F. Ventral view of the body; G. Dorsal view of the body. Scale bars: 5 mm (A – D); 10 mm (E – G). Photographs by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure2" httpUri="https://binary.pensoft.net/fig/1190525">2 A, B</figureCitation>
), slightly tapering in lateral view (Fig. 
<figureCitation id="096DB81AEE507780468FBC859AA33F01" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The holotype of Colubroelaps adleri sp. nov. in life (ZMMU Re- 18000, adult female) from Hon Tre Island, Khanh Hoa Province, Vietnam. A. Dorsal view of the head; B. Ventral view of the head; C. Lateral view of the right side of the head; D. Lateral view of the left side of the head; E. Ventral view of the tail; F. Ventral view of the body; G. Dorsal view of the body. Scale bars: 5 mm (A – D); 10 mm (E – G). Photographs by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure2" httpUri="https://binary.pensoft.net/fig/1190525">2 C, D</figureCitation>
). Rostral barely visible in dorsal aspect, triangular (Figs 
<figureCitation id="8562F87F67B30332D801582D1371FEE6" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The holotype of Colubroelaps adleri sp. nov. in life (ZMMU Re- 18000, adult female) from Hon Tre Island, Khanh Hoa Province, Vietnam. A. Dorsal view of the head; B. Ventral view of the head; C. Lateral view of the right side of the head; D. Lateral view of the left side of the head; E. Ventral view of the tail; F. Ventral view of the body; G. Dorsal view of the body. Scale bars: 5 mm (A – D); 10 mm (E – G). Photographs by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure2" httpUri="https://binary.pensoft.net/fig/1190525">2 A</figureCitation>
, 
<figureCitation id="E2D2929475AC742A9A6C13D2C9429A6D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Line drawings of head scalation of the Colubroelaps adleri sp. nov. holotype (ZMMU Re- 18000, adult female; A – C) and of the C. nguyenvansangi holotype (ZISP 25682, adult female; D – F). Line drawings show the head scales in lateral (A, D), dorsal (B, E), and ventral (C, F) aspects. Abbreviations: ACS = anterior chin shield; AT = anterior temporal; F = frontal; G = gular scale; IL = infralabial; IN = internasal; L = loreal; LPT = lower posterior temporal; M = mental; N = nasal; P = parietal; PCS = posterior chin shield; PF = prefrontal; PO = postocular; PrO = preocular; R = rostral; SL = supralabial; SO = supraocular; UPT = upper posterior temporal. Scale bar equals 5 mm. Line drawings by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure3" httpUri="https://binary.pensoft.net/fig/1190526">3 A</figureCitation>
). Eyes very small with round pupils (
<abbrev id="ABBRID0EBCAG" xlink_title="horizontal eye diameter">ED</abbrev>
<quantity id="10FA2EB923434834F9BFB6F0A2AD4EF7" metricMagnitude="-4" metricUnit="m" metricValue="5.1" unit="mm" value="0.51">0.51 mm</quantity>
, EN 
<quantity id="5BED05B98D7463F4125A325F7CD31B19" metricMagnitude="-3" metricUnit="m" metricValue="1.46" unit="mm" value="1.46">1.46 mm</quantity>
, 
<abbrev id="ABBRID0EFCAG" xlink_title="snout length">SnL</abbrev>
<quantity id="5380E95308F4735045BC055E5D31ADC5" metricMagnitude="-3" metricUnit="m" metricValue="2.78" unit="mm" value="2.78">2.78 mm</quantity>
) (Fig. 
<figureCitation id="1554239A48406FB5924A3DCE16F3766D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The holotype of Colubroelaps adleri sp. nov. in life (ZMMU Re- 18000, adult female) from Hon Tre Island, Khanh Hoa Province, Vietnam. A. Dorsal view of the head; B. Ventral view of the head; C. Lateral view of the right side of the head; D. Lateral view of the left side of the head; E. Ventral view of the tail; F. Ventral view of the body; G. Dorsal view of the body. Scale bars: 5 mm (A – D); 10 mm (E – G). Photographs by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure2" httpUri="https://binary.pensoft.net/fig/1190525">2 C, D</figureCitation>
). Single nasal on each side of head (1 / 1 nasals); nostril oval-shaped, with horizontal orienatation, completely enclosed in nasal scale, located closer to the posterior edge of nasal (Figs 
<figureCitation id="5CEB3472290CA61F3C15D3AB28BB4DF7" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The holotype of Colubroelaps adleri sp. nov. in life (ZMMU Re- 18000, adult female) from Hon Tre Island, Khanh Hoa Province, Vietnam. A. Dorsal view of the head; B. Ventral view of the head; C. Lateral view of the right side of the head; D. Lateral view of the left side of the head; E. Ventral view of the tail; F. Ventral view of the body; G. Dorsal view of the body. Scale bars: 5 mm (A – D); 10 mm (E – G). Photographs by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure2" httpUri="https://binary.pensoft.net/fig/1190525">2 C, D</figureCitation>
, 
<figureCitation id="31191163D914243142C94EE982F49807" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Line drawings of head scalation of the Colubroelaps adleri sp. nov. holotype (ZMMU Re- 18000, adult female; A – C) and of the C. nguyenvansangi holotype (ZISP 25682, adult female; D – F). Line drawings show the head scales in lateral (A, D), dorsal (B, E), and ventral (C, F) aspects. Abbreviations: ACS = anterior chin shield; AT = anterior temporal; F = frontal; G = gular scale; IL = infralabial; IN = internasal; L = loreal; LPT = lower posterior temporal; M = mental; N = nasal; P = parietal; PCS = posterior chin shield; PF = prefrontal; PO = postocular; PrO = preocular; R = rostral; SL = supralabial; SO = supraocular; UPT = upper posterior temporal. Scale bar equals 5 mm. Line drawings by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure3" httpUri="https://binary.pensoft.net/fig/1190526">3 B</figureCitation>
); single (1 / 1) loreal, small, almost triangular in shape, shorter than the eye diameter, in contact with nasal, 1 
<superScript id="A67D3689A37AB9FEF4C6F14DD05E9639">st</superScript>
and 2 
<superScript id="30C075C0B33B1BFD79D508B164B3E6C0">nd</superScript>
supralabial, preocular, prefrontal and internasal; single (1 / 1) preocular, large, elongated pentagonal in shape, subequal to the eye diameter, in contact with 2 
<superScript id="63EDD0EA8DFCFF4665340FF536F89E7C">nd</superScript>
and 3 
<superScript id="0B1E13531776D032340C9C12E8A59FBD">rd</superScript>
supralabial, the orbit, supraocular, prefrontal and loreal; single (1 / 1) postocular, trapezoidal in shape, smaller than the eye diameter, in contact with 4 
<superScript id="A957458C818A1009384F78E20D410CFE">th</superScript>
supralabial, parietal, supraocular and the orbit; single (1 / 1) supraocular, large and wide, ca. twice longer than the eye diameter, in contact with the orbit, postocular, parietal, frontal, prefrontal and preocular (Figs 
<figureCitation id="E83786741D016F03475412211FC12918" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The holotype of Colubroelaps adleri sp. nov. in life (ZMMU Re- 18000, adult female) from Hon Tre Island, Khanh Hoa Province, Vietnam. A. Dorsal view of the head; B. Ventral view of the head; C. Lateral view of the right side of the head; D. Lateral view of the left side of the head; E. Ventral view of the tail; F. Ventral view of the body; G. Dorsal view of the body. Scale bars: 5 mm (A – D); 10 mm (E – G). Photographs by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure2" httpUri="https://binary.pensoft.net/fig/1190525">2 C, D</figureCitation>
, 
<figureCitation id="3067B10B22D135A3C87705A4CB112364" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Line drawings of head scalation of the Colubroelaps adleri sp. nov. holotype (ZMMU Re- 18000, adult female; A – C) and of the C. nguyenvansangi holotype (ZISP 25682, adult female; D – F). Line drawings show the head scales in lateral (A, D), dorsal (B, E), and ventral (C, F) aspects. Abbreviations: ACS = anterior chin shield; AT = anterior temporal; F = frontal; G = gular scale; IL = infralabial; IN = internasal; L = loreal; LPT = lower posterior temporal; M = mental; N = nasal; P = parietal; PCS = posterior chin shield; PF = prefrontal; PO = postocular; PrO = preocular; R = rostral; SL = supralabial; SO = supraocular; UPT = upper posterior temporal. Scale bar equals 5 mm. Line drawings by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure3" httpUri="https://binary.pensoft.net/fig/1190526">3 B</figureCitation>
). Six supralabials on each side of the head (6 / 6), fourth one the largest, third and fourth supralabials in contact with eye, posterior edge of fourth supralabial in contact with anterior temporal, separating fifth supralabial from parietal; fourth supralabial in contact with parietal, postocular, and anterior temporal (Figs 
<figureCitation id="96950B52EE2907A67F22C3D26D0FD711" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The holotype of Colubroelaps adleri sp. nov. in life (ZMMU Re- 18000, adult female) from Hon Tre Island, Khanh Hoa Province, Vietnam. A. Dorsal view of the head; B. Ventral view of the head; C. Lateral view of the right side of the head; D. Lateral view of the left side of the head; E. Ventral view of the tail; F. Ventral view of the body; G. Dorsal view of the body. Scale bars: 5 mm (A – D); 10 mm (E – G). Photographs by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure2" httpUri="https://binary.pensoft.net/fig/1190525">2 C, D</figureCitation>
, 
