treatments-xml/data/83/21/87/832187E9FF13FF59FF746389FDF9F95C.xml

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<mods:title id="4513670784A9D4BE176620FBFE7356E4">Morphological revision of the hyperdiverse Brueelia - complex (Insecta: Phthiraptera: Ischnocera: Philopteridae) with new taxa, checklists and generic key</mods:title>
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<treatment id="832187E9FF13FF59FF746389FDF9F95C" ID-DOI="http://doi.org/10.5281/zenodo.5296961" ID-GBIF-Taxon="133100191" ID-Zenodo-Dep="5296961" LSID="urn:lsid:plazi:treatment:832187E9FF13FF59FF746389FDF9F95C" httpUri="http://treatment.plazi.org/id/832187E9FF13FF59FF746389FDF9F95C" lastPageId="154" lastPageNumber="155" pageId="153" pageNumber="154">
<subSubSection id="43926574FF13FF5AFF746389FD62FE00" box="[151,683,393,419]" pageId="153" pageNumber="154" type="nomenclature">
<paragraph id="0B3736FFFF13FF5AFF746389FD62FE00" blockId="153.[151,683,393,419]" box="[151,683,393,419]" pageId="153" pageNumber="154">
<heading id="507F8193FF13FF5AFF746389FD62FE00" bold="true" box="[151,683,393,419]" fontSize="11" level="1" pageId="153" pageNumber="154" reason="1">
<emphasis id="39FCEAEDFF13FF5AFF746389FD62FE00" bold="true" box="[151,683,393,419]" pageId="153" pageNumber="154">
<taxonomicName id="CC884D7CFF13FF5AFF746389FDD4FE00" authority="Gustafsson &amp; Bush" authorityName="Bush" authorityYear="2017" box="[151,541,393,419]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFF746389FEEAFE00" bold="true" box="[151,291,393,419]" italics="true" pageId="153" pageNumber="154">Harpactrox</emphasis>
Gustafsson &amp; Bush
</taxonomicName>
, 
<taxonomicNameLabel id="22CF5796FF13FF5AFDC96389FD62FE00" box="[554,683,393,419]" pageId="153" pageNumber="154" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="43926574FF13FF5AFF7463D0FDB4FE45" box="[151,637,464,486]" pageId="153" pageNumber="154" type="materials_examined">
<paragraph id="0B3736FFFF13FF5AFF7463D0FDB4FE45" blockId="153.[151,637,464,486]" box="[151,637,464,486]" pageId="153" pageNumber="154">
<emphasis id="39FCEAEDFF13FF5AFF7463D0FEEBFE45" bold="true" box="[151,290,464,486]" pageId="153" pageNumber="154">Type species.</emphasis>
<taxonomicName id="CC884D7CFF13FF5AFECA63D1FE37FE45" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[297,510,464,486]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="species" species="loeiensis" status="sp. nov.">
<emphasis id="39FCEAEDFF13FF5AFECA63D1FE37FE45" box="[297,510,464,486]" italics="true" pageId="153" pageNumber="154">Harpactrox loeiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22CF5796FF13FF5AFDE063D1FDB4FE45" box="[515,637,464,486]" pageId="153" pageNumber="154" rank="species">new species</taxonomicNameLabel>
</paragraph>
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<subSubSection id="43926574FF13FF5AFF74600DFB51FB29" pageId="153" pageNumber="154" type="diagnosis">
<paragraph id="0B3736FFFF13FF5AFF74600DFE65FCC9" blockId="153.[151,1437,525,2030]" pageId="153" pageNumber="154">
<emphasis id="39FCEAEDFF13FF5AFF74600DFED8FD85" bold="true" box="[151,273,525,550]" pageId="153" pageNumber="154">Diagnosis.</emphasis>
<collectingRegion id="C94CF81DFF13FF5AFEFA600EFEF2FD86" box="[281,315,526,549]" country="Maldives" name="Uthuru" pageId="153" pageNumber="154">No</collectingRegion>
other genus treated here is particularly close to 
<taxonomicName id="CC884D7CFF13FF5AFCB8600EFC14FD86" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[859,989,526,549]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFCB8600EFC14FD86" box="[859,989,526,549]" italics="true" pageId="153" pageNumber="154">Harpactrox</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22CF5796FF13FF5AFC07600DFBFAFD85" box="[996,1075,525,550]" pageId="153" pageNumber="154" rank="genus">
<emphasis id="39FCEAEDFF13FF5AFC07600DFBFAFD85" bold="true" box="[996,1075,525,550]" pageId="153" pageNumber="154">n. gen.</emphasis>
</taxonomicNameLabel>
All other genera with medianly folded parameres, proximal mesomeres that overlap with basal apodeme, and visible 
<emphasis id="39FCEAEDFF13FF5AFBAD6032FB41FDEA" box="[1102,1160,562,585]" italics="true" pageId="153" pageNumber="154">ames</emphasis>
also have 
<taxonomicName id="CC884D7CFF13FF5AFB1E6032FAEAFDEA" box="[1277,1315,562,585]" class="Insecta" family="Philopteridae" genus="Guimaraesiella" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="subSpecies" species="myiophoneae" subSpecies="aps">
<emphasis id="39FCEAEDFF13FF5AFB1E6032FAEAFDEA" box="[1277,1315,562,585]" italics="true" pageId="153" pageNumber="154">aps</emphasis>
</taxonomicName>
on at least some male tergopleurites (with the exception of some 
<taxonomicName id="CC884D7CFF13FF5AFD166055FBDAFDCD" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[757,1043,597,622]" class="Insecta" family="Philopteridae" genus="Priceiella" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Thescelovora">
<emphasis id="39FCEAEDFF13FF5AFD166055FCAAFDCD" box="[757,867,597,622]" italics="true" pageId="153" pageNumber="154">Priceiella</emphasis>
(
<emphasis id="39FCEAEDFF13FF5AFC906055FBC3FDCD" box="[883,1034,597,622]" italics="true" pageId="153" pageNumber="154">Thescelovora</emphasis>
)
</taxonomicName>
<taxonomicNameLabel id="22CF5796FF13FF5AFBF96055FB5AFDCD" box="[1050,1171,597,622]" pageId="153" pageNumber="154" rank="subGenus">
<emphasis id="39FCEAEDFF13FF5AFBF96055FB5AFDCD" bold="true" box="[1050,1171,597,622]" pageId="153" pageNumber="154">n. subgen.</emphasis>
</taxonomicNameLabel>
, 
<figureCitation id="93B32A7AFF13FF5AFB426055FAF7FDCD" box="[1185,1342,597,622]" captionStart="FIGURES 300 – 301" captionStartId="186.[151,264,1784,1806]" captionTargetBox="[164,1417,200,1754]" captionTargetId="figure@186.[151,1436,193,1762]" captionTargetPageId="186" captionText="FIGURES 300 – 301. Priceiella (Thescelovora) alliocephala n. gen., n. subgen. &amp; n. sp. ex Platylophus galericulatus ardesiacus: 300, male habitus, dorsal and ventral views. 301, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883320/files/figure.png" pageId="153" pageNumber="154">Figs 300–301</figureCitation>
), and no other genus in the 
<taxonomicName id="CC884D7CFF13FF5AFE936079FE18FD31" authorityName="Bush" authorityYear="2017" box="[368,465,633,658]" class="Insecta" family="Philopteridae" genus="Brueelia" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFF13FF5AFE936079FE18FD31" box="[368,465,633,658]" italics="true" pageId="153" pageNumber="154">Brueelia</emphasis>
</taxonomicName>
-complex have the accessory preantennal nodi found in 
<taxonomicName id="CC884D7CFF13FF5AFBB5607AFB11FD32" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[1110,1240,634,657]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="219" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFBB5607AFB11FD32" box="[1110,1240,634,657]" italics="true" pageId="153" pageNumber="154">Harpactrox</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFB0A6079FA87FD31" box="[1257,1358,633,658]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="153" pageNumber="154">Fig. 248</figureCitation>
). Only 
<taxonomicName id="CC884D7CFF13FF5AFF74609EFEE7FD16" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[151,302,670,693]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFF74609EFEE7FD16" box="[151,302,670,693]" italics="true" pageId="153" pageNumber="154">Titanomessor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22CF5796FF13FF5AFED4609DFE40FD15" box="[311,393,669,694]" pageId="153" pageNumber="154" rank="genus">
<emphasis id="39FCEAEDFF13FF5AFED4609DFE40FD15" bold="true" box="[311,393,669,694]" pageId="153" pageNumber="154">n. gen.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="93B32A7AFF13FF5AFE79609DFE34FD15" box="[410,509,669,694]" captionStart="FIGURES 212 – 216" captionStartId="135.[151,264,1883,1905]" captionTargetBox="[219,1376,221,1841]" captionTargetId="figure@135.[191,1396,193,1862]" captionTargetPageId="135" captionText="FIGURES 212 – 216. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 212, male head, dorsal and ventral views. 213, male genitalia, dorsal view. 214, mesosome, ventral view. 215, parameres, dorsal view. 216, female subgenital plate and vulval margin, ventral view. Scale for parameres same as for male genitalia." httpUri="https://zenodo.org/record/883274/files/figure.png" pageId="153" pageNumber="154">Fig. 212</figureCitation>
) has a medianly continuous dorsal preantennal suture that does not interrupt the marginal carina as in 
<taxonomicName id="CC884D7CFF13FF5AFE6E60C2FDC6FD7A" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[397,527,706,729]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="219" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFE6E60C2FDC6FD7A" box="[397,527,706,729]" italics="true" pageId="153" pageNumber="154">Harpactrox</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFDC360C1FD4DFD79" box="[544,644,705,730]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="153" pageNumber="154">Fig. 248</figureCitation>
), and only 
<taxonomicName id="CC884D7CFF13FF5AFCE760C1FC42FD79" authorityName="Ansari" authorityYear="1947" box="[772,907,705,730]" class="Insecta" family="Philopteridae" genus="Traihoriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFF13FF5AFCE760C1FC42FD79" box="[772,907,705,730]" italics="true" pageId="153" pageNumber="154">Traihoriella</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFC7F60C1FBF3FD79" box="[924,1082,705,730]" captionStart="FIGURES 265 – 269" captionStartId="166.[151,264,1947,1969]" captionTargetBox="[179,1355,202,1886]" captionTargetId="figure@166.[170,1403,193,1909]" captionTargetPageId="166" captionText="FIGURES 265 – 269. Traihoriella punjabensis Ansari, 1947 ex Psilopogon virens marshallorum: 265, male head, dorsal and ventral views. 266, male genitalia, dorsal view. 267, male mesosome, ventral view. 268, male paramere, dorsal view. 269, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883302/files/figure.png" pageId="153" pageNumber="154">Figs 266–268</figureCitation>
) have male genitalia similar to those of 
<taxonomicName id="CC884D7CFF13FF5AFEE860E6FE44FD5E" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[267,397,742,765]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="219" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFEE860E6FE44FD5E" box="[267,397,742,765]" italics="true" pageId="153" pageNumber="154">Harpactrox</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFE4660E5FD84FD5D" box="[421,589,741,766]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="153" pageNumber="154">Figs 250–252</figureCitation>
, 256–258). Like 
<taxonomicName id="CC884D7CFF13FF5AFCCA60E6FBB9FD5D" authority="Anarchonirmus" authorityName="Bush" authorityYear="2017" box="[809,1136,741,766]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFCCA60E6FC62FD5E" box="[809,939,742,765]" italics="true" pageId="153" pageNumber="154">Harpactrox</emphasis>
, 
<emphasis id="39FCEAEDFF13FF5AFC2060E5FBB9FD5D" box="[963,1136,741,766]" italics="true" pageId="153" pageNumber="154">Anarchonirmus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22CF5796FF13FF5AFB6360E5FB10FD5D" box="[1152,1241,741,766]" pageId="153" pageNumber="154" rank="genus">
<emphasis id="39FCEAEDFF13FF5AFB6360E5FB10FD5D" bold="true" box="[1152,1241,741,766]" pageId="153" pageNumber="154">n. gen.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="93B32A7AFF13FF5AFB1160E5FA93FD5D" box="[1266,1370,741,766]" captionStart="FIGURES 116 – 121" captionStartId="80.[151,264,1782,1804]" captionTargetBox="[162,1428,197,1724]" captionTargetId="figure@80.[151,1436,193,1760]" captionTargetPageId="80" captionText="FIGURES 116 – 121. Anarchonirmus albovittatus n. gen. &amp; n. sp. ex Pomatostomus temporalis strepitans: 116, male head, dorsal and ventral view; dorsal sensilla and pts not visible in the material examined. 117, female antenna, ventral view. 118, male genitalia, dorsal view. 119, male mesosome, ventral view. 120, male paramere, dorsal view. 121, female subgenital plate and vulval margin, ventral view (anterior dotted line outline pigmented part of subgenital plate, posterior dotted line extent of marginal thickening)." httpUri="https://zenodo.org/record/883224/files/figure.png" pageId="153" pageNumber="154">Fig. 116</figureCitation>
) and 
<taxonomicName id="CC884D7CFF13FF5AFF746109FE81FC81" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[151,328,777,802]" class="Insecta" family="Philopteridae" genus="Saepocephalum" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFF746109FE81FC81" box="[151,328,777,802]" italics="true" pageId="153" pageNumber="154">Saepocephalum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22CF5796FF13FF5AFEB36109FE56FC81" box="[336,415,777,802]" pageId="153" pageNumber="154" rank="genus">
<emphasis id="39FCEAEDFF13FF5AFEB36109FE56FC81" bold="true" box="[336,415,777,802]" pageId="153" pageNumber="154">n. gen.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="93B32A7AFF13FF5AFE536109FDD9FC81" box="[432,528,777,802]" captionStart="FIGURES 240 – 245" captionStartId="152.[151,264,1915,1937]" captionTargetBox="[214,1368,213,1878]" captionTargetId="figure@152.[192,1395,193,1894]" captionTargetPageId="152" captionText="FIGURES 240 – 245. Saepocephalum stephenfryi n. gen. &amp; n. sp. ex Corcorax melanoramphos: 240, male head, dorsal and ventral views. 241, female antenna, ventral view. 242, male genitalia, dorsal view. 243, male mesosome, ventral view. 244, male paramere, dorsal view. 245, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883290/files/figure.png" pageId="153" pageNumber="154">Fig. 240</figureCitation>
) both have a marginal carina that is fused to the ventral anterior plate and is not displaced or interrupted by the clypeo-labral suture. For additional comparisons, see the description of 
<taxonomicName id="CC884D7CFF13FF5AFF746151FE81FCC9" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[151,328,849,874]" class="Insecta" family="Philopteridae" genus="Saepocephalum" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="216" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFF746151FE81FCC9" box="[151,328,849,874]" italics="true" pageId="153" pageNumber="154">Saepocephalum</emphasis>
</taxonomicName>
(above).
