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<document id="297ABAFA730E55E4588997E7090AEB47" ID-CLB-Dataset="4023" ID-DOI="10.11646/zootaxa.5051.1.19" ID-GBIF-Dataset="0c8b11dc-564b-461b-9cb1-3cd61111f9b2" ID-ISSN="1175-5326" ID-Zenodo-Dep="5563851" ID-ZooBank="A607CB5B-51E7-45FD-9104-980175BBB163" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1634023045486" checkinUser="plazi" docAuthor="Mu, Fanghong &amp; Huys, Rony" docDate="2021" docId="8F5DE56C2800B10EFF2AFA26326A86F5" docLanguage="en" docName="zootaxa.5051.1.19.pdf" docOrigin="Zootaxa 5051 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Balucopsylla Rao 1972" docType="treatment" docVersion="6" lastPageNumber="497" masterDocId="73649D14280AB105FFBDFFA6366F875B" masterDocTitle="A new species of Helmutkunzia Wells &amp; Rao, 1976 from an intertidal sandy beach in Xiamen, China and proposal of Pseudobalucopsylla gen. nov. (Copepoda Harpacticoida, Miraciidae)" masterLastPageNumber="505" masterPageNumber="487" pageNumber="497" updateTime="1699459587513" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="55B7F521E66C97D20338D629D629762C">A new species of Helmutkunzia Wells &amp; Rao, 1976 from an intertidal sandy beach in Xiamen, China and proposal of Pseudobalucopsylla gen. nov. (Copepoda Harpacticoida, Miraciidae)</mods:title>
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<mods:namePart id="0B0E4B54BC51C20CDB4CF25EA2663172">Mu, Fanghong</mods:namePart>
<mods:affiliation id="D10DCFA339C833B72C50D792ABB79E4E">College of Marine Life Sciences, Ocean University of China, 5 Yushan Road, Qingdao 266003, Shandong Province, China. Department of Life Sciences, The Natural History Museum, Cromwell Road, London SW 7 5 BD, U. K.</mods:affiliation>
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<mods:namePart id="FCF44DFBDA0464ACBD8EE78B2D970B5B">Huys, Rony</mods:namePart>
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<treatment id="8F5DE56C2800B10EFF2AFA26326A86F5" ID-DOI="http://doi.org/10.5281/zenodo.5563803" ID-GBIF-Taxon="188648304" ID-Zenodo-Dep="5563803" LSID="urn:lsid:plazi:treatment:8F5DE56C2800B10EFF2AFA26326A86F5" httpUri="http://treatment.plazi.org/id/8F5DE56C2800B10EFF2AFA26326A86F5" lastPageId="11" lastPageNumber="497" pageId="10" pageNumber="497">
<subSubSection id="4FEE07F12800B10FFF2AFA26344482C1" box="[151,555,1407,1434]" pageId="10" pageNumber="497" type="nomenclature">
<paragraph id="074B547A2800B10FFF2AFA26344482C1" blockId="10.[151,555,1407,1434]" box="[151,555,1407,1434]" pageId="10" pageNumber="497">
<heading id="5C03E3162800B10FFF2AFA26344482C1" bold="true" box="[151,555,1407,1434]" fontSize="11" level="1" pageId="10" pageNumber="497" reason="1">
<emphasis id="358088682800B10FFF2AFA26344482C1" bold="true" box="[151,555,1407,1434]" pageId="10" pageNumber="497">
Notes on
<taxonomicName id="C0F42FF92800B10FFEB6FAD9344482C1" authority="Rao, 1972" authorityName="Rao" authorityYear="1972" box="[267,555,1407,1434]" class="Hexanauplia" family="Miraciidae" genus="Balucopsylla" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="497" phylum="Arthropoda" rank="genus">
<emphasis id="358088682800B10FFEB6FAD937C882C1" bold="true" box="[267,423,1407,1434]" italics="true" pageId="10" pageNumber="497">Balucopsylla</emphasis>
<bibRefCitation id="6365298B2800B10FFE12FA26344482C1" author="Rao, G. C." box="[431,555,1408,1434]" pageId="10" pageNumber="497" pagination="305 - 319" refId="ref13271" refString="Rao, G. C. (1972) Some new interstitial harpacticoid copepods from Andhra coast, India. Cahiers de Biologie marine, 13, 305 - 319. https: // doi. org / 10.21411 / CBM. A. 6 C 3 F 88 E 8" type="journal article" year="1972">Rao, 1972</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="4FEE07F12800B10EFF2AFA6F326A86F5" lastPageId="11" lastPageNumber="498" pageId="10" pageNumber="497" type="description">
