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<document id="CD41D0FF841CC6642360E2C5B93A3430" ID-DOI="10.3853/j.0067-1975.62.2010.1554" ID-ISSN="2201-4349" ID-Zenodo-Dep="5239086" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629574225023" checkinUser="felipe" docAuthor="Köhler, Frank" docDate="2010" docId="03F987C5FF8DF742036616FC49A6CDD8" docLanguage="en" docName="RecAustMus.62.2.217-284.pdf" docOrigin="Records of the Australian Museum 62 (2)" docSource="http://dx.doi.org/10.3853/j.0067-1975.62.2010.1554" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Amplirhagada lamarckiana Köhler 2010, n.sp." docType="treatment" docVersion="3" lastPageNumber="247" masterDocId="FFC0FFBDFF91F75C027510454D15CE74" masterDocTitle="Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae)" masterLastPageNumber="284" masterPageNumber="217" pageNumber="245" updateTime="1699078283413" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods id="20A19F1ADF7FB2E42754B93F10CCADEE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B5D7A3E317AE00DB2F22CFE610C0F906">
<mods:title id="65EE2901B0019C64E8C80A67EC19FEC2">Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae)</mods:title>
</mods:titleInfo>
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<mods:role id="14F322E73341C61529DD2C573658F0B1">
<mods:roleTerm id="016F44627E771221364A9B75395392DF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D62B1F0BD07EB76E2EEDD8BEB048DE21">Köhler, Frank</mods:namePart>
</mods:name>
<mods:typeOfResource id="83C9710F30004BEC02789E1A483E9D3E">text</mods:typeOfResource>
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<mods:title id="60B1E32F06093454BBDEA4620C0A9752">Records of the Australian Museum</mods:title>
</mods:titleInfo>
<mods:part id="27F2B1E9611EA8E10D5A084B17FD202F">
<mods:date id="FEFDF78888A57FD0B676669E83A12798">2010</mods:date>
<mods:detail id="C2141E5D55EE8564ED5B27D925E170C8" type="pubDate">
<mods:number id="C6BA54CC79401ED693661AA7028E6FA2">2010-11-24</mods:number>
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<mods:detail id="9CDA6ACE30710F395505ACB6C3303376" type="volume">
<mods:number id="163B60FCD31ADFECB5D6CFDDFC370533">62</mods:number>
</mods:detail>
<mods:detail id="230AF133A94689657A8111B0E71410F9" type="issue">
<mods:number id="618B971282BE017BE4EB4DE50EE5BF43">2</mods:number>
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<mods:start id="E08A3F2FC11E063AA1169C7B059B56E9">217</mods:start>
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<mods:url id="15324ED68830EC04869CBCD1A3D88FB5">http://dx.doi.org/10.3853/j.0067-1975.62.2010.1554</mods:url>
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<mods:classification id="14B49CD092289A3D076EB895CA5ABAED">journal article</mods:classification>
<mods:identifier id="9A55D5C189CCBA7765D065E84F366FE5" type="DOI">10.3853/j.0067-1975.62.2010.1554</mods:identifier>
<mods:identifier id="6772E9814420AC2F7207DE8A3BA6FF75" type="ISSN">2201-4349</mods:identifier>
<mods:identifier id="6680ED6D2B20F23D243AFFB4D977323F" type="Zenodo-Dep">5239086</mods:identifier>
<mods:identifier id="41245284C56DDD6DA3D85CC097B86355" type="ZooBank">749A271A-6942-4D4C-B0DF-B968462BFF0C</mods:identifier>
</mods:mods>
<treatment id="03F987C5FF8DF742036616FC49A6CDD8" ID-DOI="http://doi.org/10.5281/zenodo.5239037" ID-Zenodo-Dep="5239037" LSID="urn:lsid:plazi:treatment:03F987C5FF8DF742036616FC49A6CDD8" httpUri="http://treatment.plazi.org/id/03F987C5FF8DF742036616FC49A6CDD8" lastPageId="30" lastPageNumber="247" pageId="28" pageNumber="245">
<subSubSection id="C34A6558FF8DF740036616FC4FB8C8A6" box="[275,685,1720,1747]" pageId="28" pageNumber="245" type="nomenclature">