<figureCitation id="7DBB36EC880E1022943C7FCD961E0FA2" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Line drawings of head scalation of the Colubroelaps adleri sp. nov. holotype (ZMMU Re- 18000, adult female; A – C) and of the C. nguyenvansangi holotype (ZISP 25682, adult female; D – F). Line drawings show the head scales in lateral (A, D), dorsal (B, E), and ventral (C, F) aspects. Abbreviations: ACS = anterior chin shield; AT = anterior temporal; F = frontal; G = gular scale; IL = infralabial; IN = internasal; L = loreal; LPT = lower posterior temporal; M = mental; N = nasal; P = parietal; PCS = posterior chin shield; PF = prefrontal; PO = postocular; PrO = preocular; R = rostral; SL = supralabial; SO = supraocular; UPT = upper posterior temporal. Scale bar equals 5 mm. Line drawings by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure3" httpUri="https://binary.pensoft.net/fig/1190526">3 A</figureCitation>
); seven (7 / 7) infralabials, the anterior-most pair in contact with each other behind the mental (Figs 
<figureCitation id="964DDD9ECFD72C805D4C7936ABFFCABB" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The holotype of Colubroelaps adleri sp. nov. in life (ZMMU Re- 18000, adult female) from Hon Tre Island, Khanh Hoa Province, Vietnam. A. Dorsal view of the head; B. Ventral view of the head; C. Lateral view of the right side of the head; D. Lateral view of the left side of the head; E. Ventral view of the tail; F. Ventral view of the body; G. Dorsal view of the body. Scale bars: 5 mm (A – D); 10 mm (E – G). Photographs by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure2" httpUri="https://binary.pensoft.net/fig/1190525">2 B</figureCitation>
, 
<figureCitation id="937769281DB06051495A34DCF6BE389A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Line drawings of head scalation of the Colubroelaps adleri sp. nov. holotype (ZMMU Re- 18000, adult female; A – C) and of the C. nguyenvansangi holotype (ZISP 25682, adult female; D – F). Line drawings show the head scales in lateral (A, D), dorsal (B, E), and ventral (C, F) aspects. Abbreviations: ACS = anterior chin shield; AT = anterior temporal; F = frontal; G = gular scale; IL = infralabial; IN = internasal; L = loreal; LPT = lower posterior temporal; M = mental; N = nasal; P = parietal; PCS = posterior chin shield; PF = prefrontal; PO = postocular; PrO = preocular; R = rostral; SL = supralabial; SO = supraocular; UPT = upper posterior temporal. Scale bar equals 5 mm. Line drawings by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure3" httpUri="https://binary.pensoft.net/fig/1190526">3 C</figureCitation>
); the first four pairs of infralabials in contact with the anterior pair of chin shields (Figs 
<figureCitation id="B4FAE58979CC5672EE8BD10A31C34E12" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The holotype of Colubroelaps adleri sp. nov. in life (ZMMU Re- 18000, adult female) from Hon Tre Island, Khanh Hoa Province, Vietnam. A. Dorsal view of the head; B. Ventral view of the head; C. Lateral view of the right side of the head; D. Lateral view of the left side of the head; E. Ventral view of the tail; F. Ventral view of the body; G. Dorsal view of the body. Scale bars: 5 mm (A – D); 10 mm (E – G). Photographs by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure2" httpUri="https://binary.pensoft.net/fig/1190525">2 B</figureCitation>
, 
<figureCitation id="0BA9F271763C53B8751202833D350089" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Line drawings of head scalation of the Colubroelaps adleri sp. nov. holotype (ZMMU Re- 18000, adult female; A – C) and of the C. nguyenvansangi holotype (ZISP 25682, adult female; D – F). Line drawings show the head scales in lateral (A, D), dorsal (B, E), and ventral (C, F) aspects. Abbreviations: ACS = anterior chin shield; AT = anterior temporal; F = frontal; G = gular scale; IL = infralabial; IN = internasal; L = loreal; LPT = lower posterior temporal; M = mental; N = nasal; P = parietal; PCS = posterior chin shield; PF = prefrontal; PO = postocular; PrO = preocular; R = rostral; SL = supralabial; SO = supraocular; UPT = upper posterior temporal. Scale bar equals 5 mm. Line drawings by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure3" httpUri="https://binary.pensoft.net/fig/1190526">3 C</figureCitation>
); fourth and fifth infralabials in contact with the posterior pair of chin shields (Figs 
<figureCitation id="C1594D5A97854C39DA91E77C0518431A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The holotype of Colubroelaps adleri sp. nov. in life (ZMMU Re- 18000, adult female) from Hon Tre Island, Khanh Hoa Province, Vietnam. A. Dorsal view of the head; B. Ventral view of the head; C. Lateral view of the right side of the head; D. Lateral view of the left side of the head; E. Ventral view of the tail; F. Ventral view of the body; G. Dorsal view of the body. Scale bars: 5 mm (A – D); 10 mm (E – G). Photographs by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure2" httpUri="https://binary.pensoft.net/fig/1190525">2 B</figureCitation>
, 
<figureCitation id="2F5A7DACEE9357318939981A5BEF04C2" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Line drawings of head scalation of the Colubroelaps adleri sp. nov. holotype (ZMMU Re- 18000, adult female; A – C) and of the C. nguyenvansangi holotype (ZISP 25682, adult female; D – F). Line drawings show the head scales in lateral (A, D), dorsal (B, E), and ventral (C, F) aspects. Abbreviations: ACS = anterior chin shield; AT = anterior temporal; F = frontal; G = gular scale; IL = infralabial; IN = internasal; L = loreal; LPT = lower posterior temporal; M = mental; N = nasal; P = parietal; PCS = posterior chin shield; PF = prefrontal; PO = postocular; PrO = preocular; R = rostral; SL = supralabial; SO = supraocular; UPT = upper posterior temporal. Scale bar equals 5 mm. Line drawings by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure3" httpUri="https://binary.pensoft.net/fig/1190526">3 C</figureCitation>
). Three gular scales aligned between the chin shields and the first preventral (Figs 
<figureCitation id="321F850ECCD0DD6F63313972A776CE83" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The holotype of Colubroelaps adleri sp. nov. in life (ZMMU Re- 18000, adult female) from Hon Tre Island, Khanh Hoa Province, Vietnam. A. Dorsal view of the head; B. Ventral view of the head; C. Lateral view of the right side of the head; D. Lateral view of the left side of the head; E. Ventral view of the tail; F. Ventral view of the body; G. Dorsal view of the body. Scale bars: 5 mm (A – D); 10 mm (E – G). Photographs by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure2" httpUri="https://binary.pensoft.net/fig/1190525">2 B</figureCitation>
, 
<figureCitation id="02A42F22A994400D640B945ABB7E6B97" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Line drawings of head scalation of the Colubroelaps adleri sp. nov. holotype (ZMMU Re- 18000, adult female; A – C) and of the C. nguyenvansangi holotype (ZISP 25682, adult female; D – F). Line drawings show the head scales in lateral (A, D), dorsal (B, E), and ventral (C, F) aspects. Abbreviations: ACS = anterior chin shield; AT = anterior temporal; F = frontal; G = gular scale; IL = infralabial; IN = internasal; L = loreal; LPT = lower posterior temporal; M = mental; N = nasal; P = parietal; PCS = posterior chin shield; PF = prefrontal; PO = postocular; PrO = preocular; R = rostral; SL = supralabial; SO = supraocular; UPT = upper posterior temporal. Scale bar equals 5 mm. Line drawings by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure3" httpUri="https://binary.pensoft.net/fig/1190526">3 C</figureCitation>
). One pair of enlarged internasals, in contact with each other; one pair of enlarged prefrontals, in contact with each other; one wide, pentagonal frontal shield; one pair of wide, triangular parietals, in contact with each other, anteriorly separated by the protruding posterior edge of frontal (Figs 
<figureCitation id="C260D69D390DFCB31C637781090CAFEC" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The holotype of Colubroelaps adleri sp. nov. in life (ZMMU Re- 18000, adult female) from Hon Tre Island, Khanh Hoa Province, Vietnam. A. Dorsal view of the head; B. Ventral view of the head; C. Lateral view of the right side of the head; D. Lateral view of the left side of the head; E. Ventral view of the tail; F. Ventral view of the body; G. Dorsal view of the body. Scale bars: 5 mm (A – D); 10 mm (E – G). Photographs by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure2" httpUri="https://binary.pensoft.net/fig/1190525">2 A</figureCitation>
, 
<figureCitation id="F1E7AEB2D0F0C6A655786A01BC6A99C6" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Line drawings of head scalation of the Colubroelaps adleri sp. nov. holotype (ZMMU Re- 18000, adult female; A – C) and of the C. nguyenvansangi holotype (ZISP 25682, adult female; D – F). Line drawings show the head scales in lateral (A, D), dorsal (B, E), and ventral (C, F) aspects. Abbreviations: ACS = anterior chin shield; AT = anterior temporal; F = frontal; G = gular scale; IL = infralabial; IN = internasal; L = loreal; LPT = lower posterior temporal; M = mental; N = nasal; P = parietal; PCS = posterior chin shield; PF = prefrontal; PO = postocular; PrO = preocular; R = rostral; SL = supralabial; SO = supraocular; UPT = upper posterior temporal. Scale bar equals 5 mm. Line drawings by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure3" httpUri="https://binary.pensoft.net/fig/1190526">3 B</figureCitation>
), posteriorly barely extending beyond posterior edges of upper posterior temporals (Fig. 
