456 lines
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456 lines
62 KiB
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<document id="1FABA8EDAC61472E615FB982DCC2D287" ID-CLB-Dataset="7485" ID-DOI="10.11646/zootaxa.4949.2.3" ID-GBIF-Dataset="402f8d2b-37a5-4903-81cc-2c8c9de96dfc" ID-ISSN="1175-5326" ID-Zenodo-Dep="4636371" ID-ZooBank="CDB4068A-320B-4290-A0A8-17FEC1E2F2C1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616667413903" checkinUser="plazi" docAuthor="Grismer, L. Lee, Geissler, Peter, Neang, Thy, Hartmann, Timo, Wagner, Philipp & Poyarkov, Nikolay A." docDate="2021" docId="03E487BCFFA2FF84FF744D64FD1EFCBA" docLanguage="en" docName="zootaxa.4949.2.3.pdf" docOrigin="Zootaxa 4949 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyrtodactylus kulenensis Grismer & Geissler & Neang & Hartmann & Wagner & Poyarkov 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="281" masterDocId="FFDDFFC4FFB3FF90FFE34B42FFFBFFAF" masterDocTitle="Molecular phylogenetics, PCA, and MFA recover a new species of Cyrtodactylus (Squamata: Gekkonidae) from an isolated sandstone massif in northwestern Cambodia" masterLastPageNumber="288" masterPageNumber="261" pageNumber="278" updateTime="1698934985181" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="E498E34F56CCAA747ECABF0355EF76E4">
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<mods:title id="6C10A7CBA0F34C40B41597610BB92282">Molecular phylogenetics, PCA, and MFA recover a new species of Cyrtodactylus (Squamata: Gekkonidae) from an isolated sandstone massif in northwestern Cambodia</mods:title>
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</mods:role>
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<mods:namePart id="979692333763BA14004A28CD0A052443">Grismer, L. Lee</mods:namePart>
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<mods:nameIdentifier id="DB803F91189F344212051367A8A59D38" type="ORCID">0000-0002-2369-6405</mods:nameIdentifier>
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<mods:affiliation id="697A376693CCCAE4F5B96DC537439FB7">Herpetology Laboratory, Department of Biology, La Sierra University, 4500 Riverwalk Parkway, Riverside, California 92515, USA. lgrismer @ lasierra. edu; https: // orcid. org / 0000 - 0002 - 2369 - 6405</mods:affiliation>
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<mods:nameIdentifier id="1CAF59FA236A86FFD3B7C5B6590D469C" type="email">lgrismer@lasierra.edu</mods:nameIdentifier>
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<mods:namePart id="7D23106E0200E3972506AC292BD10F4E">Geissler, Peter</mods:namePart>
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<mods:affiliation id="E8B796276005F62149B2F3AC961EA679">Museum Natur und Mensch, Gerberau 32, 79098 Freiburg, GERMANY.</mods:affiliation>
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</mods:name>
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</mods:role>
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<mods:namePart id="26756EBEF83472DA89240F57555F68D3">Neang, Thy</mods:namePart>
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<mods:nameIdentifier id="5270B7C6F8B762333DDFCB8562BA2152" type="ORCID">0000-0002-2713-1639</mods:nameIdentifier>
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<mods:affiliation id="62E4FFEFBB4654C8CC6AB75A730EC555">Wild Earth Allies, 77 a, Street Beton, Bayap Village, Sangkat Phnom Penh Thmei, Khan Sen Sok, Phnom Penh, CAMBODIA thyneang 9 @ gmail. com; https: // orcid. org / 0000 - 0002 - 2713 - 1639</mods:affiliation>
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<mods:nameIdentifier id="A73DE89E1AB38DBB4A4AEBD823EC8A97" type="email">thyneang9@gmail.com</mods:nameIdentifier>
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</mods:role>
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<mods:namePart id="CE63D899B2240E06DAB61D02269BA157">Hartmann, Timo</mods:namePart>
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<mods:affiliation id="BC36FC967320EFF269D3358F51D2762D">Zoologisches Forschungsmusem Alexander Koenig, Adenauerallee 160, D- 5311 Bonn, GERMANY</mods:affiliation>
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</mods:name>
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</mods:role>
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<mods:namePart id="3A3EDEDFD435AB923D81F2E0501302DF">Wagner, Philipp</mods:namePart>
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<mods:affiliation id="0B92385FD42CF77A7803C34F6D663561">Department of Research and Conservation, Allwetterzoo, Sentruper Str. 315, 48161 Münster, GERMANY & Angkor Centre for Conservation of Biodiversity (ACCB), Kbal Spean, Phnom Kulen National Park, Siem Reap, CAMBODIA</mods:affiliation>
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</mods:role>
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<mods:namePart id="8DFC28C5927F7EDF50C1313072CB2508">Poyarkov, Nikolay A.</mods:namePart>
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<mods:nameIdentifier id="D09F46A51760DAA264E334890AC6A9FA" type="ORCID">0000-0002-7576-2283</mods:nameIdentifier>
