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<document ID-DOI= "http://dx.doi.org/10.3897/evolsyst.7.100968" ID-GBIF-Dataset= "c9a664a6-ec1d-449b-a4f5-cc42b8bfd0e6" ID-Pensoft-Pub= "2535-0730-1-117" ID-Pensoft-UUID= "CF83BFADF9D057EEB3403335CD1B4B1C" ID-ZooBank= "704B4BBCF9274DDBAAC496D7CED95318" ModsDocID= "2535-0730-7-1-117" checkinTime= "1682113244139" checkinUser= "pensoft" docAuthor= "Dellape, Pablo M. & Melo, Maria C." docDate= "2023" docId= "80289A50C4A55CC898F31E943A86920B" docLanguage= "en" docName= "EvolutSyst 7(1): 117-122" docOrigin= "Evolutionary Systematics 7 (1)" docPubDate= "2023-04-21" docSource= "http://dx.doi.org/10.3897/evolsyst.7.100968" docTitle= "Malleusocoris minimus Dellapé & Melo 2023, sp. nov." docType= "treatment" docUuid= "AABC95F6-3347-4B66-A7B6-E4F684F59C47" docUuidSource= "ZooBank" docVersion= "4" id= "CF83BFADF9D057EEB3403335CD1B4B1C" lastPageNumber= "117" masterDocId= "CF83BFADF9D057EEB3403335CD1B4B1C" masterDocTitle= "Malleusocoris, a new South American genus of Myodochini (Hemiptera, Rhyparochromidae) with modified antennae, along with some new synonymies and new combinations for misplaced taxa" masterLastPageNumber= "122" masterPageNumber= "117" pageNumber= "117" updateTime= "1732567350262" updateUser= "ExternalLinkService" >
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<mods:title id= "236EFF17DF3A8F54C9545E9F3F979315" > Malleusocoris, a new South American genus of Myodochini (Hemiptera, Rhyparochromidae) with modified antennae, along with some new synonymies and new combinations for misplaced taxa</mods:title>
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<mods:namePart id= "4D15C1D7027AB88A2FCB17DBE35F51C3" > Dellape, Pablo M.</mods:namePart>
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<mods:affiliation id= "C741346995F780713E8997A93D99888A" > Division Entomologia, Museo de la Plata, Universidad Nacional de La Plata. Paseo del Bosque s / n, B 1900 FWA, La Plata, Buenos Aires, Argentina & Consejo Nacional de Investigaciones Cientificas y Tecnicas (CONICET), Buenos Aires, Argentina</mods:affiliation>
<mods:nameIdentifier id= "392EC1931413FE963EC821B81EC02D3B" type= "email" > pdellape@fcnym.unlp.edu.ar</mods:nameIdentifier>
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<mods:namePart id= "DB585570FC7B61981B07CB6C3067568A" > Melo, Maria C.</mods:namePart>
<mods:affiliation id= "859F959650F38B57AFF1C1AC84D8B343" > Division Entomologia, Museo de la Plata, Universidad Nacional de La Plata. Paseo del Bosque s / n, B 1900 FWA, La Plata, Buenos Aires, Argentina & Consejo Nacional de Investigaciones Cientificas y Tecnicas (CONICET), Buenos Aires, Argentina</mods:affiliation>
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<mods:title id= "3844F87B009F42D2494CFED5A78EDB0B" > Evolutionary Systematics</mods:title>
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<mods:part id= "6AC925CD325FDDA2F6BA6CFBB9E0206D" >
<mods:date id= "896F1FD9CF1DB683FD93F0789346FD3A" > 2023</mods:date>
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<treatment id= "80289A50C4A55CC898F31E943A86920B" ID-GBIF-Taxon= "209565777" LSID= "urn:lsid:zoobank.org:act:AABC95F6-3347-4B66-A7B6-E4F684F59C47" httpUri= "http://treatment.plazi.org/id/80289A50C4A55CC898F31E943A86920B" lastPageNumber= "117" pageId= "0" pageNumber= "117" scope_family= "Rhyparochromidae" scope_order= "Hemiptera" scope_subFamily= "Rhyparochrominae" scope_tribe= "Myodochini" >
<subSubSection id= "48A36BCAAF454C3583710AD445CA706E" pageId= "0" pageNumber= "117" type= "nomenclature" >
<paragraph id= "A3CD1CAB0EBC00BB930D04F76760ADEB" pageId= "0" pageNumber= "117" >