</paragraph>
<paragraph id="0B3736FFFF13FF5AFF246176FB51FB29" blockId="153.[151,1437,525,2030]" pageId="153" pageNumber="154">
<taxonomicName id="CC884D7CFF13FF5AFF246176FE80FC2E" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[199,329,886,909]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="219" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFF246176FE80FC2E" box="[199,329,886,909]" italics="true" pageId="153" pageNumber="154">Harpactrox</emphasis>
</taxonomicName>
is separated from 
<taxonomicName id="CC884D7CFF13FF5AFDFA6176FD79FC2E" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[537,688,886,909]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFDFA6176FD79FC2E" box="[537,688,886,909]" italics="true" pageId="153" pageNumber="154">Titanomessor</emphasis>
</taxonomicName>
by the following characters: marginal carina displaced at osculum and clypeo-labral suture reaches anterior margin of head in 
<taxonomicName id="CC884D7CFF13FF5AFCA3619AFC1EFC12" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[832,983,922,945]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFCA3619AFC1EFC12" box="[832,983,922,945]" italics="true" pageId="153" pageNumber="154">Titanomessor</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFC0B6199FB85FC11" box="[1000,1100,921,946]" captionStart="FIGURES 212 – 216" captionStartId="135.[151,264,1883,1905]" captionTargetBox="[219,1376,221,1841]" captionTargetId="figure@135.[191,1396,193,1862]" captionTargetPageId="135" captionText="FIGURES 212 – 216. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 212, male head, dorsal and ventral views. 213, male genitalia, dorsal view. 214, mesosome, ventral view. 215, parameres, dorsal view. 216, female subgenital plate and vulval margin, ventral view. Scale for parameres same as for male genitalia." httpUri="https://zenodo.org/record/883274/files/figure.png" pageId="153" pageNumber="154">Fig. 212</figureCitation>
), but not in 
<taxonomicName id="CC884D7CFF13FF5AFB3B619AFA93FC12" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[1240,1370,922,945]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="219" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFB3B619AFA93FC12" box="[1240,1370,922,945]" italics="true" pageId="153" pageNumber="154">Harpactrox</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFA896199FF0DFC75" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="153" pageNumber="154">Figs 248</figureCitation>
, 255, 260). In 
<emphasis id="39FCEAEDFF13FF5AFE8461BEFD9FFC75" box="[359,598,958,982]" italics="true" pageId="153" pageNumber="154">
<taxonomicName id="CC884D7CFF13FF5AFE8461BEFDE5FC75" box="[359,556,958,982]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="species" species="mts">Titanomessor mts</taxonomicName>
<quantity id="CC709B1AFF13FF5AFDCF61BEFD9FFC75" box="[556,598,958,982]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" metricValueMax="1.27" metricValueMin="0.762" pageId="153" pageNumber="154" unit="in" value="4.0" valueMax="5.0" valueMin="3.0">3–5</quantity>
</emphasis>
in males and 
<emphasis id="39FCEAEDFF13FF5AFD1061BEFC8AFC75" box="[755,835,958,982]" italics="true" pageId="153" pageNumber="154">
<taxonomicName id="CC884D7CFF13FF5AFD1061BEFCD0FC75" box="[755,793,958,982]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
<quantity id="CC709B1AFF13FF5AFCFA61BEFC8AFC75" box="[793,835,958,982]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="10.16" metricValueMin="7.62" pageId="153" pageNumber="154" unit="in" value="3.5" valueMax="4.0" valueMin="3.0">3–4</quantity>
</emphasis>
in females are mesosetae (
<figureCitation id="93B32A7AFF13FF5AFB9161BDFAD9FC75" box="[1138,1296,957,982]" captionStart="FIGURES 210 – 211" captionStartId="134.[151,264,1837,1859]" captionTargetBox="[193,1373,229,1811]" captionTargetId="figure@134.[151,1436,193,1815]" captionTargetPageId="134" captionText="FIGURES 210 – 211. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 210, male habitus, dorsal and ventral views. 211, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883272/files/figure.png" pageId="153" pageNumber="154">Figs 210–211</figureCitation>
), whereas in 
<taxonomicName id="CC884D7CFF13FF5AFF7461E2FED0FC5A" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[151,281,994,1017]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="219" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFF7461E2FED0FC5A" box="[151,281,994,1017]" italics="true" pageId="153" pageNumber="154">Harpactrox</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFECB61E1FE47FC59" box="[296,398,993,1018]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="153" pageNumber="154">Figs 248</figureCitation>
, 
<figureCitation id="93B32A7AFF13FF5AFE7961E1FE0CFC59" box="[410,453,993,1018]" captionStart="FIGURES 260 – 262" captionStartId="161.[151,264,1970,1992]" captionTargetBox="[194,1356,212,1926]" captionTargetId="figure@161.[163,1404,193,1927]" captionTargetPageId="161" captionText="FIGURES 260 – 262. Harpactrox pontifrons n. gen. &amp; n. sp. ex Harpactes ardens ardens: 260, female head, dorsal and ventral views. 261, female habitus, dorsal and ventral views. 262, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883298/files/figure.png" pageId="153" pageNumber="154">260</figureCitation>
) all 
<taxonomicName id="CC884D7CFF13FF5AFE1A61E2FDE9FC5A" box="[505,544,994,1017]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="species" species="mts">
<emphasis id="39FCEAEDFF13FF5AFE1A61E2FDE9FC5A" box="[505,544,994,1017]" italics="true" pageId="153" pageNumber="154">mts</emphasis>
</taxonomicName>
are microsetae in both sexes [except 
<taxonomicName id="CC884D7CFF13FF5AFC2B61E2FBBBFC59" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[968,1138,993,1018]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="species" species="geminodus" status="sp. nov.">
<emphasis id="39FCEAEDFF13FF5AFC2B61E2FC23FC5A" box="[968,1002,994,1017]" italics="true" pageId="153" pageNumber="154">Ha</emphasis>
. 
<emphasis id="39FCEAEDFF13FF5AFC1A61E1FBBBFC59" box="[1017,1138,993,1018]" italics="true" pageId="153" pageNumber="154">geminodus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22CF5796FF13FF5AFB9961E1FB70FC59" box="[1146,1209,993,1018]" pageId="153" pageNumber="154" rank="species">
<emphasis id="39FCEAEDFF13FF5AFB9961E1FB70FC59" bold="true" box="[1146,1209,993,1018]" pageId="153" pageNumber="154">n. sp.</emphasis>
</taxonomicNameLabel>
(Fig. 255) in which 
<emphasis id="39FCEAEDFF13FF5AFF746606FF05FBBD" box="[151,204,1030,1054]" italics="true" pageId="153" pageNumber="154">
<taxonomicName id="CC884D7CFF13FF5AFF746606FF77FBBD" box="[151,190,1030,1054]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
3
</emphasis>
is slightly longer than other 
<taxonomicName id="CC884D7CFF13FF5AFDFB6606FDF6FBBE" box="[536,575,1030,1053]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="species" species="mts">
<emphasis id="39FCEAEDFF13FF5AFDFB6606FDF6FBBE" box="[536,575,1030,1053]" italics="true" pageId="153" pageNumber="154">mts</emphasis>
</taxonomicName>
]. Parameral heads are blunt and not folded medianly in 
<taxonomicName id="CC884D7CFF13FF5AFB256606FA94FBBE" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[1222,1373,1030,1053]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFB256606FA94FBBE" box="[1222,1373,1030,1053]" italics="true" pageId="153" pageNumber="154">Titanomessor</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFA8D6605FF0DFBE1" captionStart="FIGURES 212 – 216" captionStartId="135.[151,264,1883,1905]" captionTargetBox="[219,1376,221,1841]" captionTargetId="figure@135.[191,1396,193,1862]" captionTargetPageId="135" captionText="FIGURES 212 – 216. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 212, male head, dorsal and ventral views. 213, male genitalia, dorsal view. 214, mesosome, ventral view. 215, parameres, dorsal view. 216, female subgenital plate and vulval margin, ventral view. Scale for parameres same as for male genitalia." httpUri="https://zenodo.org/record/883274/files/figure.png" pageId="153" pageNumber="154">Fig. 215</figureCitation>
), but parameral heads are folded medianly in 
<taxonomicName id="CC884D7CFF13FF5AFD01662AFCADFBE2" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[738,868,1066,1089]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="219" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFD01662AFCADFBE2" box="[738,868,1066,1089]" italics="true" pageId="153" pageNumber="154">Harpactrox</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFC946629FC16FBE1" box="[887,991,1065,1090]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="153" pageNumber="154">Figs 251</figureCitation>
, 258). Mesosomal lobes are wide and gonopore is ventral and open both distally and proximally in 
<taxonomicName id="CC884D7CFF13FF5AFCA3664EFC1EFBC6" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[832,983,1102,1125]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFCA3664EFC1EFBC6" box="[832,983,1102,1125]" italics="true" pageId="153" pageNumber="154">Titanomessor</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFC05664DFB81FBC5" box="[998,1096,1101,1126]" captionStart="FIGURES 212 – 216" captionStartId="135.[151,264,1883,1905]" captionTargetBox="[219,1376,221,1841]" captionTargetId="figure@135.[191,1396,193,1862]" captionTargetPageId="135" captionText="FIGURES 212 – 216. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 212, male head, dorsal and ventral views. 213, male genitalia, dorsal view. 214, mesosome, ventral view. 215, parameres, dorsal view. 216, female subgenital plate and vulval margin, ventral view. Scale for parameres same as for male genitalia." httpUri="https://zenodo.org/record/883274/files/figure.png" pageId="153" pageNumber="154">Fig. 214</figureCitation>
), whereas in 
<taxonomicName id="CC884D7CFF13FF5AFB39664EFA95FBC6" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[1242,1372,1102,1125]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="219" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFB39664EFA95FBC6" box="[1242,1372,1102,1125]" italics="true" pageId="153" pageNumber="154">Harpactrox</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFA89664DFF0DFB29" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="153" pageNumber="154">Figs 250</figureCitation>
, 257) the lobes are slender and the gonopore is terminal, and closed at least proximally.
