treatments-xml/data/03/C7/7E/03C77E5CFF9F6858FF0AFEBBFB1F3F2B.xml

<document id="3BFD2550E17E7F727459B8C642F9A2E0" ID="10.11646/zootaxa.4244.2.4SLASH10561" ID-CLB-Dataset="36240" ID-DOI="10.11646/zootaxa.4244.2.4" ID-GBIF-Dataset="7d329373-9dc5-4e22-b9b6-11060962e0ca" ID-ISSN="1175-5326" ID-Zenodo-Dep="400866" ID-ZooBank="E82A88DA-82A4-42E5-8ED0-4D9CB6EB77C5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1489999543611" checkinUser="plazi" docAuthor="White, William T., Last, Peter R. &amp; Mana, Ralph R." docDate="2017" docId="03C77E5CFF9F6858FF0AFEBBFB1F3F2B" docLanguage="en" docName="zootaxa.4244.2.4SLASH10561.pdf" docOrigin="Zootaxa 4244 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Notoraja sereti White, Last &amp; Mana, 2017, n. sp." docType="treatment" docVersion="7" lastPageNumber="229" masterDocId="FFFE0624FF9E6852FF9DFFF0FFC13A7E" masterDocTitle="A new species of velvet skate, Notoraja sereti n. sp. (Rajiformes: Arhynchobatidae) from Papua New Guinea" masterLastPageNumber="230" masterPageNumber="219" pageNumber="220" updateTime="1698438156648" updateUser="ExternalLinkService">
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<mods:title id="AFD973BFA465ADBD4126A8D2A1C66669">A new species of velvet skate, Notoraja sereti n. sp. (Rajiformes: Arhynchobatidae) from Papua New Guinea</mods:title>
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<mods:namePart id="5BCBA11D93A7DA8266ECDB16FC431A10">White, William T.</mods:namePart>
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<mods:namePart id="2123DCE636AC6E97E0005243F79F4037">Last, Peter R.</mods:namePart>
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<mods:namePart id="B19107F3A1CDD827D9E3419A3D1D0DF1">Mana, Ralph R.</mods:namePart>
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<mods:date id="B093197767F9FD6B142DD9FB3B5EF72C">2017</mods:date>
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<treatment id="03C77E5CFF9F6858FF0AFEBBFB1F3F2B" ID-DOI="http://doi.org/10.5281/zenodo.6052754" ID-GBIF-Taxon="127848045" ID-Zenodo-Dep="6052754" LSID="urn:lsid:plazi:treatment:03C77E5CFF9F6858FF0AFEBBFB1F3F2B" httpUri="http://treatment.plazi.org/id/03C77E5CFF9F6858FF0AFEBBFB1F3F2B" lastPageId="10" lastPageNumber="229" pageId="1" pageNumber="220">
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<heading id="D0997826FF9F6853FF0AFEBBFE553B18" bold="true" box="[151,404,331,358]" fontSize="11" level="1" pageId="1" pageNumber="220" reason="1">
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<taxonomicName id="4C6EB4C9FF9F6853FF0AFEBBFE8A3B1B" ID-CoL="74BHH" box="[151,331,331,357]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="1" pageNumber="220" phylum="Chordata" rank="species" species="sereti" status="sp. nov.">
<emphasis id="B91A1358FF9F6853FF0AFEBBFE8A3B1B" bold="true" box="[151,331,331,357]" italics="true" pageId="1" pageNumber="220">Notoraja sereti</emphasis>
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<taxonomicNameLabel id="A229AE23FF9F6853FECFFEBCFE553B18" box="[338,404,332,358]" pageId="1" pageNumber="220" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="8BD1CF4AFF9F6853FF0AFE9FFE9B3BD5" blockId="1.[151,404,331,427]" pageId="1" pageNumber="220">
Papuan Velvet Skate 
<figureCitation id="1355D3CFFF9F6853FF0AFE62FF3A3BD5" box="[151,251,402,427]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStartId-0="1.[151,250,1787,1809]" captionStartId-1="2.[151,250,1888,1910]" captionStartId-2="3.[151,250,788,810]" captionStartId-3="3.[151,250,1532,1554]" captionStartId-4="4.[151,250,1750,1772]" captionStartId-5="6.[151,250,1564,1586]" captionStartId-6="6.[151,250,1826,1848]" captionStartId-7="7.[151,250,1842,1864]" captionTargetBox-0="[224,1370,833,1751]" captionTargetBox-1="[158,1419,497,1827]" captionTargetBox-2="[340,1247,193,766]" captionTargetBox-3="[166,1411,909,1502]" captionTargetBox-4="[343,1227,249,1677]" captionTargetBox-5="[340,1247,891,1543]" captionTargetBox-6="[340,1230,1650,1794]" captionTargetBox-7="[339,1246,381,1811]" captionTargetId-0="figure@1.[188,1398,798,1766]" captionTargetId-1="figure@2.[154,1432,481,1867]" captionTargetId-2="figure@3.[340,1247,193,766]" captionTargetId-3="figure@3.[151,1436,890,1511]" captionTargetId-4="figure@4.[304,1283,223,1728]" captionTargetId-5="figure@6.[340,1247,891,1543]" captionTargetId-6="figure@6.[340,1247,1636,1804]" captionTargetId-7="figure@7.[319,1267,365,1839]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="3" captionTargetPageId-4="4" captionTargetPageId-5="6" captionTargetPageId-6="6" captionTargetPageId-7="7" captionText-0="FIGURE 1 Dorsal view of the holotype of Notoraja sereti n. sp. (NTUM 10067, female 459 mm TL) immediately postcapture." captionText-1="FIGURE 2. Preserved holotype of Notoraja sereti n. sp. (NTUM 10067, female 459 mm TL): (A) dorsal view; (B) ventral view." captionText-2="FIGURE 3. Dorsal view of the adolescent male paratype of Notoraja sereti n. sp. (NTUM 10330, 363 mm TL) immediately post-capture." captionText-3="FIGURE 4. Dorsal view of the adolescent male paratype of Notoraja sereti n. sp. (NTUM 10330, 363 mm TL) after preservation." captionText-4="FIGURE 5. Dorsal head of Notoraja sereti n. sp.: (A) holotype, NTUM 10067, female 459 mm TL; (B) paratype, NTUM 10330, subadult male 363 mm TL." captionText-5="FIGURE 6. Oronasal region of the holotype of Notoraja sereti n. sp. (NTUM 10067, female 459 mm TL)." captionText-6="FIGURE 7. Dorsal view of the tail tip of the holotype of Notoraja sereti n. sp. (NTUM 10067, female 459 mm TL)." captionText-7="FIGURE 8. Dermal denticles of the holotype of Notoraja sereti n. sp. (NTUM 10067, female 459 mm TL): (A) dorsal surface; (B) ventral surface)." httpUri-0="https://zenodo.org/record/400867/files/figure.png" httpUri-1="https://zenodo.org/record/400868/files/figure.png" httpUri-2="https://zenodo.org/record/400869/files/figure.png" httpUri-3="https://zenodo.org/record/400870/files/figure.png" httpUri-4="https://zenodo.org/record/400871/files/figure.png" httpUri-5="https://zenodo.org/record/400872/files/figure.png" httpUri-6="https://zenodo.org/record/400873/files/figure.png" httpUri-7="https://zenodo.org/record/400874/files/figure.png" pageId="1" pageNumber="220">Figs 1–8</figureCitation>
; 
<tableCitation id="C6ECFAF1FF9F6853FE9AFE62FE9B3BD5" box="[263,346,402,427]" captionStart="TABLE 1" captionStartId="5.[151,239,508,531]" captionText="TABLE 1. Morphometric data for the three type specimens of Notoraja sereti n. sp. from Papua New Guinea." pageId="1" pageNumber="220">Table 1</tableCitation>
</paragraph>
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<taxonomicName id="4C6EB4C9FF9F6853FF0AFE2BFEC73B8E" ID-CoL="54JY" box="[151,262,475,496]" class="Elasmobranchii" family="Arhynchobatidae" genus="Insentiraja" kingdom="Animalia" order="Rajiformes" pageId="1" pageNumber="220" phylum="Chordata" rank="genus">Insentiraja</taxonomicName>
subtilispinosa—
</emphasis>
<treatmentCitation id="0ACFE95BFF9F6853FE32FE2AFD533B8E" author="Fricke" box="[431,658,474,496]" page="15" pageId="1" pageNumber="220" year="2014">
<bibRefCitation id="EFFFB2BBFF9F6853FE32FE2AFDAD3B8E" author="Fricke" box="[431,620,474,496]" pageId="1" pageNumber="220" refString="Fricke, R., Allen, G. R., Andrefouet, S., Chen, W. J., Hame, M. A., Laboute, P., Mana, R., Hui, T. H. &amp; Uyeno, D. (2014) Checklist of the marine and estuarine fishes of Madang District, Papua New Guinea, western Pacific Ocean, with 820 new records. Zootaxa, 3832 (1), 1 - 247." type="journal article" year="2014">
Fricke 
<emphasis id="B91A1358FF9F6853FE6BFE2BFDEB3B8E" box="[502,554,475,496]" italics="true" pageId="1" pageNumber="220">et al.</emphasis>
, 2014
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: 15
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.
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</paragraph>
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<materialsCitation id="3B06C517FF9F6853FF0AFDEDFB813848" ID-GBIF-Occurrence="1440684142" box="[151,1088,541,566]" collectionCode="NTUM" country="Iceland" location="Madang" pageId="1" pageNumber="220" specimenCode="NTUM 10067" specimenCount="1" stateProvince="Madang" typeStatus="holotype">
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<typeStatus id="54D571E8FF9F6853FF0AFDEDFEC7384B" box="[151,262,541,565]" pageId="1" pageNumber="220" type="holotype">Holotype</typeStatus>
.
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<specimenCode id="DBC86731FF9F6853FE88FDEEFE7A3848" box="[277,443,541,566]" collectionCode="NTUM" pageId="1" pageNumber="220">
<collectionCode id="ED7F578FFF9F6853FE88FDEEFEA9384B" box="[277,360,542,565]" pageId="1" pageNumber="220">NTUM</collectionCode>
10067
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, female 
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TL, southeast of Sek 
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, 
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, 
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<collectingCountry id="F3798FDAFF9F6853FBD2FDEEFAF33848" box="[1103,1330,541,566]" name="Papua New Guinea" pageId="1" pageNumber="220">Papua New Guinea</collectingCountry>
, 
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, 
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, 
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cruise, R/ 
<collectorName id="269BAA9CFF9F6853FDBAFDB1FDB23824" box="[551,627,577,602]" pageId="1" pageNumber="220">V Alis</collectorName>
, station CP 
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<quantity id="4C9662AFFF9F6853FD69FDB1FCBE3827" box="[756,895,577,602]" metricMagnitude="6" metricUnit="m" metricValue="3.96798" pageId="1" pageNumber="220" unit="m" value="3967980.0">3967, 980 m</quantity>
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depth, 
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<date id="FFD0E98AFF9F6853FC4FFDB1FB5F3827" box="[978,1182,577,602]" pageId="1" pageNumber="220" value="2012-12-02">2 December 2012</date>
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.