<paragraph id="8BEF36D3FF8DF740036616FC4FB8C8A6" blockId="28.[275,685,1720,1747]" box="[275,685,1720,1747]" pageId="28" pageNumber="245">
<heading id="D0A781BFFF8DF740036616FC4FB8C8A6" box="[275,685,1720,1747]" centered="true" fontSize="11" level="2" pageId="28" pageNumber="245" reason="2">
<emphasis id="B924EAC1FF8DF740036616FC4FB8C8A6" bold="true" box="[275,685,1720,1747]" pageId="28" pageNumber="245">
<taxonomicName id="4C504D50FF8DF740036616FC4F7CC8A7" ID-CoL="D7RG" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[275,617,1721,1747]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="245" phylum="Mollusca" rank="species" species="lamarckiana" status="sp. nov.">
<emphasis id="B924EAC1FF8DF740036616FC4F7CC8A7" bold="true" box="[275,617,1721,1747]" italics="true" pageId="28" pageNumber="245">Amplirhagada lamarckiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFF8DF740000416FD4FB8C8A6" box="[625,685,1720,1746]" pageId="28" pageNumber="245" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34A6558FF8DF74002D316B84F65C9A2" pageId="28" pageNumber="245" type="materials_examined">
<paragraph id="8BEF36D3FF8DF74002D316B84CC4C918" blockId="28.[166,794,1789,1900]" pageId="28" pageNumber="245">
<materialsCitation id="3B383C8EFF8DF74002D316B84CD8C918" collectingDate="1988-07-14" collectionCode="V" collectorName="Kessner &amp; A. Longbottom" county="Kimberley" latitude="-14.780556" location="York Sound" longLatPrecision="21" longitude="125.023056" municipality="Bonaparte Archipelago" pageId="28" pageNumber="245" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<emphasis id="B924EAC1FF8DF74002D316B84C50C961" bold="true" box="[166,325,1789,1813]" pageId="28" pageNumber="245">
<typeStatus id="54EB8871FF8DF74002D316B84DF7C961" box="[166,226,1789,1813]" pageId="28" pageNumber="245">Type</typeStatus>
locality
</emphasis>
(
<figureCitation id="136B2A56FF8DF740032C16B84CB4C960" box="[345,417,1789,1812]" captionStart="Figure 1" captionStartId="1.[121,184,1433,1454]" captionTargetBox="[121,1421,142,1408]" captionTargetId="figure-129@1.[121,1421,138,1414]" captionTargetPageId="1" captionText="Figure 1. Distribution of Amplirhagada species in the Kimberley region, northwestern Australia. (1) A. euroa n.sp., Adolphus Island. (2) A. solemiana n.sp., Middle Osborn Island. (3) A. indistincta n.sp., Southwest Osborn Island, Kidney Island. (4) A. combeana, Cassini Island. (5) A. mckenziei n.sp., Oliver Island. (6) A. ponderi n.sp., Kingsmill Island. (7) A. montesquieuana n.sp., Fenelon Island. (8) A. descartesana n.sp., Descartes Island. (9) A. katerana, Katers Island. (10) A. puescheli n.sp., unnamed island near Prudhoe Island. (11) A. decora n.sp., A. kessneri n.sp., Bigge Island. (12) A. berthierana n.sp., Berthier Island. (13) A. lamarckiana n.sp., Lamarck Island.(14) A. anderdonensis n.sp., unnamed island, Anderdon Islands. (15) A. tricenaria n.sp., Prince Frederick Harbour, north of Hunter River mouth. (16) A. regia n.sp., A. boongareensis n.sp., A. gibsoni n.sp., and Amplirhagada sp., Boongaree Island. (17) A. yorkensis n.sp., Coronation Island. (18) A. buffonensis n.sp., Buffon Island. (19) A. uwinsensis n.sp., Uwins Island. (20) A. sphaeroidea n.sp., St. Andrews Island. (21) A. basilica n.sp., and A. camdenensis n.sp., Augustus Island. (22) A. kimberleyana n.sp., NW of Wilson Point. (23) A. gemina n.sp., 1.5 km SE of Hall Point. (24) A. dubitabile n.sp., Steep Island. (25) A. napierana, north Napier ranges. (26) A. percita, Napier ranges. (27) A. castra, A. mitchelliana, and A. varia, Mitchell Plateau." figureDoi="http://doi.org/10.5281/zenodo.5239088" httpUri="https://zenodo.org/record/5239088/files/figure.png" pageId="28" pageNumber="245">Fig. 1</figureCitation>
).
