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<document id="C69F18CCBA4E3401B3B78BC93CFBFFE9" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D53920BFDD0FDBFCD232B3F" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius milneanus Heller 1914" docType="treatment" docVersion="10" lastPageNumber="81" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="80" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="38A26A3D32FBD990F9F7A6A1719CE8D8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AD26E770A761F7C686662DD0ECC3E9F9">
<mods:title id="DE174EB9FEF7F2E2AB1EAD882A766002">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
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<mods:namePart id="1A626FF3E751919D89DD541915FAFEFD">Telnov, Dmitry</mods:namePart>
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<mods:title id="89F9792EC63CCA6C882A4CE1226DE6A5">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
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<mods:date id="FE09646ED08A8A2BF292F58701EAD0F0">2020</mods:date>
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<mods:number id="B9D71E662DBE32FE51AF9CFF349A8933">2020-10-19</mods:number>
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<mods:number id="94F8858680D0C63C83EB130E2048AFAD">721</mods:number>
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<mods:classification id="487A96E25DDE3DEA63BE29B1E0246417">journal article</mods:classification>
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<mods:identifier id="505913CDBF6218F1C4F8279BF81C4280" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
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<treatment id="1A3787F69D53920BFDD0FDBFCD232B3F" ID-DOI="http://doi.org/10.5281/zenodo.4344257" ID-GBIF-Taxon="168648167" ID-Zenodo-Dep="4344257" LSID="urn:lsid:plazi:treatment:1A3787F69D53920BFDD0FDBFCD232B3F" httpUri="http://treatment.plazi.org/id/1A3787F69D53920BFDD0FDBFCD232B3F" lastPageId="80" lastPageNumber="81" pageId="79" pageNumber="80">
<subSubSection id="DA84656B9D539214FDD0FDBFCBA72A7E" box="[579,1009,513,540]" pageId="79" pageNumber="80" type="nomenclature">
<paragraph id="922136E09D539214FDD0FDBFCBA72A7E" blockId="79.[515,1072,513,577]" box="[579,1009,513,540]" pageId="79" pageNumber="80">
<heading id="C969818C9D539214FDD0FDBFCBA72A7E" box="[579,1009,513,540]" centered="true" fontSize="11" level="2" pageId="79" pageNumber="80" reason="2">
<taxonomicName id="559E4D639D539214FDD0FDBFCBA72A7E" ID-CoL="3XFK8" authority="Heller, 1914" authorityName="Heller" authorityYear="1914" box="[579,1009,513,540]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="milneanus">
<emphasis id="A0EAEAF29D539214FDD0FDBFCB012A7E" bold="true" box="[579,855,513,540]" italics="true" pageId="79" pageNumber="80">Maechidius milneanus</emphasis>
<bibRefCitation id="F60F4B119D539214FCCCFDBCCBA72A7E" author="Heller K. M." box="[863,1009,514,540]" pageId="79" pageNumber="80" pagination="615 - 666" refId="ref98213" refString="Heller K. M. 1914. Coleoptera. Nova Guinea. Resultats de l'Expedition scientifique neerlandaise a la Nouvelle-Guinee en 1907 et 1909 sous les auspices du Dr. H. A. Lorentz. Zoologie 9: 615 - 666." type="journal article" year="1914">Heller, 1914</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D539214FD90FD98CC662A22" box="[515,1072,550,577]" pageId="79" pageNumber="80" type="description">