<paragraph id="074B547A2800B10EFF2AFA6F326A86F5" blockId="10.[151,1437,1480,2011]" lastBlockId="11.[151,1437,151,430]" lastPageId="11" lastPageNumber="498" pageId="10" pageNumber="497">
<bibRefCitation id="6365298B2800B10FFF2AFA6F377A82B9" author="Rao, G. C." box="[151,277,1480,1507]" pageId="10" pageNumber="497" pagination="305 - 319" refId="ref13271" refString="Rao, G. C. (1972) Some new interstitial harpacticoid copepods from Andhra coast, India. Cahiers de Biologie marine, 13, 305 - 319. https: // doi. org / 10.21411 / CBM. A. 6 C 3 F 88 E 8" type="journal article" year="1972">Rao (1972)</bibRefCitation>
proposed the genus
<taxonomicName id="C0F42FF92800B10FFE44FA6E34E482B9" authorityName="Rao" authorityYear="1972" box="[505,651,1480,1506]" class="Hexanauplia" family="Miraciidae" genus="Balucopsylla" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="497" phylum="Arthropoda" rank="genus">
<emphasis id="358088682800B10FFE44FA6E34E482B9" box="[505,651,1480,1506]" italics="true" pageId="10" pageNumber="497">Balucopsylla</emphasis>
</taxonomicName>
for its
<typeStatus id="D84FEAD82800B10FFD67FA6F356582B8" box="[730,778,1481,1507]" pageId="10" pageNumber="497">type</typeStatus>
and only species,
<taxonomicName id="C0F42FF92800B10FFC6BFA6F32D682B9" authority="Rao, 1972" authorityName="Rao" authorityYear="1972" box="[982,1209,1480,1507]" class="Hexanauplia" family="Miraciidae" genus="Balucopsylla" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="497" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="358088682800B10FFC6BFA6F325282B9" box="[982,1085,1480,1506]" italics="true" pageId="10" pageNumber="497">B. similis</emphasis>
<bibRefCitation id="6365298B2800B10FFBF9FA6F32D682B9" author="Rao, G. C." box="[1092,1209,1480,1507]" pageId="10" pageNumber="497" pagination="305 - 319" refId="ref13271" refString="Rao, G. C. (1972) Some new interstitial harpacticoid copepods from Andhra coast, India. Cahiers de Biologie marine, 13, 305 - 319. https: // doi. org / 10.21411 / CBM. A. 6 C 3 F 88 E 8" type="journal article" year="1972">Rao, 1972</bibRefCitation>
</taxonomicName>
, from a sandy beach in Visakhapatnam (formerly known as Waltair) in
<collectingRegion id="C5309A982800B10FFD6AFA4A35FF815D" box="[727,912,1516,1542]" country="India" name="Andhra Pradesh" pageId="10" pageNumber="497">Andhra Pradesh</collectingRegion>
,
<collectingCountry id="7FE314EA2800B10FFC20FA4A35B5815D" box="[925,986,1516,1542]" name="India" pageId="10" pageNumber="497">India</collectingCountry>
. Rao suggested that his new taxon was most closely related to a group of other genera that exhibit reduced leg armature, including
<taxonomicName id="C0F42FF92800B10FFB1DF9B633738171" authorityName="Sars" authorityYear="1905" box="[1184,1308,1552,1578]" class="Hexanauplia" family="Miraciidae" genus="Schizopera" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="497" phylum="Arthropoda" rank="genus">
<emphasis id="358088682800B10FFB1DF9B633738171" box="[1184,1308,1552,1578]" italics="true" pageId="10" pageNumber="497">Schizopera</emphasis>
</taxonomicName>
,
<taxonomicName id="C0F42FF92800B10FFA96F9B6375E8115" authority="Wilson, 1932" authorityName="Wilson" authorityYear="1932" class="Hexanauplia" family="Miraciidae" genus="Goffinella" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="497" phylum="Arthropoda" rank="genus">
<emphasis id="358088682800B10FFA96F9B633F28171" box="[1323,1437,1552,1578]" italics="true" pageId="10" pageNumber="497">Goffinella</emphasis>