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<materialsCitation id="3B06C517FF9F6853FF5AFD95FBFE3800" ID-GBIF-Occurrence="1440684146" box="[199,1087,613,638]" collectionCode="NTUM" country="Iceland" location="Madang" pageId="1" pageNumber="220" specimenCode="NTUM 10042" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<emphasis id="B91A1358FF9F6853FF5AFD95FE873803" bold="true" box="[199,326,613,637]" pageId="1" pageNumber="220">
<typeStatus id="54D571E8FF9F6853FF5AFD95FE803803" box="[199,321,613,637]" pageId="1" pageNumber="220" type="paratype">Paratypes</typeStatus>
.
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<specimenCode id="DBC86731FF9F6853FECDFD96FE363800" box="[336,503,614,638]" collectionCode="NTUM" pageId="1" pageNumber="220">
<collectionCode id="ED7F578FFF9F6853FECDFD96FE623803" box="[336,419,614,637]" pageId="1" pageNumber="220">NTUM</collectionCode>
10042
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, female 
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TL, east of Sek 
<collectingCountry id="F3798FDAFF9F6853FC17FD95FC103800" box="[906,977,613,638]" name="Iceland" pageId="1" pageNumber="220">Island</collectingCountry>
, 
<collectingRegion id="49AA01A8FF9F6853FC7CFD95FBFE3800" box="[993,1087,613,638]" country="Papua New Guinea" name="Madang" pageId="1" pageNumber="220">Madang</collectingRegion>
</materialsCitation>
, 
<materialsCitation id="3B06C517FF9F6853FBD3FD96FB2B38DC" ID-GBIF-Occurrence="1440684143" collectingDate="2012-11-30" collectorName="Papua Niugini &amp; V Alis" country="Papua New Guinea" elevation="980" latitude="-5.1" location="Papua New Guinea" longLatPrecision="1305" longitude="145.88333" pageId="1" pageNumber="220" specimenCount="1" typeStatus="paratype">
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, 
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, 
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, 
<collectorName id="269BAA9CFF9F6853FE89FD7AFE7538DC" box="[276,436,649,674]" pageId="1" pageNumber="220">Papua Niugini</collectorName>
cruise, R/ 
<collectorName id="269BAA9CFF9F6853FDB5FD7AFDB538DC" box="[552,628,649,674]" pageId="1" pageNumber="220">V Alis</collectorName>
, station CP3963, 960– 
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<quantity id="4C9662AFFF9F6853FCEEFD79FC7B38DF" box="[883,954,649,674]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="1" pageNumber="220" unit="m" value="980.0">980 m</quantity>
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depth, 
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<date id="FFD0E98AFF9F6853FB93FD7AFB2B38DC" box="[1038,1258,649,674]" pageId="1" pageNumber="220" value="2012-11-30">30 November 2012</date>
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; 
<materialsCitation id="3B06C517FF9F6853FB68FD7AFC7E3894" ID-GBIF-Occurrence="1440684145" collectingDate="2012-12-14" collectionCode="NTUM" collectorName="Papua Niugini &amp; V Alis" country="Papua New Guinea" elevation="820" latitude="-5.35" location="Astrolabe Bay" longLatPrecision="1305" longitude="145.88333" pageId="1" pageNumber="220" specimenCode="NTUM 10330" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<specimenCode id="DBC86731FF9F6853FB68FD7AFA5938DC" box="[1269,1432,650,674]" collectionCode="NTUM" pageId="1" pageNumber="220">
<collectionCode id="ED7F578FFF9F6853FB68FD7AFA8938DF" box="[1269,1352,650,673]" pageId="1" pageNumber="220">NTUM</collectionCode>
10330
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, adolescent male 
<quantity id="4C9662AFFF9F6853FEC4FD5DFE7B38BB" box="[345,442,685,710]" metricMagnitude="-1" metricUnit="m" metricValue="3.63" pageId="1" pageNumber="220" unit="mm" value="363.0">363 mm</quantity>
TL, 
<location id="8EB19991FF9F6853FE65FD5DFD5F38B8" LSID="urn:lsid:plazi:treatment:03C77E5CFF9F6858FF0AFEBBFB1F3F2B:8EB19991FF9F6853FE65FD5DFD5F38B8" box="[504,670,685,710]" country="Papua New Guinea" latitude="-5.35" longLatPrecision="1305" longitude="145.88333" name="Astrolabe Bay" pageId="1" pageNumber="220" stateProvince="Madang">Astrolabe Bay</location>
, 
<collectingRegion id="49AA01A8FF9F6853FD30FD5DFCCD38B8" box="[685,780,685,710]" country="Papua New Guinea" name="Madang" pageId="1" pageNumber="220">Madang</collectingRegion>
, 
<collectingCountry id="F3798FDAFF9F6853FC86FD5EFC3F38B8" box="[795,1022,685,710]" name="Papua New Guinea" pageId="1" pageNumber="220">Papua New Guinea</collectingCountry>
, 
<geoCoordinate id="EE5AA98DFF9F6853FB90FD5DFBA238B8" box="[1037,1123,685,710]" direction="south" orientation="latitude" pageId="1" pageNumber="220" precision="925" value="-5.35">5°21' S</geoCoordinate>
, 
<geoCoordinate id="EE5AA98DFF9F6853FBEEFD5DFB2938BB" box="[1139,1256,685,710]" direction="east" orientation="longitude" pageId="1" pageNumber="220" precision="925" value="145.88333">145°53' E</geoCoordinate>
, 
<collectorName id="269BAA9CFF9F6853FB6AFD5EFA5D38B8" box="[1271,1436,685,710]" pageId="1" pageNumber="220">Papua Niugini</collectorName>
cruise, R/ 
<collectorName id="269BAA9CFF9F6853FE9EFD22FE8F3894" box="[259,334,721,746]" pageId="1" pageNumber="220">V Alis</collectorName>
, station CP4027, 800– 
<elevation id="00432879FF9F6853FDD1FD22FD523897" box="[588,659,722,746]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="1" pageNumber="220" unit="m" value="820.0">
<quantity id="4C9662AFFF9F6853FDD1FD22FD523897" box="[588,659,722,746]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="1" pageNumber="220" unit="m" value="820.0">820 m</quantity>
</elevation>
depth, 
<collectingDate id="EF941062FF9F6853FD7BFD22FC7E3894" box="[742,959,721,746]" pageId="1" pageNumber="220" value="2012-12-14">
<date id="FFD0E98AFF9F6853FD7BFD22FC7E3894" box="[742,959,721,746]" pageId="1" pageNumber="220" value="2012-12-14">14 December 2012</date>
</collectingDate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF119FC2FF9F6853FF0AF90BFF283D4E" httpUri="https://zenodo.org/record/400867/files/figure.png" pageId="1" pageNumber="220" targetBox="[224,1370,833,1751]" targetPageId="1">
<paragraph id="8BD1CF4AFF9F6853FF0AF90BFF283D4E" blockId="1.[151,1436,1787,1840]" pageId="1" pageNumber="220">
<emphasis id="B91A1358FF9F6853FF0AF90BFECE3D6F" bold="true" box="[151,271,1787,1809]" pageId="1" pageNumber="220">FIGURE 1</emphasis>
Dorsal view of the holotype of 
<taxonomicName id="4C6EB4C9FF9F6853FDF4F90CFCC43D6F" box="[617,773,1788,1809]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="1" pageNumber="220" phylum="Chordata" rank="species" species="sereti" status="sp. nov.">
<emphasis id="B91A1358FF9F6853FDF4F90CFCC43D6F" box="[617,773,1788,1809]" italics="true" pageId="1" pageNumber="220">Notoraja sereti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229AE23FF9F6853FC92F90BFC8D3D6F" box="[783,844,1787,1809]" pageId="1" pageNumber="220" rank="species">
<emphasis id="B91A1358FF9F6853FC92F90BFC8D3D6F" bold="true" box="[783,844,1787,1809]" pageId="1" pageNumber="220">n. sp.</emphasis>
</taxonomicNameLabel>
(NTUM 10067, female 459 mm TL) immediately postcapture.
</paragraph>
</caption>
<subSubSection id="C3749CC1FF9F6850FF5AF8ACFB5E3BD3" lastPageId="2" lastPageNumber="221" pageId="1" pageNumber="220" type="diagnosis">
<paragraph id="8BD1CF4AFF9F6850FF5AF8ACFB5E3BD3" blockId="1.[151,1436,1884,2018]" lastBlockId="2.[151,1437,151,429]" lastPageId="2" lastPageNumber="221" pageId="1" pageNumber="220">
<emphasis id="B91A1358FF9F6853FF5AF8ACFE803D0B" bold="true" box="[199,321,1884,1909]" pageId="1" pageNumber="220">Diagnosis.</emphasis>
A medium-size species of 
<taxonomicName id="4C6EB4C9FF9F6853FDE8F8AEFD1B3D0B" box="[629,730,1886,1909]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="1" pageNumber="220" phylum="Chordata" rank="genus">
<emphasis id="B91A1358FF9F6853FDE8F8AEFD1B3D0B" box="[629,730,1886,1909]" italics="true" pageId="1" pageNumber="220">Notoraja</emphasis>
</taxonomicName>
with the following combination of characters: disc wider than long, width 47.1–50.3% TL, length 42.0–43.6% TL; short head (dorsal head length 15.2–16.2% TL; ventral head length 21.6–22.9% TL); relatively short snout (preorbital length 10.1–11.1% TL; ventral snout length 11.2–12.1% TL); interspiracular distance 5.3–5.9 TL, internasal distance 6.2–6.8% TL, preorbital length 2.4–3.0 times orbit length and 2.9–3.6 times interorbital; two dorsal fins, interdorsal space 1.1–1.9% TL; tail width at pelvic-fin axil 0.9-1.6 times its height; single, small preorbital thorn; dorsal and most of ventral surface of disc entirely velvety, covered with fine denticles; median disc and pelvic fins naked in adolescent male; anterior pelvic-fin lobe naked in females (skin naked or with sparse denticles ventrally on mid belly, chin, nasal curtain, and around cloaca); tail long and slender, entirely velvety and without thorn-like denticles; lateral tail folds moderately well developed; nasal lobes expanded, width of nasal curtain 7.9–8.3% TL; anterior pelvic-fin lobe slightly shorter or similar length to posterior lobe; dorsal and ventral surfaces greyish brown to dark bluish grey; total pectoral-fin radials 58–60; monospondylous centra 23–25, total diplospondylous centra 103–106, total centra 126–131.