<collectingRegion id="4994F831FF8DF74003C916B84F98C960" box="[444,653,1789,1812]" country="Australia" name="Western Australia" pageId="28" pageNumber="245">Western Australia</collectingRegion>
,
<collectingCounty id="628E4E5FFF8DF74000E916B84E03C960" box="[668,790,1789,1812]" pageId="28" pageNumber="245">Kimberley</collectingCounty>
,
<collectingMunicipality id="6B8BACA9FF8DF74002D3175E4CA0C946" box="[166,437,1819,1842]" pageId="28" pageNumber="245">Bonaparte Archipelago</collectingMunicipality>
,
<location id="8E8F6008FF8DF74003B4175E4F5BC946" LSID="urn:lsid:plazi:treatment:03F987C5FF8DF742036616FC49A6CDD8:8E8F6008FF8DF74003B4175E4F5BC946" box="[449,590,1819,1842]" county="Kimberley" latitude="-14.780556" longLatPrecision="21" longitude="125.023056" municipality="Bonaparte Archipelago" name="York Sound" pageId="28" pageNumber="245" stateProvince="Western Australia">York Sound</location>
,
<location id="8E8F6008FF8DF740002B175E4E00C946" LSID="urn:lsid:plazi:treatment:03F987C5FF8DF742036616FC49A6CDD8:8E8F6008FF8DF740002B175E4E00C946" box="[606,789,1819,1842]" county="Kimberley" latitude="-14.780556" longLatPrecision="21" longitude="125.023056" municipality="Bonaparte Archipelago" name="Lamarck Island" pageId="28" pageNumber="245" stateProvince="Western Australia">Lamarck Island</location>
,
<geoCoordinate id="EE645014FF8DF74002D3177D4C31C93B" box="[166,292,1848,1871]" degrees="14" direction="south" minutes="46" orientation="latitude" pageId="28" pageNumber="245" precision="15" seconds="50" value="-14.780556">14°46'50&quot;S</geoCoordinate>
<geoCoordinate id="EE645014FF8DF740035B177D4CA5C93B" box="[302,432,1848,1871]" degrees="125" direction="east" minutes="1" orientation="longitude" pageId="28" pageNumber="245" precision="15" seconds="23" value="125.023056">125°1'23&quot;E</geoCoordinate>
; KC-042 (leg.
<collectionCode id="ED41AE16FF8DF7400028177D4F64C93B" box="[605,625,1848,1871]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="28" pageNumber="245" type="Museum">V</collectionCode>
.
<collectorName id="26A55305FF8DF740000A177D4FC2C93B" box="[639,727,1848,1871]" pageId="28" pageNumber="245">Kessner</collectorName>
&amp;
<collectorName id="26A55305FF8DF7400175177D4C38C918" pageId="28" pageNumber="245">A. Longbottom</collectorName>
,
<date id="FFEE1013FF8DF740034217104CD0C918" box="[311,453,1877,1900]" pageId="28" pageNumber="245" value="1988-07-14">
<collectingDate id="EFAAE9FBFF8DF740034217104CD0C918" box="[311,453,1877,1900]" pageId="28" pageNumber="245" value="1988-07-14">14 July 1988</collectingDate>
</date>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEF36D3FF8DF74002D317D64F65C9A2" blockId="28.[166,793,1939,2006]" pageId="28" pageNumber="245">
<emphasis id="B924EAC1FF8DF74002D317D64C3AC9D2" bold="true" box="[166,303,1939,1958]" pageId="28" pageNumber="245">Type material</emphasis>
.
<materialsCitation id="3B383C8EFF8DF740034B17D64F95C9D2" accessionNumber="S34612" box="[318,640,1939,1958]" collectionCode="WAM" pageId="28" pageNumber="245" specimenCount="1" typeStatus="holotype">
<typeStatus id="54EB8871FF8DF740034B17D64C87C9D2" box="[318,402,1939,1958]" pageId="28" pageNumber="245" type="holotype">Holotype</typeStatus>
<collectionCode id="ED41AE16FF8DF74003EF17D64CC5C9D2" box="[410,464,1939,1958]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="28" pageNumber="245" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FF8DF74003AD17D64F08C9D2" box="[472,541,1939,1958]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34612" pageId="28" pageNumber="245" type="EnaNcbi">S34612</accessionNumber>
(Pl. 1.18)
</materialsCitation>
.