<bibRefCitation id="6365298B2800B10FFF2AF992375E8115" author="Wilson, C. B." box="[151,305,1588,1614]" pageId="10" pageNumber="497" pagination="1 - 635" refId="ref14379" refString="Wilson, C. B. (1932) The copepods of the Woods Hole region, Massachusetts. Bulletin of the United States national Museum, 158, 1 - 635, pls. 1 - 41. https: // doi. org / 10.5479 / si. 03629236.158. i" type="journal article" year="1932">Wilson, 1932</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="C0F42FF92800B10FFE81F99337A18115" authorityName="Pennak" authorityYear="1942" box="[316,462,1589,1614]" class="Hexanauplia" family="Miraciidae" genus="Psammotopa" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="497" phylum="Arthropoda" rank="genus">
<emphasis id="358088682800B10FFE81F99337A18115" box="[316,462,1589,1614]" italics="true" pageId="10" pageNumber="497">Psammotopa</emphasis>
</taxonomicName>
and
<taxonomicName id="C0F42FF92800B10FFDBBF99334BB8115" authorityName="Geddes" authorityYear="1968" box="[518,724,1589,1614]" class="Hexanauplia" family="Miraciidae" genus="Protopsammotopa" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="497" phylum="Arthropoda" rank="genus">
<emphasis id="358088682800B10FFDBBF99334BB8115" box="[518,724,1589,1614]" italics="true" pageId="10" pageNumber="497">Protopsammotopa</emphasis>
</taxonomicName>
. Justification for its distinct generic status was primarily based on the 3-segmented condition of the P1 endopod in conjunction with the absence of sexual dimorphism on the P2 endopod; this combination is not displayed by any of the genera mentioned above (P1 endopod 2-segmented in
<taxonomicName id="C0F42FF92800B10FFAD4F9DA36B081E1" authorityName="Wilson" authorityYear="1932" class="Hexanauplia" family="Miraciidae" genus="Goffinella" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="497" phylum="Arthropoda" rank="genus">
<emphasis id="358088682800B10FFAD4F9DA36B081E1" italics="true" pageId="10" pageNumber="497">Goffinella</emphasis>
</taxonomicName>
,
<taxonomicName id="C0F42FF92800B10FFF53F90737EF81E1" authorityName="Pennak" authorityYear="1942" box="[238,384,1697,1722]" class="Hexanauplia" family="Miraciidae" genus="Psammotopa" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="497" phylum="Arthropoda" rank="genus">
<emphasis id="358088682800B10FFF53F90737EF81E1" box="[238,384,1697,1722]" italics="true" pageId="10" pageNumber="497">Psammotopa</emphasis>
</taxonomicName>
,
<taxonomicName id="C0F42FF92800B10FFE32F907343281E1" authorityName="Geddes" authorityYear="1968" box="[399,605,1697,1722]" class="Hexanauplia" family="Miraciidae" genus="Protopsammotopa" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="497" phylum="Arthropoda" rank="genus">
<emphasis id="358088682800B10FFE32F907343281E1" box="[399,605,1697,1722]" italics="true" pageId="10" pageNumber="497">Protopsammotopa</emphasis>
</taxonomicName>
; P2 endopod
<specimenCount id="11F29FF32800B10FFD41F907357E81E0" box="[764,785,1697,1723]" pageId="10" pageNumber="497" type="male">♂</specimenCount>
modified in
<taxonomicName id="C0F42FF92800B10FFC19F906324F81E1" authorityName="Sars" authorityYear="1905" box="[932,1056,1696,1722]" class="Hexanauplia" family="Miraciidae" genus="Schizopera" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="497" phylum="Arthropoda" rank="genus">
<emphasis id="358088682800B10FFC19F906324F81E1" box="[932,1056,1696,1722]" italics="true" pageId="10" pageNumber="497">Schizopera</emphasis>
</taxonomicName>
and
<taxonomicName id="C0F42FF92800B10FFBE4F907334881E1" authorityName="Geddes" authorityYear="1968" box="[1113,1319,1697,1722]" class="Hexanauplia" family="Miraciidae" genus="Protopsammotopa" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="497" phylum="Arthropoda" rank="genus">
<emphasis id="358088682800B10FFBE4F907334881E1" box="[1113,1319,1697,1722]" italics="true" pageId="10" pageNumber="497">Protopsammotopa</emphasis>
</taxonomicName>
).