</paragraph>
</subSubSection>
<caption id="DF119FC2FF9C6850FF0AF890FF0F3DEB" httpUri="https://zenodo.org/record/400868/files/figure.png" pageId="2" pageNumber="221" targetBox="[158,1419,497,1827]" targetPageId="2">
<paragraph id="8BD1CF4AFF9C6850FF0AF890FF0F3DEB" blockId="2.[151,1435,1888,1941]" pageId="2" pageNumber="221">
<emphasis id="B91A1358FF9C6850FF0AF890FED53D08" bold="true" box="[151,276,1888,1910]" pageId="2" pageNumber="221">FIGURE 2.</emphasis>
Preserved holotype of 
<taxonomicName id="4C6EB4C9FF9C6850FD9AF891FD633D08" box="[519,674,1889,1910]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="221" phylum="Chordata" rank="species" species="sereti" status="sp. nov.">
<emphasis id="B91A1358FF9C6850FD9AF891FD633D08" box="[519,674,1889,1910]" italics="true" pageId="2" pageNumber="221">Notoraja sereti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229AE23FF9C6850FD36F890FD273D08" box="[683,742,1888,1910]" pageId="2" pageNumber="221" rank="species">
<emphasis id="B91A1358FF9C6850FD36F890FD273D08" bold="true" box="[683,742,1888,1910]" pageId="2" pageNumber="221">n. sp.</emphasis>
</taxonomicNameLabel>
(NTUM 10067, female 459 mm TL): (A) dorsal view; (B) ventral view.
</paragraph>
</caption>
<caption id="DF119FC2FF9D6851FF0AFCE4FEDA3936" httpUri="https://zenodo.org/record/400869/files/figure.png" pageId="3" pageNumber="222" targetBox="[340,1247,193,766]" targetPageId="3">
<paragraph id="8BD1CF4AFF9D6851FF0AFCE4FEDA3936" blockId="3.[151,1436,788,840]" pageId="3" pageNumber="222">
<emphasis id="B91A1358FF9D6851FF0AFCE4FED23954" bold="true" box="[151,275,788,810]" pageId="3" pageNumber="222">FIGURE 3.</emphasis>
Dorsal view of the adolescent male paratype of 
<taxonomicName id="4C6EB4C9FF9D6851FC95FCE5FC633954" box="[776,930,789,810]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="3" pageNumber="222" phylum="Chordata" rank="species" species="sereti" status="sp. nov.">
<emphasis id="B91A1358FF9D6851FC95FCE5FC633954" box="[776,930,789,810]" italics="true" pageId="3" pageNumber="222">Notoraja sereti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229AE23FF9D6851FC34FCE4FC253954" box="[937,996,788,810]" pageId="3" pageNumber="222" rank="species">
<emphasis id="B91A1358FF9D6851FC34FCE4FC253954" bold="true" box="[937,996,788,810]" pageId="3" pageNumber="222">n. sp.</emphasis>
</taxonomicNameLabel>
(NTUM 10330, 363 mm TL) immediately post-capture.
</paragraph>
</caption>
<caption id="DF119FC2FF9D6851FF0AFA0CFEDA3C4F" httpUri="https://zenodo.org/record/400870/files/figure.png" pageId="3" pageNumber="222" targetBox="[166,1411,909,1502]" targetPageId="3">
<paragraph id="8BD1CF4AFF9D6851FF0AFA0CFEDA3C4F" blockId="3.[151,1435,1532,1585]" pageId="3" pageNumber="222">
<emphasis id="B91A1358FF9D6851FF0AFA0CFED93C6C" bold="true" box="[151,280,1532,1554]" pageId="3" pageNumber="222">FIGURE 4.</emphasis>
Dorsal view of the adolescent male paratype of 
<taxonomicName id="4C6EB4C9FF9D6851FCAFFA0DFC113C6C" box="[818,976,1533,1554]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="3" pageNumber="222" phylum="Chordata" rank="species" species="sereti" status="sp. nov.">
<emphasis id="B91A1358FF9D6851FCAFFA0DFC113C6C" box="[818,976,1533,1554]" italics="true" pageId="3" pageNumber="222">Notoraja sereti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229AE23FF9D6851FC41FA0CFBDB3C6C" box="[988,1050,1532,1554]" pageId="3" pageNumber="222" rank="species">
<emphasis id="B91A1358FF9D6851FC41FA0CFBDB3C6C" bold="true" box="[988,1050,1532,1554]" pageId="3" pageNumber="222">n. sp.</emphasis>
</taxonomicNameLabel>
(NTUM 10330, 363 mm TL) after preservation.
</paragraph>
</caption>
<subSubSection id="C3749CC1FF9D685AFF5AF9ADFB80388E" lastPageId="8" lastPageNumber="227" pageId="3" pageNumber="222" type="description">
<paragraph id="8BD1CF4AFF9D6851FF5AF9ADFF2E3D31" blockId="3.[151,1437,1629,2015]" pageId="3" pageNumber="222">
<emphasis id="B91A1358FF9D6851FF5AF9ADFE983C08" bold="true" box="[199,345,1629,1654]" pageId="3" pageNumber="222">Description.</emphasis>
Disc heart-shaped, 1.17 (1.10–1.12) times as broad as long; maximum angle in front of spiracles 102° (96–100°); anterior margin nearly straight in females, weakly undulate in adolescent male 
<typeStatus id="54D571E8FF9D6851FB78F973FA883CE5" box="[1253,1353,1667,1691]" pageId="3" pageNumber="222" type="paratype">paratype</typeStatus>
; apices broadly rounded; posterior margin moderately convex. Axis of greatest width 62% (61–64%) of disc length. Preorbital snout length 2.43 (2.98–2.99) times orbit length, 2.93 (3.28-3.58) times interorbital width; preoral snout length 1.76 (1.81–1.91) times internarial distance. Orbit diameter 1.21 (1.10–1.20) times interorbital distance, and 2.04 (1.70–1.78) times length of spiracles. Nasal lobes moderately expanded and rounded, posterior margin weakly fringed.
</paragraph>
<paragraph id="8BD1CF4AFF9D6851FF5AF8AAFB9D3DA1" blockId="3.[151,1437,1629,2015]" pageId="3" pageNumber="222">Mouth wide, its width more than 80 (80–81)% of maximum width of nasal curtain. Upper jaw weakly indented at symphysis; upper and lower jaws slightly convex on either side of symphysis. Teeth with relatively long, pointed cusps arranged in quincunx. Distance between first gill slits 1.82 (1.83) times larger than distance between nostrils; distance between fifth gill slits 1.22 (1.07) times larger than distance between nostrils.</paragraph>
<caption id="DF119FC2FF9A6856FF0AF926FE373D74" httpUri="https://zenodo.org/record/400871/files/figure.png" pageId="4" pageNumber="223" targetBox="[343,1227,249,1677]" targetPageId="4">
<paragraph id="8BD1CF4AFF9A6856FF0AF926FE373D74" blockId="4.[151,1436,1750,1802]" pageId="4" pageNumber="223">
<emphasis id="B91A1358FF9A6856FF0AF926FECE3C92" bold="true" box="[151,271,1750,1772]" pageId="4" pageNumber="223">FIGURE 5</emphasis>
. Dorsal head of 
<taxonomicName id="4C6EB4C9FF9A6856FE5DF927FD9D3C92" box="[448,604,1751,1772]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="4" pageNumber="223" phylum="Chordata" rank="species" species="sereti" status="sp. nov.">
<emphasis id="B91A1358FF9A6856FE5DF927FD9D3C92" box="[448,604,1751,1772]" italics="true" pageId="4" pageNumber="223">Notoraja sereti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229AE23FF9A6856FDF8F926FD613C92" box="[613,672,1750,1772]" pageId="4" pageNumber="223" rank="species">
<emphasis id="B91A1358FF9A6856FDF8F926FD613C92" bold="true" box="[613,672,1750,1772]" pageId="4" pageNumber="223">n. sp.</emphasis>
</taxonomicNameLabel>
: (A) holotype, NTUM 10067, female 459 mm TL; (B) paratype, NTUM 10330, subadult male 363 mm TL.
</paragraph>
</caption>
<paragraph id="8BD1CF4AFF9A6857FF5AF8C2FE8D3BAE" blockId="4.[151,1437,1842,2011]" lastBlockId="5.[151,1436,151,464]" lastPageId="5" lastPageNumber="224" pageId="4" pageNumber="223">Pelvic fins deeply incised with lobes connected by radials and membranes, anterior lobe moderately long, reaching to about posterior quarter of posterior lobe in females, finger-like with blunt tip (longer and more acute in adolescent male); posterior lobe with convex lateral margins, its posterior margin crenate due to extension of posterior radials. Tail narrowly oval at base, variably depressed over length; moderately convex dorsally, very weakly convex ventrally; tapering gradually posteriorly; tail width at axils of pelvic fins 1.69 (1.50–2.57) times width at its midlength, 2.29 (2.48–3.72) times its width at dorsal-fin origin respectively; length from rear of cloaca 1.45 (1.41–1.51) times distance from tip of snout to rear of cloaca; width at first dorsal-fin origin 2.13 (1.30–1.40) times its height; lateral skin folds originating at about anterior third of tail, extending to distal half of epichordal caudal-fin lobe; folds slightly broadening distally, but always narrower than tail width. Dorsal fins strongly raked, of similar shape and size; relatively short and rather low with convex anterior margins, straight or slightly convex posterior margins and broadly rounded rear tips; separated by a short interspace, 50% (36–60%) length of first dorsal-fin base. Epichordal caudal-fin lobe developed, separated by narrow interspace from and distinctly longer than second dorsal-fin base; hypochordal caudal lobe very low, originating near end of lateral fold, confluent with epichordal lobe.</paragraph>
<caption id="DF119FC2FF9B6857FF0AFE0CFAE4386D" box="[151,1317,508,531]" pageId="5" pageNumber="224">
<paragraph id="8BD1CF4AFF9B6857FF0AFE0CFAE4386D" blockId="5.[151,1317,508,531]" box="[151,1317,508,531]" pageId="5" pageNumber="224">
<emphasis id="B91A1358FF9B6857FF0AFE0CFECB386D" bold="true" box="[151,266,508,531]" pageId="5" pageNumber="224">TABLE 1.</emphasis>
Morphometric data for the three type specimens of 
<taxonomicName id="4C6EB4C9FF9B6857FCA9FE0CFC14386D" box="[820,981,508,531]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="224" phylum="Chordata" rank="species" species="sereti" status="sp. nov.">
<emphasis id="B91A1358FF9B6857FCA9FE0CFC14386D" box="[820,981,508,531]" italics="true" pageId="5" pageNumber="224">Notoraja sereti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229AE23FF9B6857FC41FE0CFBCE386D" box="[988,1039,508,531]" pageId="5" pageNumber="224" rank="species">n.sp.</taxonomicNameLabel>
from Papua New Guinea.