<taxonomicName id= "F6D4589590C4A2B709635051FE226E76" ID-CoL= "C7Q68" LSID= "https://zoobank.org/AABC95F6-3347-4B66-A7B6-E4F684F59C47" authority= "Dellapé & Melo, 2023" authorityName= "Dellapé & Melo" authorityYear= "2023" class= "Insecta" family= "Rhyparochromidae" genus= "Malleusocoris" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Malleusocoris minimus" order= "Hemiptera" pageId= "0" pageNumber= "117" phylum= "Arthropoda" rank= "species" species= "minimus" status= "sp. nov." > Malleusocoris minimus</taxonomicName>
<taxonomicNameLabel id= "A14107F9F4FF526DAA37136952224631" pageId= "0" pageNumber= "117" > sp. nov.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id= "3AE265EBD53D9BC60B62A73962F11430" pageId= "0" pageNumber= "117" type= "description" >
<paragraph id= "BE52B06B1BD7CE9560005B8A45F68314" pageId= "0" pageNumber= "117" >
<figureCitation id= "B93ECA05DBF09ED2D6072C158E15B86E" captionStart= "Figure 1" captionStartId= "F1" captionText= "Figure 1. Malleusocoris minimus gen. nov., sp. nov. Habitus of male holotype. Scale bar: 1 mm." httpUri= "https://binary.pensoft.net/fig/841070" pageId= "0" pageNumber= "117" > Figs 1</figureCitation>
<figureCitation id= "F504340960CCF12E62CFC7554FA6D86F" captionStart= "Figures 2– 5" captionStartId= "F2" captionText= "Figures 2 - 5. Malleusocoris minimus sp. nov. Male genitalia, of male holotype. 2. Pygophore, dorsal view; 3. Pygophore, lateral view; 4. Right paramere, outer view; 5. Right paramere, inner view. Scale bars: 0.25 mm." httpUri= "https://binary.pensoft.net/fig/841071" pageId= "0" pageNumber= "117" > , 2-5</figureCitation>
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</paragraph>
</subSubSection>
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<subSubSection id= "3EE3022C964371F8CFF760CD782728DF" pageId= "0" pageNumber= "117" type= "world catalog lygaeoidea species file link" >
<paragraph id= "EC0E1102AA8CA48A6BC8D8FB87C8AC90" pageId= "0" pageNumber= "117" >
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World catalog
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<taxonomicName id= "D0BCF8CC9FB364C96FCB9BF248C71BEE" authorityName= "Dellapé & Melo" authorityYear= "2023" class= "Insecta" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "" order= "Hemiptera" pageId= "0" pageNumber= "117" phylum= "Arthropoda" rank= "superFamily" superFamily= "Lygaeoidea" > Lygaeoidea</taxonomicName>
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Species File link.
</paragraph>
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<paragraph id= "1B425013A2AD1106CE62FF0A8C0D4CAA" pageId= "0" pageNumber= "117" > http://lsid.speciesfile.org/urn:lsid:Lygaeoidea.speciesfile.org:TaxonName:519163.</paragraph>
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</subSubSection>
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<subSubSection id= "2091E0FA690C38A10670AE1C089E9473" pageId= "0" pageNumber= "117" type= "type material" >
<paragraph id= "86C707F213CEDB4BA3222044ECBB63DD" pageId= "0" pageNumber= "117" > Type material.</paragraph>
<paragraph id= "E215C909C672D0D59BC14FE08A38821D" pageId= "0" pageNumber= "117" >
<emphasis id= "7B5A92EAD66792A022A3032C0595D8E5" bold= "true" italics= "true" pageId= "0" pageNumber= "117" > Holotype</emphasis>
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♂ (Fig.
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<figureCitation id= "9F9BFCB9571D4E8B1384DD80760BF548" captionStart= "Figure 1" captionStartId= "F1" captionText= "Figure 1. Malleusocoris minimus gen. nov., sp. nov. Habitus of male holotype. Scale bar: 1 mm." httpUri= "https://binary.pensoft.net/fig/841070" pageId= "0" pageNumber= "117" > 1</figureCitation>
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), Brazil, Mato Grosso / State Pantanal / 25 km. S of
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<normalizedToken id= "47CFA784A7D0488B0A0F954F78C09ACD" originalValue= "Poconé" > Pocone</normalizedToken>
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// Pousada form / SAO Cristovao / 29.8.2000 lgt J.