<paragraph id="922136E09D539214FD90FD98CC662A22" blockId="79.[515,1072,513,577]" box="[515,1072,550,577]" pageId="79" pageNumber="80">
<figureCitation id="0AA52A659D539214FD90FD98CA0B2A22" box="[515,605,550,576]" captionStart="Figs 58–61" captionStartId="75.[189,241,1928,1954]" captionTargetBox="[336,1249,266,1914]" captionTargetId="figure-62@75.[333,1249,257,1923]" captionTargetPageId="75" captionText="Figs 58–61. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 58. M. maleo sp. nov., holotype, ♂ (IRSN). 59. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 60. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 61. M. merdeka sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122152" httpUri="https://zenodo.org/record/4122152/files/figure.png" pageId="79" pageNumber="80">Figs 60</figureCitation>
,
<figureCitation id="0AA52A659D539214FDFAFD98CA852A22" box="[617,723,550,576]" captionStart="Figs 145–156" captionStartId="141.[189,241,1787,1813]" captionTargetBox="[328,1254,265,1740]" captionTargetId="figure-124@141.[328,1260,265,1749]" captionTargetPageId="141" captionText="Figs 145–156. Papuan Maechidius Macleay, 1819, head, dorsal view. 145. M. merdeka sp. nov., holotype, ♂ (BMNH). 146. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 147. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 148. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 149. Same, ♂ (SNSD). 150. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 151. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 152. M. nepenthephilus sp. nov., holotype, ♂ (NME). 153. Same, paratype, ♀ (DTC). 154. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 155. Same, paratype, ♀ (BMNH). 156. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122181" httpUri="https://zenodo.org/record/4122181/files/figure.png" pageId="79" pageNumber="80">148–149</figureCitation>
,
<figureCitation id="0AA52A659D539214FD4CFD98CB582A22" box="[735,782,550,576]" captionStart="Figs 226–241" captionStartId="147.[189,241,1681,1707]" captionTargetBox="[391,1192,268,1647]" captionTargetId="figure-170@147.[391,1196,265,1656]" captionTargetPageId="147" captionText="Figs 226–241. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 226. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 227. M. lobaticeps Frey, 1969, ♂ (DTC). 228. M. longipes sp. nov., holotype, ♂ (BMNH). 229. M. luniceps Fairmaire, 1883, ♂ (BMNH). 230. M. mailu sp. nov., holotype, ♂ (NHMB). 231. M. maleo sp. nov., holotype, ♂ (IRSN). 232. M. merdeka sp. nov., holotype, ♂ (BMNH). 233. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 234. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 235. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 236. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 237. M. nepenthephilus sp. nov., holotype, ♂ (NME). 238. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 239. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 240. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 241. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122193" httpUri="https://zenodo.org/record/4122193/files/figure.png" pageId="79" pageNumber="80">233</figureCitation>
,
