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<document ID-DOI="http://dx.doi.org/10.3897/asp.81.e102683" ID-Pensoft-Pub="1864-8312-81-581" ID-Pensoft-UUID="E6F5D6150A225AFBB372E719EBBEDB68" ID-ZooBank="CA56E9F0B4264014B20BD7C455FD65A7" ModsDocID="1864-8312-81-581" checkinTime="1686864336769" checkinUser="pensoft" docAuthor="Wang, Ya-Lian, Peng, Zi-Qi, Jin, Feng-Liang, Chen, Hong-Wei &amp; Huang, Jia" docDate="2023" docId="FFFDE484DEAE54B8B7A74024C471F59E" docLanguage="en" docName="ArthropodSystPhyl 81: 581-610" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 81" docPubDate="2023-06-15" docSource="http://dx.doi.org/10.3897/asp.81.e102683" docTitle="Stegana (Orthostegana) obscurala Peng &amp; Chen 2023, sp. nov." docType="treatment" docUuid="89E07D8B-C1BF-42AF-AF97-26E1D29ED9BF" docUuidSource="ZooBank" docVersion="1" id="E6F5D6150A225AFBB372E719EBBEDB68" lastPageNumber="581" masterDocId="E6F5D6150A225AFBB372E719EBBEDB68" masterDocTitle="Integrative species delimitation in the morphologically conservative Stegana subgen. Orthostegana (Diptera, Drosophilidae)" masterLastPageNumber="610" masterPageNumber="581" pageNumber="581" updateTime="1686864336769" updateUser="pensoft">
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<mods:title id="51798092DE5A2F377958964CCDAFCEEF">Integrative species delimitation in the morphologically conservative Stegana subgen. Orthostegana (Diptera, Drosophilidae)</mods:title>
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<mods:namePart id="33AE5A639C4F64C1D386E1256B37DF66">Wang, Ya-Lian</mods:namePart>
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<mods:affiliation id="DE6160FB6AF28E732879A8CEECF086D8">Department of Entomology, South China Agricultural University, 483 Wushan-lu, Guangzhou 510642, Guangdong, P. R. China</mods:affiliation>
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<mods:namePart id="1499E0F9D966C44A72A032EC0073EBAA">Peng, Zi-Qi</mods:namePart>
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<mods:affiliation id="54DD1ECAF267057ADBC22151362103B1">Department of Entomology, South China Agricultural University, 483 Wushan-lu, Guangzhou 510642, Guangdong, P. R. China</mods:affiliation>
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<mods:namePart id="AC36666397094C51DBD18A8DCFA12F29">Jin, Feng-Liang</mods:namePart>
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<mods:affiliation id="552ECD1952BF729ED9A0B5CA9C4FD504">Department of Entomology, South China Agricultural University, 483 Wushan-lu, Guangzhou 510642, Guangdong, P. R. China</mods:affiliation>
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<mods:namePart id="D0A23A99834C446EB8AEDB6C2B9A8D0F">Chen, Hong-Wei</mods:namePart>
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<mods:affiliation id="E6244AC5BF98AF5FD585463935185EC1">Department of Entomology, South China Agricultural University, 483 Wushan-lu, Guangzhou 510642, Guangdong, P. R. China</mods:affiliation>
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<mods:namePart id="3E7A48E4722EB7FADB9836F0173D98F6">Huang, Jia</mods:namePart>
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<mods:affiliation id="C2CEBB195CDE923F6DB812CA9C718A23">Department of Entomology, South China Agricultural University, 483 Wushan-lu, Guangzhou 510642, Guangdong, P. R. China</mods:affiliation>
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<mods:title id="DFB5BDE684086916D7305E37DE9912BB">Arthropod Systematics &amp; amp; Phylogeny</mods:title>
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<mods:date id="0A0E36A87B45C66DBB3BBA28C39FA27A">2023</mods:date>
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<treatment id="FFFDE484DEAE54B8B7A74024C471F59E" LSID="urn:lsid:zoobank.org:act:89E07D8B-C1BF-42AF-AF97-26E1D29ED9BF" httpUri="http://treatment.plazi.org/id/FFFDE484DEAE54B8B7A74024C471F59E" lastPageNumber="581" pageId="0" pageNumber="581">
<subSubSection id="41EC20F7D9695D0CB3C34343EFF01635" pageId="0" pageNumber="581" type="nomenclature">
<paragraph id="2E05163D7F080A99BFEC1A91B4A5507C" pageId="0" pageNumber="581">
3.2.1.8.