<materialsCitation id="3B383C8EFF8DF74000FE17D64F79C9A2" accessionNumber="S41485" collectionCode="WAM" pageId="28" pageNumber="245" specimenCode="FMNH 219163" specimenCount="1" typeStatus="paratype">
<typeStatus id="54EB8871FF8DF74000FE17D64FF7C9D2" box="[651,738,1939,1958]" pageId="28" pageNumber="245" type="paratype">Paratypes</typeStatus>
AMS C463755 (5 preserved specimens),
<specimenCode id="DBF69EA8FF8DF740038B17EE4F9BC9CA" box="[510,654,1963,1982]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="28" pageNumber="245" type="Museum">FMNH 219163</specimenCode>
(15 preserved specimens),
<collectionCode id="ED41AE16FF8DF740036617864C5DC9A2" box="[275,328,1987,2006]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="28" pageNumber="245" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FF8DF740033817864C9AC9A2" box="[333,399,1987,2006]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S41485" pageId="28" pageNumber="245" type="EnaNcbi">S41485</accessionNumber>
(8 preserved specimens)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A6558FF8DF742013216FD49A6CDD8" lastPageId="30" lastPageNumber="247" pageId="28" pageNumber="245" type="etymology">
<paragraph id="8BEF36D3FF8DF740013216FD4EFDC899" blockId="28.[839,1466,1720,1773]" pageId="28" pageNumber="245">
<emphasis id="B924EAC1FF8DF740013216FD4ED1C8A4" bold="true" box="[839,964,1720,1744]" pageId="28" pageNumber="245">Etymology</emphasis>
. In reference to Lamarck Island, where this species occurs.
</paragraph>
<paragraph id="8BEF36D3FF8DF741013217554CF4C919" blockId="28.[839,1468,1808,2008]" lastBlockId="29.[121,749,1673,1901]" lastPageId="29" lastPageNumber="246" pageId="28" pageNumber="245">
<emphasis id="B924EAC1FF8DF740013217554E94C95C" bold="true" box="[839,897,1808,1832]" pageId="28" pageNumber="245">Shell</emphasis>
(
<figureCitation id="136B2A56FF8DF74001E01754490AC95C" box="[917,1055,1809,1832]" captionStart="Figure 42" captionStartId="29.[121,184,1541,1562]" captionTargetBox="[121,1421,151,1526]" captionTargetId="figure-9@29.[121,1421,151,1526]" captionTargetPageId="29" captionText="Figure 42. SEM photographs of Amplirhagada lamarckiana n.sp., holotype WAM S34612. (A–C) Shell: (A) apical whorl viewed from above (scale 200 µm); (B) sculpture on second to forth whorl viewed from above (scale 200 µm); (C) sculpture on upper part of last whorl (scale 200 µm).(D) Jaw (scale 100 µm). (E–F) Radula: (E) central and lateral teeth (scale 20 µm); (F) middle marginal teeth (scale 10 µm)." figureDoi="http://doi.org/10.5281/zenodo.5239173" httpUri="https://zenodo.org/record/5239173/files/figure.png" pageId="28" pageNumber="245">Fig. 42A–C</figureCitation>
, Pl. 1.18). Semi-globose to broadly conical, with medium high spire; thin (translucent). Periphery evenly rounded to slightly angulate; upper sector of whorls rather flattened, basal sector rounded. Umbilicus forming a chink, 90–100 percent concealed by columellar reflection. Background colour light brownish horn, banded. Sub-sutural and mid-whorl bands diffuse to well defined, light brown, thin, visible on last whorl(s) only; ventral colour horn; outer lip colour same as shell; inner lip translucent. Protoconch c.