<bibRefCitation id="6365298B2800B10FFA82F90637788184" author="Wells, J. B. J. &amp; Rao, G. C." pageId="10" pageNumber="497" pagination="79 - 90" refId="ref14236" refString="Wells, J. B. J. &amp; Rao, G. C. (1976) The relationship of the genus Schizopera Sars within the family Diosaccidae (Copepoda: Harpacticoida). Zoological Journal of the Linnean Society, 58, 79 - 90. https: // doi. org / 10.1111 / j. 1096 - 3642.1976. tb 00821. x" type="journal article" year="1976">Wells &amp; Rao (1976)</bibRefCitation>
further expanded this genus group by including
<taxonomicName id="C0F42FF92800B10FFC83F96235D08185" authorityName="Wells" authorityYear="1967" box="[830,959,1732,1758]" class="Hexanauplia" family="Miraciidae" genus="Actopsyllus" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="497" phylum="Arthropoda" rank="genus">
<emphasis id="358088682800B10FFC83F96235D08185" box="[830,959,1732,1758]" italics="true" pageId="10" pageNumber="497">Actopsyllus</emphasis>
</taxonomicName>
,
<taxonomicName id="C0F42FF92800B10FFC73F96232188185" authorityName="Por" authorityYear="1968" box="[974,1143,1732,1758]" class="Hexanauplia" family="Miraciidae" genus="Schizoperoides" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="497" phylum="Arthropoda" rank="genus">
<emphasis id="358088682800B10FFC73F96232188185" box="[974,1143,1732,1758]" italics="true" pageId="10" pageNumber="497">Schizoperoides</emphasis>
</taxonomicName>
and their newly proposed genus
<taxonomicName id="C0F42FF92800B10FFF5CF94E37168059" authorityName="Wells &amp; Rao" authorityYear="1976" box="[225,377,1768,1794]" class="Hexanauplia" family="Miraciidae" genus="Eoschizopera" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="497" phylum="Arthropoda" rank="genus">
<emphasis id="358088682800B10FFF5CF94E37168059" box="[225,377,1768,1794]" italics="true" pageId="10" pageNumber="497">Eoschizopera</emphasis>
</taxonomicName>
, stating that in this lineage the shared reduced leg setation, with at most two outer spines on P2–P4 exp-3, appears to be congruent with the similarity in female genital field morphology. They recognized an evolutionary series
<emphasis id="358088682800B10FFECEF896351D8011" box="[371,882,1840,1867]" italics="true" pageId="10" pageNumber="497">
<taxonomicName id="C0F42FF92800B10FFECEF89634648011" authorityName="Wells &amp; Rao" authorityYear="1976" box="[371,523,1840,1866]" class="Hexanauplia" family="Miraciidae" genus="Eoschizopera" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="497" phylum="Arthropoda" rank="genus">Eoschizopera</taxonomicName>
–
<taxonomicName id="C0F42FF92800B10FFD95F89634AB8011" authorityName="Wells &amp; Rao" authorityYear="1976" box="[552,708,1840,1866]" class="Hexanauplia" family="Miraciidae" genus="Helmutkunzia" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="497" phylum="Arthropoda" rank="genus">Helmutkunzia</taxonomicName>
–
<taxonomicName id="C0F42FF92800B10FFD5DF896351D8011" authorityName="Rao" authorityYear="1972" box="[736,882,1840,1866]" class="Hexanauplia" family="Miraciidae" genus="Balucopsylla" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="497" phylum="Arthropoda" rank="genus">Balucopsylla</taxonomicName>
</emphasis>