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<normalizedToken id= "F2EAEA293BE8F4D6F7EAF60EE7C3A575" originalValue= "Růžička" > Ruzicka</normalizedToken>
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// MLP / HE-10661 (MLP);
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<emphasis id= "CD582C3AA969E30C1B2D0F6990E0B4F4" bold= "true" italics= "true" pageId= "0" pageNumber= "117" > paratypes</emphasis>
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1 ♂ 2 ♀♀, same data as for holotype // MLP / HE-10662-10664 (MLP); 1 ♂ 5 ♀♀, same data as for holotype (NMPC); 1 ♀, 29.8.2000 lgt J.
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<normalizedToken id= "8D6F01B0C3F66E39FE4AA1ECD7A800A4" originalValue= "Růžička" > Ruzicka</normalizedToken>
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/ Pousada form / SAO Cristovao (NMPC); 1 ♂, Brazil, Mato Grosso state, / Pantanal, 20 km. S of
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<normalizedToken id= "61D0B29B8A56AF71FC43D38C98304A00" originalValue= "Poconé" > Pocone</normalizedToken>
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, / Pousada [farm] POUSO ALEGRE, / 30.viii.2000, Jan
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<normalizedToken id= "22228E64AA41080E87C73578ED0ED22E" originalValue= "Růžička" > Ruzicka</normalizedToken>
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leg., B9 // ca. 100 m a.s.l., on light at / evening and night (20.00-23.00 of / local time-GMT/UTC +04.00), / light bulbs near the farm, end of / dry season (NMPC); 1 ♂, Argentina - Misiones / PN
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<normalizedToken id= "8F4D04DA8BFAEBC2EE80ED83BD316E2E" originalValue= "Iguazú" > Iguazu</normalizedToken>
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/ X-1980 T. Luz / D.J. Carpintero // MLP / HE-10665 (MLP).
</paragraph>
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<caption id= "082047A1765A0EA81EAE44446E387616" httpUri= "https://binary.pensoft.net/fig/841070" pageId= "0" pageNumber= "117" start= "Figure 1" startId= "F1" >
<paragraph id= "D3654CEB410D812024F70DBC9CC68A6A" pageId= "0" pageNumber= "117" >
<emphasis id= "31AB3D6B8F2EEE48D97BF07F263CAE3C" bold= "true" pageId= "0" pageNumber= "117" > Figure 1.</emphasis>
<taxonomicName id= "6284DAA6ADF0EEC042F2397A383893E0" authorityName= "Dellapé & Melo" authorityYear= "2023" class= "Insecta" family= "Rhyparochromidae" genus= "Malleusocoris" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Malleusocoris minimus" order= "Hemiptera" pageId= "0" pageNumber= "117" phylum= "Arthropoda" rank= "species" species= "minimus" >
<emphasis id= "654F694A4572DA2A36F0608D38C6C392" italics= "true" pageId= "0" pageNumber= "117" > Malleusocoris minimus</emphasis>
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</taxonomicName>
gen. nov., sp. nov. Habitus of male holotype. Scale bar: 1 mm.
</paragraph>
</caption>
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<subSubSection id= "BE8363239E05F7104F93B268A6F8DBF2" pageId= "0" pageNumber= "117" type= "description" >
<paragraph id= "EDEC1788C43E6887C7FB1A18FC9AF443" pageId= "0" pageNumber= "117" > Description.</paragraph>
<paragraph id= "96BBF9E692B1EC3D6AD73E27C7C7432E" pageId= "0" pageNumber= "117" >
<emphasis id= "4E8E3EE3D63D9DA1E418073437C79A13" bold= "true" pageId= "0" pageNumber= "117" > Holotype male</emphasis>
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(Fig.