<quantity id="4CA89B36FF8CF74102E616864DC4C8AE" box="[147,209,1731,1754]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="29" pageNumber="246" unit="mm" value="3.0">3 mm</quantity>
in diameter, comprising 1.7 whorls, with fine and indistinct lirae. Teleoconch with rather strong axial growth lines. Angle of aperture 45°; outer lip rounded, sharp to moderately thick, slightly expanded, reflected, basal node of lip absent or weak; palatal node absent. Parietal wall of inner lip absent or inconspicuous.
</paragraph>
<caption id="DF2F665BFF8CF741020C1640489BC83B" ID-DOI="http://doi.org/10.5281/zenodo.5239173" ID-Zenodo-Dep="5239173" httpUri="https://zenodo.org/record/5239173/files/figure.png" pageId="29" pageNumber="246" startId="29.[121,184,1541,1562]" targetBox="[121,1421,151,1526]" targetPageId="29">
<paragraph id="8BEF36D3FF8CF741020C1640489BC83B" blockId="29.[121,1422,1541,1615]" pageId="29" pageNumber="246">
Figure 42. SEM photographs of
<taxonomicName id="4C504D50FF8CF74103C616404FA8C86E" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[435,701,1541,1562]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="246" phylum="Mollusca" rank="species" species="lamarckiana" status="sp. nov.">
<emphasis id="B924EAC1FF8CF74103C616404FA8C86E" box="[435,701,1541,1562]" italics="true" pageId="29" pageNumber="246">Amplirhagada lamarckiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFF8CF74100B616404FE6C86E" box="[707,755,1541,1562]" pageId="29" pageNumber="246" rank="species">n.sp.</taxonomicNameLabel>
, holotype WAM
<accessionNumber id="9403AB30FF8CF74101EC16404EF0C86E" box="[921,997,1541,1562]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34612" pageId="29" pageNumber="246" type="EnaNcbi">S34612</accessionNumber>
. (
<emphasis id="B924EAC1FF8CF741018216404934C86E" box="[1015,1057,1541,1562]" italics="true" pageId="29" pageNumber="246">A–C</emphasis>
) Shell: (
<emphasis id="B924EAC1FF8CF741060216404993C86E" box="[1143,1158,1541,1562]" italics="true" pageId="29" pageNumber="246">A</emphasis>
) apical whorl viewed from above (scale 200 µm); (
<emphasis id="B924EAC1FF8CF741032216654C73C841" box="[343,358,1568,1589]" italics="true" pageId="29" pageNumber="246">B</emphasis>
) sculpture on second to forth whorl viewed from above (scale 200 µm); (
<emphasis id="B924EAC1FF8CF741066D1665493DC841" box="[1048,1064,1568,1589]" italics="true" pageId="29" pageNumber="246">C</emphasis>
) sculpture on upper part of last whorl (scale 200 µm). (
<emphasis id="B924EAC1FF8CF7410362167F4C3DC83B" box="[279,296,1594,1615]" italics="true" pageId="29" pageNumber="246">D</emphasis>
) Jaw (scale 100 µm). (
<emphasis id="B924EAC1FF8CF7410388167F4F33C83B" box="[509,550,1594,1615]" italics="true" pageId="29" pageNumber="246">E–F</emphasis>
) Radula: (
<emphasis id="B924EAC1FF8CF74100FF167F4F8CC83B" box="[650,665,1594,1615]" italics="true" pageId="29" pageNumber="246">E</emphasis>
) central and lateral teeth (scale 20 µm); (
<emphasis id="B924EAC1FF8CF741066C167F493DC83B" box="[1049,1064,1594,1615]" italics="true" pageId="29" pageNumber="246">F</emphasis>
) middle marginal teeth (scale 10 µm).
</paragraph>
</caption>
<paragraph id="8BEF36D3FF8CF741020C17D54CDDC996" blockId="29.[121,749,1936,2018]" pageId="29" pageNumber="246">
<emphasis id="B924EAC1FF8CF741020C17D54C58C9DC" bold="true" box="[121,333,1936,1960]" pageId="29" pageNumber="246">Pallial morphology</emphasis>
. Pallial cavity deep, extending
<emphasis id="B924EAC1FF8CF74100FA17D54F82C9EA" bold="true" box="[655,663,1936,1950]" pageId="29" pageNumber="246">
<superScript id="7C259B9BFF8CF74100FA17D54F82C9EA" attach="right" box="[655,663,1936,1950]" fontSize="6" pageId="29" pageNumber="246">3</superScript>
</emphasis>
⁄
<emphasis id="B924EAC1FF8CF74100EE17D24FB6C9D1" bold="true" box="[667,675,1943,1957]" pageId="29" pageNumber="246">4</emphasis>
whorl. Pigmentation on mantle mottled, black. Kidney extending more than half of pallial cavity.
