300 lines
19 KiB
XML
300 lines
19 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.222.87779" ID-Pensoft-Pub="1314-2003-222-75" ID-Pensoft-UUID="348E6053A6D55787993694B23682FDE9" ModsDocID="1314-2003-222-75" checkinTime="1679617121532" checkinUser="pensoft" docAuthor="Maas, Paul J. M., Maas-van de Kamer, Hiltje, Andre, Thiago, Skinner, David, Valderrama, Eugenio & Specht, Chelsea D." docDate="2023" docId="09336D8339175178B7FE6E7E4AC55868" docLanguage="en" docName="PhytoKeys 222: 75-127" docOrigin="PhytoKeys 222" docPubDate="2023-03-22" docSource="http://dx.doi.org/10.3897/phytokeys.222.87779" docTitle="Costus alleniopsis Maas & D. Skinner 2023, sp. nov." docType="treatment" docVersion="1" id="348E6053A6D55787993694B23682FDE9" lastPageNumber="75" masterDocId="348E6053A6D55787993694B23682FDE9" masterDocTitle="Eighteen new species of Neotropical Costaceae (Zingiberales)" masterLastPageNumber="127" masterPageNumber="75" pageNumber="75" updateTime="1679617121532" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Eighteen new species of Neotropical Costaceae (Zingiberales)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Maas, Paul J. M.</mods:namePart>
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<mods:affiliation>Naturalis Biodiversity Centre, Botany, P. O. Box 9517, 2300 RA Leiden, Netherlands</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Maas-van de Kamer, Hiltje</mods:namePart>
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<mods:affiliation>Naturalis Biodiversity Centre, Botany, P. O. Box 9517, 2300 RA Leiden, Netherlands</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Andre, Thiago</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4148-3662</mods:nameIdentifier>
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<mods:affiliation>Universidade de Brasilia, Departamento de Botanica, Campus Universitario Darcy Ribeiro, Asa Norte, Brasilia (DF), Brazil</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Skinner, David</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4585-925X</mods:nameIdentifier>
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<mods:affiliation>Le Jardin Ombrage, Tallahassee, (Private botanical garden, Botanic Gardens Conservation International - BGCI - registration ID 50148), Florida, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Valderrama, Eugenio</mods:namePart>
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<mods:affiliation>Cornell University, Section of Plant Biology and the L. H. Bailey Hortorium, School of Integrative Plant Science, Ithaca, NY, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Specht, Chelsea D.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7746-512X</mods:nameIdentifier>
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<mods:affiliation>Cornell University, Section of Plant Biology and the L. H. Bailey Hortorium, School of Integrative Plant Science, Ithaca, NY, USA</mods:affiliation>
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<mods:nameIdentifier type="email">cdspecht@cornell.edu</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-03-22</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>222</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>75</mods:start>
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<mods:end>127</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.222.87779</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.222.87779</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-222-75</mods:identifier>
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<mods:identifier type="Pensoft-UUID">348E6053A6D55787993694B23682FDE9</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:09336D8339175178B7FE6E7E4AC55868" httpUri="http://treatment.plazi.org/id/09336D8339175178B7FE6E7E4AC55868" lastPageNumber="75" pageId="0" pageNumber="75">
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<subSubSection pageId="0" pageNumber="75" type="nomenclature">
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<paragraph pageId="0" pageNumber="75">
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<taxonomicName LSID="09336D83-3917-5178-B7FE-6E7E4AC55868" authority="Maas & D. Skinner" authorityName="Maas & D. Skinner" authorityYear="2023" class="Liliopsida" family="Costaceae" genus="Costus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Costus alleniopsis" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="alleniopsis" status="sp. nov.">Costus alleniopsis Maas & D.Skinner</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="75">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="75" type="diagnosis">
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<paragraph pageId="0" pageNumber="75">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="75">
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<taxonomicName authorityName="Maas & D. Skinner" authorityYear="2023" class="Liliopsida" family="Costaceae" genus="Costus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Costus alleniopsis" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="alleniopsis">
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<emphasis italics="true" pageId="0" pageNumber="75">Costus alleniopsis</emphasis>
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</taxonomicName>
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sp. nov. (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Costus alleniopsis Maas & D. Skinner A inflorescence B bracts and flower C plant in habitat D ligule and leaf base E ligule and leaf base of C. allenii for comparison F leaf plicae and indument G leaf plicae and indument of Costus allenii for comparison. Photos of C. alleniopsis in habitat, Veraguas, Panama by Dave Skinner." figureDoi="10.3897/phytokeys.222.87779.figure3" httpUri="https://binary.pensoft.net/fig/828342" pageId="0" pageNumber="75">3</figureCitation>
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) resembles
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<taxonomicName class="Liliopsida" family="Costaceae" kingdom="Plantae" lsidName="C. allenii" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="allenii">
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<emphasis italics="true" pageId="0" pageNumber="75">C. allenii</emphasis>
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</taxonomicName>
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but differs by distinctly plicate leaves and a longer ligule.