<figureCitation id="0AA52A659D539214FC8AFD98CB1E2A23" box="[793,840,550,577]" captionStart="Figs 317–332" captionStartId="154.[189,241,1681,1707]" captionTargetBox="[401,1187,265,1655]" captionTargetId="figure-172@154.[401,1187,265,1669]" captionTargetPageId="154" captionText="Figs 317–332. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 317. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 318. M. lobaticeps Frey, 1969, ♂ (DTC). 319. M. longipes sp. nov., holotype, ♂ (BMNH). 320. M. luniceps Fairmaire, 1883, ♂ (BMNH). 321. M. mailu sp. nov., holotype, ♂ (NHMB). 322. M. maleo sp. nov., holotype, ♂ (IRSN). 323. M. merdeka sp. nov., holotype, ♂ (BMNH). 324. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 325. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 326. M. muticusArrow, 1941, syntype, ♂ (BMNH). 327. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 328. M. nepenthephilus sp. nov., holotype, ♂ (NME). 329. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 330. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 331. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 332. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122210" httpUri="https://zenodo.org/record/4122210/files/figure.png" pageId="79" pageNumber="80">325</figureCitation>
, 398, 494, 670–672
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D539214FF2EFDD6CB072C7B" pageId="79" pageNumber="80" type="materials_examined">
<paragraph id="922136E09D539214FF2EFDD6C93A2AE0" blockId="79.[189,364,616,642]" box="[189,364,616,642]" pageId="79" pageNumber="80">
<heading id="C969818C9D539214FF2EFDD6C93A2AE0" bold="true" box="[189,364,616,642]" fontSize="11" level="3" pageId="79" pageNumber="80" reason="3">
<emphasis id="A0EAEAF29D539214FF2EFDD6C93A2AE0" bold="true" box="[189,364,616,642]" pageId="79" pageNumber="80">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D539214FF79FD26CA3A2AD0" blockId="79.[189,1399,663,762]" box="[234,620,663,690]" pageId="79" pageNumber="80">
<emphasis id="A0EAEAF29D539214FF79FD26C9302AD0" bold="true" box="[234,358,664,690]" pageId="79" pageNumber="80">
<typeStatus id="4D2588429D539214FF79FD26C9302AD0" box="[234,358,664,690]" pageId="79" pageNumber="80" type="lectotype">Lectotype</typeStatus>
</emphasis>
[herewith designated]
</paragraph>
<paragraph id="922136E09D539214FF2EFD02CCCC2A9B" blockId="79.[189,1399,663,762]" pageId="79" pageNumber="80">
<materialsCitation id="22F63CBD9D539214FF2EFD02CCCC2A9B" ID-GBIF-Occurrence="2898878349" collectionCode="SNSD" country="Papua New Guinea" location="PAPUA NEW GUINEA" pageId="79" pageNumber="80" specimenCount="2" specimenCount-female="1" typeStatus="typus">
<collectingCountry id="EA8976709D539214FF2EFD02C9852AB7" box="[189,467,699,726]" name="Papua New Guinea" pageId="79" pageNumber="80">PAPUA NEW GUINEA</collectingCountry>
•
<specimenCount id="8498FD699D539214FE75FD02C9AB2AB4" box="[486,509,700,726]" pageId="79" pageNumber="80" type="female">♀</specimenCount>
; “Neu Guinea Milne-Bay [p, blue label] // 1910 [p] 18 [h] [blue label]//
<typeStatus id="4D2588429D539214FAB6FD05CD252AB7" box="[1317,1395,699,725]" pageId="79" pageNumber="80">Typus</typeStatus>
<specimenCount id="8498FD699D539214FAE0FD05CD212AB7" box="[1395,1399,699,725]" pageId="79" pageNumber="80" type="male">♂</specimenCount>
[p]
<taxonomicName id="559E4D639D539214FF74FD61C9362A9B" authorityName="Heller" authorityYear="1914" box="[231,352,735,761]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="milneanus">
<emphasis id="A0EAEAF29D539214FF74FD61C9362A9B" box="[231,352,735,761]" italics="true" pageId="79" pageNumber="80">milneanus</emphasis>
</taxonomicName>
H. [h, red label] // Staatl. Museum für Tierkunde.Dresden [p]“;