<figureCitation id="7752CBA450D5451D2A4770921EDF4AE0" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Line drawings of head scalation of the Colubroelaps adleri sp. nov. holotype (ZMMU Re- 18000, adult female; A – C) and of the C. nguyenvansangi holotype (ZISP 25682, adult female; D – F). Line drawings show the head scales in lateral (A, D), dorsal (B, E), and ventral (C, F) aspects. Abbreviations: ACS = anterior chin shield; AT = anterior temporal; F = frontal; G = gular scale; IL = infralabial; IN = internasal; L = loreal; LPT = lower posterior temporal; M = mental; N = nasal; P = parietal; PCS = posterior chin shield; PF = prefrontal; PO = postocular; PrO = preocular; R = rostral; SL = supralabial; SO = supraocular; UPT = upper posterior temporal. Scale bar equals 5 mm. Line drawings by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure3" httpUri="https://binary.pensoft.net/fig/1190526">3 B</figureCitation>
). Single (1 / 1) anterior temporal, quadrangular in shape; two (2 / 2) posterior temporals, lower one the largest, two and a half times larger than upper posterior temporal (Fig. 
<figureCitation id="1DCD3B7F0C8708C53119E852CE7A85E4" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Line drawings of head scalation of the Colubroelaps adleri sp. nov. holotype (ZMMU Re- 18000, adult female; A – C) and of the C. nguyenvansangi holotype (ZISP 25682, adult female; D – F). Line drawings show the head scales in lateral (A, D), dorsal (B, E), and ventral (C, F) aspects. Abbreviations: ACS = anterior chin shield; AT = anterior temporal; F = frontal; G = gular scale; IL = infralabial; IN = internasal; L = loreal; LPT = lower posterior temporal; M = mental; N = nasal; P = parietal; PCS = posterior chin shield; PF = prefrontal; PO = postocular; PrO = preocular; R = rostral; SL = supralabial; SO = supraocular; UPT = upper posterior temporal. Scale bar equals 5 mm. Line drawings by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure3" httpUri="https://binary.pensoft.net/fig/1190526">3 A, B</figureCitation>
), upper one rectangular-shaped, narrow, notably protruding beyond the upper edge of lower posterior temporal (Figs 
<figureCitation id="C633C667DFF0B4A1C0DE1B76D5509C8F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The holotype of Colubroelaps adleri sp. nov. in life (ZMMU Re- 18000, adult female) from Hon Tre Island, Khanh Hoa Province, Vietnam. A. Dorsal view of the head; B. Ventral view of the head; C. Lateral view of the right side of the head; D. Lateral view of the left side of the head; E. Ventral view of the tail; F. Ventral view of the body; G. Dorsal view of the body. Scale bars: 5 mm (A – D); 10 mm (E – G). Photographs by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure2" httpUri="https://binary.pensoft.net/fig/1190525">2 C, D</figureCitation>
, 
<figureCitation id="E11FFDE213144D6BBFF3F74580F3CCB3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Line drawings of head scalation of the Colubroelaps adleri sp. nov. holotype (ZMMU Re- 18000, adult female; A – C) and of the C. nguyenvansangi holotype (ZISP 25682, adult female; D – F). Line drawings show the head scales in lateral (A, D), dorsal (B, E), and ventral (C, F) aspects. Abbreviations: ACS = anterior chin shield; AT = anterior temporal; F = frontal; G = gular scale; IL = infralabial; IN = internasal; L = loreal; LPT = lower posterior temporal; M = mental; N = nasal; P = parietal; PCS = posterior chin shield; PF = prefrontal; PO = postocular; PrO = preocular; R = rostral; SL = supralabial; SO = supraocular; UPT = upper posterior temporal. Scale bar equals 5 mm. Line drawings by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure3" httpUri="https://binary.pensoft.net/fig/1190526">3 B</figureCitation>
), in dorsal aspect notably protruding beyond the posterior edge of parietal (Fig. 
<figureCitation id="6EFF2D9E5A9F182CCED9308BEEBD3408" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Line drawings of head scalation of the Colubroelaps adleri sp. nov. holotype (ZMMU Re- 18000, adult female; A – C) and of the C. nguyenvansangi holotype (ZISP 25682, adult female; D – F). Line drawings show the head scales in lateral (A, D), dorsal (B, E), and ventral (C, F) aspects. Abbreviations: ACS = anterior chin shield; AT = anterior temporal; F = frontal; G = gular scale; IL = infralabial; IN = internasal; L = loreal; LPT = lower posterior temporal; M = mental; N = nasal; P = parietal; PCS = posterior chin shield; PF = prefrontal; PO = postocular; PrO = preocular; R = rostral; SL = supralabial; SO = supraocular; UPT = upper posterior temporal. Scale bar equals 5 mm. Line drawings by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure3" httpUri="https://binary.pensoft.net/fig/1190526">3 B</figureCitation>
). Dorsal scales in 14–14 – 14 rows. Dorsal scales rhomboid, tile-shaped, all smooth, and of the same size (Fig. 
<figureCitation id="001CA501E28B0791AABE6CD59413B301" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The holotype of Colubroelaps adleri sp. nov. in life (ZMMU Re- 18000, adult female) from Hon Tre Island, Khanh Hoa Province, Vietnam. A. Dorsal view of the head; B. Ventral view of the head; C. Lateral view of the right side of the head; D. Lateral view of the left side of the head; E. Ventral view of the tail; F. Ventral view of the body; G. Dorsal view of the body. Scale bars: 5 mm (A – D); 10 mm (E – G). Photographs by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure2" httpUri="https://binary.pensoft.net/fig/1190525">2 G</figureCitation>
). Ventrals 234; cloacal plate divided (Fig. 
<figureCitation id="CB51923E70FA43535C5FD5D175CAD6D5" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The holotype of Colubroelaps adleri sp. nov. in life (ZMMU Re- 18000, adult female) from Hon Tre Island, Khanh Hoa Province, Vietnam. A. Dorsal view of the head; B. Ventral view of the head; C. Lateral view of the right side of the head; D. Lateral view of the left side of the head; E. Ventral view of the tail; F. Ventral view of the body; G. Dorsal view of the body. Scale bars: 5 mm (A – D); 10 mm (E – G). Photographs by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure2" httpUri="https://binary.pensoft.net/fig/1190525">2 E</figureCitation>
); 30 subcaudals, all divided (Fig. 
<figureCitation id="205FF5A15005CCA6216436E672F4A897" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The holotype of Colubroelaps adleri sp. nov. in life (ZMMU Re- 18000, adult female) from Hon Tre Island, Khanh Hoa Province, Vietnam. A. Dorsal view of the head; B. Ventral view of the head; C. Lateral view of the right side of the head; D. Lateral view of the left side of the head; E. Ventral view of the tail; F. Ventral view of the body; G. Dorsal view of the body. Scale bars: 5 mm (A – D); 10 mm (E – G). Photographs by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure2" httpUri="https://binary.pensoft.net/fig/1190525">2 E, F</figureCitation>
).
</paragraph>
<caption id="B2F858F89A0DCEFC5FC4EC046AFB2D69" ID-DOI="10.3897/herpetozoa.37.e137809.figure2" ID-arpha="810A7A31-BA9B-58AB-B8FE-115E23766DF9" httpUri="https://binary.pensoft.net/fig/1190525" startId="F2">
<paragraph id="36731B1AD0A41199F2AA6772E515B653">
<label id="E3C1CD8493FF6F31F669DC1C3D85460A">Figure 2.</label>
</paragraph>
<paragraph id="A709F56547FECF495F6C9AD39F85ADA4">
The holotype of 
<taxonomicName id="367CE04E4200DDCF4D530BA63DFCB48E" authorityName="Poyarkov &amp; Bragin &amp; Nguyen" authorityYear="2024" class="Squamata" family="Colubridae" genus="Colubroelaps" kingdom="Animalia" phylum="Chordata" rank="species" species="adleri" status="sp. nov.">
<emphasis id="3612FB9CB4A1D6742740162098961A1A" italics="true">Colubroelaps adleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6606E546DDC8CB150A969FB214B486FC" rank="species">sp. nov.</taxonomicNameLabel>
in life (ZMMU Re- 18000, adult female) from Hon Tre Island, Khanh Hoa Province, Vietnam. 