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<figureCitation id= "C4F6B2CF915D1C51534D6FE5381462C3" captionStart= "Figure 1" captionStartId= "F1" captionText= "Figure 1. Malleusocoris minimus gen. nov., sp. nov. Habitus of male holotype. Scale bar: 1 mm." httpUri= "https://binary.pensoft.net/fig/841070" pageId= "0" pageNumber= "117" > 1</figureCitation>
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). Total length 2.84. Head length 0.48, head width 0.58, interocular space 0.32, interocellar space 0.18; antennal segments length: scape 0.22, pedicel 0.32, basiflagellomere 0.20, distiflagellomere 0.50, scape width 0.08, distiflagellomere width 0.12. Anterior pronotal lobe length 0.32, width 0.60, posterior pronotal lobe length 0.24, width 0.92.
</paragraph>
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<paragraph id= "762D627ECB58573D5C0CF55D0C327C04" pageId= "0" pageNumber= "117" >
<emphasis id= "8EBA81BBEADEC50E08FC5795D618DD33" bold= "true" italics= "true" pageId= "0" pageNumber= "117" > Head</emphasis>
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brown, with abundant short, whitish, decumbent setae. Antenniferous tubercles subparallel.
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<emphasis id= "A37FDA2C2B6EA585C9C6BA21257EA10D" bold= "true" italics= "true" pageId= "0" pageNumber= "117" > Antennae</emphasis>
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: Scape surpassing apex of head by about half its length; pedicel and basiflagellomere widened distally; scape, pedicel and basiflagellomere light brown, distiflagellomere fusiform and enlarged; much darker and slightly paler apically; with abundant short whitish decumbent setae.
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<emphasis id= "DC12BFE13D11DC52274876AB2056C733" bold= "true" italics= "true" pageId= "0" pageNumber= "117" > Labium</emphasis>
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attaining mesosternum, segment I extending to half the length of eyes, remote from prosternum.
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<emphasis id= "CBE074542F0D075299F7F2E96A5AC6E0" bold= "true" pageId= "0" pageNumber= "117" >
<emphasis id= "456CB93E0D25EBE112FE85551B00E11F" italics= "true" pageId= "0" pageNumber= "117" > Thorax</emphasis>
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.
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<emphasis id= "8941C21D0F7D0DEE46619FA8238BF86F" italics= "true" pageId= "0" pageNumber= "117" > Pronotum</emphasis>
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</emphasis>
with a faint median carina, more evident on transverse impression. Anterior pronotal lobe brown, posterior lobe paler with five longitudinal darker stripes, median macula reaching transverse impression; with abundant short whitish decumbent setae.
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<emphasis id= "0D1911FEB8E8C688EE5160A68BD4B336" bold= "true" italics= "true" pageId= "0" pageNumber= "117" > Scutellum</emphasis>
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with a median carina on posterior half; coarsely punctate; brown with apex whitish; with same setae as on pronotum.
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<emphasis id= "C3FCB77129F119C181F7E5FD80383988" bold= "true" italics= "true" pageId= "0" pageNumber= "117" > Hemelytra</emphasis>
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irregularly pale brown, with paler margins interrupted by a pale brown spot at level of apex of claval commissure, apex of corium pale brown; with whitish setae on punctures, shorter than those of head, pronotum and scutellum.
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<emphasis id= "E7A1D0BA15249CDC6867AA3D600471FB" bold= "true" italics= "true" pageId= "0" pageNumber= "117" > Legs</emphasis>
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light brown, with short whitish setae. Forefemur with a few minute spines in inner row.
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<emphasis id= "27DFC94C0732E4D7D2BE8239CB5F6A12" bold= "true" pageId= "0" pageNumber= "117" >
<emphasis id= "9A508E3BAFF287B0561AAD0C0F2CCD0A" italics= "true" pageId= "0" pageNumber= "117" > Abdomen</emphasis>
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.