<taxonomicName id="E4495931E976DA01BFA706F48D110325" LSID="https://zoobank.org/89E07D8B-C1BF-42AF-AF97-26E1D29ED9BF" authority="Peng &amp; Chen" authorityName="Peng &amp; Chen" authorityYear="2023" class="Insecta" family="Drosophilidae" genus="Stegana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stegana (Orthostegana) obscurala" order="Diptera" pageId="0" pageNumber="581" phylum="Arthropoda" rank="species" species="obscurala" status="sp. nov." subGenus="Orthostegana">Stegana (Orthostegana) obscurala Peng &amp; Chen</taxonomicName>
<taxonomicNameLabel id="D217DE1D350ABCF8560D058779B0C865" pageId="0" pageNumber="581">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="8FF84A55C55526720F6CE1711F255AEC" pageId="0" pageNumber="581" type="description">
<paragraph id="C39258E5A366403C814520D4A0FCEBA8" pageId="0" pageNumber="581">
<figureCitation id="3AD1B99F63159C2E0E5CCA4005E5EEF9" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Head, thorax, and abdomen in dorsal and lateral views. A, B Stegana (Orthostegana) mohnihei Peng &amp; Chen sp. nov. (female); C, D S. (O.) obscurala Peng &amp; Chen sp. nov. (male); E, F S. (O.) obscurala Peng &amp; Chen sp. nov. (female); G, H S. (O.) pinguitia Peng &amp; Chen sp. nov. (male)." figureDoi="10.3897/asp.81.e102683.figure8" httpUri="https://binary.pensoft.net/fig/864475" pageId="0" pageNumber="581">Figs 8C-F</figureCitation>
<figureCitation id="1243C0A97C9FCACBA779F47C577F263D" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Mesotarsus of males in ventral view. A Stegana (Orthostegana) latipalpula Peng &amp; Chen sp. nov.; B S. (O.) macrostephana Peng &amp; Chen sp. nov.; C S. (O.) mohnihei Peng &amp; Chen sp. nov.; D S. (O.) obscurala Peng &amp; Chen sp. nov.; E S. (O.) pinguitia Peng &amp; Chen sp. nov." figureDoi="10.3897/asp.81.e102683.figure10" httpUri="https://binary.pensoft.net/fig/864477" pageId="0" pageNumber="581">, 10D</figureCitation>
<figureCitation id="6A047DAF4BE6F2BAC18CCE79D41270E6" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Stegana (Orthostegana) obscurala Peng &amp; Chen sp. nov., male (A - D) and female terminalia (E). A Epandrium, surstylus, and cercus in lateral view; B Surstylus in ventral view; C, D Hypandrium, phallapodeme, pregonite, aedeagus, and aedeagal sheath in ventral and lateral views; E Eighth sternite in ventral view. Scale bars = 0.1 mm." figureDoi="10.3897/asp.81.e102683.figure19" httpUri="https://binary.pensoft.net/fig/864486" pageId="0" pageNumber="581">, 19</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="5FA8EF2203AB34A4F8D2BD94A831CE3B" pageId="0" pageNumber="581" type="diagnosis">
<paragraph id="D433A4E89A47967D40749E1742D51BD4" pageId="0" pageNumber="581">Diagnosis.</paragraph>
<paragraph id="B46BE1332375C66F8A54216FAEB3F026" pageId="0" pageNumber="581">
This species closely resembles
<taxonomicName id="470B1538D08669E82D7331AB3CA71A5A" lsidName="S. (O.) aini" pageId="0" pageNumber="581" rank="species" species="aini" subGenus="O.">S. (O.) aini</taxonomicName>
<emphasis id="2FC825EF071D4DE1270853E6ADD7A92A" bold="true" pageId="0" pageNumber="581">sp. nov.</emphasis>
in the shape of both male and female terminalia (Figs