<emphasis id="C145CFB077D27585BF296D560D22FCCC" bold="true">A.</emphasis>
Dorsal view of the head; 
<emphasis id="A3E970EF4F3E79C40A22B2C783875CF5" bold="true">B.</emphasis>
Ventral view of the head; 
<emphasis id="208DFA479343C297844A4C5648B3E242" bold="true">C.</emphasis>
Lateral view of the right side of the head; 
<emphasis id="7474F8A05CA0A09708D4191ADE76E84D" bold="true">D.</emphasis>
Lateral view of the left side of the head; 
<emphasis id="6DFB1A1850DCED7369A054C337893D3E" bold="true">E.</emphasis>
Ventral view of the tail; 
<emphasis id="FF73F442FFFF0F85DB599BCEFD06ABE4" bold="true">F.</emphasis>
Ventral view of the body; 
<emphasis id="ED65A71B84C22A88E8B394A34FE37F67" bold="true">G.</emphasis>
Dorsal view of the body. Scale bars: 5 mm (
<emphasis id="D151D3816BDF7995787E8196728D7877" bold="true">A – D</emphasis>
); 10 mm (
<emphasis id="52670FB5274513FEA85C83C0A5016420" bold="true">E – G</emphasis>
). Photographs by A. M. Bragin.
</paragraph>
</caption>
<caption id="1C790E804C65251042FFA775640257CC" ID-DOI="10.3897/herpetozoa.37.e137809.figure3" ID-arpha="2EEF69AB-7EB4-5B86-8D20-70E3BDC5D5CB" httpUri="https://binary.pensoft.net/fig/1190526" startId="F3">
<paragraph id="6CCDC6B3F6A3526C73BE34BF037455DF">
<label id="CDB845C5AD5D3E967E423AF02086F9B6">Figure 3.</label>
</paragraph>
<paragraph id="21723F754AC4A0DE51FCF8F4DC5C311F">
Line drawings of head scalation of the 
<taxonomicName id="1D3B8ACDAEDC5B3E3E55C4B3ED5AB5C5" authorityName="Poyarkov &amp; Bragin &amp; Nguyen" authorityYear="2024" class="Squamata" family="Colubridae" genus="Colubroelaps" kingdom="Animalia" phylum="Chordata" rank="species" species="adleri" status="sp. nov.">
<emphasis id="161C3A33914DAB050F08FCD170F94108" italics="true">Colubroelaps adleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="540D344263EC0CE7F1DA0CA7CE9994F3" rank="species">sp. nov.</taxonomicNameLabel>
holotype (ZMMU Re- 18000, adult female; 
<emphasis id="027B2FF0F563CA4304CC0920AD6084D1" bold="true">A – C</emphasis>
) and of the 
<taxonomicName id="CE16C27714824F56DB6AA7BC4243D500" authorityName="Orlov, Kharin, Ananjeva, Nguyen &amp; Nguyen" authorityYear="2009" class="Squamata" family="Colubridae" genus="Colubroelaps" kingdom="Animalia" phylum="Chordata" rank="species" species="nguyenvansangi">
<emphasis id="70963318CADA4A5CC0CB7039C46338CD" italics="true">C. nguyenvansangi</emphasis>
</taxonomicName>
holotype (ZISP 25682, adult female; 
<emphasis id="219C5CC1995D14F49CB0953A46310A32" bold="true">D – F</emphasis>
). Line drawings show the head scales in lateral (
<emphasis id="8CFFA5CC77B51D6ADD704F22267F676D" bold="true">A, D</emphasis>
), dorsal (
<emphasis id="5BBC12EA91B5766D292EDC50CAD0C842" bold="true">B, E</emphasis>
), and ventral (
<emphasis id="F1C6778A9B586702877D6EC43ED29F40" bold="true">C, F</emphasis>
) aspects. Abbreviations: ACS = anterior chin shield; 
<abbrev id="ABBRID0EJIAG" xlink_title="number of anterior temporals">AT</abbrev>
= anterior temporal; F = frontal; G = gular scale; 
<abbrev id="ABBRID0ENIAG" xlink_title="number of infralabial scales">IL</abbrev>
= infralabial; IN = internasal; L = loreal; LPT = lower posterior temporal; M = mental; N = nasal; P = parietal; PCS = posterior chin shield; PF = prefrontal; 
<abbrev id="ABBRID0ERIAG" xlink_title="number of postoculars">PO</abbrev>
= postocular; 
<abbrev id="ABBRID0EVIAG" xlink_title="number of preoculars">PrO</abbrev>
= preocular; R = rostral; 
<abbrev id="ABBRID0EZIAG" xlink_title="number of supralabial scales">SL</abbrev>
= supralabial; SO = supraocular; UPT = upper posterior temporal. Scale bar equals 5 mm. Line drawings by A. M. Bragin.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0ECJAG" type="Coloration in life">
<paragraph id="AC48ACE39570B28036A8EE9994D445EC">
<heading id="0A79A30DB5877B754EFD11ED0CB884AE" reason="title">Coloration in life.</heading>
</paragraph>
<paragraph id="83D20E2C2CF07A2E44A4BAF4D9944F7E">
Body glossy but lacking the metal / bluish iridescence (Figs 
<figureCitation id="BE4DE039992A9AE762E55A16F7C22F1A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The holotype of Colubroelaps adleri sp. nov. in life (ZMMU Re- 18000, adult female) from Hon Tre Island, Khanh Hoa Province, Vietnam. A. Dorsal view of the head; B. Ventral view of the head; C. Lateral view of the right side of the head; D. Lateral view of the left side of the head; E. Ventral view of the tail; F. Ventral view of the body; G. Dorsal view of the body. Scale bars: 5 mm (A – D); 10 mm (E – G). Photographs by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure2" httpUri="https://binary.pensoft.net/fig/1190525">2 A – G</figureCitation>
, 
<figureCitation id="15BCCD3C980DE45858E292A7360A33D5" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Two species of the genus Colubroelaps in life. A. Colubroelaps adleri sp. nov., holotype ZMMU Re- 18000, adult female from Dam Bay Research Station, Hon Tre Island, Khanh Hoa Province, Vietnam; B. Colubroelaps nguyenvansangi, ZMMU Re- 15485, adult male from Cat Tien NP., Tan Phu District, Dong Nai Province, Vietnam. Photographs by: A. M. Bragin (A); E. A. Galoyan (B)." figureDoi="10.3897/herpetozoa.37.e137809.figure4" httpUri="https://binary.pensoft.net/fig/1190527">4 A</figureCitation>
). Dorsal background coloration pale brown, with narrow and interrupted dark stripe along spine (Figs 
<figureCitation id="334B146AB24820483730404F4DD4B943" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The holotype of Colubroelaps adleri sp. nov. in life (ZMMU Re- 18000, adult female) from Hon Tre Island, Khanh Hoa Province, Vietnam. A. Dorsal view of the head; B. Ventral view of the head; C. Lateral view of the right side of the head; D. Lateral view of the left side of the head; E. Ventral view of the tail; F. Ventral view of the body; G. Dorsal view of the body. Scale bars: 5 mm (A – D); 10 mm (E – G). Photographs by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure2" httpUri="https://binary.pensoft.net/fig/1190525">2 G</figureCitation>
, 
<figureCitation id="72A424D4BA6739CF85C716A909FC7905" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Two species of the genus Colubroelaps in life. A. Colubroelaps adleri sp. nov., holotype ZMMU Re- 18000, adult female from Dam Bay Research Station, Hon Tre Island, Khanh Hoa Province, Vietnam; B. Colubroelaps nguyenvansangi, ZMMU Re- 15485, adult male from Cat Tien NP., Tan Phu District, Dong Nai Province, Vietnam. Photographs by: A. M. Bragin (A); E. A. Galoyan (B)." figureDoi="10.3897/herpetozoa.37.e137809.figure4" httpUri="https://binary.pensoft.net/fig/1190527">4 A</figureCitation>
), formed by small, ca. 1 scale in size, dark brown to black, diamond-shaped blotches (Fig. 
<figureCitation id="8D15009158C9EEDB84BB251CD112C03C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The holotype of Colubroelaps adleri sp. nov. in life (ZMMU Re- 18000, adult female) from Hon Tre Island, Khanh Hoa Province, Vietnam. A. Dorsal view of the head; B. Ventral view of the head; C. Lateral view of the right side of the head; D. Lateral view of the left side of the head; E. Ventral view of the tail; F. Ventral view of the body; G. Dorsal view of the body. Scale bars: 5 mm (A – D); 10 mm (E – G). Photographs by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure2" httpUri="https://binary.pensoft.net/fig/1190525">2 A</figureCitation>
); body flanks dark gray, nearly black, with indistinct light gray mottling along the scale edges; ventral surface of body and tail off-white with light greenish tint (Fig. 
<figureCitation id="FD0A0DD364F616F9CF29036AFB5B012B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The holotype of Colubroelaps adleri sp. nov. in life (ZMMU Re- 18000, adult female) from Hon Tre Island, Khanh Hoa Province, Vietnam. A. Dorsal view of the head; B. Ventral view of the head; C. Lateral view of the right side of the head; D. Lateral view of the left side of the head; E. Ventral view of the tail; F. Ventral view of the body; G. Dorsal view of the body. Scale bars: 5 mm (A – D); 10 mm (E – G). Photographs by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure2" httpUri="https://binary.pensoft.net/fig/1190525">2 E, F</figureCitation>
). Head black to dark brown with rostral, nasals, prefrontals, preoculars, loreal, the first two supralabials, as well as the anterior part of supraoculars and frontal shield dirty yellowish-brown with dark brown irregular spots (Fig. 