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<emphasis id= "8C48032C1865355013DCBA40712B418C" italics= "true" pageId= "0" pageNumber= "117" > Male genitalia</emphasis>
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</emphasis>
: Pygophore (Figs
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<figureCitation id= "B97E1E282F6251CC71ADFCBC5D9336FC" captionStart= "Figures 2– 5" captionStartId= "F2" captionText= "Figures 2 - 5. Malleusocoris minimus sp. nov. Male genitalia, of male holotype. 2. Pygophore, dorsal view; 3. Pygophore, lateral view; 4. Right paramere, outer view; 5. Right paramere, inner view. Scale bars: 0.25 mm." httpUri= "https://binary.pensoft.net/fig/841071" pageId= "0" pageNumber= "117" > 2</figureCitation>
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,
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<figureCitation id= "1411E49961FAB57B592A7B43AC514D44" captionStart= "Figures 2– 5" captionStartId= "F2" captionText= "Figures 2 - 5. Malleusocoris minimus sp. nov. Male genitalia, of male holotype. 2. Pygophore, dorsal view; 3. Pygophore, lateral view; 4. Right paramere, outer view; 5. Right paramere, inner view. Scale bars: 0.25 mm." httpUri= "https://binary.pensoft.net/fig/841071" pageId= "0" pageNumber= "117" > 3</figureCitation>
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) broadly rounded in dorsal view, aperture quadrangular anterior to the sub-quadrangular inner projections; declivent posteriorly in lateral view. Parameres (Figs
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<figureCitation id= "B09FF66A21EC735C4AAE257516235995" captionStart= "Figures 2– 5" captionStartId= "F2" captionText= "Figures 2 - 5. Malleusocoris minimus sp. nov. Male genitalia, of male holotype. 2. Pygophore, dorsal view; 3. Pygophore, lateral view; 4. Right paramere, outer view; 5. Right paramere, inner view. Scale bars: 0.25 mm." httpUri= "https://binary.pensoft.net/fig/841071" pageId= "0" pageNumber= "117" > 4</figureCitation>
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,
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<figureCitation id= "5562E21AC211122066120EA344E0F358" captionStart= "Figures 2– 5" captionStartId= "F2" captionText= "Figures 2 - 5. Malleusocoris minimus sp. nov. Male genitalia, of male holotype. 2. Pygophore, dorsal view; 3. Pygophore, lateral view; 4. Right paramere, outer view; 5. Right paramere, inner view. Scale bars: 0.25 mm." httpUri= "https://binary.pensoft.net/fig/841071" pageId= "0" pageNumber= "117" > 5</figureCitation>
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) with long and curved blade.
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<emphasis id= "7409011531778AF5C1B1C1F699499D9A" bold= "true" italics= "true" pageId= "0" pageNumber= "117" > Aedeagus</emphasis>
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weakly sclerotized, conjunctiva and vesica unspined, ejaculatory reservoir well developed, wings well projected laterally.
</paragraph>
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<caption id= "704CB5ED7C59FED93250E988799D3EF9" httpUri= "https://binary.pensoft.net/fig/841071" pageId= "0" pageNumber= "117" start= "Figures 2– 5" startId= "F2" >
<paragraph id= "4F773887F5F98554F509794BD85BAED7" pageId= "0" pageNumber= "117" >
<emphasis id= "9DE0ECCCDABD1B3CDAE9998FBB2EC8CE" bold= "true" pageId= "0" pageNumber= "117" > Figures 2-5.</emphasis>
<taxonomicName id= "6A4570E917B3D3E6DFB534DE46F2CC27" authorityName= "Dellapé & Melo" authorityYear= "2023" class= "Insecta" family= "Rhyparochromidae" genus= "Malleusocoris" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Malleusocoris minimus" order= "Hemiptera" pageId= "0" pageNumber= "117" phylum= "Arthropoda" rank= "species" species= "minimus" >
<emphasis id= "1F7138A45EBFA06D87CC893D8B369A02" italics= "true" pageId= "0" pageNumber= "117" > Malleusocoris minimus</emphasis>
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</taxonomicName>
sp. nov. Male genitalia, of male holotype.
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<emphasis id= "762997978FE1732BE68B163DFCA4384C" bold= "true" pageId= "0" pageNumber= "117" > 2.</emphasis>
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Pygophore, dorsal view;
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<emphasis id= "A4B2729B85CF08B101EEFD7C5B8B5181" bold= "true" pageId= "0" pageNumber= "117" > 3.</emphasis>
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Pygophore, lateral view;
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<emphasis id= "512ADF5F00F1A704D050AC6D358A79DB" bold= "true" pageId= "0" pageNumber= "117" > 4.</emphasis>
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Right paramere, outer view;
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<emphasis id= "A17B3F354739E470F6D45938F62ABE5B" bold= "true" pageId= "0" pageNumber= "117" > 5.</emphasis>
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Right paramere, inner view. Scale bars: 0.25 mm.