<figureCitation id="87A1B982E13E95C5636C26619225D094" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Stegana (Orthostegana) aini Peng &amp; Chen sp. nov., male (A - D) and female terminalia (E). A Epandrium, surstylus, and cercus in lateral view; B Surstylus in ventral view; C, D Hypandrium, phallapodeme, pregonite, aedeagus, and aedeagal sheath in ventral and lateral views; E Eighth sternite in ventral view. Scale bars = 0.1 mm." figureDoi="10.3897/asp.81.e102683.figure12" httpUri="https://binary.pensoft.net/fig/864479" pageId="0" pageNumber="581">12</figureCitation>
,
<figureCitation id="930117BA5834D291CA545B1493103DB4" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Stegana (Orthostegana) obscurala Peng &amp; Chen sp. nov., male (A - D) and female terminalia (E). A Epandrium, surstylus, and cercus in lateral view; B Surstylus in ventral view; C, D Hypandrium, phallapodeme, pregonite, aedeagus, and aedeagal sheath in ventral and lateral views; E Eighth sternite in ventral view. Scale bars = 0.1 mm." figureDoi="10.3897/asp.81.e102683.figure19" httpUri="https://binary.pensoft.net/fig/864486" pageId="0" pageNumber="581">19</figureCitation>
), but can be distinguished from the latter by having the pleura with a distinct brown to dark brown longitudinal stripe above (Fig.
<figureCitation id="90A16FB9401CE6345C039C21B58FFDB9" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Head, thorax, and abdomen in dorsal and lateral views. A, B Stegana (Orthostegana) mohnihei Peng &amp; Chen sp. nov. (female); C, D S. (O.) obscurala Peng &amp; Chen sp. nov. (male); E, F S. (O.) obscurala Peng &amp; Chen sp. nov. (female); G, H S. (O.) pinguitia Peng &amp; Chen sp. nov. (male)." figureDoi="10.3897/asp.81.e102683.figure8" httpUri="https://binary.pensoft.net/fig/864475" pageId="0" pageNumber="581">8D, F</figureCitation>
); the mesotarsus ventrally with 2 rows of black, apically blunt, stout spines; metatarsus with 12/6+1 spines; 2nd to 4th segments with 6/1, 3/1, and 2/1 spines, respectively (Fig.
<figureCitation id="0A1DEC8007E3BE86D8102344D7EB35AB" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Mesotarsus of males in ventral view. A Stegana (Orthostegana) latipalpula Peng &amp; Chen sp. nov.; B S. (O.) macrostephana Peng &amp; Chen sp. nov.; C S. (O.) mohnihei Peng &amp; Chen sp. nov.; D S. (O.) obscurala Peng &amp; Chen sp. nov.; E S. (O.) pinguitia Peng &amp; Chen sp. nov." figureDoi="10.3897/asp.81.e102683.figure10" httpUri="https://binary.pensoft.net/fig/864477" pageId="0" pageNumber="581">10D</figureCitation>
); pregonite with
<emphasis id="C88489E154A725C40EFABBECAE4FF51B" italics="true" pageId="0" pageNumber="581">ca.</emphasis>
3 sensilla distally (Fig.
<figureCitation id="D8861EFF464FCB7D634E1A77EC635AB3" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Stegana (Orthostegana) obscurala Peng &amp; Chen sp. nov., male (A - D) and female terminalia (E). A Epandrium, surstylus, and cercus in lateral view; B Surstylus in ventral view; C, D Hypandrium, phallapodeme, pregonite, aedeagus, and aedeagal sheath in ventral and lateral views; E Eighth sternite in ventral view. Scale bars = 0.1 mm." figureDoi="10.3897/asp.81.e102683.figure19" httpUri="https://binary.pensoft.net/fig/864486" pageId="0" pageNumber="581">19C, D</figureCitation>
).