</paragraph>
</subSubSection>
<subSubSection id="43926574FF13FF59FF246696FBF1FC6B" lastPageId="154" lastPageNumber="155" pageId="153" pageNumber="154" type="description">
<paragraph id="0B3736FFFF13FF5AFF246696FE30F999" blockId="153.[151,1437,525,2030]" pageId="153" pageNumber="154">
<taxonomicName id="CC884D7CFF13FF5AFF246696FE80FB0E" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[199,329,1174,1197]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="219" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFF246696FE80FB0E" box="[199,329,1174,1197]" italics="true" pageId="153" pageNumber="154">Harpactrox</emphasis>
</taxonomicName>
and 
<taxonomicName id="CC884D7CFF13FF5AFE9D6695FDCCFB0D" authorityName="Ansari" authorityYear="1947" box="[382,517,1173,1198]" class="Insecta" family="Philopteridae" genus="Traihoriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFF13FF5AFE9D6695FDCCFB0D" box="[382,517,1173,1198]" italics="true" pageId="153" pageNumber="154">Traihoriella</emphasis>
</taxonomicName>
can be separated by the following non-genitalic characters: 
<taxonomicName id="CC884D7CFF13FF5AFB466696FB05FB0E" box="[1189,1228,1174,1197]" class="Insecta" family="Philopteridae" genus="Guimaraesiella" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="subSpecies" species="myiophoneae" subSpecies="aps">
<emphasis id="39FCEAEDFF13FF5AFB466696FB05FB0E" box="[1189,1228,1174,1197]" italics="true" pageId="153" pageNumber="154">aps</emphasis>
</taxonomicName>
present on at least some segments in male 
<taxonomicName id="CC884D7CFF13FF5AFE4F66B9FDFDFB71" authorityName="Ansari" authorityYear="1947" box="[428,564,1209,1234]" class="Insecta" family="Philopteridae" genus="Traihoriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFF13FF5AFE4F66B9FDFDFB71" box="[428,564,1209,1234]" italics="true" pageId="153" pageNumber="154">Traihoriella</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFDA666B9FD67FB71" box="[581,686,1209,1234]" captionStart="FIGURES 263 – 264" captionStartId="165.[151,264,1976,1998]" captionTargetBox="[196,1407,327,1954]" captionTargetId="figure@165.[151,1436,303,1993]" captionTargetPageId="165" captionText="FIGURES 263 – 264. Traihoriella punjabensis Ansari, 1947 ex Psilopogon virens marshallorum: 263, male habitus, dorsal and ventral views. 264, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883300/files/figure.png" pageId="153" pageNumber="154">Figs 263</figureCitation>
, 
<figureCitation id="93B32A7AFF13FF5AFD5866BAFD20FB71" box="[699,745,1210,1234]" captionStart="FIGURES 270 – 271" captionStartId="168.[151,264,1980,2002]" captionTargetBox="[204,1408,582,1960]" captionTargetId="figure@168.[151,1436,569,1974]" captionTargetPageId="168" captionText="FIGURES 270 – 271. Traihoriella laticeps (Piaget, 1880) n. comb. ex Aulacorhynchus prasinus atrogularis: 270 male habitus, dorsal and ventral views. 271, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883304/files/figure.png" pageId="153" pageNumber="154">270</figureCitation>
; 
<tableCitation id="460A0344FF13FF5AFD1466B9FC87FB71" box="[759,846,1209,1234]" captionStart="TABLE 2" captionStartId="30.[151,239,796,819]" captionText="TABLE 2. Summary of abdominal chaetotaxy of segments II – VIII for the Brueelia - complex, including both genera and subgenera. In genera where chaetotaxy differs between species group (marked with *), only that of the type species is given here, and more detailed accounts are given separately under the respective genus entries and in Tables 3 – 12. Trichoid setae of segment VIII are present in all genera, and are not listed. Abbreviations: ps = paratergal seta; aps = accessory post-spiracular seta; psps = principal post-spiracular seta; tps = tergal posterior seta; ss = sutural seta; sts = sternal seta." pageId="153" pageNumber="154">Table 2</tableCitation>
), but never in 
<taxonomicName id="CC884D7CFF13FF5AFC1966BAFBB5FB72" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[1018,1148,1210,1233]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="219" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFC1966BAFBB5FB72" box="[1018,1148,1210,1233]" italics="true" pageId="153" pageNumber="154">Harpactrox</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFB6E66B9FB3FFB71" box="[1165,1270,1209,1234]" captionStart="FIGURES 246 – 247" captionStartId="155.[151,264,1922,1944]" captionTargetBox="[199,1341,238,1835]" captionTargetId="figure@155.[164,1388,195,1874]" captionTargetPageId="155" captionText="FIGURES 246 – 247. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 246, male habitus, dorsal and ventral views; only examined male with distorted abdomen, and shape of abdomen here reproduced as closely as possible; anterior sternites and some ventral setae missing in only available male specimen, and here included tentatively. 247, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883292/files/figure.png" pageId="153" pageNumber="154">Figs 246</figureCitation>
, 
<figureCitation id="93B32A7AFF13FF5AFAE066B9FAF8FB71" box="[1283,1329,1209,1234]" captionStart="FIGURES 253 – 254" captionStartId="158.[151,264,1955,1977]" captionTargetBox="[207,1386,225,1934]" captionTargetId="figure@158.[151,1436,193,1934]" captionTargetPageId="158" captionText="FIGURES 253 – 254. Harpactrox geminodus n. gen. &amp; n. sp. ex Harpactes erythrocephalus erythrocephalus: 253, male habitus, dorsal and ventral views. 254, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883296/files/figure.png" pageId="153" pageNumber="154">253</figureCitation>
); clypeolabral suture reaches anterior margin of head and marginal carina is displaced dorsally and ventrally at osculum in 
<taxonomicName id="CC884D7CFF13FF5AFF746701FED7FAB9" authorityName="Ansari" authorityYear="1947" box="[151,286,1281,1306]" class="Insecta" family="Philopteridae" genus="Traihoriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFF13FF5AFF746701FED7FAB9" box="[151,286,1281,1306]" italics="true" pageId="153" pageNumber="154">Traihoriella</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFED36701FE50FAB9" box="[304,409,1281,1306]" captionStart="FIGURES 265 – 269" captionStartId="166.[151,264,1947,1969]" captionTargetBox="[179,1355,202,1886]" captionTargetId="figure@166.[170,1403,193,1909]" captionTargetPageId="166" captionText="FIGURES 265 – 269. Traihoriella punjabensis Ansari, 1947 ex Psilopogon virens marshallorum: 265, male head, dorsal and ventral views. 266, male genitalia, dorsal view. 267, male mesosome, ventral view. 268, male paramere, dorsal view. 269, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883302/files/figure.png" pageId="153" pageNumber="154">Figs 265</figureCitation>
, 
<figureCitation id="93B32A7AFF13FF5AFE456702FE1DFAB9" box="[422,468,1282,1306]" captionStart="FIGURES 272 – 276" captionStartId="169.[151,264,1950,1972]" captionTargetBox="[176,1364,204,1880]" captionTargetId="figure@169.[170,1398,193,1909]" captionTargetPageId="169" captionText="FIGURES 272 – 276. Traihoriella laticeps (Piaget, 1880) n. comb. ex Aulacorhynchus prasinus atrogularis: 272, male head, dorsal and ventral views. 273, male genitalia, dorsal view. 274, male mesosome, ventral view. 275, male paramere, dorsal view. 276, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883306/files/figure.png" pageId="153" pageNumber="154">272</figureCitation>
), but not in 
<taxonomicName id="CC884D7CFF13FF5AFD866702FD2EFABA" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[613,743,1282,1305]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="219" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFD866702FD2EFABA" box="[613,743,1282,1305]" italics="true" pageId="153" pageNumber="154">Harpactrox</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFD1A6701FCA8FAB9" box="[761,865,1281,1306]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="153" pageNumber="154">Figs 248</figureCitation>
, 255, 260); dorsal preantennal suture is absent in 
<taxonomicName id="CC884D7CFF13FF5AFF746725FED7FA9D" authorityName="Ansari" authorityYear="1947" box="[151,286,1317,1342]" class="Insecta" family="Philopteridae" genus="Traihoriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFF13FF5AFF746725FED7FA9D" box="[151,286,1317,1342]" italics="true" pageId="153" pageNumber="154">Traihoriella</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFECD6725FE5DFA9D" box="[302,404,1317,1342]" captionStart="FIGURES 265 – 269" captionStartId="166.[151,264,1947,1969]" captionTargetBox="[179,1355,202,1886]" captionTargetId="figure@166.[170,1403,193,1909]" captionTargetPageId="166" captionText="FIGURES 265 – 269. Traihoriella punjabensis Ansari, 1947 ex Psilopogon virens marshallorum: 265, male head, dorsal and ventral views. 266, male genitalia, dorsal view. 267, male mesosome, ventral view. 268, male paramere, dorsal view. 269, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883302/files/figure.png" pageId="153" pageNumber="154">Figs 265</figureCitation>
, 
<figureCitation id="93B32A7AFF13FF5AFE436726FE07FA9D" box="[416,462,1318,1342]" captionStart="FIGURES 272 – 276" captionStartId="169.[151,264,1950,1972]" captionTargetBox="[176,1364,204,1880]" captionTargetId="figure@169.[170,1398,193,1909]" captionTargetPageId="169" captionText="FIGURES 272 – 276. Traihoriella laticeps (Piaget, 1880) n. comb. ex Aulacorhynchus prasinus atrogularis: 272, male head, dorsal and ventral views. 273, male genitalia, dorsal view. 274, male mesosome, ventral view. 275, male paramere, dorsal view. 276, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883306/files/figure.png" pageId="153" pageNumber="154">272</figureCitation>
), but present in 
<taxonomicName id="CC884D7CFF13FF5AFD606726FCCCFA9E" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[643,773,1318,1341]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="219" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFD606726FCCCFA9E" box="[643,773,1318,1341]" italics="true" pageId="153" pageNumber="154">Harpactrox</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFCF76725FCBEFA9D" box="[788,887,1317,1342]" captionStart="FIGURES 184 – 188" captionStartId="121.[151,264,1915,1937]" captionTargetBox="[198,1349,204,1875]" captionTargetId="figure@121.[192,1395,193,1894]" captionTargetPageId="121" captionText="FIGURES 184 – 188. Turdinirmus merulensis (Denny, 1842) ex Turdus merula: 184, male head, dorsal and ventral views. 185, male genitalia, dorsal view. 186, male mesosome, ventral view. 187, male paramere, dorsal view. 188, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883260/files/figure.png" pageId="153" pageNumber="154">Fig. 184</figureCitation>
); 
<emphasis id="39FCEAEDFF13FF5AFC6F6726FC08FA9D" box="[908,961,1318,1342]" italics="true" pageId="153" pageNumber="154">
<taxonomicName id="CC884D7CFF13FF5AFC6F6726FC7AFA9D" box="[908,947,1318,1342]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
3
</emphasis>
are macrosetae in 
<taxonomicName id="CC884D7CFF13FF5AFB766725FAD5FA9D" authorityName="Ansari" authorityYear="1947" box="[1173,1308,1317,1342]" class="Insecta" family="Philopteridae" genus="Traihoriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFF13FF5AFB766725FAD5FA9D" box="[1173,1308,1317,1342]" italics="true" pageId="153" pageNumber="154">Traihoriella</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFACF6725FA47FA9D" box="[1324,1422,1317,1342]" captionStart="FIGURES 196 – 197" captionStartId="127.[151,264,1947,1969]" captionTargetBox="[187,1385,227,1925]" captionTargetId="figure@127.[151,1436,193,1925]" captionTargetPageId="127" captionText="FIGURES 196 – 197. Maculinirmus mundus (Nitzsch [in Giebel], 1866) ex Oriolus oriolus oriolus: 196, male habitus, dorsal and ventral views. 197, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883266/files/figure.png" pageId="153" pageNumber="154">Fig. 197</figureCitation>
), but microsetae in 
<taxonomicName id="CC884D7CFF13FF5AFE84674AFE20FAC2" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[359,489,1354,1377]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="219" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFE84674AFE20FAC2" box="[359,489,1354,1377]" italics="true" pageId="153" pageNumber="154">Harpactrox</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFE196749FDABFAC1" box="[506,610,1353,1378]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="153" pageNumber="154">Figs 248</figureCitation>
, 255, 260). 