</paragraph>
</caption>
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<paragraph id= "5F9AE7275A49DEB96495F32BCB865761" pageId= "0" pageNumber= "117" > Paratypes similar to holotype.</paragraph>
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</subSubSection>
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<subSubSection id= "AA093B8250985698E3ABE5AF4465A291" pageId= "0" pageNumber= "117" type= "description" >
<paragraph id= "8800D870C6386EEC4AC7473CC533C804" pageId= "0" pageNumber= "117" >
<normalizedToken id= "682BB1C116C746B93F7FF760DB0EA947" originalValue= "Paratypes’ " > Paratypes'</normalizedToken>
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measurements
</paragraph>
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<paragraph id= "9CF2EAF881E466DD5BB92BB0C20B6316" pageId= "0" pageNumber= "117" >
<emphasis id= "424D280B88649A5DDA587FE85D977F65" bold= "true" pageId= "0" pageNumber= "117" > (min-max, mean). Males</emphasis>
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(n = 3): Total length 2.76-2.84, 2.80. Head length 0.46-0.50, 0.49, head width 0.56-0.60, 0.58, interocular width 0.30-0.32, 0.31, interocellar width 0.18-0.20, 0.19. Antennal segments length: scape 0.20-0.22, 0.21, pedicel 0.30-0.34, 0.32, basiflagellomere 0.18-0.20, 0.19, distiflagellomere 0.42-0.48, 0.46; scape width 0.06-0.08, 0.07, distiflagellomere width 0.12. Anterior pronotal lobe length 0.30-0.34, 0.31, width 0.60-0.66, 0.63; posterior pronotal lobe length, 0.26-0.28, 0.27; width 0.90-0.96, 0.93.
</paragraph>
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<paragraph id= "81A7217D5D6950B28EA90A85A57DFACB" pageId= "0" pageNumber= "117" >
<emphasis id= "C4BD13101E7E650FB446EC322689E572" bold= "true" pageId= "0" pageNumber= "117" > Females</emphasis>
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(n = 5): Total length 2.92-3.32, 3.11. Head length 0.48-0.56, 0.54, head width 0.62-0.64, 0.64, interocular width 0.34-0.36, 0.34, interocellar width 0.20-0.24, 0.21. Antennal segments length: scape 0.22-0.26, 0.24, pedicel 0.32-0.38, 0.34, basiflagellomere 0.18-0.20, 0.19, distiflagellomere 0.48-0.50, 0.49; scape width 0.08, distiflagellomere width 0.12-0.14, 0.13. Anterior pronotal lobe length 0.30-0.38, 0.34, width 0.66-0.72, 0.69; posterior pronotal lobe length, 0.28-0.30, 0.29; width 1.00-1.08, 1.03.
</paragraph>
</subSubSection>
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<subSubSection id= "93F2A3EE79A0A150D0483FDCFBE2C949" pageId= "0" pageNumber= "117" type= "etymology" >
<paragraph id= "672E83373E93B76F065449AFCDE9DA73" pageId= "0" pageNumber= "117" > Etymology.</paragraph>
<paragraph id= "605F38886F29223FCC76F9CE6EA0F279" pageId= "0" pageNumber= "117" >
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The specific epithet
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<emphasis id= "C1E2FB953FE6DA804CF56608E2AAE05F" italics= "true" pageId= "0" pageNumber= "117" > minimus</emphasis>
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is an adjective and refers to the small size of this new species.
</paragraph>
</subSubSection>
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<subSubSection id= "16A2E83C6FD4B965DC4D4415AF3F07E3" pageId= "0" pageNumber= "117" type= "distribution" >
<paragraph id= "4CBD6EBCBB66960CA8ED5C1736B98276" pageId= "0" pageNumber= "117" > Distribution.</paragraph>
<paragraph id= "76E871DDF3BFA796317A615DC689F4CF" pageId= "0" pageNumber= "117" >
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Brazil, Mato Grosso, Pantanal; and Argentina, Misiones, PN
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<normalizedToken id= "243AB21D3D90960B320AADEFDC5B17D2" originalValue= "Iguazú" > Iguazu</normalizedToken>
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.
</paragraph>
</subSubSection>
</treatment>
</document>