</paragraph>
<caption id="7499229722D495D28566675ABEEA515B" doi="10.3897/asp.81.e102683.figure19" httpUri="https://binary.pensoft.net/fig/864486" pageId="0" pageNumber="581" start="Figure 19" startId="F19">
<paragraph id="4C391A5331B5CA08D5737B3EAA7E3643" pageId="0" pageNumber="581">
<emphasis id="A14A2F1087CAD527EE02904A7638EC9E" bold="true" pageId="0" pageNumber="581">Figure 19.</emphasis>
<taxonomicName id="1138B108FBA118553D9CEEE4CFB8CD4F" authorityName="Peng &amp; Chen" authorityYear="2023" class="Insecta" family="Drosophilidae" genus="Stegana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stegana (Orthostegana) obscurala" order="Diptera" pageId="0" pageNumber="581" phylum="Arthropoda" rank="species" species="obscurala" subGenus="Orthostegana">Stegana (Orthostegana) obscurala</taxonomicName>
Peng &amp; Chen
<emphasis id="5B9AB153ED8B25AB2236EEE5D86F667D" bold="true" pageId="0" pageNumber="581">sp. nov.</emphasis>
, male (
<emphasis id="0A38F56A4763E4EE0FB96BCB5B65BA38" bold="true" pageId="0" pageNumber="581">A</emphasis>
-
<emphasis id="8F74193E13977764BE271759D8D0FC5F" bold="true" pageId="0" pageNumber="581">D</emphasis>
) and female terminalia (
<emphasis id="F4D4AE256CEE14408C921177A2BD341F" bold="true" pageId="0" pageNumber="581">E</emphasis>
).
<emphasis id="C6DBA514A1FB640C8FF221C775BDD749" bold="true" pageId="0" pageNumber="581">A</emphasis>
Epandrium, surstylus, and cercus in lateral view;
<emphasis id="AC5D9F7A58C46B4401118B2264F66090" bold="true" pageId="0" pageNumber="581">B</emphasis>
Surstylus in ventral view;
<emphasis id="4B7241F9A012708C7BC3F947A179817C" bold="true" pageId="0" pageNumber="581">C</emphasis>
,
<emphasis id="6D77C2726A1EDC1865FC90DD52AFED59" bold="true" pageId="0" pageNumber="581">D</emphasis>
Hypandrium, phallapodeme, pregonite, aedeagus, and aedeagal sheath in ventral and lateral views;
<emphasis id="4FD0D241E2A9671A55964544D00C6795" bold="true" pageId="0" pageNumber="581">E</emphasis>
Eighth sternite in ventral view. Scale bars = 0.1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="535994A790BB335B93DEDACEAE5AD544" pageId="0" pageNumber="581" type="description">
<paragraph id="C8BB88A53A908C52EE3367493D528ECE" pageId="0" pageNumber="581">Description.</paragraph>
<paragraph id="6C202CF0106B2450DB49B618445213F9" pageId="0" pageNumber="581">
<emphasis id="3EDFAB9EB95E99A99CEFB02F076291CB" bold="true" pageId="0" pageNumber="581">
Male and female.
<emphasis id="D2310ABD7B3DF0DD6E56CBA5DFDB5EFD" italics="true" pageId="0" pageNumber="581">Head</emphasis>
</emphasis>
: Ocellar triangle dark brown, with 2 small setae posterior to ocellar setae. Postocellar seta absent. Frons brown, with numerous minute interfrontal setulae. Pedicel and first flagellomere yellow, with black pubescence. Face brown, with narrowed, black band on lower margin. Gena brownish. Clypeus mostly brownish. Palpus brownish.
<emphasis id="C20A3600DD497A37FF3991CDFE62B0FE" bold="true" italics="true" pageId="0" pageNumber="581">Thorax</emphasis>
: Mesonotum mostly yellow to brown (Fig.