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</paragraph>
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<caption doi="10.3897/phytokeys.222.87779.figure3" httpUri="https://binary.pensoft.net/fig/828342" pageId="0" pageNumber="75" start="Figure 3" startId="F3">
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<paragraph pageId="0" pageNumber="75">
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<emphasis bold="true" pageId="0" pageNumber="75">Figure 3.</emphasis>
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<taxonomicName authorityName="Maas & D. Skinner" authorityYear="2023" class="Liliopsida" family="Costaceae" genus="Costus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Costus alleniopsis" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="alleniopsis">
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<emphasis italics="true" pageId="0" pageNumber="75">Costus alleniopsis</emphasis>
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</taxonomicName>
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Maas & D. Skinner
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<emphasis bold="true" pageId="0" pageNumber="75">A</emphasis>
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inflorescence
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<emphasis bold="true" pageId="0" pageNumber="75">B</emphasis>
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bracts and flower
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<emphasis bold="true" pageId="0" pageNumber="75">C</emphasis>
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plant in habitat
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<emphasis bold="true" pageId="0" pageNumber="75">D</emphasis>
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ligule and leaf base
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<emphasis bold="true" pageId="0" pageNumber="75">E</emphasis>
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ligule and leaf base of
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<taxonomicName class="Liliopsida" family="Costaceae" kingdom="Plantae" lsidName="C. allenii" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="allenii">
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<emphasis italics="true" pageId="0" pageNumber="75">C. allenii</emphasis>
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</taxonomicName>
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for comparison
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<emphasis bold="true" pageId="0" pageNumber="75">F</emphasis>
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leaf plicae and indument
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<emphasis bold="true" pageId="0" pageNumber="75">G</emphasis>
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leaf plicae and indument of
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<taxonomicName class="Liliopsida" family="Costaceae" genus="Costus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Costus allenii" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="allenii">
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<emphasis italics="true" pageId="0" pageNumber="75">Costus allenii</emphasis>
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</taxonomicName>
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for comparison. Photos of
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<taxonomicName class="Liliopsida" family="Costaceae" kingdom="Plantae" lsidName="C. alleniopsis" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="alleniopsis">
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<emphasis italics="true" pageId="0" pageNumber="75">C. alleniopsis</emphasis>
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</taxonomicName>
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in habitat, Veraguas, Panama by Dave Skinner.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="75" type="materials_examined">
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<paragraph pageId="0" pageNumber="75">
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<typeStatus>Type</typeStatus>
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.
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</paragraph>
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<paragraph pageId="0" pageNumber="75">
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<materialsCitation collectingDate="1974-09-08" collectorName="Maas, Dressler" country="Panama" elevation="725" location="Rio Segundo Brazo" specimenCount="1" stateProvince="Veraguas" typeStatus="holotype">
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<collectingCountry name="Panama">Panama</collectingCountry>
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,
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<collectingRegion country="Panama" name="Veraguas">Veraguas</collectingRegion>
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:
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<location LSID="urn:lsid:plazi:treatment:09336D8339175178B7FE6E7E4AC55868:D38188CC7F4B69D96C9BB38864438F5A" country="Panama" name="Rio Segundo Brazo" stateProvince="Veraguas">
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<normalizedToken originalValue="Río">Rio</normalizedToken>
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Segundo Brazo
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</location>
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,
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<quantity metricMagnitude="2" metricUnit="m" metricValue="7.25" metricValueMax="7.5" metricValueMin="7.0" unit="m" value="725.0" valueMax="750.0" valueMin="700.0">
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<elevation metricMagnitude="2" metricUnit="m" metricValue="7.25" metricValueMax="7.5" metricValueMin="7.0" unit="m" value="725.0" valueMax="750.0" valueMin="700.0">700-750 m</elevation>
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</quantity>
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,
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<collectingDate value="1974-09-08">8 Sep 1974</collectingDate>
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,
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<emphasis italics="true" pageId="0" pageNumber="75">
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<collectorName>Maas</collectorName>
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&
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<collectorName>Dressler</collectorName>
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1652
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</emphasis>
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(
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<typeStatus>holotype</typeStatus>
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U1231561; isotypes F, MO-2193025)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="75" type="description">