<figureCitation id="28E1E8DE46EE1014D999BBB34F0FB788" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The holotype of Colubroelaps adleri sp. nov. in life (ZMMU Re- 18000, adult female) from Hon Tre Island, Khanh Hoa Province, Vietnam. A. Dorsal view of the head; B. Ventral view of the head; C. Lateral view of the right side of the head; D. Lateral view of the left side of the head; E. Ventral view of the tail; F. Ventral view of the body; G. Dorsal view of the body. Scale bars: 5 mm (A – D); 10 mm (E – G). Photographs by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure2" httpUri="https://binary.pensoft.net/fig/1190525">2 A – D</figureCitation>
). Ventral surfaces of the head white with chocolate-brown blotches (Fig. 
<figureCitation id="E5B7E880B5FEC0829719225591CCAADC" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The holotype of Colubroelaps adleri sp. nov. in life (ZMMU Re- 18000, adult female) from Hon Tre Island, Khanh Hoa Province, Vietnam. A. Dorsal view of the head; B. Ventral view of the head; C. Lateral view of the right side of the head; D. Lateral view of the left side of the head; E. Ventral view of the tail; F. Ventral view of the body; G. Dorsal view of the body. Scale bars: 5 mm (A – D); 10 mm (E – G). Photographs by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure2" httpUri="https://binary.pensoft.net/fig/1190525">2 B</figureCitation>
). After one year in preservative, the background dorsal color faded to beige-gray, and yellowish parts faded to off-white; overall, the pattern of dark markings remained unchanged.
</paragraph>
<caption id="A901C576C594EB9151D80243EF59686E" ID-DOI="10.3897/herpetozoa.37.e137809.figure4" ID-arpha="AC407FDA-D605-5FA9-95EF-32B09A1A0590" httpUri="https://binary.pensoft.net/fig/1190527" startId="F4">
<paragraph id="6405345780E561392506C86F8714DF12">
<label id="26D7C89BD5C67C572CBAE3D1D26F4347">Figure 4.</label>
</paragraph>
<paragraph id="BFC603D54C46FFFF08EC94D3636FEEC1">
Two species of the genus 
<taxonomicName id="B7F3157ED920C0439E54862A2057C76C" authorityName="Orlov, Kharin, Ananjeva, Nguyen &amp; Nguyen" authorityYear="2009" class="Squamata" family="Colubridae" genus="Colubroelaps" kingdom="Animalia" phylum="Chordata" rank="genus">
<emphasis id="CFD8ED619C3B4CA11A3004309706B158" italics="true">Colubroelaps</emphasis>
</taxonomicName>
in life. 
<emphasis id="265A83DCBA4D0CD776494FD81354B4B1" bold="true">A.</emphasis>
<taxonomicName id="9046A8A0AF8DEAC8CBB926D23F67052C" authorityName="Poyarkov &amp; Bragin &amp; Nguyen" authorityYear="2024" class="Squamata" family="Colubridae" genus="Colubroelaps" kingdom="Animalia" phylum="Chordata" rank="species" species="adleri" status="sp. nov.">
<emphasis id="8F635EE9D7612D59238380E89A244188" italics="true">Colubroelaps adleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CDEF11A27063A4C9CFB51A953FDE191C" rank="species">sp. nov.</taxonomicNameLabel>
, holotype ZMMU Re- 18000, adult female from Dam Bay Research Station, Hon Tre Island, Khanh Hoa Province, Vietnam; 
<emphasis id="7323199947969B81F9E79F58E0C845F8" bold="true">B.</emphasis>
<taxonomicName id="4362687FA77D007E11CC9398F73B8CEC" authority=", ZMMU" authorityName="ZMMU" class="Squamata" family="Colubridae" genus="Colubroelaps" kingdom="Animalia" phylum="Chordata" rank="species" species="nguyenvansangi">
<emphasis id="E77D7E98E1CFEF7D47FF1B0E5D009301" italics="true">Colubroelaps nguyenvansangi</emphasis>
, ZMMU
</taxonomicName>
Re- 15485, adult male from Cat Tien 
<abbrev id="ABBRID0ERLAG" xlink_title="National Park">NP</abbrev>
., Tan Phu District, Dong Nai Province, Vietnam. Photographs by: A. M. Bragin (
<emphasis id="82A7D56ADAF44A1F52CF22493D64DED0" bold="true">A</emphasis>
); E. A. Galoyan (
<emphasis id="7718008413D8729B6EB5827AD215720B" bold="true">B</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0E5LAG" type="comparisons">
<paragraph id="71925603CD6D6D23B9C3F1CB11FB6814">
<heading id="77C8F207385AB3D28D2013771DE12859" reason="title">Comparisons.</heading>
</paragraph>
<paragraph id="16BCE5731A229A15CE4E9318CAF2E73C">
The main differences between the new species and 
<taxonomicName id="AAA1D6A80090D38784F442937DF91910" class="Squamata" family="Colubridae" genus="Colubroelaps" kingdom="Animalia" phylum="Chordata" rank="species" species="nguyenvansangi">
<emphasis id="EB5C109987F43CBFD7CEEF5EFFB2D12B" italics="true">C. nguyenvansangi</emphasis>
</taxonomicName>
are summarized in Table 
<tableCitation id="CB283E6FC407F799D4F1E5435F1DB703" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological data on the type series of Colubroelaps adleri sp. nov. and C. nguyenvansangi; diagnostic differences from the new species are marked in bold. Notes: for abbreviations, see the Materials and methods section; “ * ”: data on the holotype of C. nguyenvansangi as reported by Orlov et al. (2009), when different from our observations." httpUri="http://table.plazi.org/id/8E5F0BED38B5CEE14412962DA7D57026" tableUuid="8E5F0BED38B5CEE14412962DA7D57026">1</tableCitation>
. 
<taxonomicName id="9625685C64D415E7FA17DA67984C9AAF" authorityName="Poyarkov &amp; Bragin &amp; Nguyen" authorityYear="2024" class="Squamata" family="Colubridae" genus="Colubroelaps" kingdom="Animalia" phylum="Chordata" rank="species" species="adleri" status="sp. nov.">
<emphasis id="0413B9298D55935610B0E1F1B5B6C89D" italics="true">Colubroelaps adleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7A94B49118AD7D4F6015E76AFEE6AE7A" rank="species">sp. nov.</taxonomicNameLabel>
can be easily distinguished from 
<taxonomicName id="78904DFDCBA45808BE0D39E4DB84D908" class="Squamata" family="Colubridae" genus="Colubroelaps" kingdom="Animalia" phylum="Chordata" rank="species" species="nguyenvansangi">
<emphasis id="53B8845C25D5EC188CB50511B6D2F900" italics="true">C. nguyenvansangi</emphasis>
</taxonomicName>
by having notably shorter tail (
<abbrev id="ABBRID0EJNAG" xlink_title="tail length">TaL</abbrev>
/ 
<abbrev id="ABBRID0ENNAG" xlink_title="total length">TL</abbrev>
<quantity id="E2DC2D66A95E2F30059CCE857EF7F16C" metricMagnitude="-3" metricUnit="m" metricValue="2.54" unit="in" value="0.1">0.10 in</quantity>
a single female vs. 
<quantity id="E1EE5D681219990123CE1CA98ECE9CC8" metricMagnitude="-3" metricUnit="m" metricValue="5.334" unit="in" value="0.21">0.21 in</quantity>
females and 
<quantity id="A77499C8A08FCF0E5EE225D4845ABAA0" metricMagnitude="-3" metricUnit="m" metricValue="5.587999999999999" metricValueMax="5.842" metricValueMin="5.334" unit="in" value="0.22" valueMax="0.23" valueMin="0.21">0.21–0.23 in</quantity>
males; however, this character should be taken cautiously: the tail of 
<taxonomicName id="3EC2B5E27482395AD460B7F3CF67EF19" authorityName="Poyarkov &amp; Bragin &amp; Nguyen" authorityYear="2024" class="Squamata" family="Colubridae" genus="Colubroelaps" kingdom="Animalia" phylum="Chordata" rank="species" species="adleri" status="sp. nov.">
<emphasis id="7F5AAB429A1C7673581DA5D0072C484C" italics="true">Colubroelaps adleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EC4CA84D2641175181545F342721E367" rank="species">sp. nov.</taxonomicNameLabel>
<typeStatus id="462836A57D161DF5D4ABF3DF66301FE0" type="holotype">holotype</typeStatus>
may be incomplete; cases of caudal autotomy were earlier reported for the members of 
<taxonomicName id="0E49915EB64518A1061F79E6D3EBCFC7" authorityName="Dunn" authorityYear="1928" family="Sibynophiidae" higherTaxonomySource="GBIF" kingdom="Animalia" phylum="Chordata" rank="family">Sibynophiidae</taxonomicName>
and likely may also occur in 
<taxonomicName id="7008CA36C0D1269181D6C5ADF989DC00" authorityName="Orlov, Kharin, Ananjeva, Nguyen &amp; Nguyen" authorityYear="2009" class="Squamata" family="Colubridae" genus="Colubroelaps" kingdom="Animalia" phylum="Chordata" rank="genus">
<emphasis id="1CEBAB751283FAA2D50C33EC399B1110" italics="true">Colubroelaps</emphasis>
</taxonomicName>
; see 
<bibRefCitation id="C36085E015EF079C7F3F49BFFCDFAF7D" author="Mendelson JR III" journalOrPublisher="The University of Texas at Arlington, Unpublished Master Thesis" refId="B14" refString="Mendelson JR III (1991) Tail breakage in Coniophanes fissidens and other tropical colubrid snakes. The University of Texas at Arlington, Unpublished Master Thesis, 120 pp." title="Tail breakage in Coniophanes fissidens and other tropical colubrid snakes." year="1991">Mendelson, 1991</bibRefCitation>
), lower number of body scale rows (DSR 14–14 – 14 vs. 16–16 – 16); lower number of ventrals (
<abbrev id="ABBRID0EIOAG" xlink_title="number of ventral scales">VEN</abbrev>
<quantity id="4F5B6884E095E43F8B3B4E698BF37D6A" metricMagnitude="0" metricUnit="m" metricValue="5.9436" unit="in" value="234.0">234 in</quantity>
a single female vs. 