</paragraph>
</caption>
<paragraph id="8BD1CF4AFF9B6857FC95FDDEFB3039C6" blockId="5.[159,1428,558,2027]" pageId="5" pageNumber="224">
<typeStatus id="54D571E8FF9B6857FC95FDDEFCA0383A" box="[776,865,558,580]" pageId="5" pageNumber="224" type="holotype">holotype</typeStatus>
<typeStatus id="54D571E8FF9B6857FC41FDDFFAE9383A" box="[988,1320,559,580]" pageId="5" pageNumber="224" type="paratype">paratype paratype</typeStatus>
female subadult male female Total length (mm) 459 363 458 Disc width 50.3 47.1 47.8 Disc length (direct) 43.0 42.0 43.6 Snout to maximum width 26.6 25.7 27.9 Snout length (preorbital direct) 10.1 10.6 11.1 Snout to spiracle 15.0 15.0 14.8 Head (dorsal length) 15.2 16.0 16.2 Orbit diameter 4.2 3.6 3.7
</paragraph>
<paragraph id="8BD1CF4AFF9B6857FF02FC3CFB30399C" blockId="5.[159,1428,558,2027]" box="[159,1265,972,994]" pageId="5" pageNumber="224">Orbit and spiracle length 4.8 4.9 4.6</paragraph>
<paragraph id="8BD1CF4AFF9B6857FF02FC05FB303E75" blockId="5.[159,1428,558,2027]" box="[159,1265,1013,1035]" pageId="5" pageNumber="224">Spiracle length (main pore) 2.0 2.1 2.1</paragraph>
<paragraph id="8BD1CF4AFF9B6857FF02FBEEFB303E4A" blockId="5.[159,1428,558,2027]" box="[159,1265,1054,1076]" pageId="5" pageNumber="224">Distance between orbits 3.5 3.0 3.4</paragraph>
<paragraph id="8BD1CF4AFF9B6857FF02FBB8FB303E20" blockId="5.[159,1428,558,2027]" box="[159,1265,1096,1118]" pageId="5" pageNumber="224">Distance between spiracles 5.8 5.3 5.9</paragraph>
<paragraph id="8BD1CF4AFF9B6857FF02FB81FB303F01" blockId="5.[159,1428,558,2027]" pageId="5" pageNumber="224">Distance-snout to cloaca 40.9 39.8 41.0 Cloaca to D1 47.3 46.6 47.0 Cloaca to D2 51.7 51.1 50.4 Cloaca to caudal origin 55.4 55.7 53.8 Distance-cloaca to caudal-fin tip 59.4 60.1 58.0 Ventral snout length (pre upper jaw) 11.9 11.2 12.1 Prenasal length 8.7 8.2 8.9</paragraph>
<paragraph id="8BD1CF4AFF9B6857FF02FA62FB303FAC" blockId="5.[159,1428,558,2027]" pageId="5" pageNumber="224">Ventral head length (to fifth gill) 21.6 22.2 22.9 Mouth width 6.7 6.4 6.6</paragraph>
<paragraph id="8BD1CF4AFF9B6857FF02FA15FB303F85" blockId="5.[159,1428,558,2027]" box="[159,1265,1509,1531]" pageId="5" pageNumber="224">Distance between nostrils 6.8 6.2 6.3</paragraph>
<paragraph id="8BD1CF4AFF9B6857FF02F9FEFB303C5A" blockId="5.[159,1428,558,2027]" box="[159,1265,1550,1572]" pageId="5" pageNumber="224">Nasal curtain length 4.3 4.6 4.7</paragraph>
<paragraph id="8BD1CF4AFF9B6857FF02F9C8FB303C30" blockId="5.[159,1428,558,2027]" box="[159,1265,1592,1614]" pageId="5" pageNumber="224">Nasal curtain (total width) 8.3 7.9 8.2</paragraph>
<paragraph id="8BD1CF4AFF9B6857FF02F991FB303C09" blockId="5.[159,1428,558,2027]" box="[159,1265,1633,1655]" pageId="5" pageNumber="224">Nasal curtain (min width) 4.4 3.9 4.1</paragraph>
<paragraph id="8BD1CF4AFF9B6857FF02F97AFB303CDE" blockId="5.[159,1428,558,2027]" box="[159,1265,1674,1696]" pageId="5" pageNumber="224">Nasal curtain (lobe width) 2.1 2.0 2.2</paragraph>
<paragraph id="8BD1CF4AFF9B6857FF02F944FB303CB4" blockId="5.[159,1428,558,2027]" box="[159,1265,1716,1738]" pageId="5" pageNumber="224">Width of first gill opening 1.5 1.3 1.4</paragraph>
<paragraph id="8BD1CF4AFF9B6857FF02F92DFB303C8D" blockId="5.[159,1428,558,2027]" box="[159,1265,1757,1779]" pageId="5" pageNumber="224">Width of fifth gill opening 1.5 0.9 1.0</paragraph>
<paragraph id="8BD1CF4AFF9B6857FF02F8F6FB1F3D62" blockId="5.[159,1428,558,2027]" box="[159,1246,1798,1820]" pageId="5" pageNumber="224">Distance between first gill openings 12.4 11.4 –</paragraph>
<paragraph id="8BD1CF4AFF9B6857FF02F8C0FB1F3D38" blockId="5.[159,1428,558,2027]" box="[159,1246,1840,1862]" pageId="5" pageNumber="224">Distance between fifth gill openings 8.3 6.7 –</paragraph>
<paragraph id="8BD1CF4AFF9B6857FF02F8A9FB1F3D11" blockId="5.[159,1428,558,2027]" box="[159,1246,1881,1903]" pageId="5" pageNumber="224">Clasper (post cloacal length) – 16.4 –</paragraph>
<paragraph id="8BD1CF4AFF9B6857FF02F872FB3C3DBC" blockId="5.[159,1428,558,2027]" pageId="5" pageNumber="224">Length of anterior pelvic lobe 12.7 13.7 13.1 Length of posterior pelvic lobe 13.4 13.7 13.5</paragraph>
<paragraph id="8BD1CF4AFF9B6855FBC3F825FA923A86" blockId="5.[159,1428,558,2027]" lastBlockId="7.[151,1436,151,320]" lastPageId="7" lastPageNumber="226" pageId="5" pageNumber="224">
<emphasis id="B91A1358FF9B6857FBC3F825FA553D95" box="[1118,1428,2005,2027]" italics="true" pageId="5" pageNumber="224">......continued on the next page</emphasis>
<typeStatus id="54D571E8FF986854FC95FF39FCA03AA1" box="[776,865,201,223]" pageId="6" pageNumber="225" type="holotype">holotype</typeStatus>
<typeStatus id="54D571E8FF986854FC41FF3AFAE93AA1" box="[988,1320,202,223]" pageId="6" pageNumber="225" type="paratype">paratype paratype</typeStatus>
female subadult male female Pelvic base width 8.4 7.4 7.6 Tail at axil pelvic fins (width) 3.6 3.9 2.7 Tail at axil pelvic fins (height) 2.6 2.4 2.9 Tail at midlength (width) 2.1 1.5 1.8 Tail at midlength (height) 1.3 1.3 1.3 Tail at D1 origin (width) 1.6 1.0 1.1 Tail at 0 1 origin (height) 0.7 0.7 0.8 D1 base length 3.3 3.2 3.1 D1 height 1.5 1.2 1.5 D1 origin to caudal-fin tip 12.3 13.8 12.0 D2 origin to caudal-fin tip 7.7 8.9 8.2 Caudal-fin length 4.4 4.6 4.0 lnterdorsal distance 1.7 1.9 1.1 Dorsal surface of disc velvety, entirely covered with fine, densely spaced, dermal denticles; denticles bristlelike, erect, with almost straight pointed crowns. Ventral surface of disc similarly covered on most surfaces, except median disc (oronasal region, branchial region and belly, 
<figureCitation id="1355D3CFFF986854FC84F857FCA83DBE" box="[793,873,1959,1984]" captionStart="FIGURE 10" captionStartId="9.[151,250,1934,1956]" captionTargetBox="[229,1362,199,1898]" captionTargetId="figure@9.[210,1377,193,1913]" captionTargetPageId="9" captionText="FIGURE 10. Denticle pattern in Notoraja sereti n. sp. highlighting areas lacking denticles: (A) ventral head of adolescent male paratype (NTUM 10330, 363 mm TL); (B) belly of female paratype (NTUM 10042, 458 mm TL)." httpUri="https://zenodo.org/record/400876/files/figure.png" pageId="6" pageNumber="225">Fig. 10</figureCitation>
b) and entire pelvic fins of adolescent male naked; anterior pelvic-fin lobe naked in females (skin naked or with sparse denticles ventrally on mid belly, chin, nasal curtain, and around cloaca). A single, small, but conspicuous preorbital thorn present; no other orbital thorns or thorns on disc of 
<typeStatus id="54D571E8FF996855FEC1FF4DFE5B3AAB" box="[348,410,189,213]" pageId="7" pageNumber="226">types</typeStatus>
. Tail velvety, densely covered with fine, dermal denticles; no enlarged thorns; a few small, variably spaced thornlets on midline of tail but these very small, barely taller than largest denticles adjacent.