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<paragraph pageId="0" pageNumber="75">Description.</paragraph>
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<paragraph pageId="0" pageNumber="75">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="75">Herb</emphasis>
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0.5-1.2 m tall.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="75">Leaves</emphasis>
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sheaths 10-20 mm diam; ligule obliquely truncate, 12-20 mm long, often reddish; petiole 10-25 mm long; sheaths, ligule and petiole densely brownish villose; lamina ovate to obovate, 25-32 by 13-16 cm, abaxially sometimes reddish, 10-12-plicate, adaxially and abaxially densely brownish villose, base cordate, acute, or obtuse, apex acuminate (acumen 10-15 mm long).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="75">Inflorescence</emphasis>
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ovoid, 6-7
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<normalizedToken originalValue="×">x</normalizedToken>
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3-5 cm, terminating a leafy shoot or rarely terminating a leafless shoot; bracts rather densely to densely brownish villose, bracteole, calyx, ovary, and capsule rather densely to densely puberulous to glabrous.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="75">Flowers</emphasis>
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adaxially oriented to erect; bracts green, rarely red, coriaceous, broadly ovate to ovate, 3-5
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<normalizedToken originalValue="×">x</normalizedToken>
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2-5 cm, apex obtuse, callus 5-7 mm long; bracteole boat-shaped, 20-30 mm long; calyx pink to red, 8-10 mm long, lobes shallowly triangular, 2-3 mm long; corolla yellow, 60-70 mm long, glabrous, lobes narrowly elliptic, 50-60 mm long; labellum yellow, distal edge horizontally spreading, broadly obovate, 50-70
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<normalizedToken originalValue="×">x</normalizedToken>
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40-50 mm, lateral lobes striped with red, middle lobe reflexed, with yellow honey mark, margin crenulate; stamen yellow, 40-50
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<normalizedToken originalValue="×">x</normalizedToken>
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12-15 mm, not exceeding the labellum, apex red, obtuse to irregularly dentate, anther 7-10 mm long.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="75">Capsule</emphasis>
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ellipsoid, 15-20 mm long.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="75" type="distribution">
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<paragraph pageId="0" pageNumber="75">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="75">
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Panama (
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<normalizedToken originalValue="Coclé">Cocle</normalizedToken>
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,
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<normalizedToken originalValue="Colón">Colon</normalizedToken>
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, Veraguas) (Fig.
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Distribution maps for new species of Costaceae: Chamaecostus - Costus douglasdalyi A Chamaecostus manausensis Maas & H. Maas B Costus alfredoi Maas & H. Maas C Costus alleniopsis Maas & D. Skinner D Costus alticolus Maas & H. Maas E Costus antioquiensis Maas & H. Maas F Costus callosus Maas & H. Maas G Costus cochabambae Maas & H. Maas H Costus convexus Maas & D. Skinner I Costus douglasdalyi Maas & H. Maas." figureDoi="10.3897/phytokeys.222.87779.figure21" httpUri="https://binary.pensoft.net/fig/828360" pageId="0" pageNumber="75">21C</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="75" type="habitat">
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<paragraph pageId="0" pageNumber="75">Habitat and ecology.</paragraph>
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<paragraph pageId="0" pageNumber="75">In forests, at elevations of 200-1300 m. Flowering in the rainy season.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="75" type="etymology">
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<paragraph pageId="0" pageNumber="75">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="75">
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This species strongly resembles
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<taxonomicName class="Liliopsida" family="Costaceae" kingdom="Plantae" lsidName="C. allenii" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="allenii">
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<emphasis italics="true" pageId="0" pageNumber="75">C. allenii</emphasis>
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</taxonomicName>
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, and the Greek word "
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<emphasis italics="true" pageId="0" pageNumber="75">opsis</emphasis>
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" (= likeness) refers to the resemblance with that species.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="75" type="notes">
|
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<paragraph pageId="0" pageNumber="75">Notes.</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="75">
|
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|
|
<taxonomicName authorityName="Maas & D. Skinner" authorityYear="2023" class="Liliopsida" family="Costaceae" genus="Costus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Costus alleniopsis" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="alleniopsis">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Costus alleniopsis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
sp. nov. is restricted to Panama. It is closest to
|
|||
|
|
<taxonomicName class="Liliopsida" family="Costaceae" kingdom="Plantae" lsidName="C. allenii" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="allenii">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">C. allenii</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
Maas, differing by a longer ligule (12-20 mm vs. 2-10 mm) and distinctly plicate leaves. They both share adaxial oriented to erect flowers, a feature not often seen in the genus
|
|||
|
|
<taxonomicName class="Liliopsida" family="Costaceae" genus="Costus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Costus" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="genus">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Costus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="0" pageNumber="75" type="paratypes">
|
|||
|
|
<paragraph pageId="0" pageNumber="75">Paratypes.</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="75">
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="75">
|
|||
|
|
Panama.