<quantity id="0F849F10F4BAE45CB20A85F7B87E1DA7" metricMagnitude="0" metricUnit="m" metricValue="6.7818" unit="in" value="267.0">267 in</quantity>
female, 
<quantity id="4AB3788ED44A173C7E1479DAAA61CED9" metricMagnitude="0" metricUnit="m" metricValue="7.2898" metricValueMax="7.4168" metricValueMin="7.1628" unit="in" value="287.0" valueMax="292.0" valueMin="282.0">282–292 in</quantity>
males); much lower number of subcaudals (
<abbrev id="ABBRID0EMOAG" xlink_title="number of subcaudal scales, not including the terminal pointed scute">SC</abbrev>
<quantity id="7216FD8854E70FE32F9F2449DF4B7460" metricMagnitude="-1" metricUnit="m" metricValue="7.62" unit="in" value="30.0">30 in</quantity>
a single female vs. 
<quantity id="7E55E2191F1BBBF444A3D4980E0B3934" metricMagnitude="0" metricUnit="m" metricValue="2.0574" unit="in" value="81.0">81 in</quantity>
female, 
<quantity id="F46E90635FFF89848B5A5338CB3DCBE2" metricMagnitude="0" metricUnit="m" metricValue="2.1971" metricValueMax="2.2098" metricValueMin="2.1844" unit="in" value="86.5" valueMax="87.0" valueMin="86.0">86–87 in</quantity>
males). Furthermore, the new species can be easily diagnosed from 
<taxonomicName id="027B452245C95308F4E3488988E36606" class="Squamata" family="Colubridae" genus="Colubroelaps" kingdom="Animalia" phylum="Chordata" rank="species" species="nguyenvansangi">
<emphasis id="CE1D3C16ED903420C6D6071E709903D7" italics="true">C. nguyenvansangi</emphasis>
</taxonomicName>
by having three gular scales between the chin shields and the first preventral (vs. two) (Fig. 
<figureCitation id="F7F0045C20B3E916D9C1449768A7F07E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Line drawings of head scalation of the Colubroelaps adleri sp. nov. holotype (ZMMU Re- 18000, adult female; A – C) and of the C. nguyenvansangi holotype (ZISP 25682, adult female; D – F). Line drawings show the head scales in lateral (A, D), dorsal (B, E), and ventral (C, F) aspects. Abbreviations: ACS = anterior chin shield; AT = anterior temporal; F = frontal; G = gular scale; IL = infralabial; IN = internasal; L = loreal; LPT = lower posterior temporal; M = mental; N = nasal; P = parietal; PCS = posterior chin shield; PF = prefrontal; PO = postocular; PrO = preocular; R = rostral; SL = supralabial; SO = supraocular; UPT = upper posterior temporal. Scale bar equals 5 mm. Line drawings by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure3" httpUri="https://binary.pensoft.net/fig/1190526">3 C, F</figureCitation>
); by fourth infralabial in contact with parietal and anterior temporal, separating fifth infralabial from parietal and postocular (vs. fifth infralabial in contact with parietal and postocular) (Fig. 
<figureCitation id="B78558195B0A83F0ACCC27F2601C5DC6" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Line drawings of head scalation of the Colubroelaps adleri sp. nov. holotype (ZMMU Re- 18000, adult female; A – C) and of the C. nguyenvansangi holotype (ZISP 25682, adult female; D – F). Line drawings show the head scales in lateral (A, D), dorsal (B, E), and ventral (C, F) aspects. Abbreviations: ACS = anterior chin shield; AT = anterior temporal; F = frontal; G = gular scale; IL = infralabial; IN = internasal; L = loreal; LPT = lower posterior temporal; M = mental; N = nasal; P = parietal; PCS = posterior chin shield; PF = prefrontal; PO = postocular; PrO = preocular; R = rostral; SL = supralabial; SO = supraocular; UPT = upper posterior temporal. Scale bar equals 5 mm. Line drawings by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure3" httpUri="https://binary.pensoft.net/fig/1190526">3 A, D</figureCitation>
); by upper posterior temporal being elongated and much narrower than lower one, noticeably protruding beyond the level of the posterior edge of parietals (vs. upper posterior temporal small, as long as lower posterior temporal, not protruding beyond the level of the posterior edge of parietals) (Fig. 
<figureCitation id="12EAA85DEA2C6452EA627B2F4C7FDC6D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Line drawings of head scalation of the Colubroelaps adleri sp. nov. holotype (ZMMU Re- 18000, adult female; A – C) and of the C. nguyenvansangi holotype (ZISP 25682, adult female; D – F). Line drawings show the head scales in lateral (A, D), dorsal (B, E), and ventral (C, F) aspects. Abbreviations: ACS = anterior chin shield; AT = anterior temporal; F = frontal; G = gular scale; IL = infralabial; IN = internasal; L = loreal; LPT = lower posterior temporal; M = mental; N = nasal; P = parietal; PCS = posterior chin shield; PF = prefrontal; PO = postocular; PrO = preocular; R = rostral; SL = supralabial; SO = supraocular; UPT = upper posterior temporal. Scale bar equals 5 mm. Line drawings by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure3" httpUri="https://binary.pensoft.net/fig/1190526">3 A – E</figureCitation>
). 
<taxonomicName id="1513B0EB4E037170793DDD15CFA9917F" authorityName="Poyarkov &amp; Bragin &amp; Nguyen" authorityYear="2024" class="Squamata" family="Colubridae" genus="Colubroelaps" kingdom="Animalia" phylum="Chordata" rank="species" species="adleri" status="sp. nov.">
<emphasis id="28BFE7BF80E4278B5DAAB9975C0791CC" italics="true">Colubroelaps adleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="39B54B80E763B0E17E18F6FAF3F39E2F" rank="species">sp. nov.</taxonomicNameLabel>
can be further diagnosed from 
<taxonomicName id="278ACD96B7400E700CBAC7405DD19EFE" class="Squamata" family="Colubridae" genus="Colubroelaps" kingdom="Animalia" phylum="Chordata" rank="species" species="nguyenvansangi">
<emphasis id="F1B0845458932AB710B3C7B7CBA9BFC5" italics="true">C. nguyenvansangi</emphasis>
</taxonomicName>
by having pale brown dorsum with narrow and interrupted dark stripe along spine (vs. reddish-brown dorsum with narrow and continuous dark stripe along spine) (Fig. 
<figureCitation id="F5AC2E648A7C53E56BD3A38A6D425BBD" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Two species of the genus Colubroelaps in life. A. Colubroelaps adleri sp. nov., holotype ZMMU Re- 18000, adult female from Dam Bay Research Station, Hon Tre Island, Khanh Hoa Province, Vietnam; B. Colubroelaps nguyenvansangi, ZMMU Re- 15485, adult male from Cat Tien NP., Tan Phu District, Dong Nai Province, Vietnam. Photographs by: A. M. Bragin (A); E. A. Galoyan (B)." figureDoi="10.3897/herpetozoa.37.e137809.figure4" httpUri="https://binary.pensoft.net/fig/1190527">4 A, B</figureCitation>
); by lacking the metallic iridescence on dorsum and body flanks (vs. body with bluish metallic iridescence) (Fig. 
<figureCitation id="89B5550A35653651BCD7F4520B6D6AF1" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Two species of the genus Colubroelaps in life. A. Colubroelaps adleri sp. nov., holotype ZMMU Re- 18000, adult female from Dam Bay Research Station, Hon Tre Island, Khanh Hoa Province, Vietnam; B. Colubroelaps nguyenvansangi, ZMMU Re- 15485, adult male from Cat Tien NP., Tan Phu District, Dong Nai Province, Vietnam. Photographs by: A. M. Bragin (A); E. A. Galoyan (B)." figureDoi="10.3897/herpetozoa.37.e137809.figure4" httpUri="https://binary.pensoft.net/fig/1190527">4 A, B</figureCitation>
); and by having dirty yellowish-brown anterior part of head with irregular dark brown blotches (vs. uniform light yellow to white blotch on anterior part of head lacking dark markings) (Fig. 