in which the latter represented the most morphologically reduced taxon. The next record of the genus is the one by
<bibRefCitation id="6365298B2800B10FFCC4F8F232668035" author="Wells, J. B. J." box="[889,1033,1876,1902]" pageId="10" pageNumber="497" pagination="91 - 93" refId="ref14120" refString="Wells, J. B. J. (1978) A preliminary report on the littoral copepods of Moce Island, Lau group, Fiji. In: Lau-Tonga 1977. Report from the Royal Society of New Zealand Expedition. Bulletin of the Royal Society of New Zealand, 17, 91 - 93." type="journal article" year="1978">Wells (1978)</bibRefCitation>
who reported
<taxonomicName id="C0F42FF92800B10FFB15F8F2330F8034" box="[1192,1376,1876,1903]" class="Hexanauplia" family="Miraciidae" genus="Balucopsylla" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="497" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="358088682800B10FFB15F8F233558035" box="[1192,1338,1876,1902]" italics="true" pageId="10" pageNumber="497">Balucopsylla</emphasis>
sp.
</taxonomicName>
from the intertidal zone of Moce island,
<collectingCountry id="7FE314EA2800B10FFD97F8DE343E80C9" box="[554,593,1912,1938]" name="Fiji" pageId="10" pageNumber="497">Fiji</collectingCountry>
but this species remained undescribed.
<bibRefCitation id="6365298B2800B10FFBA7F8DE336E80C9" author="Wells, J. B. J. &amp; Rao, G. C." box="[1050,1281,1912,1939]" pageId="10" pageNumber="497" pagination="1 - 385" refId="ref14303" refString="Wells, J. B. J. &amp; Rao, G. C. (1987) Littoral Harpacticoida (Crustacea: Copepoda) from Andaman and Nicobar Islands. Memoirs of the Zoological Survey of India, 16 (4), 1 - 385." type="journal article" year="1987">Wells &amp; Rao (1987)</bibRefCitation>
subsequently added a second species,
<taxonomicName id="C0F42FF92800B10FFE12F83B352F80ED" authority="Wells &amp; Rao, 1987" authorityName="Wells &amp; Rao" authorityYear="1987" box="[431,832,1948,1975]" class="Hexanauplia" family="Miraciidae" genus="Balucopsylla" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="497" phylum="Arthropoda" rank="species" species="triarticulata">
<emphasis id="358088682800B10FFE12F83B343680ED" box="[431,601,1948,1974]" italics="true" pageId="10" pageNumber="497">B. triarticulata</emphasis>
<bibRefCitation id="6365298B2800B10FFDDFF83A352F80ED" author="Wells, J. B. J. &amp; Rao, G. C." box="[610,832,1948,1975]" pageId="10" pageNumber="497" pagination="1 - 385" refId="ref14303" refString="Wells, J. B. J. &amp; Rao, G. C. (1987) Littoral Harpacticoida (Crustacea: Copepoda) from Andaman and Nicobar Islands. Memoirs of the Zoological Survey of India, 16 (4), 1 - 385." type="journal article" year="1987">Wells &amp; Rao, 1987</bibRefCitation>
</taxonomicName>
, from the Andaman and Nicobar island chain, which appeared more primitive than the
<typeStatus id="D84FEAD82800B10FFDAAF86734288080" box="[535,583,1985,2011]" pageId="10" pageNumber="497">type</typeStatus>
species in the segmentation of the antennary exopod, yet displayed a more reduced armature on the endopods of P2 and P3. Despite these differences