</paragraph>
<caption id="DF119FC2FF986854FF02FF50FE4C3AC8" box="[159,397,160,182]" pageId="6" pageNumber="225">
<paragraph id="8BD1CF4AFF986854FF02FF50FE4C3AC8" blockId="6.[159,1320,160,802]" box="[159,397,160,182]" pageId="6" pageNumber="225">
<emphasis id="B91A1358FF986854FF02FF50FECD3AC8" bold="true" box="[159,268,160,182]" pageId="6" pageNumber="225">TABLE 1.</emphasis>
(Continued)
</paragraph>
</caption>
<caption id="DF119FC2FF986854FF0AF9ECFB153C4C" box="[151,1236,1564,1586]" httpUri="https://zenodo.org/record/400872/files/figure.png" pageId="6" pageNumber="225" targetBox="[340,1247,891,1543]" targetPageId="6">
<paragraph id="8BD1CF4AFF986854FF0AF9ECFB153C4C" blockId="6.[151,1236,1564,1586]" box="[151,1236,1564,1586]" pageId="6" pageNumber="225">
<emphasis id="B91A1358FF986854FF0AF9ECFED33C4C" bold="true" box="[151,274,1564,1586]" pageId="6" pageNumber="225">FIGURE 6.</emphasis>
Oronasal region of the holotype of 
<taxonomicName id="4C6EB4C9FF986854FDE1F9EDFCD43C4C" box="[636,789,1565,1586]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="6" pageNumber="225" phylum="Chordata" rank="species" species="sereti" status="sp. nov.">
<emphasis id="B91A1358FF986854FDE1F9EDFCD43C4C" box="[636,789,1565,1586]" italics="true" pageId="6" pageNumber="225">Notoraja sereti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229AE23FF986854FC81F9ECFC943C4C" box="[796,853,1564,1586]" pageId="6" pageNumber="225" rank="species">
<emphasis id="B91A1358FF986854FC81F9ECFC943C4C" bold="true" box="[796,853,1564,1586]" pageId="6" pageNumber="225">n. sp.</emphasis>
</taxonomicNameLabel>
(NTUM 10067, female 459 mm TL).
</paragraph>
</caption>
<caption id="DF119FC2FF986854FF0AF8D2FAF43D46" box="[151,1333,1826,1848]" httpUri="https://zenodo.org/record/400873/files/figure.png" pageId="6" pageNumber="225" targetBox="[340,1230,1650,1794]" targetPageId="6">
<paragraph id="8BD1CF4AFF986854FF0AF8D2FAF43D46" blockId="6.[151,1333,1826,1848]" box="[151,1333,1826,1848]" pageId="6" pageNumber="225">
<emphasis id="B91A1358FF986854FF0AF8D2FECC3D46" bold="true" box="[151,269,1826,1848]" pageId="6" pageNumber="225">FIGURE 7</emphasis>
. Dorsal view of the tail tip of the holotype of 
<taxonomicName id="4C6EB4C9FF986854FD40F8D3FCB73D46" box="[733,886,1827,1848]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="6" pageNumber="225" phylum="Chordata" rank="species" species="sereti" status="sp. nov.">
<emphasis id="B91A1358FF986854FD40F8D3FCB73D46" box="[733,886,1827,1848]" italics="true" pageId="6" pageNumber="225">Notoraja sereti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229AE23FF986854FCE0F8D2FC773D46" box="[893,950,1826,1848]" pageId="6" pageNumber="225" rank="species">
<emphasis id="B91A1358FF986854FCE0F8D2FC773D46" bold="true" box="[893,950,1826,1848]" pageId="6" pageNumber="225">n. sp.</emphasis>
</taxonomicNameLabel>
(NTUM 10067, female 459 mm TL).
</paragraph>
</caption>
<paragraph id="8BD1CF4AFF996855FF5AFEF5FE9C3B41" blockId="7.[151,1436,151,320]" pageId="7" pageNumber="226">
No adult males available; adolescent male 
<typeStatus id="54D571E8FF996855FD37FEF5FCCB3B63" box="[682,778,261,285]" pageId="7" pageNumber="226" type="paratype">paratype</typeStatus>
with relatively long and very slender clasper, post cloacal length 16.4% TL.
</paragraph>
<caption id="DF119FC2FF996855FF0AF8C2FEA53D19" httpUri="https://zenodo.org/record/400874/files/figure.png" pageId="7" pageNumber="226" targetBox="[339,1246,381,1811]" targetPageId="7">
<paragraph id="8BD1CF4AFF996855FF0AF8C2FEA53D19" blockId="7.[151,1436,1842,1895]" pageId="7" pageNumber="226">
<emphasis id="B91A1358FF996855FF0AF8C2FED33D36" bold="true" box="[151,274,1842,1864]" pageId="7" pageNumber="226">FIGURE 8.</emphasis>
Dermal denticles of the holotype of 
<taxonomicName id="4C6EB4C9FF996855FD19F8C3FCDD3D36" box="[644,796,1843,1864]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="7" pageNumber="226" phylum="Chordata" rank="species" species="sereti" status="sp. nov.">
<emphasis id="B91A1358FF996855FD19F8C3FCDD3D36" box="[644,796,1843,1864]" italics="true" pageId="7" pageNumber="226">Notoraja sereti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229AE23FF996855FCBEF8C2FC9A3D36" box="[803,859,1842,1864]" pageId="7" pageNumber="226" rank="species">
<emphasis id="B91A1358FF996855FCBEF8C2FC9A3D36" bold="true" box="[803,859,1842,1864]" pageId="7" pageNumber="226">n. sp.</emphasis>
</taxonomicNameLabel>
(NTUM 10067, female 459 mm TL): (A) dorsal surface; (B) ventral surface).
</paragraph>
</caption>
<paragraph id="8BD1CF4AFF99685AFF5AF864FF103AAA" blockId="7.[151,1436,1940,2037]" lastBlockId="8.[151,1436,151,789]" lastPageId="8" lastPageNumber="227" pageId="7" pageNumber="226">Tooth rows in upper jaw 43 (39–41); lower jaw 39 (37). Pectoral propterygial radials 27 (26–27); mesopterygial radials 11 (10-11); metapterygial radials 20–21 (22-23); total radials 58–59 (58–60). Monospondylous centra 25 (23–24); diplospondylous predorsal centra 70 (66–71); predorsal centra 95 (90–94); post-first dorsal-fin origin centra 36 (32–38); total diplospondylous centra 106 (103–104); total centra 131 (126– 128).</paragraph>
<paragraph id="8BD1CF4AFF96685AFF5AFF2FFF2C381E" blockId="8.[151,1436,151,789]" pageId="8" pageNumber="227">
<emphasis id="B91A1358FF96685AFF5AFF2FFEE13A86" bold="true" box="[199,288,223,248]" pageId="8" pageNumber="227">Colour.</emphasis>
<typeStatus id="54D571E8FF96685AFEBAFF2FFE4E3A86" box="[295,399,223,248]" pageId="8" pageNumber="227" type="holotype">Holotype</typeStatus>
and female 
<typeStatus id="54D571E8FF96685AFD81FF10FDBD3A86" box="[540,636,224,248]" pageId="8" pageNumber="227" type="paratype">paratype</typeStatus>
(fresh): Dorsal surface uniformly medium brown, slightly paler above orbits and at pectoral insertions; anterior third of tail darker brown with a narrow, darker brownish median stripe originating anterior to dark brown marking (about level with pelvic-fin insertions); dorsal and caudal fins medium brown with darker edges. Ventral surface of disc similar in colour to dorsal disc; anterior third of tail slightly darker brown. When preserved, dorsal coloration slightly paler brown; dark brown dorsal midline obvious on anterior tail, but darker anterior marking on tail more obvious. Adolescent male 
<typeStatus id="54D571E8FF96685AFC18FE65FC243BD3" box="[901,997,405,429]" pageId="8" pageNumber="227" type="paratype">paratype</typeStatus>
(fresh): dorsal surface bluish grey with paler anterior pelvic-fin lobes; most of dorsal midline and tail distinctly paler except near tip. When preserved, dorsal coloration dark brown, darkest distally near disc apices; dark midline on anterior tail obvious; dark marking on anterior tail less obvious; ventral surface of disc and tail dark, slightly paler than dorsal surface in adolescent male but similar in females; adolescent male translucent on belly and branchial region, noticeably darker near disc margin.
</paragraph>
<paragraph id="8BD1CF4AFF96685AFF5AFD9BFB80388E" blockId="8.[151,1436,151,789]" pageId="8" pageNumber="227">
<emphasis id="B91A1358FF96685AFF5AFD9BFF3F38FA" bold="true" box="[199,254,619,644]" pageId="8" pageNumber="227">Size.</emphasis>
Known from the three 
<typeStatus id="54D571E8FF96685AFD9FFD9DFDF338FB" box="[514,562,621,645]" pageId="8" pageNumber="227">type</typeStatus>
specimens consisting of two females (459 and 
<quantity id="4C9662AFFF96685AFBDCFD9CFB5F38FA" box="[1089,1182,620,645]" metricMagnitude="-1" metricUnit="m" metricValue="4.58" pageId="8" pageNumber="227" unit="mm" value="458.0">458 mm</quantity>
TL) and an adolescent male (
<quantity id="4C9662AFFF96685AFF43FD7FFEFF38D9" box="[222,318,655,680]" metricMagnitude="-1" metricUnit="m" metricValue="3.63" pageId="8" pageNumber="227" unit="mm" value="363.0">363 mm</quantity>
TL). The clasper of the adolescent male (postcloacal length 16.4% TL) was possibly about threequarters of full extension (maximum length of adult claspers of 
<taxonomicName id="4C6EB4C9FF96685AFCEFFD45FC3938B2" box="[882,1016,692,716]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="227" phylum="Chordata" rank="species" species="sapphira">
<emphasis id="B91A1358FF96685AFCEFFD45FC3938B2" box="[882,1016,692,716]" italics="true" pageId="8" pageNumber="227">N. sapphira</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C6EB4C9FF96685AFBACFD45FB5F38B2" box="[1073,1182,693,716]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="227" phylum="Chordata" rank="species" species="azurea">
<emphasis id="B91A1358FF96685AFBACFD45FB5F38B2" box="[1073,1182,693,716]" italics="true" pageId="8" pageNumber="227">N. azurea</emphasis>
</taxonomicName>
22.5% and 22.6% TL respectively). Based on the adolescent male size, the females are most likely adults.