<figureCitation id="66F209C3B53306D1E6DED2194ABE039E" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Head, thorax, and abdomen in dorsal and lateral views. A, B Stegana (Orthostegana) mohnihei Peng &amp; Chen sp. nov. (female); C, D S. (O.) obscurala Peng &amp; Chen sp. nov. (male); E, F S. (O.) obscurala Peng &amp; Chen sp. nov. (female); G, H S. (O.) pinguitia Peng &amp; Chen sp. nov. (male)." figureDoi="10.3897/asp.81.e102683.figure8" httpUri="https://binary.pensoft.net/fig/864475" pageId="0" pageNumber="581">8C, E</figureCitation>
). Scutellum yellow in males (Fig.
<figureCitation id="70567EAB05464849E17082B4BF7CB2A3" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Head, thorax, and abdomen in dorsal and lateral views. A, B Stegana (Orthostegana) mohnihei Peng &amp; Chen sp. nov. (female); C, D S. (O.) obscurala Peng &amp; Chen sp. nov. (male); E, F S. (O.) obscurala Peng &amp; Chen sp. nov. (female); G, H S. (O.) pinguitia Peng &amp; Chen sp. nov. (male)." figureDoi="10.3897/asp.81.e102683.figure8" httpUri="https://binary.pensoft.net/fig/864475" pageId="0" pageNumber="581">8C</figureCitation>
), dark brown in females (Fig.
<figureCitation id="F877AA014DC83E76B01904B64ABB1637" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Head, thorax, and abdomen in dorsal and lateral views. A, B Stegana (Orthostegana) mohnihei Peng &amp; Chen sp. nov. (female); C, D S. (O.) obscurala Peng &amp; Chen sp. nov. (male); E, F S. (O.) obscurala Peng &amp; Chen sp. nov. (female); G, H S. (O.) pinguitia Peng &amp; Chen sp. nov. (male)." figureDoi="10.3897/asp.81.e102683.figure8" httpUri="https://binary.pensoft.net/fig/864475" pageId="0" pageNumber="581">8E</figureCitation>
). Halter mostly brownish, with a yellow patch at knob (Fig.
<figureCitation id="B8F902B92C6B91A168C59DAC5A1ED36C" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Head, thorax, and abdomen in dorsal and lateral views. A, B Stegana (Orthostegana) mohnihei Peng &amp; Chen sp. nov. (female); C, D S. (O.) obscurala Peng &amp; Chen sp. nov. (male); E, F S. (O.) obscurala Peng &amp; Chen sp. nov. (female); G, H S. (O.) pinguitia Peng &amp; Chen sp. nov. (male)." figureDoi="10.3897/asp.81.e102683.figure8" httpUri="https://binary.pensoft.net/fig/864475" pageId="0" pageNumber="581">8D-F</figureCitation>
). Legs yellowish (Fig.
<figureCitation id="C56DB129F8D4B811786AD000487C83C2" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Mesotarsus of males in ventral view. A Stegana (Orthostegana) latipalpula Peng &amp; Chen sp. nov.; B S. (O.) macrostephana Peng &amp; Chen sp. nov.; C S. (O.) mohnihei Peng &amp; Chen sp. nov.; D S. (O.) obscurala Peng &amp; Chen sp. nov.; E S. (O.) pinguitia Peng &amp; Chen sp. nov." figureDoi="10.3897/asp.81.e102683.figure10" httpUri="https://binary.pensoft.net/fig/864477" pageId="0" pageNumber="581">10D</figureCitation>
).
<emphasis id="D5D531C95162164D2635A1238B9F1652" bold="true" italics="true" pageId="0" pageNumber="581">Abdominal tergites</emphasis>
: brown to black.
<emphasis id="408F01469490F532D49E19914DF32879" bold="true" italics="true" pageId="0" pageNumber="581">Male terminalia</emphasis>
: Epandrium mostly pubescent except for anterior and ventral margins, with
<emphasis id="F8881422F5256C28AC1849244EA5895B" italics="true" pageId="0" pageNumber="581">ca.</emphasis>
20 setae on dorsal to posterior portions per side (Fig.