<bibRefCitation id="6365298B2801B10EFC4AFF3132B087EA" author="Wells, J. B. J. &amp; Rao, G. C." box="[1015,1247,151,177]" pageId="11" pageNumber="498" pagination="1 - 385" refId="ref14303" refString="Wells, J. B. J. &amp; Rao, G. C. (1987) Littoral Harpacticoida (Crustacea: Copepoda) from Andaman and Nicobar Islands. Memoirs of the Zoological Survey of India, 16 (4), 1 - 385." type="journal article" year="1987">Wells &amp; Rao (1987)</bibRefCitation>
considered both species congeneric based on their shared absence of sexual dimorphism in the P2 endopod. Their claim that the 3-segmented antennary exopod in
<taxonomicName id="C0F42FF92801B10EFDA6FF7934AB87A2" authorityName="Wells &amp; Rao" authorityYear="1987" box="[539,708,223,249]" class="Hexanauplia" family="Miraciidae" genus="Balucopsylla" kingdom="Animalia" order="Harpacticoida" pageId="11" pageNumber="498" phylum="Arthropoda" rank="species" species="triarticulata">
<emphasis id="358088682801B10EFDA6FF7934AB87A2" box="[539,708,223,249]" italics="true" pageId="11" pageNumber="498">B. triarticulata</emphasis>
</taxonomicName>
reinforces their previously suggested close relationship between
<taxonomicName id="C0F42FF92801B10EFF2AFEA5375C8646" authorityName="Wells &amp; Rao" authorityYear="1976" box="[151,307,259,285]" class="Hexanauplia" family="Miraciidae" genus="Helmutkunzia" kingdom="Animalia" order="Harpacticoida" pageId="11" pageNumber="498" phylum="Arthropoda" rank="genus">
<emphasis id="358088682801B10EFF2AFEA5375C8646" box="[151,307,259,285]" italics="true" pageId="11" pageNumber="498">Helmutkunzia</emphasis>
</taxonomicName>
and
<taxonomicName id="C0F42FF92801B10EFED1FEA534828645" authority="(Wells &amp; Rao 1976)" baseAuthorityName="Wells &amp; Rao" baseAuthorityYear="1976" box="[364,749,259,286]" class="Hexanauplia" family="Miraciidae" genus="Balucopsylla" kingdom="Animalia" order="Harpacticoida" pageId="11" pageNumber="498" phylum="Arthropoda" rank="genus">
<emphasis id="358088682801B10EFED1FEA537918646" box="[364,510,259,285]" italics="true" pageId="11" pageNumber="498">Balucopsylla</emphasis>
(
<bibRefCitation id="6365298B2801B10EFDB3FEA5348B8645" author="Wells, J. B. J. &amp; Rao, G. C." box="[526,740,259,286]" pageId="11" pageNumber="498" pagination="79 - 90" refId="ref14236" refString="Wells, J. B. J. &amp; Rao, G. C. (1976) The relationship of the genus Schizopera Sars within the family Diosaccidae (Copepoda: Harpacticoida). Zoological Journal of the Linnean Society, 58, 79 - 90. https: // doi. org / 10.1111 / j. 1096 - 3642.1976. tb 00821. x" type="journal article" year="1976">Wells &amp; Rao 1976</bibRefCitation>
)
</taxonomicName>
is futile since it is based on a plesiomorphic character state. Morphological comparison of the
<typeStatus id="D84FEAD82801B10EFDA7FE813425861A" box="[538,586,295,321]" pageId="11" pageNumber="498">type</typeStatus>
species,
<taxonomicName id="C0F42FF92801B10EFD0FFE813574861A" authorityName="Rao" authorityYear="1972" box="[690,795,295,321]" class="Hexanauplia" family="Miraciidae" genus="Balucopsylla" kingdom="Animalia" order="Harpacticoida" pageId="11" pageNumber="498" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="358088682801B10EFD0FFE813574861A" box="[690,795,295,321]" italics="true" pageId="11" pageNumber="498">B. similis</emphasis>
</taxonomicName>
, with