<collectionCode id="F48FAE259D539214FBDBFD61CCC12A9B" box="[1096,1175,735,761]" collectionName="SNSD" pageId="79" pageNumber="80">SNSD</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="922136E09D539214FF79FC9CC9C12B5E" blockId="79.[189,1399,802,900]" box="[234,407,802,828]" pageId="79" pageNumber="80">
<emphasis id="A0EAEAF29D539214FF79FC9CC9C12B5E" bold="true" box="[234,407,802,828]" pageId="79" pageNumber="80">
<typeStatus id="4D2588429D539214FF79FC9CC9C12B5E" box="[234,407,802,828]" pageId="79" pageNumber="80" type="paralectotype">Paralectotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D539214FF2EFCF8CB2F2BE1" blockId="79.[189,1399,802,900]" pageId="79" pageNumber="80">
<materialsCitation id="22F63CBD9D539214FF2EFCF8CB202BE1" ID-GBIF-Occurrence="2898878417" collectionCode="SNSD" country="Papua New Guinea" location="PAPUA NEW GUINEA" pageId="79" pageNumber="80" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<collectingCountry id="EA8976709D539214FF2EFCF8C9BE2B3D" box="[189,488,837,864]" name="Papua New Guinea" pageId="79" pageNumber="80">PAPUA NEW GUINEA</collectingCountry>
•
<specimenCount id="8498FD699D539214FD9CFCFBCA122B02" box="[527,580,837,864]" pageId="79" pageNumber="80" type="female">1 ♀</specimenCount>
; “Neu
<collectingCountry id="EA8976709D539214FD34FCFBCAAA2B3D" box="[679,764,837,863]" name="Guinea-Bissau" pageId="79" pageNumber="80">Guinea</collectingCountry>
Milne-Bay [p, blue label]// 1910 [p] 18 [h] [blue label]// Staatl. Museum für Tierkunde.Dresden [p]”;
<collectionCode id="F48FAE259D539214FCB4FCD7CB202BE1" box="[807,886,873,899]" collectionName="SNSD" pageId="79" pageNumber="80">SNSD</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="922136E09D539214FF2EFC12C9302BA7" blockId="79.[189,358,939,966]" box="[189,358,939,966]" pageId="79" pageNumber="80">
<heading id="C969818C9D539214FF2EFC12C9302BA7" bold="true" box="[189,358,939,966]" fontSize="11" level="3" pageId="79" pageNumber="80" reason="3">
<emphasis id="A0EAEAF29D539214FF2EFC12C9302BA7" bold="true" box="[189,358,939,966]" pageId="79" pageNumber="80">New material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D539214FF2EFC65CB072C7B" blockId="79.[189,1398,986,1049]" pageId="79" pageNumber="80">
<materialsCitation id="22F63CBD9D539214FF2EFC65CB182C7B" ID-GBIF-Occurrence="2898878434" collectionCode="MUCHE, SNSD" collectorName="Milnebay N. Guinea" country="Papua New Guinea" location="PAPUA NEW GUINEA" pageId="79" pageNumber="80" specimenCount="15" specimenCount-female="1">
<collectingCountry id="EA8976709D539214FF2EFC65C9B22B97" box="[189,484,987,1013]" name="Papua New Guinea" pageId="79" pageNumber="80">PAPUA NEW GUINEA</collectingCountry>
•
<specimenCount id="8498FD699D539214FD9AFC64CA252B97" box="[521,627,986,1013]" pageId="79" pageNumber="80">15 specs</specimenCount>
(♂♀); “
<collectorName id="3F6B53369D539214FD4CFC65CB872B97" box="[735,977,987,1013]" pageId="79" pageNumber="80">Milnebay N.Guinea</collectorName>
// W.H.
<collectionCode id="F48FAE259D539214FBB3FC65CCD82B97" box="[1056,1166,987,1013]" pageId="79" pageNumber="80">MUCHE</collectionCode>
Radeberg, Ankauf [p] // Staatl.Museum für Tierkunde Dresden [p]”;
<collectionCode id="F48FAE259D539214FD6CFC41CB182C7B" box="[767,846,1023,1049]" collectionName="SNSD" pageId="79" pageNumber="80">SNSD</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D539214FF2EFBFFCA502D00" pageId="79" pageNumber="80" type="discussion">
<paragraph id="922136E09D539214FF2EFBFFC9792C39" blockId="79.[189,303,1089,1115]" box="[189,303,1089,1115]" pageId="79" pageNumber="80">