<figureCitation id="649292330067EB0B9E533C8D6819E05C" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Stegana (Orthostegana) obscurala Peng &amp; Chen sp. nov., male (A - D) and female terminalia (E). A Epandrium, surstylus, and cercus in lateral view; B Surstylus in ventral view; C, D Hypandrium, phallapodeme, pregonite, aedeagus, and aedeagal sheath in ventral and lateral views; E Eighth sternite in ventral view. Scale bars = 0.1 mm." figureDoi="10.3897/asp.81.e102683.figure19" httpUri="https://binary.pensoft.net/fig/864486" pageId="0" pageNumber="581">19A</figureCitation>
). Surstylus separated from epandrium, with
<emphasis id="BB58F11E25BDEEC54054583BEA135354" italics="true" pageId="0" pageNumber="581">ca.</emphasis>
2 long and 10 short peg-shaped prensisetae and a few setae near posteroventral margin (Fig.
<figureCitation id="5624C83298974C5C00ADF5D4BAB4EC20" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Stegana (Orthostegana) obscurala Peng &amp; Chen sp. nov., male (A - D) and female terminalia (E). A Epandrium, surstylus, and cercus in lateral view; B Surstylus in ventral view; C, D Hypandrium, phallapodeme, pregonite, aedeagus, and aedeagal sheath in ventral and lateral views; E Eighth sternite in ventral view. Scale bars = 0.1 mm." figureDoi="10.3897/asp.81.e102683.figure19" httpUri="https://binary.pensoft.net/fig/864486" pageId="0" pageNumber="581">19B</figureCitation>
). Pregonites separated and small, slightly expanded distally (Fig.
<figureCitation id="8A489BC31FB6C0FAF841DFD9AB907136" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Stegana (Orthostegana) obscurala Peng &amp; Chen sp. nov., male (A - D) and female terminalia (E). A Epandrium, surstylus, and cercus in lateral view; B Surstylus in ventral view; C, D Hypandrium, phallapodeme, pregonite, aedeagus, and aedeagal sheath in ventral and lateral views; E Eighth sternite in ventral view. Scale bars = 0.1 mm." figureDoi="10.3897/asp.81.e102683.figure19" httpUri="https://binary.pensoft.net/fig/864486" pageId="0" pageNumber="581">19C, D</figureCitation>
). Aedeagus pubescent distally, with some hairs apically (Fig.
<figureCitation id="E54AFC542C0789CD4ACCA80A1A708C3A" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Stegana (Orthostegana) obscurala Peng &amp; Chen sp. nov., male (A - D) and female terminalia (E). A Epandrium, surstylus, and cercus in lateral view; B Surstylus in ventral view; C, D Hypandrium, phallapodeme, pregonite, aedeagus, and aedeagal sheath in ventral and lateral views; E Eighth sternite in ventral view. Scale bars = 0.1 mm." figureDoi="10.3897/asp.81.e102683.figure19" httpUri="https://binary.pensoft.net/fig/864486" pageId="0" pageNumber="581">19C, D</figureCitation>
).
<emphasis id="C5D1A517661FB281F7F8330C5360ADFB" bold="true" italics="true" pageId="0" pageNumber="581">Female terminalia</emphasis>
: Eighth sternite with 4 rows of asymmetrical peg-shaped prensisetae subposteromedially and posterolaterally, and with 3 sensilla distally per side (Fig.