<collectingRegion id="C94CF81DFF13FF5AFD0C6749FCE0FAC1" box="[751,809,1353,1378]" country="Maldives" name="Maale" pageId="153" pageNumber="154">Male</collectingRegion>
genitalia of 
<taxonomicName id="CC884D7CFF13FF5AFC5E674AFBF6FAC2" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[957,1087,1354,1377]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="219" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFC5E674AFBF6FAC2" box="[957,1087,1354,1377]" italics="true" pageId="153" pageNumber="154">Harpactrox</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFBB36749FB38FAC1" box="[1104,1265,1353,1378]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="153" pageNumber="154">Figs 249–251</figureCitation>
, 256–258) and 
<taxonomicName id="CC884D7CFF13FF5AFF74676DFED7FA25" authorityName="Ansari" authorityYear="1947" box="[151,286,1389,1414]" class="Insecta" family="Philopteridae" genus="Traihoriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFF13FF5AFF74676DFED7FA25" box="[151,286,1389,1414]" italics="true" pageId="153" pageNumber="154">Traihoriella</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFECE676DFE03FA25" box="[301,458,1389,1414]" captionStart="FIGURES 265 – 269" captionStartId="166.[151,264,1947,1969]" captionTargetBox="[179,1355,202,1886]" captionTargetId="figure@166.[170,1403,193,1909]" captionTargetPageId="166" captionText="FIGURES 265 – 269. Traihoriella punjabensis Ansari, 1947 ex Psilopogon virens marshallorum: 265, male head, dorsal and ventral views. 266, male genitalia, dorsal view. 267, male mesosome, ventral view. 268, male paramere, dorsal view. 269, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883302/files/figure.png" pageId="153" pageNumber="154">Figs 266–268</figureCitation>
, 
<figureCitation id="93B32A7AFF13FF5AFE37676DFDFEFA25" box="[468,567,1389,1414]" captionStart="FIGURES 272 – 276" captionStartId="169.[151,264,1950,1972]" captionTargetBox="[176,1364,204,1880]" captionTargetId="figure@169.[170,1398,193,1909]" captionTargetPageId="169" captionText="FIGURES 272 – 276. Traihoriella laticeps (Piaget, 1880) n. comb. ex Aulacorhynchus prasinus atrogularis: 272, male head, dorsal and ventral views. 273, male genitalia, dorsal view. 274, male mesosome, ventral view. 275, male paramere, dorsal view. 276, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883306/files/figure.png" pageId="153" pageNumber="154">273–275</figureCitation>
) are very similar, and as more species of each genus are discovered, it may be impossible to separate these two genera on male genitalia alone, however they are separable on the following characters: 
<emphasis id="39FCEAEDFF13FF5AFEFB67B6FE9BFA6E" box="[280,338,1462,1485]" italics="true" pageId="153" pageNumber="154">ames</emphasis>
microsetae are present in 
<taxonomicName id="CC884D7CFF13FF5AFD9967B6FD35FA6E" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[634,764,1462,1485]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="219" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFD9967B6FD35FA6E" box="[634,764,1462,1485]" italics="true" pageId="153" pageNumber="154">Harpactrox</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFCE867B5FCBBFA6D" box="[779,882,1461,1486]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="153" pageNumber="154">Figs 250</figureCitation>
, 257), but absent in 
<taxonomicName id="CC884D7CFF13FF5AFBBB67B5FB16FA6D" authorityName="Ansari" authorityYear="1947" box="[1112,1247,1461,1486]" class="Insecta" family="Philopteridae" genus="Traihoriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFF13FF5AFBBB67B5FB16FA6D" box="[1112,1247,1461,1486]" italics="true" pageId="153" pageNumber="154">Traihoriella</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFB0C67B5FA9CFA6D" box="[1263,1365,1461,1486]" captionStart="FIGURES 265 – 269" captionStartId="166.[151,264,1947,1969]" captionTargetBox="[179,1355,202,1886]" captionTargetId="figure@166.[170,1403,193,1909]" captionTargetPageId="166" captionText="FIGURES 265 – 269. Traihoriella punjabensis Ansari, 1947 ex Psilopogon virens marshallorum: 265, male head, dorsal and ventral views. 266, male genitalia, dorsal view. 267, male mesosome, ventral view. 268, male paramere, dorsal view. 269, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883302/files/figure.png" pageId="153" pageNumber="154">Figs 267</figureCitation>
, 
<figureCitation id="93B32A7AFF13FF5AFA8267B6FA46FA6D" box="[1377,1423,1462,1486]" captionStart="FIGURES 272 – 276" captionStartId="169.[151,264,1950,1972]" captionTargetBox="[176,1364,204,1880]" captionTargetId="figure@169.[170,1398,193,1909]" captionTargetPageId="169" captionText="FIGURES 272 – 276. Traihoriella laticeps (Piaget, 1880) n. comb. ex Aulacorhynchus prasinus atrogularis: 272, male head, dorsal and ventral views. 273, male genitalia, dorsal view. 274, male mesosome, ventral view. 275, male paramere, dorsal view. 276, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883306/files/figure.png" pageId="153" pageNumber="154">274</figureCitation>
); 
<emphasis id="39FCEAEDFF13FF5AFF7467DAFF0FFA51" box="[151,198,1498,1522]" italics="true" pageId="153" pageNumber="154">pst2</emphasis>
microsetae in 
<taxonomicName id="CC884D7CFF13FF5AFE9367DAFE3BFA52" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[368,498,1498,1521]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="219" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFE9367DAFE3BFA52" box="[368,498,1498,1521]" italics="true" pageId="153" pageNumber="154">Harpactrox</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFDE067D9FDA3FA51" box="[515,618,1497,1522]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="153" pageNumber="154">Figs 251</figureCitation>
, 258), but sensilla in 
<taxonomicName id="CC884D7CFF13FF5AFC8567D9FC24FA51" authorityName="Ansari" authorityYear="1947" box="[870,1005,1497,1522]" class="Insecta" family="Philopteridae" genus="Traihoriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFF13FF5AFC8567D9FC24FA51" box="[870,1005,1497,1522]" italics="true" pageId="153" pageNumber="154">Traihoriella</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFC1D67D9FBAFFA51" box="[1022,1126,1497,1522]" captionStart="FIGURES 265 – 269" captionStartId="166.[151,264,1947,1969]" captionTargetBox="[179,1355,202,1886]" captionTargetId="figure@166.[170,1403,193,1909]" captionTargetPageId="166" captionText="FIGURES 265 – 269. Traihoriella punjabensis Ansari, 1947 ex Psilopogon virens marshallorum: 265, male head, dorsal and ventral views. 266, male genitalia, dorsal view. 267, male mesosome, ventral view. 268, male paramere, dorsal view. 269, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883302/files/figure.png" pageId="153" pageNumber="154">Figs 268</figureCitation>
, 
<figureCitation id="93B32A7AFF13FF5AFB9767D9FB6BFA51" box="[1140,1186,1497,1522]" captionStart="FIGURES 272 – 276" captionStartId="169.[151,264,1950,1972]" captionTargetBox="[176,1364,204,1880]" captionTargetId="figure@169.[170,1398,193,1909]" captionTargetPageId="169" captionText="FIGURES 272 – 276. Traihoriella laticeps (Piaget, 1880) n. comb. ex Aulacorhynchus prasinus atrogularis: 272, male head, dorsal and ventral views. 273, male genitalia, dorsal view. 274, male mesosome, ventral view. 275, male paramere, dorsal view. 276, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883306/files/figure.png" pageId="153" pageNumber="154">275</figureCitation>
). Neither 
<taxonomicName id="CC884D7CFF13FF5AFAF767D9FA52FA51" authorityName="Ansari" authorityYear="1947" box="[1300,1435,1497,1522]" class="Insecta" family="Philopteridae" genus="Traihoriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFF13FF5AFAF767D9FA52FA51" box="[1300,1435,1497,1522]" italics="true" pageId="153" pageNumber="154">Traihoriella</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFF7C67FDFECDF9B5" box="[159,260,1533,1558]" captionStart="FIGURES 265 – 269" captionStartId="166.[151,264,1947,1969]" captionTargetBox="[179,1355,202,1886]" captionTargetId="figure@166.[170,1403,193,1909]" captionTargetPageId="166" captionText="FIGURES 265 – 269. Traihoriella punjabensis Ansari, 1947 ex Psilopogon virens marshallorum: 265, male head, dorsal and ventral views. 266, male genitalia, dorsal view. 267, male mesosome, ventral view. 268, male paramere, dorsal view. 269, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883302/files/figure.png" pageId="153" pageNumber="154">Figs 269</figureCitation>
, 
<figureCitation id="93B32A7AFF13FF5AFEED67FDFEF0F9B5" box="[270,313,1533,1558]" captionStart="FIGURES 272 – 276" captionStartId="169.[151,264,1950,1972]" captionTargetBox="[176,1364,204,1880]" captionTargetId="figure@169.[170,1398,193,1909]" captionTargetPageId="169" captionText="FIGURES 272 – 276. Traihoriella laticeps (Piaget, 1880) n. comb. ex Aulacorhynchus prasinus atrogularis: 272, male head, dorsal and ventral views. 273, male genitalia, dorsal view. 274, male mesosome, ventral view. 275, male paramere, dorsal view. 276, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883306/files/figure.png" pageId="153" pageNumber="154">276</figureCitation>
) nor 
<taxonomicName id="CC884D7CFF13FF5AFE9767FEFE3FF9B6" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[372,502,1534,1557]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="219" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFE9767FEFE3FF9B6" box="[372,502,1534,1557]" italics="true" pageId="153" pageNumber="154">Harpactrox</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFDE067FDFDA0F9B5" box="[515,617,1533,1558]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="153" pageNumber="154">Figs 252</figureCitation>
, 259, 262) have cross-pieces, and in both genera the 
<emphasis id="39FCEAEDFF13FF5AFB5167FEFB1EF9B6" box="[1202,1239,1534,1557]" italics="true" pageId="153" pageNumber="154">vos</emphasis>
are at least partly situated on the subgenital plate.