</paragraph>
<paragraph id="8BEF36D3FF8CF742016F16CD4ECACF0B" blockId="29.[794,1422,1672,2018]" lastBlockId="30.[839,1467,154,383]" lastPageId="30" lastPageNumber="247" pageId="29" pageNumber="246">
<emphasis id="B924EAC1FF8CF741016F16CD4905C8D4" bold="true" box="[794,1040,1672,1696]" pageId="29" pageNumber="246">Genital morphology</emphasis>
(
<figureCitation id="136B2A56FF8CF741065016CC49A3C8D4" box="[1061,1206,1673,1696]" captionStart-0="Figure 43" captionStart-1="Figure 44" captionStartId-0="30.[166,229,616,637]" captionStartId-1="30.[166,228,1931,1952]" captionTargetBox-0="[183,790,129,601]" captionTargetBox-1="[190,781,748,1873]" captionTargetId-0="figure-8@30.[166,793,125,609]" captionTargetId-1="figure-32@30.[167,794,731,1885]" captionTargetPageId-0="30" captionTargetPageId-1="30" captionText-0="Figure 43. Genitalia of Amplirhagada lamarckiana n.sp., holotype WAM S34612 (14 July, scale 10 mm). Compare with Fig. 3 for labelling of structures." captionText-1="Figure 44. Interior ofpenial chamber ofAmplirhagada lamarckiana n.sp., paratype FMNH 219163 (14 July, scale 5 mm).Compare with Fig. 4 for labelling of structures." figureDoi-0="http://doi.org/10.5281/zenodo.5239175" figureDoi-1="http://doi.org/10.5281/zenodo.5239177" httpUri-0="https://zenodo.org/record/5239175/files/figure.png" httpUri-1="https://zenodo.org/record/5239177/files/figure.png" pageId="29" pageNumber="246">Figs. 43–44</figureCitation>
). Penis straight to curved, of about same length as anterior part of oviduct. Vas deferens coils before entering penis. Penial retractor muscle shorter than penis. Penial sheath delicate. Penial verge tiny (barely visible), slender to spatulate, with pointed tip. Penial wall small, arranged in rows over entire length of inner penial wall. Main stimulatory pilaster elongated, sculptured by horizontal ridges that support little hooks, comprising apical to median portion of penial chamber. Two to three additional pilasters are covered by undifferentiated pustules. Vas deferens entering penial sheath half way up. Vagina moderate, posteriorly inflated; inner vaginal wall with smooth longitudinal pilasters. Spermathecal duct moderately thick, internally with smooth longitudinal pilasters. Spermathecal head globular, connected with oviduct by connective tissue, internally smooth, with delicate wall. Free oviduct comprises more or less half of anterior part of oviduct, straight. Spermoviduct longer than anterior part of oviduct. Talon embedded in albumen gland at junction with spermoviduct.
</paragraph>
<paragraph id="8BEF36D3FF8FF742013211E74E86CCB5" blockId="30.[839,1467,418,705]" pageId="30" pageNumber="247">
<emphasis id="B924EAC1FF8FF742013211E7492BCFCE" bold="true" box="[839,1086,418,442]" pageId="30" pageNumber="247">Radular morphology</emphasis>
(
<figureCitation id="136B2A56FF8FF742062511E749C1CFCD" box="[1104,1236,418,441]" captionStart="Figure 42" captionStartId="29.[121,184,1541,1562]" captionTargetBox="[121,1421,151,1526]" captionTargetId="figure-9@29.[121,1421,151,1526]" captionTargetPageId="29" captionText="Figure 42. SEM photographs of Amplirhagada lamarckiana n.sp., holotype WAM S34612. (A–C) Shell: (A) apical whorl viewed from above (scale 200 µm); (B) sculpture on second to forth whorl viewed from above (scale 200 µm); (C) sculpture on upper part of last whorl (scale 200 µm).(D) Jaw (scale 100 µm). (E–F) Radula: (E) central and lateral teeth (scale 20 µm); (F) middle marginal teeth (scale 10 µm)." figureDoi="http://doi.org/10.5281/zenodo.5239173" httpUri="https://zenodo.org/record/5239173/files/figure.png" pageId="30" pageNumber="247">Fig. 42E–F</figureCitation>
). Rectangular. Tooth formula C+14+4+18. With 157 rows of teeth, 27.9 rows per mm (n = 1). Central teeth with bluntly pointed, triangular to ovate mesocones, shorter than base of tooth; ectocones vestigial. Lateral teeth with sharply pointed, triangular mesocones, length equal to base of tooth; ectocones and endocones vestigial. Marginal teeth with triangular mesocones; ectocones shorter and narrower than mesocones, occasionally divided into two denticles, endocones reduced in size.