</paragraph>
</subSubSection>
<subSubSection id="C3749CC1FF96685AFF5AFD0BFB86396A" box="[199,1095,763,789]" pageId="8" pageNumber="227" type="distribution">
<paragraph id="8BD1CF4AFF96685AFF5AFD0BFB86396A" blockId="8.[151,1436,151,789]" box="[199,1095,763,789]" pageId="8" pageNumber="227">
<emphasis id="B91A1358FF96685AFF5AFD0BFEA1396A" bold="true" box="[199,352,763,788]" pageId="8" pageNumber="227">Distribution.</emphasis>
Off 
<collectingRegion id="49AA01A8FF96685AFE09FD0CFE33396B" box="[404,498,764,789]" country="Papua New Guinea" name="Madang" pageId="8" pageNumber="227">Madang</collectingRegion>
, 
<collectingCountry id="F3798FDAFF96685AFE63FD0DFD16396B" box="[510,727,764,789]" name="Papua New Guinea" pageId="8" pageNumber="227">Papua New Guinea</collectingCountry>
at depths of 
<quantity id="4C9662AFFF96685AFCFAFD0CFC27396A" box="[871,998,764,788]" metricMagnitude="2" metricUnit="m" metricValue="8.9" metricValueMax="9.8" metricValueMin="8.0" pageId="8" pageNumber="227" unit="m" value="890.0" valueMax="980.0" valueMin="800.0">800–980 m</quantity>
(
<figureCitation id="1355D3CFFF96685AFC68FD0CFBF8396A" box="[1013,1081,764,789]" captionStart="FIGURE 9" captionStartId="8.[151,250,1862,1884]" captionTargetBox="[151,1435,841,1841]" captionTargetId="figure@8.[151,1436,841,1841]" captionTargetPageId="8" captionText="FIGURE 9. Map showing the capture locations of the type specimens of Notoraja sereti n. sp. (red star denotes holotype, yellow circles paratypes) in Astrolabe Bay, Papua New Guinea. Inset map shows location of main map in context of the whole country (Image © NASA, TerraMetrics, Google Earth)." httpUri="https://zenodo.org/record/400875/files/figure.png" pageId="8" pageNumber="227">Fig. 9</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF119FC2FF96685AFF0AF8B6FD0B3DE7" httpUri="https://zenodo.org/record/400875/files/figure.png" pageId="8" pageNumber="227" targetBox="[151,1435,841,1841]" targetPageId="8">
<paragraph id="8BD1CF4AFF96685AFF0AF8B6FD0B3DE7" blockId="8.[151,1436,1862,1945]" pageId="8" pageNumber="227">
<emphasis id="B91A1358FF96685AFF0AF8B6FECE3D22" bold="true" box="[151,271,1862,1884]" pageId="8" pageNumber="227">FIGURE 9</emphasis>
. Map showing the capture locations of the type specimens of 
<taxonomicName id="4C6EB4C9FF96685AFC03F8B7FBFB3D22" box="[926,1082,1863,1884]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="227" phylum="Chordata" rank="species" species="sereti" status="sp. nov.">
<emphasis id="B91A1358FF96685AFC03F8B7FBFB3D22" box="[926,1082,1863,1884]" italics="true" pageId="8" pageNumber="227">Notoraja sereti</emphasis>
</taxonomicName>
<emphasis id="B91A1358FF96685AFBDEF8B6FBBE3D22" bold="true" box="[1091,1151,1862,1884]" pageId="8" pageNumber="227">
<taxonomicNameLabel id="A229AE23FF96685AFBDEF8B6FBBE3D22" box="[1091,1151,1862,1884]" pageId="8" pageNumber="227" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(red star denotes holotype, yellow circles paratypes) in Astrolabe Bay, Papua New Guinea. Inset map shows location of main map in context of the whole country (Image © NASA, TerraMetrics, Google Earth).
</paragraph>
</caption>
<caption id="DF119FC2FF97685BFF0AF87EFBBA3DBD" httpUri="https://zenodo.org/record/400876/files/figure.png" pageId="9" pageNumber="228" targetBox="[229,1362,199,1898]" targetPageId="9">
<paragraph id="8BD1CF4AFF97685BFF0AF87EFBBA3DBD" blockId="9.[151,1435,1934,1987]" pageId="9" pageNumber="228">
<emphasis id="B91A1358FF97685BFF0AF87EFEDE3DDA" bold="true" box="[151,287,1934,1956]" pageId="9" pageNumber="228">FIGURE 10.</emphasis>
Denticle pattern in 
<taxonomicName id="4C6EB4C9FF97685BFE78F87FFDBF3DDA" box="[485,638,1935,1956]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="9" pageNumber="228" phylum="Chordata" rank="species" species="sereti" status="sp. nov.">
<emphasis id="B91A1358FF97685BFE78F87FFDBF3DDA" box="[485,638,1935,1956]" italics="true" pageId="9" pageNumber="228">Notoraja sereti</emphasis>
</taxonomicName>
<emphasis id="B91A1358FF97685BFD19F87EFD7C3DDA" bold="true" box="[644,701,1934,1956]" pageId="9" pageNumber="228">
<taxonomicNameLabel id="A229AE23FF97685BFD19F87EFD7C3DDA" box="[644,701,1934,1956]" pageId="9" pageNumber="228" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
highlighting areas lacking denticles: (A) ventral head of adolescent male paratype (NTUM 10330, 363 mm TL); (B) belly of female paratype (NTUM 10042, 458 mm TL).
</paragraph>
</caption>
<subSubSection id="C3749CC1FF946858FF5AFF68FE5838FB" pageId="10" pageNumber="229" type="etymology">
<paragraph id="8BD1CF4AFF946858FF5AFF68FDEC3A86" blockId="10.[151,1437,151,1365]" pageId="10" pageNumber="229">
<emphasis id="B91A1358FF946858FF5AFF68FE8F3ACE" bold="true" box="[199,334,152,176]" pageId="10" pageNumber="229">Etymology.</emphasis>
Named after the highly respected French ichthyologist, Dr. Bernard Séret, who has contributed greatly to the taxonomy of sharks and rays, and in particular to our knowledge of skates of the genus 
<taxonomicName id="4C6EB4C9FF946858FAADFF4DFA543AAA" box="[1328,1429,189,212]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="229" phylum="Chordata" rank="genus">
<emphasis id="B91A1358FF946858FAADFF4DFA543AAA" box="[1328,1429,189,212]" italics="true" pageId="10" pageNumber="229">Notoraja</emphasis>
</taxonomicName>
. English name: Papuan Velvet Skate.
</paragraph>
<paragraph id="8BD1CF4AFF946858FF5AFEF3FE5838FB" blockId="10.[151,1437,151,1365]" pageId="10" pageNumber="229">
<emphasis id="B91A1358FF946858FF5AFEF3FEAC3B62" bold="true" box="[199,365,259,284]" pageId="10" pageNumber="229">Comparisons.</emphasis>
The new species is assigned to the genus 
<taxonomicName id="4C6EB4C9FF946858FCD1FEF5FC733B62" box="[844,946,261,284]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="229" phylum="Chordata" rank="genus">
<emphasis id="B91A1358FF946858FCD1FEF5FC733B62" box="[844,946,261,284]" italics="true" pageId="10" pageNumber="229">Notoraja</emphasis>
</taxonomicName>
which is particularly diverse in this part of the Indo–Pacific. The dorsal and most of the ventral surfaces of 
<taxonomicName id="4C6EB4C9FF946858FC15FED8FBF53B41" box="[904,1076,296,319]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="sereti">
<emphasis id="B91A1358FF946858FC15FED8FBF53B41" box="[904,1076,296,319]" italics="true" pageId="10" pageNumber="229">Notoraja sereti</emphasis>
</taxonomicName>
are entirely covered with fine, densely spaced, dermal denticles which is shared by the velcro skates, 
<taxonomicName id="4C6EB4C9FF946858FC57FEBDFAD63B1A" authority="Seret &amp; Last, 2012" authorityName="Seret &amp; Last" authorityYear="2012" box="[970,1303,332,356]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="alisae">
<emphasis id="B91A1358FF946858FC57FEBDFBEF3B1A" box="[970,1070,332,356]" italics="true" pageId="10" pageNumber="229">N. alisae</emphasis>
<bibRefCitation id="EFFFB2BBFF946858FBAAFEBCFAD63B1A" author="Seret" box="[1079,1303,332,356]" pageId="10" pageNumber="229" refString="Seret, B. &amp; Last, P. R. (2012) New deep water skates of the genus Notoraja Ishiyama, 1958 (Rajoidei, Arhynchobatidae) from the southwest Pacific. Zoosystema, 34 (2), 319 - 341." type="journal article" year="2012">Séret &amp; Last, 2012</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="4C6EB4C9FF946858FAB9FEBDFEB63BF6" authority="Seret &amp; Last, 2012" authorityName="Seret &amp; Last" authorityYear="2012" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="fijiensis">
<emphasis id="B91A1358FF946858FAB9FEBDFA5D3B1A" box="[1316,1436,332,356]" italics="true" pageId="10" pageNumber="229">N. fijiensis</emphasis>
<bibRefCitation id="EFFFB2BBFF946858FF0AFE80FEB63BF6" author="Seret" box="[151,375,368,392]" pageId="10" pageNumber="229" refString="Seret, B. &amp; Last, P. R. (2012) New deep water skates of the genus Notoraja Ishiyama, 1958 (Rajoidei, Arhynchobatidae) from the southwest Pacific. Zoosystema, 34 (2), 319 - 341." type="journal article" year="2012">Séret &amp; Last, 2012</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="4C6EB4C9FF946858FE1BFE81FD343BF6" authority="Seret &amp; Last, 2012" authorityName="Seret &amp; Last" authorityYear="2012" box="[390,757,368,392]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="inusitata">
<emphasis id="B91A1358FF946858FE1BFE81FDC83BF9" box="[390,521,368,392]" italics="true" pageId="10" pageNumber="229">N. inusitata</emphasis>
<bibRefCitation id="EFFFB2BBFF946858FD8EFE80FD343BF6" author="Seret" box="[531,757,368,392]" pageId="10" pageNumber="229" refString="Seret, B. &amp; Last, P. R. (2012) New deep water skates of the genus Notoraja Ishiyama, 1958 (Rajoidei, Arhynchobatidae) from the southwest Pacific. Zoosystema, 34 (2), 319 - 341." type="journal article" year="2012">Séret &amp; Last, 2012</bibRefCitation>
</taxonomicName>
; and 
<taxonomicName id="4C6EB4C9FF946858FCA8FE81FB203BF6" authority="Seret &amp; Last, 2012" authorityName="Seret &amp; Last" authorityYear="2012" box="[821,1249,368,392]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="longiventralis">
<emphasis id="B91A1358FF946858FCA8FE81FC373BF6" box="[821,1014,368,392]" italics="true" pageId="10" pageNumber="229">N. longiventralis</emphasis>
<bibRefCitation id="EFFFB2BBFF946858FC62FE80FB203BF6" author="Seret" box="[1023,1249,368,392]" pageId="10" pageNumber="229" refString="Seret, B. &amp; Last, P. R. (2012) New deep water skates of the genus Notoraja Ishiyama, 1958 (Rajoidei, Arhynchobatidae) from the southwest Pacific. Zoosystema, 34 (2), 319 - 341." type="journal article" year="2012">Séret &amp; Last, 2012</bibRefCitation>
</taxonomicName>
. In contrast, the remaining species in the genus, i.e. 