</paragraph>
<paragraph id="0B3736FFFF13FF5AFF246445FBBCF801" blockId="153.[151,1437,525,2030]" pageId="153" pageNumber="154">
The ventral preantennal structure of 
<taxonomicName id="CC884D7CFF13FF5AFD826446FD2AF9FE" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[609,739,1606,1629]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="219" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFD826446FD2AF9FE" box="[609,739,1606,1629]" italics="true" pageId="153" pageNumber="154">Harpactrox</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFD126445FC9EF9FD" box="[753,855,1605,1630]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="153" pageNumber="154">Figs 248</figureCitation>
, 255, 260) is similar to that of 
<taxonomicName id="CC884D7CFF13FF5AFB516445FA96F9FD" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[1202,1375,1605,1630]" class="Insecta" family="Philopteridae" genus="Anarchonirmus" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFB516445FA96F9FD" box="[1202,1375,1605,1630]" italics="true" pageId="153" pageNumber="154">Anarchonirmus</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFA8D6445FF0AF921" captionStart="FIGURES 116 – 121" captionStartId="80.[151,264,1782,1804]" captionTargetBox="[162,1428,197,1724]" captionTargetId="figure@80.[151,1436,193,1760]" captionTargetPageId="80" captionText="FIGURES 116 – 121. Anarchonirmus albovittatus n. gen. &amp; n. sp. ex Pomatostomus temporalis strepitans: 116, male head, dorsal and ventral view; dorsal sensilla and pts not visible in the material examined. 117, female antenna, ventral view. 118, male genitalia, dorsal view. 119, male mesosome, ventral view. 120, male paramere, dorsal view. 121, female subgenital plate and vulval margin, ventral view (anterior dotted line outline pigmented part of subgenital plate, posterior dotted line extent of marginal thickening)." httpUri="https://zenodo.org/record/883224/files/figure.png" pageId="153" pageNumber="154">Fig. 116</figureCitation>
), but there are few other similarities. Female subgenital plate flares into a medianly displaced cross-piece in 
<taxonomicName id="CC884D7CFF13FF5AFF74648DFE8DF905" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[151,324,1677,1702]" class="Insecta" family="Philopteridae" genus="Anarchonirmus" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFF74648DFE8DF905" box="[151,324,1677,1702]" italics="true" pageId="153" pageNumber="154">Anarchonirmus</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFEB7648DFE7AF905" box="[340,435,1677,1702]" captionStart="FIGURES 116 – 121" captionStartId="80.[151,264,1782,1804]" captionTargetBox="[162,1428,197,1724]" captionTargetId="figure@80.[151,1436,193,1760]" captionTargetPageId="80" captionText="FIGURES 116 – 121. Anarchonirmus albovittatus n. gen. &amp; n. sp. ex Pomatostomus temporalis strepitans: 116, male head, dorsal and ventral view; dorsal sensilla and pts not visible in the material examined. 117, female antenna, ventral view. 118, male genitalia, dorsal view. 119, male mesosome, ventral view. 120, male paramere, dorsal view. 121, female subgenital plate and vulval margin, ventral view (anterior dotted line outline pigmented part of subgenital plate, posterior dotted line extent of marginal thickening)." httpUri="https://zenodo.org/record/883224/files/figure.png" pageId="153" pageNumber="154">Fig. 121</figureCitation>
) but not in 
<taxonomicName id="CC884D7CFF13FF5AFDD4648EFD70F906" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[567,697,1678,1701]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="219" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFDD4648EFD70F906" box="[567,697,1678,1701]" italics="true" pageId="153" pageNumber="154">Harpactrox</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFD2B648DFCE7F905" box="[712,814,1677,1702]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="153" pageNumber="154">Figs 252</figureCitation>
, 259, 262); antennae are sexually dimorphic and dorsal preantennal suture is absent in 
<taxonomicName id="CC884D7CFF13FF5AFE0C64B1FD55F969" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[495,668,1713,1738]" class="Insecta" family="Philopteridae" genus="Anarchonirmus" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFE0C64B1FD55F969" box="[495,668,1713,1738]" italics="true" pageId="153" pageNumber="154">Anarchonirmus</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFD4864B1FC8AF969" box="[683,835,1713,1738]" captionStart-0="FIGURES 114 – 115" captionStart-1="FIGURES 116 – 121" captionStartId-0="79.[151,264,1928,1950]" captionStartId-1="80.[151,264,1782,1804]" captionTargetBox-0="[171,1378,205,1884]" captionTargetBox-1="[162,1428,197,1724]" captionTargetId-0="figure@79.[158,1402,193,1884]" captionTargetId-1="figure@80.[151,1436,193,1760]" captionTargetPageId-0="79" captionTargetPageId-1="80" captionText-0="FIGURES 114 – 115. Anarchonirmus albovittatus n. gen. &amp; n. sp. ex Pomatostomus temporalis strepitans: 114, male habitus, dorsal and ventral views. 115, female habitus, dorsal and ventral views. (dotted lines on sternites outline densely pigmented areas)." captionText-1="FIGURES 116 – 121. Anarchonirmus albovittatus n. gen. &amp; n. sp. ex Pomatostomus temporalis strepitans: 116, male head, dorsal and ventral view; dorsal sensilla and pts not visible in the material examined. 117, female antenna, ventral view. 118, male genitalia, dorsal view. 119, male mesosome, ventral view. 120, male paramere, dorsal view. 121, female subgenital plate and vulval margin, ventral view (anterior dotted line outline pigmented part of subgenital plate, posterior dotted line extent of marginal thickening)." httpUri-0="https://zenodo.org/record/883222/files/figure.png" httpUri-1="https://zenodo.org/record/883224/files/figure.png" pageId="153" pageNumber="154">Figs 115-117</figureCitation>
), but antennae are monomorphic and suture is present in 
<taxonomicName id="CC884D7CFF13FF5AFF5064D6FEFCF94E" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[179,309,1750,1773]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="219" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFF5064D6FEFCF94E" box="[179,309,1750,1773]" italics="true" pageId="153" pageNumber="154">Harpactrox</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFEA064D5FE60F94D" box="[323,425,1749,1774]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="153" pageNumber="154">Figs 248</figureCitation>
, 255, 260); parameral heads are blunt and parameral blades are slender with a distinct heel distal to mesosome in 
<taxonomicName id="CC884D7CFF13FF5AFE7B64F9FD8CF8B1" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[408,581,1785,1810]" class="Insecta" family="Philopteridae" genus="Anarchonirmus" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFE7B64F9FD8CF8B1" box="[408,581,1785,1810]" italics="true" pageId="153" pageNumber="154">Anarchonirmus</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFDB564F9FD70F8B1" box="[598,697,1785,1810]" captionStart="FIGURES 116 – 121" captionStartId="80.[151,264,1782,1804]" captionTargetBox="[162,1428,197,1724]" captionTargetId="figure@80.[151,1436,193,1760]" captionTargetPageId="80" captionText="FIGURES 116 – 121. Anarchonirmus albovittatus n. gen. &amp; n. sp. ex Pomatostomus temporalis strepitans: 116, male head, dorsal and ventral view; dorsal sensilla and pts not visible in the material examined. 117, female antenna, ventral view. 118, male genitalia, dorsal view. 119, male mesosome, ventral view. 120, male paramere, dorsal view. 121, female subgenital plate and vulval margin, ventral view (anterior dotted line outline pigmented part of subgenital plate, posterior dotted line extent of marginal thickening)." httpUri="https://zenodo.org/record/883224/files/figure.png" pageId="153" pageNumber="154">Fig. 120</figureCitation>
), but parameral heads are folded medianly and parameral blades are broad with no such heel in 
<taxonomicName id="CC884D7CFF13FF5AFDF0651EFD5CF896" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[531,661,1822,1845]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="219" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFDF0651EFD5CF896" box="[531,661,1822,1845]" italics="true" pageId="153" pageNumber="154">Harpactrox</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFD4A651DFCDAF895" box="[681,787,1821,1846]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="153" pageNumber="154">Figs 251</figureCitation>
, 258). In both genera the gonopore is terminal, but in 
<taxonomicName id="CC884D7CFF13FF5AFF746541FE8DF8F9" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[151,324,1857,1882]" class="Insecta" family="Philopteridae" genus="Anarchonirmus" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFF746541FE8DF8F9" box="[151,324,1857,1882]" italics="true" pageId="153" pageNumber="154">Anarchonirmus</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFEB66541FE7CF8F9" box="[341,437,1857,1882]" captionStart="FIGURES 116 – 121" captionStartId="80.[151,264,1782,1804]" captionTargetBox="[162,1428,197,1724]" captionTargetId="figure@80.[151,1436,193,1760]" captionTargetPageId="80" captionText="FIGURES 116 – 121. Anarchonirmus albovittatus n. gen. &amp; n. sp. ex Pomatostomus temporalis strepitans: 116, male head, dorsal and ventral view; dorsal sensilla and pts not visible in the material examined. 117, female antenna, ventral view. 118, male genitalia, dorsal view. 119, male mesosome, ventral view. 120, male paramere, dorsal view. 121, female subgenital plate and vulval margin, ventral view (anterior dotted line outline pigmented part of subgenital plate, posterior dotted line extent of marginal thickening)." httpUri="https://zenodo.org/record/883224/files/figure.png" pageId="153" pageNumber="154">Fig. 119</figureCitation>
) the mesosomal lobes are large and serrated, and extend distal to gonopore, whereas in 
<taxonomicName id="CC884D7CFF13FF5AFF746566FED0F8DE" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[151,281,1894,1917]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="219" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFF746566FED0F8DE" box="[151,281,1894,1917]" italics="true" pageId="153" pageNumber="154">Harpactrox</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFEC96565FE5BF8DD" box="[298,402,1893,1918]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="153" pageNumber="154">Figs 250</figureCitation>
, 257) the lobes are small and smooth, and do not extend distal to gonopore. 
<collectingRegion id="C94CF81DFF13FF5AFAE86566FAE4F8DE" box="[1291,1325,1894,1917]" country="Maldives" name="Uthuru" pageId="153" pageNumber="154">No</collectingRegion>
<emphasis id="39FCEAEDFF13FF5AFAD46566FAB8F8DE" box="[1335,1393,1894,1917]" italics="true" pageId="153" pageNumber="154">ames</emphasis>
are visible in 
<taxonomicName id="CC884D7CFF13FF5AFEE66589FE7BF801" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[261,434,1929,1954]" class="Insecta" family="Philopteridae" genus="Anarchonirmus" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFEE66589FE7BF801" box="[261,434,1929,1954]" italics="true" pageId="153" pageNumber="154">Anarchonirmus</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFE216589FDEBF801" box="[450,546,1929,1954]" captionStart="FIGURES 116 – 121" captionStartId="80.[151,264,1782,1804]" captionTargetBox="[162,1428,197,1724]" captionTargetId="figure@80.[151,1436,193,1760]" captionTargetPageId="80" captionText="FIGURES 116 – 121. Anarchonirmus albovittatus n. gen. &amp; n. sp. ex Pomatostomus temporalis strepitans: 116, male head, dorsal and ventral view; dorsal sensilla and pts not visible in the material examined. 117, female antenna, ventral view. 118, male genitalia, dorsal view. 119, male mesosome, ventral view. 120, male paramere, dorsal view. 121, female subgenital plate and vulval margin, ventral view (anterior dotted line outline pigmented part of subgenital plate, posterior dotted line extent of marginal thickening)." httpUri="https://zenodo.org/record/883224/files/figure.png" pageId="153" pageNumber="154">Fig. 119</figureCitation>
), but these are visible in 
<taxonomicName id="CC884D7CFF13FF5AFCDA658AFC72F802" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[825,955,1930,1953]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="153" pageNumber="219" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF13FF5AFCDA658AFC72F802" box="[825,955,1930,1953]" italics="true" pageId="153" pageNumber="154">Harpactrox</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF13FF5AFC296589FBF9F801" box="[970,1072,1929,1954]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="153" pageNumber="154">Figs 250</figureCitation>
, 257).
</paragraph>
<paragraph id="0B3736FFFF13FF5AFF2465B1FA54F84D" blockId="153.[151,1437,525,2030]" pageId="153" pageNumber="154">
<emphasis id="39FCEAEDFF13FF5AFF2465B1FE98F869" bold="true" box="[199,337,1969,1994]" pageId="153" pageNumber="154">Description</emphasis>
. 
<emphasis id="39FCEAEDFF13FF5AFEBC65B1FE13F86B" bold="true" box="[351,474,1969,1994]" italics="true" pageId="153" pageNumber="154">Both sexes</emphasis>
. Head indented-dome shaped, often broader than long (
<figureCitation id="93B32A7AFF13FF5AFBA865B1FB78F86A" box="[1099,1201,1969,1994]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="153" pageNumber="154">Figs 248</figureCitation>
, 255, 260). Marginal carina not interrupted laterally or medianly, continuous with ventral anterior plate, not displaced at osculum.