<taxonomicName id="C0F42FF92801B10EFCD9FE813262861A" authorityName="Wells &amp; Rao" authorityYear="1987" box="[868,1037,295,321]" class="Hexanauplia" family="Miraciidae" genus="Balucopsylla" kingdom="Animalia" order="Harpacticoida" pageId="11" pageNumber="498" phylum="Arthropoda" rank="species" species="triarticulata">
<emphasis id="358088682801B10EFCD9FE813262861A" box="[868,1037,295,321]" italics="true" pageId="11" pageNumber="498">B. triarticulata</emphasis>
</taxonomicName>
and related forms subsequently reported by
<bibRefCitation id="6365298B2801B10EFEB4FEED37CB863E" author="Mielke, W." box="[265,420,331,357]" pageId="11" pageNumber="498" pagination="55 - 60" refId="ref12915" refString="Mielke, W. (1994) New records of two copepod species (Crustacea) from the Pacific coast of Costa Rica. Microfauna marina, 9, 55 - 60." type="journal article" year="1994">Mielke (1994</bibRefCitation>
,
<bibRefCitation id="6365298B2801B10EFE12FEED3784863E" author="Mielke, W." box="[431,491,331,357]" pageId="11" pageNumber="498" pagination="115 - 152" refId="ref12980" refString="Mielke, W. (1997) Interstitial Fauna of Galapagos. XXXIX. Copepoda, part 7. Microfauna marina, 11, 115 - 152." type="journal article" year="1997">1997</bibRefCitation>
), reveals significant differences warranting a subdivision of the genus
<taxonomicName id="C0F42FF92801B10EFABEFEED33FA863E" baseAuthorityName="Wells &amp; Rao" baseAuthorityYear="1976" box="[1283,1429,331,357]" class="Hexanauplia" family="Miraciidae" genus="Balucopsylla" kingdom="Animalia" order="Harpacticoida" pageId="11" pageNumber="498" phylum="Arthropoda" rank="genus">
<emphasis id="358088682801B10EFABEFEED33FA863E" box="[1283,1429,331,357]" italics="true" pageId="11" pageNumber="498">Balucopsylla</emphasis>
</taxonomicName>
. The generic concept of the latter is here restricted to accommodate
<taxonomicName id="C0F42FF92801B10EFC35FEC9359F86D2" authorityName="Rao" authorityYear="1972" box="[904,1008,367,393]" class="Hexanauplia" family="Miraciidae" genus="Balucopsylla" kingdom="Animalia" order="Harpacticoida" pageId="11" pageNumber="498" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="358088682801B10EFC35FEC9359F86D2" box="[904,1008,367,393]" italics="true" pageId="11" pageNumber="498">B. similis</emphasis>
</taxonomicName>
as the
<typeStatus id="D84FEAD82801B10EFB83FEC9320186D2" box="[1086,1134,367,393]" pageId="11" pageNumber="498">type</typeStatus>
and only species while the remaining members are assigned to
<taxonomicName id="C0F42FF92801B10EFD96FE35356286F6" authority="Mu &amp; Huys, 2021" authorityName="Mu &amp; Huys" authorityYear="2021" box="[555,781,403,429]" class="Hexanauplia" family="Miraciidae" genus="Pseudobalucopsylla" kingdom="Animalia" order="Harpacticoida" pageId="11" pageNumber="498" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="358088682801B10EFD96FE35356286F6" box="[555,781,403,429]" italics="true" pageId="11" pageNumber="498">Pseudobalucopsylla</emphasis>
</taxonomicName>
<emphasis id="358088682801B10EFCAEFE35351386F6" bold="true" box="[787,892,403,429]" pageId="11" pageNumber="498">
<taxonomicNameLabel id="2EB335132801B10EFCAEFE35351386F6" box="[787,892,403,429]" pageId="11" pageNumber="498" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
(see below)
</paragraph>
</subSubSection>
</treatment>
</document>
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