<taxonomicName id="4C6EB4C9FF946858FDB9FE65FC723BD2" authority="McEachran &amp; Last, 2008" authorityName="McEachran &amp; Last" authorityYear="2008" box="[548,947,404,429]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="azurea">
<emphasis id="B91A1358FF946858FDB9FE65FD4E3BD2" box="[548,655,405,428]" italics="true" pageId="10" pageNumber="229">N. azurea</emphasis>
<bibRefCitation id="EFFFB2BBFF946858FD0BFE64FC723BD2" author="McEachran" box="[662,947,404,429]" pageId="10" pageNumber="229" refString="McEachran, J. D. &amp; Last, P. R. (2008) New deepwater skates of the genus Notoraja (Rajoidei: Arhynchobatidae) from southern Australia and the eastern Indian Ocean. CSIRO Marine and Atmospheric Research Paper, No. 0 21, 155 - 172." type="journal article" year="2008">McEachran &amp; Last, 2008</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="4C6EB4C9FF946858FC23FE65FAB63BD3" authority="Last &amp; McEachran, 2006" authorityName="Last &amp; McEachran" authorityYear="2006" box="[958,1399,404,429]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="hirticauda">
<emphasis id="B91A1358FF946858FC23FE65FB923BD2" box="[958,1107,404,428]" italics="true" pageId="10" pageNumber="229">N. hirticauda</emphasis>
<bibRefCitation id="EFFFB2BBFF946858FBC7FE65FAB63BD3" author="Last" box="[1114,1399,404,429]" pageId="10" pageNumber="229" refString="Last, P. R. &amp; McEachran, J. D. (2006) Notoraja hirticauda, a new species of skate (Chondrichthyes: Rajoidei) from the southeastern Indian Ocean. Memoirs of the Museum of Uictoria, 63 (1), 65 - 75." type="journal article" year="2006">Last &amp; McEachran, 2006</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="4C6EB4C9FF946858FA1FFE65FE263BAE" authority="McEachran &amp; Last, 2008" authorityName="McEachran &amp; Last" authorityYear="2008" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="lira">
<emphasis id="B91A1358FF946858FA1FFE65FF013BAE" italics="true" pageId="10" pageNumber="229">N. lira</emphasis>
<bibRefCitation id="EFFFB2BBFF946858FF5BFE47FE263BAE" author="McEachran" box="[198,487,439,464]" pageId="10" pageNumber="229" refString="McEachran, J. D. &amp; Last, P. R. (2008) New deepwater skates of the genus Notoraja (Rajoidei: Arhynchobatidae) from southern Australia and the eastern Indian Ocean. CSIRO Marine and Atmospheric Research Paper, No. 0 21, 155 - 172." type="journal article" year="2008">McEachran &amp; Last, 2008</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="4C6EB4C9FF946858FE6FFE49FC003BAE" authority="McEachran &amp; Last, 1994" authorityName="McEachran &amp; Last" authorityYear="1994" box="[498,961,439,464]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="ochroderma">
<emphasis id="B91A1358FF946858FE6FFE49FD5A3BAE" box="[498,667,440,464]" italics="true" pageId="10" pageNumber="229">N. ochroderma</emphasis>
<bibRefCitation id="EFFFB2BBFF946858FD3FFE47FC003BAE" author="McEachran" box="[674,961,439,464]" pageId="10" pageNumber="229" refString="McEachran, J. D. &amp; Last, P. R. (1994) New species of skate, Notoraja ochroderma, from off Queensland, Australia, with comments on the taxonomic limits of Notoraja (Chondrichthyes, Rajoidei). Copeia, 1994 (No. 2), 413 - 421. https: // doi. org / 10.2307 / 1446989" type="journal article" year="1994">McEachran &amp; Last, 1994</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="4C6EB4C9FF946858FC53FE49FAF33BAE" authority="Seret &amp; Last, 2009" authorityName="Seret &amp; Last" authorityYear="2009" box="[974,1330,440,464]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="sapphira">
<emphasis id="B91A1358FF946858FC53FE49FB923BAE" box="[974,1107,440,464]" italics="true" pageId="10" pageNumber="229">N. sapphira</emphasis>
<bibRefCitation id="EFFFB2BBFF946858FBC7FE48FAF33BAE" author="Seret" box="[1114,1330,440,464]" pageId="10" pageNumber="229" refString="Seret, B. &amp; Last, P. R. (2009) Notoraja sapphira n. sp. (Rajoidei: Arhynchobatidae), a new deepwater skate from the slopes of the Norfolk Ridge (south-west Pacific). Zootaxa, 2153, 24 - 34." type="journal article" year="2009">Séret &amp; Last, 2009</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="4C6EB4C9FF946858FAA3FE49FE7D3B8A" authority="McEachran &amp; Last, 2008" authorityName="McEachran &amp; Last" authorityYear="2008" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="sticta">
<emphasis id="B91A1358FF946858FAA3FE49FA5D3BB1" box="[1342,1436,440,464]" italics="true" pageId="10" pageNumber="229">N. sticta</emphasis>
<bibRefCitation id="EFFFB2BBFF946858FF0AFE2CFE7D3B8A" author="McEachran" box="[151,444,476,501]" pageId="10" pageNumber="229" refString="McEachran, J. D. &amp; Last, P. R. (2008) New deepwater skates of the genus Notoraja (Rajoidei: Arhynchobatidae) from southern Australia and the eastern Indian Ocean. CSIRO Marine and Atmospheric Research Paper, No. 0 21, 155 - 172." type="journal article" year="2008">McEachran &amp; Last, 2008</bibRefCitation>
</taxonomicName>
; and 
<taxonomicName id="4C6EB4C9FF946858FE67FE2DFCFE3B8A" authority="Hiyama, 1940" authorityName="Hiyama" authorityYear="1940" box="[506,831,476,501]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="tobitukai">
<emphasis id="B91A1358FF946858FE67FE2DFD403B8A" box="[506,641,476,500]" italics="true" pageId="10" pageNumber="229">N. tobitukai</emphasis>
(
<bibRefCitation id="EFFFB2BBFF946858FD0CFE2CFCF63B8A" author="Hiyama" box="[657,823,476,501]" pageId="10" pageNumber="229" refString="Hiyama, Y. (1940) Descriptions of two new species of fish, Raja tobitukai and Chlorophthalmus acutifrons. Japanese Journal of Zoology, 9 (1), 169 - 173." type="journal article" year="1940">Hiyama, 1940</bibRefCitation>
)
</taxonomicName>
, all have a smooth ventral surface entirely devoid of denticles. However, it should be noted that the adolescent male has a poorer coverage of denticles on the ventral surface than the larger females (see e.g. 
<figureCitation id="1355D3CFFF946858FDF9FDD4FD773842" box="[612,694,548,573]" captionStart="FIGURE 10" captionStartId="9.[151,250,1934,1956]" captionTargetBox="[229,1362,199,1898]" captionTargetId="figure@9.[210,1377,193,1913]" captionTargetPageId="9" captionText="FIGURE 10. Denticle pattern in Notoraja sereti n. sp. highlighting areas lacking denticles: (A) ventral head of adolescent male paratype (NTUM 10330, 363 mm TL); (B) belly of female paratype (NTUM 10042, 458 mm TL)." httpUri="https://zenodo.org/record/400876/files/figure.png" pageId="10" pageNumber="229">Fig. 10</figureCitation>
a), and it is likely that denticle coverage on the ventral surface is size related. Juveniles were unavailable for study but they are likely to be less well spinulated (possibly naked) than adults and adolescents.
</paragraph>
</subSubSection>
<subSubSection id="C3749CC1FF946858FF5AFD60FB1F3F2B" pageId="10" pageNumber="229" type="description">
<paragraph id="8BD1CF4AFF946858FF5AFD60FB2239FE" blockId="10.[151,1437,151,1365]" pageId="10" pageNumber="229">
<taxonomicName id="4C6EB4C9FF946858FF5AFD60FEB038D9" box="[199,369,656,679]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="sereti">
<emphasis id="B91A1358FF946858FF5AFD60FEB038D9" box="[199,369,656,679]" italics="true" pageId="10" pageNumber="229">Notoraja sereti</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C6EB4C9FF946858FD95FD61FDAC38D6" box="[520,621,656,680]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="alisae">
<emphasis id="B91A1358FF946858FD95FD61FDAC38D6" box="[520,621,656,680]" italics="true" pageId="10" pageNumber="229">N. alisae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C6EB4C9FF946858FDE6FD61FD3538D6" box="[635,756,656,680]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="fijiensis">
<emphasis id="B91A1358FF946858FDE6FD61FD3538D6" box="[635,756,656,680]" italics="true" pageId="10" pageNumber="229">N. fijiensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C6EB4C9FF946858FC9EFD61FC4738D9" box="[771,902,656,680]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="inusitata">
<emphasis id="B91A1358FF946858FC9EFD61FC4738D9" box="[771,902,656,680]" italics="true" pageId="10" pageNumber="229">N. inusitata</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C6EB4C9FF946858FC23FD61FBBD38D6" box="[958,1148,656,680]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="longiventralis">
<emphasis id="B91A1358FF946858FC23FD61FBBD38D6" box="[958,1148,656,680]" italics="true" pageId="10" pageNumber="229">N. longiventralis</emphasis>
</taxonomicName>
in having a shorter snout (preorbital length 10.1–11.1 
<emphasis id="B91A1358FF946858FE45FD45FE2E38B2" box="[472,495,693,716]" italics="true" pageId="10" pageNumber="229">vs</emphasis>
. 11.5–14.5% TL, prenasal length 8.2–8.9 
<emphasis id="B91A1358FF946858FC55FD45FC1E38B2" box="[968,991,693,716]" italics="true" pageId="10" pageNumber="229">vs</emphasis>
. 9.8–12.1% TL); a shorter head (dorsal head length 15.2–16.2 
<emphasis id="B91A1358FF946858FE39FD29FE7A388E" box="[420,443,729,752]" italics="true" pageId="10" pageNumber="229">vs</emphasis>
. 17.1–19.3% TL, ventral head length 21.6–22.9 
<emphasis id="B91A1358FF946858FC66FD29FBD3388E" box="[1019,1042,729,752]" italics="true" pageId="10" pageNumber="229">vs</emphasis>
. 22.9–25.9% TL); fewer pectoral radials (total radials 58–60 
<emphasis id="B91A1358FF946858FE51FD0DFE22396A" box="[460,483,765,788]" italics="true" pageId="10" pageNumber="229">vs</emphasis>
. 61–74) and fewer vertebral centra (predorsal diplospondylous centra 66–71 
<emphasis id="B91A1358FF946858FAD1FD0DFAA2396A" box="[1356,1379,765,788]" italics="true" pageId="10" pageNumber="229">vs</emphasis>
. 72– 82; predorsal centra 90–95 
<emphasis id="B91A1358FF946858FE5AFCD1FE1F3946" box="[455,478,801,824]" italics="true" pageId="10" pageNumber="229">vs</emphasis>
. 98–107; total centra 126–131 
<emphasis id="B91A1358FF946858FCA4FCD1FC913946" box="[825,848,801,824]" italics="true" pageId="10" pageNumber="229">vs</emphasis>
. 135–152). It differs from 
<taxonomicName id="4C6EB4C9FF946858FBE7FCD1FB1C3946" box="[1146,1245,800,824]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="alisae">
<emphasis id="B91A1358FF946858FBE7FCD1FB1C3946" box="[1146,1245,800,824]" italics="true" pageId="10" pageNumber="229">N. alisae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C6EB4C9FF946858FB76FCD1FAAC3949" box="[1259,1389,800,824]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="inusitata">
<emphasis id="B91A1358FF946858FB76FCD1FAAC3949" box="[1259,1389,800,824]" italics="true" pageId="10" pageNumber="229">N. inusitata</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C6EB4C9FF946858FF0AFCB5FE993922" box="[151,344,836,860]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="longiventralis">
<emphasis id="B91A1358FF946858FF0AFCB5FE993922" box="[151,344,836,860]" italics="true" pageId="10" pageNumber="229">N. longiventralis</emphasis>
</taxonomicName>
in disc length (42.0–43.6 
<emphasis id="B91A1358FF946858FD0AFCB5FD6F3922" box="[663,686,837,860]" italics="true" pageId="10" pageNumber="229">vs</emphasis>
. 43.8–47.8% TL). It differs from 
<taxonomicName id="4C6EB4C9FF946858FBD4FCB5FB703922" box="[1097,1201,836,860]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="alisae">
<emphasis id="B91A1358FF946858FBD4FCB5FB703922" box="[1097,1201,836,860]" italics="true" pageId="10" pageNumber="229">N. alisae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C6EB4C9FF946858FB59FCB5FA803922" box="[1220,1345,836,860]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="fijiensis">
<emphasis id="B91A1358FF946858FB59FCB5FA803922" box="[1220,1345,836,860]" italics="true" pageId="10" pageNumber="229">N. fijiensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C6EB4C9FF946858FA1FFCB5FF383901" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="inusitata">
<emphasis id="B91A1358FF946858FA1FFCB5FF383901" italics="true" pageId="10" pageNumber="229">N. inusitata</emphasis>
</taxonomicName>
in having more widely separated dorsal fins (interdorsal space 1.1–1.9 
<emphasis id="B91A1358FF946858FB84FC99FBF139FE" box="[1049,1072,873,896]" italics="true" pageId="10" pageNumber="229">vs</emphasis>
. 0.5–0.9% TL).