</paragraph>
<paragraph id="0B3736FFFF10FF59FF746297FB7FFEE3" blockId="154.[151,1437,151,1616]" pageId="154" pageNumber="155">
Clypeo-labral suture does not reach anterior margin of head. Ventral carinae not clearly continuous with marginal carina in all species. Dorsal preantennal suture transversal, encompassing 
<emphasis id="39FCEAEDFF10FF59FC3A62BCFBC9FF76" box="[985,1024,188,213]" italics="true" pageId="154" pageNumber="155">ads</emphasis>
and, in some species, the 
<emphasis id="39FCEAEDFF10FF59FAC462BCFA96FF76" box="[1319,1375,188,213]" italics="true" pageId="154" pageNumber="155">dsms</emphasis>
(Fig. 255). Head setae as in 
<figureCitation id="93B32A7AFF10FF59FE7462DFFE35FF5B" box="[407,508,223,248]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="154" pageNumber="155">Figs 248</figureCitation>
, 255, 260; 
<emphasis id="39FCEAEDFF10FF59FD9A62E0FD69FF5B" box="[633,672,224,248]" italics="true" pageId="154" pageNumber="155">as3</emphasis>
and 
<emphasis id="39FCEAEDFF10FF59FD3462E1FD37FF5B" box="[727,766,225,248]" italics="true" pageId="154" pageNumber="155">pns</emphasis>
absent. Preantennal nodi very large; small accessory nodus median to the main nodus, only connected ventrally. Coni short. Antennae monomorphic. Eye protruding. Temporal carinae not visible. All 
<taxonomicName id="CC884D7CFF10FF59FDEC6329FDFFFEE3" box="[527,566,297,320]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="154" pageNumber="155" phylum="Arthropoda" rank="species" species="mts">
<emphasis id="39FCEAEDFF10FF59FDEC6329FDFFFEE3" box="[527,566,297,320]" italics="true" pageId="154" pageNumber="155">mts</emphasis>
</taxonomicName>
of roughly equal size, short. Gular plate large, indistinct.
</paragraph>
<paragraph id="0B3736FFFF10FF59FF24634CFF2FFE73" blockId="154.[151,1437,151,1616]" pageId="154" pageNumber="155">
Prothorax rectangular (
<figureCitation id="93B32A7AFF10FF59FE30634CFDBFFEC6" box="[467,630,332,357]" captionStart="FIGURES 246 – 247" captionStartId="155.[151,264,1922,1944]" captionTargetBox="[199,1341,238,1835]" captionTargetId="figure@155.[164,1388,195,1874]" captionTargetPageId="155" captionText="FIGURES 246 – 247. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 246, male habitus, dorsal and ventral views; only examined male with distorted abdomen, and shape of abdomen here reproduced as closely as possible; anterior sternites and some ventral setae missing in only available male specimen, and here included tentatively. 247, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883292/files/figure.png" pageId="154" pageNumber="155">Figs 246–247</figureCitation>
, 
<figureCitation id="93B32A7AFF10FF59FD65634CFD23FEC6" box="[646,746,332,357]" captionStart="FIGURES 253 – 254" captionStartId="158.[151,264,1955,1977]" captionTargetBox="[207,1386,225,1934]" captionTargetId="figure@158.[151,1436,193,1934]" captionTargetPageId="158" captionText="FIGURES 253 – 254. Harpactrox geminodus n. gen. &amp; n. sp. ex Harpactes erythrocephalus erythrocephalus: 253, male habitus, dorsal and ventral views. 254, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883296/files/figure.png" pageId="154" pageNumber="155">253–254</figureCitation>
, 
<figureCitation id="93B32A7AFF10FF59FD18634CFCE0FEC6" box="[763,809,332,357]" captionStart="FIGURES 260 – 262" captionStartId="161.[151,264,1970,1992]" captionTargetBox="[194,1356,212,1926]" captionTargetId="figure@161.[163,1404,193,1927]" captionTargetPageId="161" captionText="FIGURES 260 – 262. Harpactrox pontifrons n. gen. &amp; n. sp. ex Harpactes ardens ardens: 260, female head, dorsal and ventral views. 261, female habitus, dorsal and ventral views. 262, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883298/files/figure.png" pageId="154" pageNumber="155">261</figureCitation>
); 
<emphasis id="39FCEAEDFF10FF59FCA2634DFCBBFEC7" box="[833,882,333,356]" italics="true" pageId="154" pageNumber="155">ppss</emphasis>
on postero-lateral corner. Proepimera variable. Pterothorax pentagonal; lateral margins widely divergent; posterior margin convergent to rounded median point; 
<emphasis id="39FCEAEDFF10FF59FF746395FF02FE0F" box="[151,203,405,428]" italics="true" pageId="154" pageNumber="155">mms</emphasis>
widely separated medianly. Meso- and metasterna not fused, nude. Leg chaetotaxy as in 
<figureCitation id="93B32A7AFF10FF59FB556394FAC0FE0E" box="[1206,1289,404,429]" captionStart="FIGURES 25" captionStartId="14.[151,264,1012,1034]" captionTargetBox="[173,1428,205,974]" captionTargetId="figure@14.[151,1436,193,991]" captionTargetPageId="14" captionText="FIGURES 25. Dorso-ventral views of coxae I – III, trochanters I – III, and femora I – III, showing terminology of the leg setae used here. Other parts of the thorax removed for clarity. Setae situated marginally are illustrated and named on both dorsal (lefthand) and ventral (right-hand) sides. Terminology for individual seta as used in the text obtained by combining the leg segment (larger font) and the setal name (smaller font). Abbreviations: a 1 – 5, anterior setae 1 – 5; cI – III, coxae I – III; d 1 – 3, dorsal setae 1 – 3; dm 1 – 4, distal marginal setae 1 – 4; fI – III, femur I – III; p 1 – 3, posterior setae 1 – 3; s 1 – 6, sensilla 1 – 6; tI – III, trochanter I – III; v 1 – 4, ventral setae 1 – 4." httpUri="https://zenodo.org/record/883171/files/figure.png" pageId="154" pageNumber="155">Fig. 25</figureCitation>
, except 
<emphasis id="39FCEAEDFF10FF59FA866394FA55FE0F" box="[1381,1436,404,428]" italics="true" pageId="154" pageNumber="155">fI-p2</emphasis>
absent.
</paragraph>
<paragraph id="0B3736FFFF10FF59FF2463DCFDE1FD4C" blockId="154.[151,1437,151,1616]" pageId="154" pageNumber="155">
Abdomen slenderly oval (
<figureCitation id="93B32A7AFF10FF59FE0B63DCFD4FFE56" box="[488,646,476,501]" captionStart="FIGURES 246 – 247" captionStartId="155.[151,264,1922,1944]" captionTargetBox="[199,1341,238,1835]" captionTargetId="figure@155.[164,1388,195,1874]" captionTargetPageId="155" captionText="FIGURES 246 – 247. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 246, male habitus, dorsal and ventral views; only examined male with distorted abdomen, and shape of abdomen here reproduced as closely as possible; anterior sternites and some ventral setae missing in only available male specimen, and here included tentatively. 247, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883292/files/figure.png" pageId="154" pageNumber="155">Figs 246–247</figureCitation>
, 
<figureCitation id="93B32A7AFF10FF59FD7363DCFD3FFE56" box="[656,758,476,501]" captionStart="FIGURES 253 – 254" captionStartId="158.[151,264,1955,1977]" captionTargetBox="[207,1386,225,1934]" captionTargetId="figure@158.[151,1436,193,1934]" captionTargetPageId="158" captionText="FIGURES 253 – 254. Harpactrox geminodus n. gen. &amp; n. sp. ex Harpactes erythrocephalus erythrocephalus: 253, male habitus, dorsal and ventral views. 254, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883296/files/figure.png" pageId="154" pageNumber="155">253–254</figureCitation>
, 
<figureCitation id="93B32A7AFF10FF59FCE363DCFCE7FE56" box="[768,814,476,501]" captionStart="FIGURES 260 – 262" captionStartId="161.[151,264,1970,1992]" captionTargetBox="[194,1356,212,1926]" captionTargetId="figure@161.[163,1404,193,1927]" captionTargetPageId="161" captionText="FIGURES 260 – 262. Harpactrox pontifrons n. gen. &amp; n. sp. ex Harpactes ardens ardens: 260, female head, dorsal and ventral views. 261, female habitus, dorsal and ventral views. 262, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883298/files/figure.png" pageId="154" pageNumber="155">261</figureCitation>
). Abdominal chaetotaxy as in 
<tableCitation id="460A0344FF10FF59FB6263DCFB1EFE57" box="[1153,1239,476,501]" captionStart="TABLE 2" captionStartId="30.[151,239,796,819]" captionText="TABLE 2. Summary of abdominal chaetotaxy of segments II – VIII for the Brueelia - complex, including both genera and subgenera. In genera where chaetotaxy differs between species group (marked with *), only that of the type species is given here, and more detailed accounts are given separately under the respective genus entries and in Tables 3 – 12. Trichoid setae of segment VIII are present in all genera, and are not listed. Abbreviations: ps = paratergal seta; aps = accessory post-spiracular seta; psps = principal post-spiracular seta; tps = tergal posterior seta; ss = sutural seta; sts = sternal seta." pageId="154" pageNumber="155">Table 2</tableCitation>
. 
<taxonomicName id="CC884D7CFF10FF59FB0263DDFACEFE57" box="[1249,1287,477,500]" class="Insecta" family="Philopteridae" genus="Guimaraesiella" kingdom="Animalia" order="Phthiraptera" pageId="154" pageNumber="155" phylum="Arthropoda" rank="subSpecies" species="myiophoneae" subSpecies="aps">
<emphasis id="39FCEAEDFF10FF59FB0263DDFACEFE57" box="[1249,1287,477,500]" italics="true" pageId="154" pageNumber="155">aps</emphasis>
</taxonomicName>
absent on all segments. Tergopleurites rectangular; tergopleurites II–IX+X in male, and in segments II–VIII in female narrowly to moderately separated medianly. Sternal plates rectangular, not approaching pleurites. Pleural incrassations wide, with ornate anterior ends. Re-entrant heads variable. 
<collectingRegion id="C94CF81DFF10FF59FD056047FCE9FDC3" box="[742,800,583,608]" country="Maldives" name="Maale" pageId="154" pageNumber="155">Male</collectingRegion>
subgenital plate indistinct in 
<taxonomicName id="CC884D7CFF10FF59FB8A6049FAA3FDC3" authorityName="Bush" authorityYear="2017" box="[1129,1386,583,608]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="154" pageNumber="155" phylum="Arthropoda" rank="species" species="geminodus">
<emphasis id="39FCEAEDFF10FF59FB8A6049FAA3FDC3" box="[1129,1386,583,608]" italics="true" pageId="154" pageNumber="155">Harpactrox geminodus</emphasis>
</taxonomicName>
, but trapezoidal in 
<taxonomicName id="CC884D7CFF10FF59FED8606DFE06FD26" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[315,463,620,645]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="154" pageNumber="155" phylum="Arthropoda" rank="species" species="loeiensis" status="sp. nov.">
<emphasis id="39FCEAEDFF10FF59FED8606DFE94FD27" box="[315,349,621,644]" italics="true" pageId="154" pageNumber="155">Ha</emphasis>
. 