<figureCitation id="682008FE294DD82595E40C42A3A74B82" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Stegana (Orthostegana) obscurala Peng &amp; Chen sp. nov., male (A - D) and female terminalia (E). A Epandrium, surstylus, and cercus in lateral view; B Surstylus in ventral view; C, D Hypandrium, phallapodeme, pregonite, aedeagus, and aedeagal sheath in ventral and lateral views; E Eighth sternite in ventral view. Scale bars = 0.1 mm." figureDoi="10.3897/asp.81.e102683.figure19" httpUri="https://binary.pensoft.net/fig/864486" pageId="0" pageNumber="581">19E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DF1A930CEAF245F01580305E317AD278" pageId="0" pageNumber="581" type="description">
<paragraph id="27A06D55BC175FD8B4338399752E0D6B" pageId="0" pageNumber="581">Measurements and indices.</paragraph>
<paragraph id="CA565A7BCA4E250FB0BEDA3659C3691F" pageId="0" pageNumber="581">
<emphasis id="8A8197BA0BB3DCA4978F7321D516E4FC" bold="true" pageId="0" pageNumber="581">BL</emphasis>
= 3.13 mm in holotype (range in 2♂ and 2♀ paratypes: 2.60-3.00 in ♂, 2.87-4.21 in ♀),
<emphasis id="CDA05A9138A960424CB986316CC32836" bold="true" pageId="0" pageNumber="581">ThL</emphasis>
= 1.37 mm (1.00-1.13 in ♂, 1.07-1.60 in ♀),
<emphasis id="344B2F120B3AA7651410AE0F2C69AB20" bold="true" pageId="0" pageNumber="581">WL</emphasis>
= 2.33 mm (1.90-2.17 in ♂, 2.20-2.77 in ♀),
<emphasis id="5A18BF8C7E2591CCF62D3917256BC731" bold="true" pageId="0" pageNumber="581">WW</emphasis>
= 1.03 mm (0.93-0.97 in ♂, 1.00-1.27 in ♀),
<emphasis id="F66F1DB843CA3164297FAFEBB291957A" bold="true" pageId="0" pageNumber="581">arb</emphasis>
= 6/5 (6-7/4-6),
<emphasis id="2484DC8EB38CFCE336AA4B9EE4300078" bold="true" pageId="0" pageNumber="581">avd</emphasis>
= 0.92 (0.88-0.92),
<emphasis id="6CE3296AFA5465C067D2A8A4ABB45389" bold="true" pageId="0" pageNumber="581">adf</emphasis>
= 1.71 (1.71-2.13),
<emphasis id="47499A40714D4564B299BEE597B6DF7B" bold="true" pageId="0" pageNumber="581">flw</emphasis>
= 1.86 (2.00-2.17),
<emphasis id="2FB795E29C0976A4B4F20AAD33F500E4" bold="true" pageId="0" pageNumber="581">FW/HW</emphasis>
= 0.38 (0.32-0.40),
<emphasis id="15EB12AD759BB9DD8D03DD5B6AE0F256" bold="true" pageId="0" pageNumber="581">ch/o</emphasis>
= 0.15 (0.13-0.16),
<emphasis id="E2BCE7266269B35E4CC4FFF898905987" bold="true" pageId="0" pageNumber="581">prorb</emphasis>
= 0.90 (0.82-0.90),
<emphasis id="0B080F4752B06CE38A182D2359476C08" bold="true" pageId="0" pageNumber="581">rcorb</emphasis>
= 0.50 (0.50-0.65),
<emphasis id="AC2018906000444514766AF87FEBDD31" bold="true" pageId="0" pageNumber="581">vb</emphasis>
= 0.41 (0.40-0.75),
<emphasis id="79E75E2776379AC32982F8E50B0F843F" bold="true" pageId="0" pageNumber="581">dcl</emphasis>
= 0.54 (0.38-0.50),
<emphasis id="DB1CDD8366B8E96A9A0E58886177CEE2" bold="true" pageId="0" pageNumber="581">presctl</emphasis>
= 0.39 (0.40-0.50),
<emphasis id="EC7933801AD818853175F412992BA1B7" bold="true" pageId="0" pageNumber="581">sctl</emphasis>
= 1.33 (1.33-1.50),
<emphasis id="C79CC0D778DC192D594007CD0E893D5D" bold="true" pageId="0" pageNumber="581">sterno</emphasis>
= 0.84 (0.77-0.84),
<emphasis id="56EC0C47F4E7C9456386BFA81AF61C2E" bold="true" pageId="0" pageNumber="581">orbito</emphasis>