</paragraph>
<paragraph id="8BD1CF4AFF946858FF5AFC7DFB973EBA" blockId="10.[151,1437,151,1365]" pageId="10" pageNumber="229">
<taxonomicName id="4C6EB4C9FF946858FF5AFC7DFEB139DA" box="[199,368,909,932]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="sereti">
<emphasis id="B91A1358FF946858FF5AFC7DFEB139DA" box="[199,368,909,932]" italics="true" pageId="10" pageNumber="229">Notoraja sereti</emphasis>
</taxonomicName>
further differs from 
<taxonomicName id="4C6EB4C9FF946858FDC5FC7DFD7D39DA" box="[600,700,908,932]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="alisae">
<emphasis id="B91A1358FF946858FDC5FC7DFD7D39DA" box="[600,700,908,932]" italics="true" pageId="10" pageNumber="229">N. alisae</emphasis>
</taxonomicName>
in the following characters: disc width 47.1–50.3 
<emphasis id="B91A1358FF946858FB72FC7DFAC739DA" box="[1263,1286,909,932]" italics="true" pageId="10" pageNumber="229">vs</emphasis>
. 50.9–55.5% TL; orbit and spiracle length 4.6–4.9 
<emphasis id="B91A1358FF946858FDA1FC41FD9239B6" box="[572,595,945,968]" italics="true" pageId="10" pageNumber="229">vs</emphasis>
. 5.1–5.5% TL; and distance between spiracles 5.3–5.9 
<emphasis id="B91A1358FF946858FB5FFC41FB1839B6" box="[1218,1241,945,968]" italics="true" pageId="10" pageNumber="229">vs</emphasis>
. 6.2–6.7% TL. It further differs from 
<taxonomicName id="4C6EB4C9FF946858FE1BFC25FDC33992" box="[390,514,980,1004]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="fijiensis">
<emphasis id="B91A1358FF946858FE1BFC25FDC33992" box="[390,514,980,1004]" italics="true" pageId="10" pageNumber="229">N. fijiensis</emphasis>
</taxonomicName>
in the following characters: ventral surface dark (
<emphasis id="B91A1358FF946858FBC7FC25FBB03992" box="[1114,1137,981,1004]" italics="true" pageId="10" pageNumber="229">vs</emphasis>
. creamy white); snout to spiracle 14.8–15.0 
<emphasis id="B91A1358FF946858FEF3FC09FE443E6E" box="[366,389,1017,1040]" italics="true" pageId="10" pageNumber="229">vs</emphasis>
. 15.5–16.2% TL; distance from snout to cloaca 39.8–41.0 
<emphasis id="B91A1358FF946858FBBFFC09FBF83E6E" box="[1058,1081,1017,1040]" italics="true" pageId="10" pageNumber="229">vs</emphasis>
. 35.0–38.9% TL; ventral snout length 11.2–12.1 
<emphasis id="B91A1358FF946858FEC1FBEDFEB23E4A" box="[348,371,1053,1076]" italics="true" pageId="10" pageNumber="229">vs</emphasis>
. 13.2–14.6% TL; and a wider mouth (mouth width 6.4–6.7 
<emphasis id="B91A1358FF946858FB80FBEDFBF53E4A" box="[1053,1076,1053,1076]" italics="true" pageId="10" pageNumber="229">vs</emphasis>
. 5.7–6.2% TL). 
<taxonomicName id="4C6EB4C9FF946858FB6FFBEDFA5D3E4A" box="[1266,1436,1053,1076]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="sereti">
<emphasis id="B91A1358FF946858FB6FFBEDFA5D3E4A" box="[1266,1436,1053,1076]" italics="true" pageId="10" pageNumber="229">Notoraja sereti</emphasis>
</taxonomicName>
further differs from 
<taxonomicName id="4C6EB4C9FF946858FEE8FBB1FE393E29" box="[373,504,1088,1112]" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="inusitata">
<emphasis id="B91A1358FF946858FEE8FBB1FE393E29" box="[373,504,1088,1112]" italics="true" pageId="10" pageNumber="229">N. inusitata</emphasis>
</taxonomicName>
in the following characters: snout to spiracle 14.8–15.0 
<emphasis id="B91A1358FF946858FBF7FBB1FB403E26" box="[1130,1153,1089,1112]" italics="true" pageId="10" pageNumber="229">vs</emphasis>
. 17.3% TL; nasal curtain total width 7.9–8.3 
<emphasis id="B91A1358FF946858FEEBFB95FE4C3E02" box="[374,397,1125,1148]" italics="true" pageId="10" pageNumber="229">vs</emphasis>
. 7.2% TL; and fewer radials on the mesopterygium 10–11 
<emphasis id="B91A1358FF946858FBAFFB95FB883E02" box="[1074,1097,1125,1148]" italics="true" pageId="10" pageNumber="229">vs</emphasis>
. 16. It further differs from 
<taxonomicName id="4C6EB4C9FF946858FA1FFB95FEF23EDE" class="Elasmobranchii" family="Arhynchobatidae" genus="Notoraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="longiventralis">
<emphasis id="B91A1358FF946858FA1FFB95FEF23EDE" italics="true" pageId="10" pageNumber="229">N. longiventralis</emphasis>
</taxonomicName>
in having a much shorter anterior pelvic-fin lobe (its length 12.7–13.7 
<emphasis id="B91A1358FF946858FB1CFB79FB593EDE" box="[1153,1176,1161,1184]" italics="true" pageId="10" pageNumber="229">vs</emphasis>
. 14.8–15.0% TL) and shorter distance from the cloaca to the first dorsal fin (46.6–47.3 
<emphasis id="B91A1358FF946858FCF2FB5DFC473EBA" box="[879,902,1197,1220]" italics="true" pageId="10" pageNumber="229">vs</emphasis>
. 45.1–45.9% TL).
</paragraph>
<paragraph id="8BD1CF4AFF946858FF5AFB3FFB1F3F2B" blockId="10.[151,1437,151,1365]" pageId="10" pageNumber="229">
The new species of arhynchobatid skate is distinguishable from members of the closely related genus 
<taxonomicName id="4C6EB4C9FF946858FF0AFB05FED33F72" box="[151,274,1269,1292]" class="Elasmobranchii" family="Arhynchobatidae" genus="Insentiraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="229" phylum="Chordata" rank="genus">
<emphasis id="B91A1358FF946858FF0AFB05FED33F72" box="[151,274,1269,1292]" italics="true" pageId="10" pageNumber="229">Insentiraja</emphasis>
</taxonomicName>
in having numerous small denticles on the ventral surface (
<emphasis id="B91A1358FF946858FC2AFB05FC0F3F72" box="[951,974,1269,1292]" italics="true" pageId="10" pageNumber="229">vs</emphasis>
. ventral surface smooth) and the skin on both dorsal and ventral surfaces not loose and flabby (
<emphasis id="B91A1358FF946858FC91FAE9FCE23F4E" box="[780,803,1305,1328]" italics="true" pageId="10" pageNumber="229">vs</emphasis>
. skin noticeably loose and flabby). It also differs from members of another closely-related genus 
<taxonomicName id="4C6EB4C9FF946858FDEFFACCFD2D3F2A" box="[626,748,1340,1364]" class="Elasmobranchii" family="Arhynchobatidae" genus="Brochiraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="229" phylum="Chordata" rank="genus">
<emphasis id="B91A1358FF946858FDEFFACCFD2D3F2A" box="[626,748,1340,1364]" italics="true" pageId="10" pageNumber="229">Brochiraja</emphasis>
</taxonomicName>
in lacking a bifurcate thorn on the snout tip.
</paragraph>
</subSubSection>
</treatment>
</document>