<heading id="C969818C9D539214FF2EFBFFC9792C39" bold="true" box="[189,303,1089,1115]" fontSize="11" level="3" pageId="79" pageNumber="80" reason="3">
<emphasis id="A0EAEAF29D539214FF2EFBFFC9792C39" bold="true" box="[189,303,1089,1115]" pageId="79" pageNumber="80">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D539214FF2EFBCECA502D00" blockId="79.[189,1399,1136,1378]" pageId="79" pageNumber="80">
<bibRefCitation id="F60F4B119D539214FF2EFBCEC9CC2CE9" author="Heller K. M." box="[189,410,1136,1163]" pageId="79" pageNumber="80" pagination="615 - 666" refId="ref98213" refString="Heller K. M. 1914. Coleoptera. Nova Guinea. Resultats de l'Expedition scientifique neerlandaise a la Nouvelle-Guinee en 1907 et 1909 sous les auspices du Dr. H. A. Lorentz. Zoologie 9: 615 - 666." type="journal article" year="1914">Heller (1914: 628)</bibRefCitation>
based his description of
<taxonomicName id="559E4D639D539214FD54FBCFCB3F2CE8" authorityName="Heller" authorityYear="1914" box="[711,873,1136,1162]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="milneanus">
<emphasis id="A0EAEAF29D539214FD54FBCFCB3F2CE8" box="[711,873,1136,1162]" italics="true" pageId="79" pageNumber="80">M. milneanus</emphasis>
</taxonomicName>
on an unstated number of specimens. Since two specimens with identical locality data are present in SNSD (and a number of specimens with slightly different labels – see above), the
<typeStatus id="4D2588429D539214FDDCFB06CAED2CB0" box="[591,699,1208,1234]" pageId="79" pageNumber="80" type="lectotype">lectotype</typeStatus>
designated here is the only
<typeStatus id="4D2588429D539214FB82FB06CC3A2CB0" box="[1041,1132,1208,1234]" pageId="79" pageNumber="80" type="syntype">syntype</typeStatus>
with Heller’s original “typus” handwriting on a red label, which, without confidence, may indicate that this specimen was selected by the original author as the
<typeStatus id="4D2588429D539214FDEFFABECAB12D78" box="[636,743,1280,1306]" pageId="79" pageNumber="80" type="holotype">holotype</typeStatus>
. The
<typeStatus id="4D2588429D539214FCBBFABECBC22D78" box="[808,916,1280,1306]" pageId="79" pageNumber="80" type="lectotype">lectotype</typeStatus>
designation is made in order to enhance the stability of nomenclature and fix the specimen I based my redescription on, in case other
<typeStatus id="4D2588429D539214FA83FA9ACD212D5C" box="[1296,1399,1316,1342]" pageId="79" pageNumber="80" type="syntype">syntypes</typeStatus>
are discovered in the future.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D53920BFF2EFA34CD232B3F" lastPageId="80" lastPageNumber="81" pageId="79" pageNumber="80" type="description">
<paragraph id="922136E09D539214FF2EFA34C9182DC6" blockId="79.[189,334,1418,1444]" box="[189,334,1418,1444]" pageId="79" pageNumber="80">
<heading id="C969818C9D539214FF2EFA34C9182DC6" bold="true" box="[189,334,1418,1444]" fontSize="11" level="3" pageId="79" pageNumber="80" reason="3">
<emphasis id="A0EAEAF29D539214FF2EFA34C9182DC6" bold="true" box="[189,334,1418,1444]" pageId="79" pageNumber="80">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D539214FF2EFA04CBDF2DB6" blockId="79.[189,905,1465,1492]" box="[189,905,1465,1492]" pageId="79" pageNumber="80">
MEASUREMENTS. Total body length 7.30–9.10 (
<typeStatus id="4D2588429D539214FD46FA07CB122DB6" box="[725,836,1465,1492]" pageId="79" pageNumber="80" type="lectotype">lectotype</typeStatus>
) mm.