<figureCitation id="36D750EE834B27B4062CCBAB02A72C83" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Two species of the genus Colubroelaps in life. A. Colubroelaps adleri sp. nov., holotype ZMMU Re- 18000, adult female from Dam Bay Research Station, Hon Tre Island, Khanh Hoa Province, Vietnam; B. Colubroelaps nguyenvansangi, ZMMU Re- 15485, adult male from Cat Tien NP., Tan Phu District, Dong Nai Province, Vietnam. Photographs by: A. M. Bragin (A); E. A. Galoyan (B)." figureDoi="10.3897/herpetozoa.37.e137809.figure4" httpUri="https://binary.pensoft.net/fig/1190527">4 A, B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJQAG" type="etymology">
<paragraph id="C22F1003FCF767D28A6F48703FC11AB6">
<heading id="5F6C8EDB1B9F0BE7968170A87E4678E0" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="C87F9E5D0DA40701FD1AAEC75A7BA261">
The species epithet ‘ 
<emphasis id="E495E8EE452CF6F4F52F3363ED6A03A3" italics="true">adleri</emphasis>
’ is a patronymic adjective in genitive singular. We name the new species in honor of Dr. Kraig Adler, Professor Emeritus at Cornell University (
<collectingRegion id="40CF854246219F4CA4D47BAA07CCD156" country="United States of America" name="New York">New York</collectingRegion>
, 
<collectingCountry id="4340E28D3FEAABF8A23CCC27540AB2B4" name="United States of America">USA</collectingCountry>
), in recognition of his outstanding support to the international herpetological community as well as his remarkable scientific contribution to Asian herpetology. We suggest the following common names for the new species: “ 
<emphasis id="95EEF8F65832F6E0851EDABF339083DD" italics="true">Adler’s lace snake</emphasis>
” (in English), “ 
<emphasis id="9C2ED5A1868071AF838585769941EC13" italics="true">Shnurkovaya zmeya Adlera</emphasis>
” (Шнурковая змея Адлера, in Russian), and “ 
<emphasis id="FA1C98C2BE454369E3E190F262248F8E" italics="true">R ắn h ổ nư ớc Át-Lơ</emphasis>
” (in Vietnamese).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E6QAG" type="distribution">
<paragraph id="6BEB9AD934EFA7E788CB690FA4F1BFAE">
<heading id="E392F7BB4D924016E280D6D9AE16839A" reason="title">Distribution and natural history notes.</heading>
</paragraph>
<paragraph id="411F064A51D1999721FD4450F6ECACD8">
Currently, 
<taxonomicName id="038A369A1D7B28B230BBE1635F2FB54E" authorityName="Poyarkov &amp; Bragin &amp; Nguyen" authorityYear="2024" class="Squamata" family="Colubridae" genus="Colubroelaps" kingdom="Animalia" phylum="Chordata" rank="species" species="adleri" status="sp. nov.">
<emphasis id="11745AE99A2FB1C8CEB1BBA541C10335" italics="true">Colubroelaps adleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C9D5A00F505A9EBA5AA44383F9957DC0" rank="species">sp. nov.</taxonomicNameLabel>
is known only from a single locality in secondary dry maritime evergreen forest on Hon Tre Island, 
<collectingRegion id="DFF1E6E84436FA02A3A841911B6E5F81" country="Vietnam" name="Khanh Hoa">Khanh Hoa Province</collectingRegion>
, South Central Coastal Region of 
<collectingCountry id="EADAF1FB53CC29EEA10F0A30677168C4" name="Vietnam">Vietnam</collectingCountry>
(Figs 
<figureCitation id="0585A1D07BAAC289F7C5C1833536ED87" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of the genus Colubroelaps in Vietnam. Localities: Colubroelaps adleri sp. nov. (red): (1) Hon Tre Island within Nha Trang Bay, Khanh Hoa Province (type locality); C. nguyenvansangi (green): (2) Loc Bac, Lam Dong Province (type locality); (3) Di Linh, Lam Dong Province; (4) Bu Gia Map NP, Binh Phuoc Province; (5) Bu Dang, Binh Phuoc Province; (6) Tuy Duc, Dak Nong Province; (7) Cat Tien NP, Dong Nai Province; (8) Buon Ma Thuot, Dak Lak Province; (9) Krong Buk, Dak Lak Province." figureDoi="10.3897/herpetozoa.37.e137809.figure1" httpUri="https://binary.pensoft.net/fig/1190524">1</figureCitation>
, 
<figureCitation id="EB171039CE72C270A01D3E1CBB2AFB50" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitat of Colubroelaps adleri sp. nov. A. Macrohabitat; B. Microhabitat of the new species in Dam Bay Research Station, Hon Tre Island, Nha Trang, Khanh Hoa Province, Vietnam. Photographs by A. V. Alexandrova." figureDoi="10.3897/herpetozoa.37.e137809.figure5" httpUri="https://binary.pensoft.net/fig/1190528">5</figureCitation>
). The new species is also expected to inhabit other islands of the Nha Trang Bay, though they are much smaller than Hon Tre, and on some of them, forest vegetation has been greatly destroyed. The only known specimen of 
<taxonomicName id="EDEA67D58E0586F5EC0FFFADAD0864DF" authorityName="Poyarkov &amp; Bragin &amp; Nguyen" authorityYear="2024" class="Squamata" family="Colubridae" genus="Colubroelaps" kingdom="Animalia" phylum="Chordata" rank="species" species="adleri" status="sp. nov.">
<emphasis id="02E014B694DCACC228144EBF98D020E6" italics="true">Colubroelaps adleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4E7BDDB355BA9DA4AC5898D328080532" rank="species">sp. nov.</taxonomicNameLabel>
was collected during the daytime (14 h 00) while crossing the road. The individual was collected near a garbage dump at the Dam Bay Research Station at 
<quantity id="73B2C72F478E7E3CD7CD93F73AA28914" metricMagnitude="1" metricUnit="m" metricValue="3.0" unit="m" value="30.0">30 m</quantity>
a. s. l. elevation (Fig. 
<figureCitation id="9A71FDB4591F76CB82038921E99E9DBC" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitat of Colubroelaps adleri sp. nov. A. Macrohabitat; B. Microhabitat of the new species in Dam Bay Research Station, Hon Tre Island, Nha Trang, Khanh Hoa Province, Vietnam. Photographs by A. V. Alexandrova." figureDoi="10.3897/herpetozoa.37.e137809.figure5" httpUri="https://binary.pensoft.net/fig/1190528">5 A</figureCitation>
), ca. 
<quantity id="35337AB9F0051FB2C38FBF2366F7D6CE" metricMagnitude="1" metricUnit="m" metricValue="1.0" unit="m" value="10.0">10 m</quantity>
from a dry maritime mixed low evergreen forest. The forest near the type locality is dominated by 
<taxonomicName id="79D08FC03D050FC59533283FCB30C14D" authority="Hance" authorityName="Hance" class="Magnoliopsida" family="Anacardiaceae" genus="Buchanania" kingdom="Plantae" order="Sapindales" phylum="Tracheophyta" rank="species" species="reticulata">
<emphasis id="5D28ADC71C2AFAEB66F66D4EE90050C5" italics="true">Buchanania reticulata</emphasis>
Hance
</taxonomicName>
, 
<taxonomicName id="4E4AB944C83A4CB82A7628564E5F2CE3" authority="(Roxb.) Burtt. &amp; Hill" authorityName="Burtt. &amp; Hill" baseAuthorityName="Roxb." class="Magnoliopsida" family="Anacardiaceae" genus="Choerospondias" kingdom="Plantae" order="Sapindales" phylum="Tracheophyta" rank="species" species="axillaris">
<emphasis id="91ADC3DD28539A0605072D5E4172405D" italics="true">Choerospondias axillaris</emphasis>
(Roxb.) Burtt. &amp; Hill
</taxonomicName>
, 
<taxonomicName id="150D85B12B11992655C2C476E29DBA36" authority="(Evrard &amp; Tardieu) Ph" authorityName="Ph" baseAuthorityName="Evrard &amp; Tardieu" class="Magnoliopsida" family="Anacardiaceae" genus="Pentaspadon" kingdom="Plantae" order="Sapindales" phylum="Tracheophyta" rank="species" species="annamense">
<emphasis id="CD0E90E416740A56EFAC3F4CE7EDE01E" italics="true">Pentaspadon annamense</emphasis>
(Evrard &amp; Tardieu) Ph
</taxonomicName>
ạmh., 
<taxonomicName id="ED3EE083B8E02660C6355743406F2CED" class="Magnoliopsida" family="Anacardiaceae" genus="Spondias" kingdom="Plantae" order="Sapindales" phylum="Tracheophyta" rank="species" species="pinnata">
<emphasis id="7B1B9479067F5D7774C63CB304CB4C37" italics="true">Spondias pinnata</emphasis>
</taxonomicName>
(L. f.) Kurz, and 
<taxonomicName id="DEDBB43D2DB92D6337DA70A37884DCA8" class="Magnoliopsida" family="Fabaceae" genus="Ormosia" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="5EF78DFEBE3D26EF93FB94549DE5F586" italics="true">Ormosia</emphasis>
sp.
</taxonomicName>
, including occasional trees of 
<taxonomicName id="57AE9DB216A174FD5B15662A773CA062" authority="Teijsm. ex Miq." authorityName="Miq." class="Magnoliopsida" family="Fabaceae" genus="Sindora" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="species" species="siamensis">
<emphasis id="52F4286DF61EDC9B19439A22576E985A" italics="true">Sindora siamensis</emphasis>
Teijsm. ex Miq.