|
|||
|
|
<normalizedToken originalValue="Coclé">Cocle</normalizedToken>
|
|||
|
|
</emphasis>
|
|||
|
|
: Alto Calvario, above El
|
|||
|
|
<normalizedToken originalValue="Copé">Cope</normalizedToken>
|
|||
|
|
, ca. 6 km N of El
|
|||
|
|
<normalizedToken originalValue="Copé">Cope</normalizedToken>
|
|||
|
|
, 23 Jun 1988,
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Croat 68796</emphasis>
|
|||
|
|
(MO, U); Alto Calvario, forest along Continental Divide, ca. 5 miles N of El
|
|||
|
|
<normalizedToken originalValue="Copé">Cope</normalizedToken>
|
|||
|
|
, 900-1000 m, 31 Mar 1993,
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Croat 75052</emphasis>
|
|||
|
|
(MO, U); Caribbean side of divide at El
|
|||
|
|
<normalizedToken originalValue="Copé">Cope</normalizedToken>
|
|||
|
|
, 200-400 m, 4 Feb 1983,
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Hamilton & Davidse 2778</emphasis>
|
|||
|
|
(MO); near sawmill, 8 km N of El
|
|||
|
|
<normalizedToken originalValue="Copé">Cope</normalizedToken>
|
|||
|
|
(28 km NW of
|
|||
|
|
<normalizedToken originalValue="Penonomé">Penonome</normalizedToken>
|
|||
|
|
), 600-750 m, 1 Sep 1977,
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Maas et al. 2760</emphasis>
|
|||
|
|
(U); Parque Nacional General Omar Torrijos Herrera, El
|
|||
|
|
<normalizedToken originalValue="Copé">Cope</normalizedToken>
|
|||
|
|
, Sendero Rana Dorada, 750 m, 31 May 2004,
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Maas et al. 9541</emphasis>
|
|||
|
|
(U); above El Potroso sawmill at Continental Divide, N of El
|
|||
|
|
<normalizedToken originalValue="Copé">Cope</normalizedToken>
|
|||
|
|
, 1200-1300 m, 13 May 1981,
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Sytsma & Andersson 4610</emphasis>
|
|||
|
|
(MO, U).
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="75">
|
|||
|
|
<normalizedToken originalValue="Colón">Colon</normalizedToken>
|
|||
|
|
</emphasis>
|
|||
|
|
: MPSA Concession, around Petaquilla Tower, 23 May 2012,
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Hammel et al. 26342</emphasis>
|
|||
|
|
(MO).
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="75">Veraguas</emphasis>
|
|||
|
|
: valley of
|
|||
|
|
<normalizedToken originalValue="Río">Rio</normalizedToken>
|
|||
|
|
Dos Bocas, on road between Alto Piedra (above Santa
|
|||
|
|
<normalizedToken originalValue="Fé">Fe</normalizedToken>
|
|||
|
|
) and Calovebora, 350-400 m, 29 Aug 1974,
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Croat 27455</emphasis>
|
|||
|
|
(MO, U); near the Continental Divide, ca. 15 km NW of Santa
|
|||
|
|
<normalizedToken originalValue="Fé">Fe</normalizedToken>
|
|||
|
|
, 800 m, 22 Oct 1980,
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Maas & Dressler 5006 p.p.</emphasis>
|
|||
|
|
(K, L, U); 16 km NW of Santa
|
|||
|
|
<normalizedToken originalValue="Fé">Fe</normalizedToken>
|
|||
|
|
, on road to Calovebora (Panamanian Highway 35), 300-500 m, 16 May 1975,
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Mori & Kallunki 6124</emphasis>
|
|||
|
|
(MO).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|