</paragraph>
<paragraph id="922136E09D53920BFF2EFA42CC492A8F" blockId="79.[189,1399,1531,2025]" lastBlockId="80.[189,1399,265,749]" lastPageId="80" lastPageNumber="81" pageId="79" pageNumber="80">
Dorsum and venter uniformly brown, appendages somewhat paler, forebody with greenish metallic lustre. Head convex dorsally on frons between compound eyes, subopaque dorsally and ventrally. Labroclypeus (
<figureCitation id="0AA52A659D539214FEE3F9FDCA412E3F" box="[368,535,1603,1629]" captionStart="Figs 145–156" captionStartId="141.[189,241,1787,1813]" captionTargetBox="[328,1254,265,1740]" captionTargetId="figure-124@141.[328,1260,265,1749]" captionTargetPageId="141" captionText="Figs 145–156. Papuan Maechidius Macleay, 1819, head, dorsal view. 145. M. merdeka sp. nov., holotype, ♂ (BMNH). 146. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 147. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 148. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 149. Same, ♂ (SNSD). 150. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 151. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 152. M. nepenthephilus sp. nov., holotype, ♂ (NME). 153. Same, paratype, ♀ (DTC). 154. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 155. Same, paratype, ♀ (BMNH). 156. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122181" httpUri="https://zenodo.org/record/4122181/files/figure.png" pageId="79" pageNumber="80">Figs 148–149</figureCitation>
) in both sexes very broadly shallowly emarginate anteriorly, its lateral margins strongly sinuous in both dorsal and lateral views. Anterolateral angles broadly rounded, hardly protruding, raised up at angle of nearly 80-90° to frons in lateral view. Anterior and lateral margins of labroclypeus smooth. Upper- and underside of labroclypeus with sparse moderately long setae along anterior and lateral margins. Canthus strongly obtusely angulate in dorsal view. Punctures of frons circular to hexagonal, in part annular, large and shallow. Intervening spaces glossy, variably large. Setae inconspicuous, appressed, generally not surpassing length of corresponding punctures. Antenna 7-segmented, club 3-lamellate. Pronotum transverse, flattened dorsally, subopaque dorsally and laterally. Anterior margin of pronotum broadly emarginate with protruding anterolateral angles, basal margin broadly rounded. Lateral margin of pronotum broadly rounded in dorsal view, strongly crenulate all along (
<figureCitation id="0AA52A659D539214FE39F814CA442FA7" box="[426,530,1962,1989]" captionStart="Figs 226–241" captionStartId="147.[189,241,1681,1707]" captionTargetBox="[391,1192,268,1647]" captionTargetId="figure-170@147.[391,1196,265,1656]" captionTargetPageId="147" captionText="Figs 226–241. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 226. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 227. M. lobaticeps Frey, 1969, ♂ (DTC). 228. M. longipes sp. nov., holotype, ♂ (BMNH). 229. M. luniceps Fairmaire, 1883, ♂ (BMNH). 230. M. mailu sp. nov., holotype, ♂ (NHMB). 231. M. maleo sp. nov., holotype, ♂ (IRSN). 232. M. merdeka sp. nov., holotype, ♂ (BMNH). 233. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 234. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 235. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 236. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 237. M. nepenthephilus sp. nov., holotype, ♂ (NME). 238. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 239. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 240. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 241. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122193" httpUri="https://zenodo.org/record/4122193/files/figure.png" pageId="79" pageNumber="80">Fig. 233</figureCitation>
). Lateral margin of pronotum sinuous in lateral view. Short suberect seta present between every two crenulae. With small circular vague median impression on either side of disc. Hypomeron separated from prosternum by low nearly straight carina, with moderately long setae on its anterolateral margin opposite to compound eye. Antennal pocket shallow. Punctures of pronotal disc large, circular to ovoid, dense, annular. Intervening spaces distinctly smaller than punctures. Pronotal setae inconspicuous, appressed; each seta rises from anterior margin of corresponding puncture, not surpassing its length. Scutellar shield narrowly rounded apically. Elytron subopaque, without or with vague tracks of four flat longitudinal carinae, including sutural one. Surface of elytral disc wrinkled and with sinuous (long and narrow) incision-shaped punctures. Intervening spaces microrugulose (