</paragraph>
<caption id="DF2F665BFF8FF74202D3122D4C6ACCC7" ID-DOI="http://doi.org/10.5281/zenodo.5239175" ID-Zenodo-Dep="5239175" httpUri="https://zenodo.org/record/5239175/files/figure.png" pageId="30" pageNumber="247" startId="30.[166,229,616,637]" targetBox="[183,790,129,601]" targetPageId="30">
<paragraph id="8BEF36D3FF8FF74202D3122D4C6ACCC7" blockId="30.[166,794,616,691]" pageId="30" pageNumber="247">
Figure 43. Genitalia of
<taxonomicName id="4C504D50FF8FF74203F7122C4F9CCC0A" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[386,649,617,638]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="247" phylum="Mollusca" rank="species" species="lamarckiana" status="sp. nov.">
<emphasis id="B924EAC1FF8FF74203F7122C4F9CCC0A" box="[386,649,617,638]" italics="true" pageId="30" pageNumber="247">Amplirhagada lamarckiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFF8FF74200FB122D4FA8CC09" box="[654,701,616,637]" pageId="30" pageNumber="247" rank="species">n.sp.</taxonomicNameLabel>
, holotype WAM
<accessionNumber id="9403AB30FF8FF742029C12C64C27CCEC" box="[233,306,643,664]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34612" pageId="30" pageNumber="247" type="EnaNcbi">S34612</accessionNumber>
(14 July, scale 10 mm). Compare with Fig. 3 for labelling of structures.
</paragraph>
</caption>
<paragraph id="8BEF36D3FF8FF742013212A049A6CDD8" blockId="30.[839,1467,741,940]" pageId="30" pageNumber="247">
<emphasis id="B924EAC1FF8FF742013212A0495CCC89" bold="true" box="[839,1097,741,765]" pageId="30" pageNumber="247">Comparative remarks</emphasis>
. Pustulation of inner penial wall somewhat similar to
<taxonomicName id="4C504D50FF8FF7420635134649C9CD6E" authorityName="Köhler" authorityYear="2010" box="[1088,1244,771,794]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="247" phylum="Mollusca" rank="species" species="indistincta">
<emphasis id="B924EAC1FF8FF7420635134649C9CD6E" box="[1088,1244,771,794]" italics="true" pageId="30" pageNumber="247">A. indistincta</emphasis>
</taxonomicName>
and
<taxonomicName id="4C504D50FF8FF742076E134648A6CD6E" authorityName="Köhler" authorityYear="2010" box="[1307,1459,771,794]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="247" phylum="Mollusca" rank="species" species="solemiana">
<emphasis id="B924EAC1FF8FF742076E134648A6CD6E" box="[1307,1459,771,794]" italics="true" pageId="30" pageNumber="247">A. solemiana</emphasis>
</taxonomicName>
, which lack a well-differentiated main pilaster and differ by more conical shape of shell. Main pilaster differs in shape and surface pattern from the two species from Bigge Island. Material of the present species has been labelled as “
<taxonomicName id="4C504D50FF8FF742012613D04907CDD8" box="[851,1042,917,940]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="247" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B924EAC1FF8FF742012613D04EF8CDD8" box="[851,1005,917,940]" italics="true" pageId="30" pageNumber="247">Amplirhagada</emphasis>
sp.
</taxonomicName>
68” by Solem.
</paragraph>
</subSubSection>
</treatment>
</document>
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