= 1.80 (1.67-2.00),
<emphasis id="33EF37FDB5BA1B991E84ED0F4D0481CA" bold="true" pageId="0" pageNumber="581">dcp</emphasis>
= 0.24 (0.22-0.29),
<emphasis id="89BAC707AB3756D8544654571E3FA2EF" bold="true" pageId="0" pageNumber="581">sctlp</emphasis>
= 0.90 (0.89-1.17),
<emphasis id="B97BFECFF2E7FF9DA1775480C4F7E165" bold="true" pageId="0" pageNumber="581">C</emphasis>
= 2.75 (2.20-2.50),
<emphasis id="8B3FE7F76D14AAD19EA1972C6AAE94E7" bold="true" pageId="0" pageNumber="581">4c</emphasis>
= 0.84 (0.84-1.00),
<emphasis id="0975511A732A870EAF7889F9C2123F48" bold="true" pageId="0" pageNumber="581">4v</emphasis>
= 1.79 (1.63-1.84),
<emphasis id="C1D43D383661E4CEEBFD9DA5B192D75B" bold="true" pageId="0" pageNumber="581">5x</emphasis>
= 1.29 (1.25-1.50),
<emphasis id="0A9DD6DD6C09668B7F8DAAAB8A0B0B41" bold="true" pageId="0" pageNumber="581">ac</emphasis>
= 16.00 (13.33-16.00),
<emphasis id="176A584C92D4CB258A28663C0B13F82C" bold="true" pageId="0" pageNumber="581">M</emphasis>
= 0.47 (0.42-0.53),
<emphasis id="1CFA58344B00762A088E85C648DDDAC1" bold="true" pageId="0" pageNumber="581">C3F</emphasis>
= 0.75 (0.69-0.76).
</paragraph>
</subSubSection>
<subSubSection id="7FAE91C0F159F2D987FA0A6961C5A8A7" pageId="0" pageNumber="581" type="distribution">
<paragraph id="0CA48E02EB4EAFFD728E3111D3F158DF" pageId="0" pageNumber="581">Distribution.</paragraph>
<paragraph id="89AD8C0CD8A11BEF91C8AF122DA5DDFC" pageId="0" pageNumber="581">China (Yunnan).</paragraph>
</subSubSection>
<subSubSection id="7C83A4422D78196DFC8D3678C5297A6A" pageId="0" pageNumber="581" type="type material">
<paragraph id="E5FD6AA38DC87F0A0699B8741F6F2435" pageId="0" pageNumber="581">Type material examined.</paragraph>
<paragraph id="D225431EA163F463EF6F7F8DAECC2319" pageId="0" pageNumber="581">
<emphasis id="0A6F323BFE270966FF16067F9D69F0CE" bold="true" pageId="0" pageNumber="581">Holotype</emphasis>
: ♂ (SCAU, No. 112125), China: Wangtianshu, Mengla, Yunnan,
<geoCoordinate id="60B6AE9201FD565E0F6C1FBD7B9BF647" degrees="21" direction="north" minutes="37" orientation="latitude" precision="15" seconds="10" value="21.619444">21°3710″N</geoCoordinate>
,
<geoCoordinate id="BBF9E39E1997DCC3FC65B527FFDB7CE7" degrees="101" direction="east" minutes="35" orientation="longitude" precision="15" seconds="17" value="101.58806">101°3517″E</geoCoordinate>
, altitude 570 m, 9.v.2012, swept from tussock, H.W. Chen.
<emphasis id="5C3A0747FD47C4233F4C3EA4327E01DA" bold="true" pageId="0" pageNumber="581">Paratypes</emphasis>
: 2♂, 2♀ (SCAU, Nos. 112126-29), H.W. Chen, J.J. Gao, other data same as the holotype.
</paragraph>
</subSubSection>
<subSubSection id="27DBF540121B7AEF7B687FF7BC52E1B6" pageId="0" pageNumber="581" type="etymology">
<paragraph id="89B43FA427FBE3E14C22681850CE16C3" pageId="0" pageNumber="581">Etymology.</paragraph>
<paragraph id="8C88D8E62F998511CFFE3792E2ACAB15" pageId="0" pageNumber="581">
From the Latin word &quot;
<emphasis id="FC8D7C7DFD9B3DB71DA55F8D08978F97" italics="true" pageId="0" pageNumber="581">obscurus</emphasis>
&quot; (= obscure), referring to the mostly dark-colored abdominal tergites.
</paragraph>
</subSubSection>
</treatment>
</document>