</taxonomicName>
, 
<taxonomicName id="7E3126CD41E76A6B96E80B5A8ABED2D5" authority="(S. Vidal) Corner" authorityName="Corner" baseAuthorityName="S. Vidal" class="Magnoliopsida" family="Moraceae" genus="Streblus" kingdom="Plantae" order="Rosales" phylum="Tracheophyta" rank="species" species="ilicifolius">
<emphasis id="F2C37FC597F49F8CC31A5A3FE9ADBC9A" italics="true">Streblus ilicifolius</emphasis>
(S. Vidal) Corner
</taxonomicName>
, and 
<taxonomicName id="96DC58D1127FE1FD5D57A8F4226D0812" authority="Gagnep" authorityName="Gagnep" class="Magnoliopsida" family="Pentaphylacaceae" genus="Eurya" kingdom="Plantae" order="Ericales" phylum="Tracheophyta" rank="species" species="turfosa">
<emphasis id="07CD9FC035C500A92F5376553070507C" italics="true">Eurya turfosa</emphasis>
Gagnep
</taxonomicName>
, and with an undergrowth formed primarily by 
<taxonomicName id="1FBFB23B0411940D9DB6A326BB73060A" class="Squamata" family="Teiidae" genus="Dracaena" kingdom="Animalia" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B363A69440C26BA7F5105D0DD823BE5B" italics="true">Dracaena</emphasis>
sp.
</taxonomicName>
and with occasional specimens of 
<taxonomicName id="CE3915CBCD80905270EBEAA1CD3C0EFC" authority="Miq." authorityName="Miq." class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" phylum="Tracheophyta" rank="species" species="rumphii">
<emphasis id="4EE5A912E9977A0045A5F6F3EB0655EC" italics="true">Cycas rumphii</emphasis>
Miq.
</taxonomicName>
(Fig. 
<figureCitation id="03B65B4F98412E1EE7AA61AAD846B1DD" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitat of Colubroelaps adleri sp. nov. A. Macrohabitat; B. Microhabitat of the new species in Dam Bay Research Station, Hon Tre Island, Nha Trang, Khanh Hoa Province, Vietnam. Photographs by A. V. Alexandrova." figureDoi="10.3897/herpetozoa.37.e137809.figure5" httpUri="https://binary.pensoft.net/fig/1190528">5 B</figureCitation>
) (plant identification — A. N. Kuznetsov, pers. comm.). Other species of snakes recorded in sympatry with the new species at the type locality included 
<taxonomicName id="34A3818D46544B809D67D0CA6FAFB481" authority="(Blanford, 1878)" baseAuthorityName="Blanford" baseAuthorityYear="1878" class="Squamata" family="Colubridae" genus="Lycodon" kingdom="Animalia" phylum="Chordata" rank="species" species="davisonii">
<emphasis id="AE14697CDD3BA06C203A98EC633D3D4A" italics="true">Lycodon davisonii</emphasis>
(Blanford, 1878)
</taxonomicName>
, 
<taxonomicName id="8F088645F4F89CEFB90EDD16499256B6" authority="(Boie, 1827)" baseAuthorityName="Boie" baseAuthorityYear="1827" class="Squamata" family="Colubridae" genus="Lycodon" kingdom="Animalia" phylum="Chordata" rank="species" species="capucinus">
<emphasis id="87D1C2574691E0849D5A4FDA3CDB37BA" italics="true">L. capucinus</emphasis>
(Boie, 1827)
</taxonomicName>
, 
<taxonomicName id="CC1D579D2B6B1ABF2EF0661A680F4346" authority="(Schlegel, 1837)" baseAuthorityName="Schlegel" baseAuthorityYear="1837" class="Squamata" family="Colubridae" genus="Ptyas" kingdom="Animalia" phylum="Chordata" rank="species" species="korros">
<emphasis id="B4F712AEE1542A79EA78EF7615F3E3AF" italics="true">Ptyas korros</emphasis>
(Schlegel, 1837)
</taxonomicName>
, 
<taxonomicName id="7DAEB99FAB8EBE6D950708AACE0FE0F8" authority="(Cantor, 1836)" baseAuthorityName="Cantor" baseAuthorityYear="1836" class="Squamata" family="Elapidae" genus="Ophiophagus" kingdom="Animalia" phylum="Chordata" rank="species" species="hannah">
<emphasis id="90A0607F01E60B65AE5870017C33CC68" italics="true">Ophiophagus hannah</emphasis>
(Cantor, 1836)
</taxonomicName>
, and 
<taxonomicName id="074F4E4AE2838112DF1FF0C029C289F5" authority="(Gray, 1842)" baseAuthorityName="Gray" baseAuthorityYear="1842" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" phylum="Chordata" rank="species" species="albolabris">
<emphasis id="C4FB6108C751FEA456CA44B5939A8860" italics="true">Trimeresurus albolabris</emphasis>
(Gray, 1842)
</taxonomicName>
.
</paragraph>
<caption id="209448C2BA00A75312A95FD404A8386D" ID-DOI="10.3897/herpetozoa.37.e137809.figure5" ID-arpha="9B88731E-D2CD-50B7-9D49-4CC0E7D9DAFC" httpUri="https://binary.pensoft.net/fig/1190528" startId="F5">
<paragraph id="5BA480B6F6452FE497B49FADAD214CC6">
<label id="A8CDF160970D6CE00A1F42F3E514264A">Figure 5.</label>
</paragraph>
<paragraph id="85D933724F88B7038BDB1ECCE95D70AB">
Habitat of 
<taxonomicName id="D1026D497555826A89FE5EAE4745BB8C" authorityName="Poyarkov &amp; Bragin &amp; Nguyen" authorityYear="2024" class="Squamata" family="Colubridae" genus="Colubroelaps" kingdom="Animalia" phylum="Chordata" rank="species" species="adleri" status="sp. nov.">
<emphasis id="969FD77DDF6AD67AAFF91FEDB88E18D0" italics="true">Colubroelaps adleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8CFC9295DAD18E322B9E2B127B32901A" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="67A33CC38683DF039501D001B42C69E8" bold="true">A.</emphasis>
Macrohabitat; 
<emphasis id="4DBCF1973C04C00AF8401FB91C4EF4A8" bold="true">B.</emphasis>
Microhabitat of the new species in Dam Bay Research Station, Hon Tre Island, Nha Trang, Khanh Hoa Province, Vietnam. Photographs by A. V. Alexandrova.
</paragraph>
</caption>
<paragraph id="7A13770A38755C8E84BB4EB4EE617536">
A parasitic invasion of the Acanthocephala (Kölr.) worm was found in the subcutaneous cavity between the skin and the body muscles in the posterior third of the specimen length on its dorsal side (Figs 
<figureCitation id="D9403BEB4C642635A1D2F3B715D5CAE2" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The holotype of Colubroelaps adleri sp. nov. in life (ZMMU Re- 18000, adult female) from Hon Tre Island, Khanh Hoa Province, Vietnam. A. Dorsal view of the head; B. Ventral view of the head; C. Lateral view of the right side of the head; D. Lateral view of the left side of the head; E. Ventral view of the tail; F. Ventral view of the body; G. Dorsal view of the body. Scale bars: 5 mm (A – D); 10 mm (E – G). Photographs by A. M. Bragin." figureDoi="10.3897/herpetozoa.37.e137809.figure2" httpUri="https://binary.pensoft.net/fig/1190525">2 C</figureCitation>
, 
<figureCitation id="3AD7334A0012C11BC5C1867787119563" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Two species of the genus Colubroelaps in life. A. Colubroelaps adleri sp. nov., holotype ZMMU Re- 18000, adult female from Dam Bay Research Station, Hon Tre Island, Khanh Hoa Province, Vietnam; B. Colubroelaps nguyenvansangi, ZMMU Re- 15485, adult male from Cat Tien NP., Tan Phu District, Dong Nai Province, Vietnam. Photographs by: A. M. Bragin (A); E. A. Galoyan (B)." figureDoi="10.3897/herpetozoa.37.e137809.figure4" httpUri="https://binary.pensoft.net/fig/1190527">4 A</figureCitation>
). The presence of this parasite may indicate that 
<taxonomicName id="14308171C5F6270E47163B19F64393AC" authorityName="Poyarkov &amp; Bragin &amp; Nguyen" authorityYear="2024" class="Squamata" family="Colubridae" genus="Colubroelaps" kingdom="Animalia" phylum="Chordata" rank="species" species="adleri" status="sp. nov.">
<emphasis id="DB0D22A244AE5FDA831C2C90828C9B6E" italics="true">Colubroelaps adleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2B66C3EB832542FAB0236197401E08DE" rank="species">sp. nov.</taxonomicNameLabel>
likely feeds on insects, as these animals are the intermediate hosts for acanthocephalan parasitic worms. All other aspects of the ecology of 
<taxonomicName id="FF799F2BD4199C79351DEEA7A28ABEB4" authorityName="Poyarkov &amp; Bragin &amp; Nguyen" authorityYear="2024" class="Squamata" family="Colubridae" genus="Colubroelaps" kingdom="Animalia" phylum="Chordata" rank="species" species="adleri" status="sp. nov.">
<emphasis id="07987E0EEDFFD9108CA4F14CE8B94B17" italics="true">Colubroelaps adleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3E741F5B39587449159C80DF41EB15A9" rank="species">sp. nov.</taxonomicNameLabel>
, including information on diet, preferred microhabitats, reproduction, and predators, remain unknown.
</paragraph>
</subSubSection>
</treatment>
</document>