<figureCitation id="0AA52A659D4C920BFA97FE62CD3F2995" box="[1284,1385,476,503]" captionStart="Figs 317–332" captionStartId="154.[189,241,1681,1707]" captionTargetBox="[401,1187,265,1655]" captionTargetId="figure-172@154.[401,1187,265,1669]" captionTargetPageId="154" captionText="Figs 317–332. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 317. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 318. M. lobaticeps Frey, 1969, ♂ (DTC). 319. M. longipes sp. nov., holotype, ♂ (BMNH). 320. M. luniceps Fairmaire, 1883, ♂ (BMNH). 321. M. mailu sp. nov., holotype, ♂ (NHMB). 322. M. maleo sp. nov., holotype, ♂ (IRSN). 323. M. merdeka sp. nov., holotype, ♂ (BMNH). 324. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 325. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 326. M. muticusArrow, 1941, syntype, ♂ (BMNH). 327. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 328. M. nepenthephilus sp. nov., holotype, ♂ (NME). 329. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 330. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 331. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 332. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122210" httpUri="https://zenodo.org/record/4122210/files/figure.png" pageId="80" pageNumber="81">Fig. 325</figureCitation>
). Elytral setae inconspicuous, appressed, not surpassing length of corresponding incisions. Male and female pygidium flattened dorsally, with large shallow annular punctures (
<figureCitation id="0AA52A659D4C920BFBB2FD9DCCDF2A5F" box="[1057,1161,547,573]" captionStart="Figs487–503" captionStartId="165.[189,241,1681,1707]" captionTargetBox="[422,1160,265,1655]" captionTargetId="figure-171@165.[421,1167,265,1658]" captionTargetPageId="165" captionText="Figs487–503.Papuan andWallaceanMaechidius Macleay, 1819,pygidium, dorsalview.487.M.lobaticeps Frey, 1969, ♂ (DTC). 488. M. longipes sp. nov., holotype, ♂ (BMNH). 489. M. luniceps Fairmaire, 1883, ♂ (BMNH). 490. M. mailu sp. nov., holotype, ♂ (NHMB). 491. M. maleo sp. nov., holotype, ♂ (IRSN). 492. M. merdeka sp. nov., holotype, ♂ (BMNH). 493. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 494. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 495. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 496. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 497. M. nepenthephilus sp. nov., holotype, ♂ (NME). 498. M. opatroidesArrow, 1941 comb. rest., lectotype, ♀ (BMNH). 499. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 500. Same, paratype, ♀ (BMNH). 501. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 502. Same, different aspect. 503. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122226" httpUri="https://zenodo.org/record/4122226/files/figure.png" pageId="80" pageNumber="81">Fig. 494</figureCitation>
). Intervening spaces glossy, variably large. Setae of pygidium short, inconspicuous, sparse, generally not surpassing length of corresponding punctures except several long suberect ones along posterior margin of pygidium. Protibia widened distally, with three external teeth (Fig. 398). Male protibial terminal spur very short, female one longer and very narrow. Male metatibial terminal spurs long, nearly equally long, female lower metatibial spur distinctly longer than the upper. Tarsal claws with pulvilli.
</paragraph>
<paragraph id="922136E09D4C920BFF2EFCAAC9E62B4C" blockId="80.[189,432,788,814]" box="[189,432,788,814]" pageId="80" pageNumber="81">
<heading id="C969818C9D4C920BFF2EFCAAC9E62B4C" bold="true" box="[189,432,788,814]" fontSize="11" level="3" pageId="80" pageNumber="81" reason="3">
<emphasis id="A0EAEAF29D4C920BFF2EFCAAC9E62B4C" bold="true" box="[189,432,788,814]" pageId="80" pageNumber="81">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D4C920BFF2EFCFDCD232B3F" blockId="80.[189,1397,835,861]" box="[189,1397,835,861]" pageId="80" pageNumber="81">Female lamellae of antennal club shorter; protibial terminal spur longer and more slender than in male.</paragraph>
</subSubSection>
</treatment>
</document>
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