<emphasis id="39FCEAEDFF10FF59FE8E606CFE06FD26" box="[365,463,620,645]" italics="true" pageId="154" pageNumber="155">loeiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22CF5796FF10FF59FE3B606CFDD0FD26" box="[472,537,620,645]" pageId="154" pageNumber="155" rank="species">
<emphasis id="39FCEAEDFF10FF59FE3B606CFDD0FD26" bold="true" box="[472,537,620,645]" pageId="154" pageNumber="155">n. sp.</emphasis>
</taxonomicNameLabel>
, reaching distal end of abdomen. Female subgenital plate indistinct anteriorly, approaching vulval margin, with no cross-piece. Vulval margin (
<figureCitation id="93B32A7AFF10FF59FC8A608FFC06FD0B" box="[873,975,655,680]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="154" pageNumber="155">Figs 252</figureCitation>
, 259, 262) with short 
<emphasis id="39FCEAEDFF10FF59FB206091FB26FD0B" box="[1219,1263,657,680]" italics="true" pageId="154" pageNumber="155">vms</emphasis>
, thorn-like 
<emphasis id="39FCEAEDFF10FF59FA916091FA5DFD0B" box="[1394,1428,657,680]" italics="true" pageId="154" pageNumber="155">vss</emphasis>
; 
<emphasis id="39FCEAEDFF10FF59FF7460B5FF75FD6F" box="[151,188,693,716]" italics="true" pageId="154" pageNumber="155">vos</emphasis>
located on subgenital plate (in 
<taxonomicName id="CC884D7CFF10FF59FDFD60B5FD08FD6F" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[542,705,692,716]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="154" pageNumber="155" phylum="Arthropoda" rank="species" species="pontifrons" status="sp. nov.">
<emphasis id="39FCEAEDFF10FF59FDFD60B5FD89FD6F" box="[542,576,693,716]" italics="true" pageId="154" pageNumber="155">Ha</emphasis>
. 
<emphasis id="39FCEAEDFF10FF59FDAC60B4FD08FD6F" box="[591,705,692,716]" italics="true" pageId="154" pageNumber="155">pontifrons</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22CF5796FF10FF59FD2B60B4FCC1FD6E" box="[712,776,692,717]" pageId="154" pageNumber="155" rank="species">
<emphasis id="39FCEAEDFF10FF59FD2B60B4FCC1FD6E" bold="true" box="[712,776,692,717]" pageId="154" pageNumber="155">n. sp.</emphasis>
</taxonomicNameLabel>
both on and along lateral margin of plate, 
<figureCitation id="93B32A7AFF10FF59FB0A60B4FA82FD6E" box="[1257,1355,692,717]" captionStart="FIGURES 260 – 262" captionStartId="161.[151,264,1970,1992]" captionTargetBox="[194,1356,212,1926]" captionTargetId="figure@161.[163,1404,193,1927]" captionTargetPageId="161" captionText="FIGURES 260 – 262. Harpactrox pontifrons n. gen. &amp; n. sp. ex Harpactes ardens ardens: 260, female head, dorsal and ventral views. 261, female habitus, dorsal and ventral views. 262, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883298/files/figure.png" pageId="154" pageNumber="155">Fig. 262</figureCitation>
); distal setae longer, situated median to 
<emphasis id="39FCEAEDFF10FF59FE1D60D9FDE9FD53" box="[510,544,729,752]" italics="true" pageId="154" pageNumber="155">vss</emphasis>
.
</paragraph>
<paragraph id="0B3736FFFF10FF59FF2460FCFBF1FC6B" blockId="154.[151,1437,151,1616]" pageId="154" pageNumber="155">
Basal apodeme slightly convergent distally (
<figureCitation id="93B32A7AFF10FF59FD5E60FCFCEDFCB7" box="[701,804,764,789]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="154" pageNumber="155">Figs 249</figureCitation>
, 256). Proximal mesosome oval to rounded trapezoidal, overlapping basal apodeme. Gonopore (
<figureCitation id="93B32A7AFF10FF59FD96611FFD2AFC9B" box="[629,739,799,824]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="154" pageNumber="155">Figs 250</figureCitation>
, 257) distinct, different between species, but very broad, dominating the terminal end of the mesosome, closed distally and proximally. Ventral ridges vague. Mesosomal lobes small, triangular; 1–2 
<emphasis id="39FCEAEDFF10FF59FE376169FDC7FC23" box="[468,526,873,896]" italics="true" pageId="154" pageNumber="155">ames</emphasis>
sensilla submedianly anterior to gonopore; 1–2 
<emphasis id="39FCEAEDFF10FF59FBD36169FBA3FC23" box="[1072,1130,873,896]" italics="true" pageId="154" pageNumber="155">pmes</emphasis>
submedianly just lateral to gonopore; 
<emphasis id="39FCEAEDFF10FF59FEEC618DFE80FC07" box="[271,329,909,932]" italics="true" pageId="154" pageNumber="155">pmes</emphasis>
may be microsetae or sensilla. Parameral heads (
<figureCitation id="93B32A7AFF10FF59FC8C618CFC1DFC06" box="[879,980,908,933]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="154" pageNumber="155">Figs 251</figureCitation>
, 258) folded medianly. Parameral blades lobe-like; 
<emphasis id="39FCEAEDFF10FF59FEEA61B0FEF1FC6B" box="[265,312,944,968]" italics="true" pageId="154" pageNumber="155">pst1</emphasis>
sensillus, central; 
<emphasis id="39FCEAEDFF10FF59FDEB61B0FDFEFC6B" box="[520,567,944,968]" italics="true" pageId="154" pageNumber="155">pst2</emphasis>
microseta, lateral or sublateral near distal tip.
</paragraph>
</subSubSection>
<subSubSection id="43926574FF10FF59FF2461D4FEA6FAAE" pageId="154" pageNumber="155" type="biology_ecology">
<paragraph id="0B3736FFFF10FF59FF2461D4FF36FB03" blockId="154.[151,1437,151,1616]" pageId="154" pageNumber="155">
<emphasis id="39FCEAEDFF10FF59FF2461D4FE55FC4E" bold="true" box="[199,412,980,1005]" pageId="154" pageNumber="155">
<collectingRegion id="C94CF81DFF10FF59FF2461D4FF36FC4E" box="[199,255,980,1005]" country="Afghanistan" name="Khost" pageId="154" pageNumber="155">Host</collectingRegion>
distribution.
</emphasis>
Known only from two species of Asian trogons of the genus 
<taxonomicName id="CC884D7CFF10FF59FB8061D5FA51FC4F" authority="Swainson, 1833" authorityName="Swainson" authorityYear="1833" box="[1123,1432,980,1005]" class="Aves" family="Trogonidae" genus="Harpactes" kingdom="Animalia" order="Trogoniformes" pageId="154" pageNumber="155" phylum="Chordata" rank="genus">
<emphasis id="39FCEAEDFF10FF59FB8061D5FB11FC4F" box="[1123,1240,981,1004]" italics="true" pageId="154" pageNumber="155">Harpactes</emphasis>
Swainson, 1833
</taxonomicName>
. 
<bibRefCitation id="6F194B0EFF10FF59FF7461F8FE47FBB3" box="[151,398,1015,1040]" pageId="154" pageNumber="155" refString="Hosner, P. A., Sheldon, F. H., Lim, H. C. &amp; Moyle, R. G. (2010) Phylogeny and biogeography of the Asian trogons (Aves: Trogoniformes) inferred from nuclear and mitochondrial DNA sequences. Molecular Phylogenetics and Evolution, 57, 1219 - 1225." type="journal article">
Hosner 
<emphasis id="39FCEAEDFF10FF59FF1561F9FEE7FBB3" box="[246,302,1015,1040]" italics="true" pageId="154" pageNumber="155">et al</emphasis>
. (2010)
</bibRefCitation>
showed that 
<taxonomicName id="CC884D7CFF10FF59FDD461F7FC61FBB3" authority="Bonaparte, 1854" authorityName="Bonaparte" authorityYear="1854" box="[567,936,1015,1040]" class="Aves" family="Trogonidae" genus="Apalharpactes" higherTaxonomySource="CoL" kingdom="Animalia" order="Trogoniformes" pageId="154" pageNumber="155" phylum="Chordata" rank="genus">
<emphasis id="39FCEAEDFF10FF59FDD461F7FD13FBB3" box="[567,730,1015,1040]" italics="true" pageId="154" pageNumber="155">Apalharpactes</emphasis>
Bonaparte, 1854
</taxonomicName>
is best treated as a separate genus from 
<taxonomicName id="CC884D7CFF10FF59FF74661DFEC5FB97" authorityName="Swainson" authorityYear="1833" box="[151,268,1053,1076]" class="Aves" family="Trogonidae" genus="Harpactes" kingdom="Animalia" order="Trogoniformes" pageId="154" pageNumber="155" phylum="Chordata" rank="genus">
<emphasis id="39FCEAEDFF10FF59FF74661DFEC5FB97" box="[151,268,1053,1076]" italics="true" pageId="154" pageNumber="155">Harpactes</emphasis>
</taxonomicName>
. We have not seen any material from the two host species in 
<taxonomicName id="CC884D7CFF10FF59FC1D661CFB68FB96" authorityName="Bonaparte" authorityYear="1854" box="[1022,1185,1052,1077]" class="Aves" family="Trogonidae" genus="Apalharpactes" higherTaxonomySource="CoL" kingdom="Animalia" order="Trogoniformes" pageId="154" pageNumber="155" phylum="Chordata" rank="genus">
<emphasis id="39FCEAEDFF10FF59FC1D661CFB68FB96" box="[1022,1185,1052,1077]" italics="true" pageId="154" pageNumber="155">Apalharpactes</emphasis>
</taxonomicName>
, but note that in the phylogeny of 
<bibRefCitation id="6F194B0EFF10FF59FED46640FDEAFBFB" box="[311,547,1087,1112]" pageId="154" pageNumber="155" refString="Hosner, P. A., Sheldon, F. H., Lim, H. C. &amp; Moyle, R. G. (2010) Phylogeny and biogeography of the Asian trogons (Aves: Trogoniformes) inferred from nuclear and mitochondrial DNA sequences. Molecular Phylogenetics and Evolution, 57, 1219 - 1225." type="journal article">
Hosner 
<emphasis id="39FCEAEDFF10FF59FE726641FE0CFBFB" box="[401,453,1087,1112]" italics="true" pageId="154" pageNumber="155">et al</emphasis>
. (2010)
</bibRefCitation>
, 
<taxonomicName id="CC884D7CFF10FF59FDD3663FFD1AFBFB" authorityName="Bonaparte" authorityYear="1854" box="[560,723,1087,1112]" class="Aves" family="Trogonidae" genus="Apalharpactes" higherTaxonomySource="CoL" kingdom="Animalia" order="Trogoniformes" pageId="154" pageNumber="155" phylum="Chordata" rank="genus">
<emphasis id="39FCEAEDFF10FF59FDD3663FFD1AFBFB" box="[560,723,1087,1112]" italics="true" pageId="154" pageNumber="155">Apalharpactes</emphasis>
</taxonomicName>
is more closely related to the African 
<taxonomicName id="CC884D7CFF10FF59FB77663FFF1BFBDF" authority="Swainson, 1833" authorityName="Swainson" authorityYear="1833" class="Aves" family="Trogonidae" genus="Apaloderma" kingdom="Animalia" order="Trogoniformes" pageId="154" pageNumber="155" phylum="Chordata" rank="genus">
<emphasis id="39FCEAEDFF10FF59FB77663FFAD6FBFB" box="[1172,1311,1087,1112]" italics="true" pageId="154" pageNumber="155">Apaloderma</emphasis>
Swainson, 1833
</taxonomicName>
, which is parasitised by the louse genus 
<taxonomicName id="CC884D7CFF10FF59FD7C6664FC1FFBDE" authority="Tendeiro, 1960" authorityName="Tendeiro" authorityYear="1960" box="[671,982,1124,1149]" class="Insecta" family="Philopteridae" genus="Trogoniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="154" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFF10FF59FD7C6664FCE8FBDE" box="[671,801,1124,1149]" italics="true" pageId="154" pageNumber="155">Trogoniella</emphasis>
<bibRefCitation id="6F194B0EFF10FF59FCCA6664FC1FFBDE" author="Tendeiro" box="[809,982,1124,1149]" pageId="154" pageNumber="155" refString="Tendeiro, J. (1960) Etudes sur les Mallophages africains. Junta de Investigacoes Cientificas do Ultramar. Estudios, Ensaios e Documentos (Portugal), 65, 1 - 234." type="journal article" year="1960">Tendeiro, 1960</bibRefCitation>
</taxonomicName>
, which does not belong to the 
<taxonomicName id="CC884D7CFF10FF59FAD16664FA5BFBDE" authorityName="Bush" authorityYear="2017" box="[1330,1426,1124,1149]" class="Insecta" family="Philopteridae" genus="Brueelia" kingdom="Animalia" order="Phthiraptera" pageId="154" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFF10FF59FAD16664FA5BFBDE" box="[1330,1426,1124,1149]" italics="true" pageId="154" pageNumber="155">Brueelia</emphasis>
</taxonomicName>
- complex.
</paragraph>
<paragraph id="0B3736FFFF10FF59FF2466ACFEA6FAAE" blockId="154.[151,1437,151,1616]" pageId="154" pageNumber="155">
<materialsCitation id="BBE03CA2FF10FF59FF2466ACFEA6FAAE" ID-GBIF-Occurrence="1587279384" country="Thailand" location="southern India." pageId="154" pageNumber="155" specimenCount="1" stateProvince="South-East">
<emphasis id="39FCEAEDFF10FF59FF2466ACFE72FB66" bold="true" box="[199,443,1196,1221]" pageId="154" pageNumber="155">Geographical range.</emphasis>
The three species described here occur in 
<collectingCountry id="739F766FFF10FF59FC7566ACFC33FB66" box="[918,1018,1196,1221]" name="Thailand" pageId="154" pageNumber="155">Thailand</collectingCountry>
and on the 
<collectingCountry id="739F766FFF10FF59FB9866ACFB35FB66" box="[1147,1276,1196,1221]" name="Philippines" pageId="154" pageNumber="155">Philippines</collectingCountry>
, but the genus may range throughout the distribution of 
<taxonomicName id="CC884D7CFF10FF59FD8D66D1FD2AFB4B" authorityName="Swainson" authorityYear="1833" box="[622,739,1233,1256]" class="Aves" family="Trogonidae" genus="Harpactes" kingdom="Animalia" order="Trogoniformes" pageId="154" pageNumber="155" phylum="Chordata" rank="genus">
<emphasis id="39FCEAEDFF10FF59FD8D66D1FD2AFB4B" box="[622,739,1233,1256]" italics="true" pageId="154" pageNumber="155">Harpactes</emphasis>
</taxonomicName>
trogons, which includes most of 
<collectingRegion id="C94CF81DFF10FF59FB8166CFFB14FB4B" box="[1122,1245,1231,1256]" country="Botswana" name="South East" pageId="154" pageNumber="155">South-East</collectingRegion>
Asia, 
<collectingCountry id="739F766FFF10FF59FACB66CFFA51FB4B" box="[1320,1432,1231,1256]" name="Indonesia" pageId="154" pageNumber="155">Indonesia</collectingCountry>
, and 
<location id="0E576024FF10FF59FF2566F4FEA6FAAE" LSID="urn:lsid:plazi:treatment:832187E9FF13FF59FF746389FDF9F95C:0E576024FF10FF59FF2566F4FEA6FAAE" box="[198,367,1268,1293]" country="Thailand" name="southern India." pageId="154" pageNumber="155" stateProvince="South-East">southern India.</location>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="43926574FF10FF59FF246717FD5EFADB" pageId="154" pageNumber="155" type="etymology">
<paragraph id="0B3736FFFF10FF59FF246717FD5EFADB" blockId="154.[151,1437,151,1616]" pageId="154" pageNumber="155">
<emphasis id="39FCEAEDFF10FF59FF246717FE87FA93" bold="true" box="[199,334,1303,1328]" pageId="154" pageNumber="155">Etymology.</emphasis>
<taxonomicName id="CC884D7CFF10FF59FEB56719FE11FA93" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[342,472,1305,1328]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="154" pageNumber="219" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF10FF59FEB56719FE11FA93" box="[342,472,1305,1328]" italics="true" pageId="154" pageNumber="155">Harpactrox</emphasis>
</taxonomicName>
is formed by the host genus of the 
<typeStatus id="D433885DFF10FF59FC8C6718FC56FA93" box="[879,927,1304,1328]" pageId="154" pageNumber="155">type</typeStatus>
species, 
<taxonomicName id="CC884D7CFF10FF59FBEB6719FAF5FA93" authority="Swainson, 1833" authorityName="Swainson" authorityYear="1833" box="[1032,1340,1303,1328]" class="Aves" family="Trogonidae" genus="Harpactes" kingdom="Animalia" order="Trogoniformes" pageId="154" pageNumber="155" phylum="Chordata" rank="genus">
<emphasis id="39FCEAEDFF10FF59FBEB6719FBB4FA93" box="[1032,1149,1305,1328]" italics="true" pageId="154" pageNumber="155">Harpactes</emphasis>
Swainson, 1833
</taxonomicName>
, and the Greek “- 
<emphasis id="39FCEAEDFF10FF59FF1C673DFEE2FAF7" box="[255,299,1341,1364]" italics="true" pageId="154" pageNumber="155">trox</emphasis>
” for “gnawer” or “nibbler”. The generic name of the hosts, 
<taxonomicName id="CC884D7CFF10FF59FBF5673DFB42FAF7" authorityName="Swainson" authorityYear="1833" box="[1046,1163,1341,1364]" class="Aves" family="Trogonidae" genus="Harpactes" kingdom="Animalia" order="Trogoniformes" pageId="154" pageNumber="155" phylum="Chordata" rank="genus">
<emphasis id="39FCEAEDFF10FF59FBF5673DFB42FAF7" box="[1046,1163,1341,1364]" italics="true" pageId="154" pageNumber="155">Harpactes</emphasis>
</taxonomicName>
is derved from Greek “ 
<emphasis id="39FCEAEDFF10FF59FF40675FFED8FADB" box="[163,273,1375,1400]" italics="true" pageId="154" pageNumber="155">harpaktes</emphasis>
” for “robber”. Gender: masculine.
</paragraph>
</subSubSection>
<subSubSection id="43926574FF10FF59FF246784FDF9F95C" pageId="154" pageNumber="155" type="discussion">
<paragraph id="0B3736FFFF10FF59FF246784FB89F9F3" blockId="154.[151,1437,151,1616]" pageId="154" pageNumber="155">
<emphasis id="39FCEAEDFF10FF59FF246784FEF2FA3E" bold="true" box="[199,315,1412,1437]" pageId="154" pageNumber="155">Remarks.</emphasis>
With the description of 
<taxonomicName id="CC884D7CFF10FF59FDB16785FD1DFA3F" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[594,724,1413,1436]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="154" pageNumber="219" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF10FF59FDB16785FD1DFA3F" box="[594,724,1413,1436]" italics="true" pageId="154" pageNumber="155">Harpactrox</emphasis>
</taxonomicName>
, four different genera of ischnoceran lice have been described from members of the 
<taxonomicName id="CC884D7CFF10FF59FE6A67A7FCCAFA63" authority="(Price et al. 2003)" authorityName="Price et al." authorityYear="2003" box="[393,771,1447,1472]" class="Aves" kingdom="Animalia" order="Trogoniformes" pageId="154" pageNumber="155" phylum="Chordata" rank="order">
Trogoniformes (
<bibRefCitation id="6F194B0EFF10FF59FDDD67A7FD35FA63" author="Price" box="[574,764,1447,1472]" pageId="154" pageNumber="155" refString="Price, R. D., Hellenthal, R. A. &amp; Palma, R. L. (2003) World checklist of chewing lice with host associations and keys to families and genera. In: Price, R. D., Hellenthal, R. A., Palma, R. L., Johnson, K. P. &amp; Clayton, D. H. The Chewing lice: world checklist and biological overview. Illinois Natural History Survey Special Publication 24. x + 501 pp." type="book" year="2003">
Price 
<emphasis id="39FCEAEDFF10FF59FD6367A9FD71FA63" box="[640,696,1447,1472]" italics="true" pageId="154" pageNumber="155">et al.</emphasis>
2003
</bibRefCitation>
)
</taxonomicName>
. Based on their phylogenetic position in Bush 
<emphasis id="39FCEAEDFF10FF59FAEF67A9FAF5FA63" box="[1292,1340,1447,1472]" italics="true" pageId="154" pageNumber="155">et al</emphasis>
. (2016), and some aspects of morphology, we here place 
<taxonomicName id="CC884D7CFF10FF59FCFA67CCFCB0FA46" authorityName="Bush" authorityYear="2017" box="[793,889,1484,1509]" class="Insecta" family="Philopteridae" genus="Brueelia" kingdom="Animalia" order="Phthiraptera" pageId="154" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFF10FF59FCFA67CCFCB0FA46" box="[793,889,1484,1509]" italics="true" pageId="154" pageNumber="155">Brueelia</emphasis>
</taxonomicName>
-complex lice from New World trogons in 
<emphasis id="39FCEAEDFF10FF59FF7467EFFE8CF9AB" box="[151,325,1519,1544]" italics="true" pageId="154" pageNumber="155">Guimaraesiella</emphasis>
. This placement, however, is tenuous, as there are also morphological similarities between 
<emphasis id="39FCEAEDFF10FF59FF746414FE8CF98E" box="[151,325,1556,1581]" italics="true" pageId="154" pageNumber="155">Guimaraesiella</emphasis>
from trogons and 
<taxonomicName id="CC884D7CFF10FF59FDC16415FD6DF98F" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[546,676,1557,1580]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="154" pageNumber="219" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFF10FF59FDC16415FD6DF98F" box="[546,676,1557,1580]" italics="true" pageId="154" pageNumber="155">Harpactrox</emphasis>
</taxonomicName>
, especially in the preantennal area; future studies with additional material may find that these lice are more closely related than is presently assumed.
</paragraph>
<paragraph id="0B3736FFFF10FF59FF74647AFE98F930" blockId="154.[151,568,1658,1791]" box="[151,337,1658,1683]" pageId="154" pageNumber="155">Included species</paragraph>
<paragraph id="0B3736FFFF10FF59FF74649FFDF9F95C" blockId="154.[151,568,1658,1791]" pageId="154" pageNumber="155">
* 
<taxonomicName id="CC884D7CFF10FF59FF46649FFE6FF914" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[165,422,1694,1719]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="154" pageNumber="155" phylum="Arthropoda" rank="species" species="geminodus" status="sp. nov.">
<emphasis id="39FCEAEDFF10FF59FF46649FFE6FF914" box="[165,422,1694,1719]" italics="true" pageId="154" pageNumber="155">Harpactrox geminodus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22CF5796FF10FF59FE4D649EFDF1F914" box="[430,568,1694,1719]" pageId="154" pageNumber="155" rank="species">
<emphasis id="39FCEAEDFF10FF59FE4D649EFDF1F914" bold="true" box="[430,568,1694,1719]" pageId="154" pageNumber="155">new species</emphasis>
</taxonomicNameLabel>
* 
<taxonomicName id="CC884D7CFF10FF59FF4664C4FE46F978" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[165,399,1730,1755]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="154" pageNumber="155" phylum="Arthropoda" rank="species" species="loeiensis" status="sp. nov.">
<emphasis id="39FCEAEDFF10FF59FF4664C4FE46F978" box="[165,399,1730,1755]" italics="true" pageId="154" pageNumber="155">Harpactrox loeiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22CF5796FF10FF59FE7564C3FDD4F978" box="[406,541,1730,1755]" pageId="154" pageNumber="155" rank="species">new species</taxonomicNameLabel>
* 
<taxonomicName id="CC884D7CFF10FF59FF4664E7FE56F95C" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[165,415,1767,1791]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="154" pageNumber="155" phylum="Arthropoda" rank="species" species="pontifrons" status="sp. nov.">
<emphasis id="39FCEAEDFF10FF59FF4664E7FE56F95C" box="[165,415,1767,1791]" italics="true" pageId="154" pageNumber="155">Harpactrox pontifrons</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22CF5796FF10FF59FE4564E6FDF9F95C" box="[422,560,1766,1791]" pageId="154" pageNumber="155" rank="species">
<emphasis id="39FCEAEDFF10FF59FE4564E6FDF9F95C" bold="true" box="[422,560,1766,1791]" pageId="154" pageNumber="155">new species</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>