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<mods:affiliation id="D9CBB0E7B66BCDD954167AEAFFECE5C6">Department of Vertebrate Zoology, Biological Faculty, M. V. Lomonosov Moscow State University, Moscow 119234, RUSSIA. n. poyarkov @ gmail. com; https: // orcid. org / 0000 - 0002 - 7576 - 2283 & Joint Russian-Vietnamese Tropical Research and Technological Center, Nghia Do, Cau Giay, Hanoi, VIETNAM</mods:affiliation>
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<mods:nameIdentifier id="343C240664BB50DA646EBEEFC42881F6" type="email">n.poyarkov@gmail.com</mods:nameIdentifier>
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<mods:title id="924B060E0635FD48B7C7D360E73B18DC">Zootaxa</mods:title>
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<mods:part id="62C46DBDE283ECDFA700BC802E7F36F6">
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<mods:date id="3451A26C1F23A245CEF84A63632EC376">2021</mods:date>
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<mods:number id="1B7E22EE5B52D6A33F1BEEA7AF754165">2021-03-25</mods:number>
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<mods:number id="6D0C8EB09CFF822BD154369F49F4519D">4949</mods:number>
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<mods:start id="AB502DA28B7DAC38195FA952B999FCC7">261</mods:start>
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<mods:identifier id="24CAF8948F1A15988B3E7B9896D80F89" type="DOI">10.11646/zootaxa.4949.2.3</mods:identifier>
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<mods:identifier id="1A9C110DE2B3360E82C7EEEA740B0AFD" type="Zenodo-Dep">4636371</mods:identifier>
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<treatment id="03E487BCFFA2FF84FF744D64FD1EFCBA" ID-DOI="http://doi.org/10.5281/zenodo.4665033" ID-GBIF-Taxon="180876864" ID-Zenodo-Dep="4665033" LSID="urn:lsid:plazi:treatment:03E487BCFFA2FF84FF744D64FD1EFCBA" httpUri="http://treatment.plazi.org/id/03E487BCFFA2FF84FF744D64FD1EFCBA" lastPageId="20" lastPageNumber="281" pageId="17" pageNumber="278">
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<subSubSection id="C3576521FFA2FF81FF744D64FCC9F9CB" pageId="17" pageNumber="278" type="nomenclature">
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<paragraph id="8BF236AAFFA2FF81FF744D64FDD1F9EE" blockId="17.[151,818,1574,1673]" box="[151,554,1574,1601]" pageId="17" pageNumber="278">
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<heading id="D0BA81C6FFA2FF81FF744D64FDD1F9EE" bold="true" box="[151,554,1574,1601]" fontSize="11" level="1" pageId="17" pageNumber="278" reason="1">
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<emphasis id="B939EAB8FFA2FF81FF744D64FDD1F9EE" bold="true" box="[151,554,1574,1601]" pageId="17" pageNumber="278">
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<taxonomicName id="4C4D4D29FFA2FF81FF744D64FE3DF9EE" authority="Grismer & Geissler & Neang & Hartmann & Wagner & Poyarkov, 2021" authorityName="Grismer & Geissler & Neang & Hartmann & Wagner & Poyarkov" authorityYear="2021" box="[151,454,1574,1601]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="278" phylum="Chordata" rank="species" species="kulenensis" status="sp. nov.">
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<emphasis id="B939EAB8FFA2FF81FF744D64FE3DF9EE" bold="true" box="[151,454,1574,1601]" italics="true" pageId="17" pageNumber="278">Cyrtodactylus kulenensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20A57C3FFA2FF81FE2E4D65FDD1F9EE" box="[461,554,1575,1601]" pageId="17" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF236AAFFA2FF81FF744D08FCC9F9CB" blockId="17.[151,818,1574,1673]" box="[151,818,1610,1637]" pageId="17" pageNumber="278">Suggested Common Name: Phnom Kulen Bent-toed Gecko</paragraph>
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</subSubSection>
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<subSubSection id="C3576521FFA2FF81FF744D2CFD46F964" pageId="17" pageNumber="278" type="description">
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<paragraph id="8BF236AAFFA2FF81FF744D2CFF10F926" blockId="17.[151,818,1574,1673]" box="[151,235,1646,1673]" pageId="17" pageNumber="278">
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(
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<figureCitation id="13762A2FFFA2FF81FF434D2CFF18F926" box="[160,227,1646,1673]" captionStart="FIGURE 7" captionStartId="19.[151,250,1279,1303]" captionTargetBox="[173,1421,189,1250]" captionTargetId="figure-17@19.[166,1421,181,1256]" captionTargetPageId="19" captionText="FIGURE 7. Holotype of adult male Cyrtodactylus kulenensis sp. nov. (ZFMK 92573. A. Dorsal view. B. Ventral view of femoral and precloacal regions. C. Ventral view of left foot. D. Lateral view of head. E. Dorsal view of head. F. Ventral view of gular region." figureDoi="http://doi.org/10.5281/zenodo.4636385" httpUri="https://zenodo.org/record/4636385/files/figure.png" pageId="17" pageNumber="278">Fig. 7</figureCitation>
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)
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</paragraph>
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<paragraph id="8BF236AAFFA2FF81FF744DF1FD46F964" blockId="17.[151,701,1715,1739]" box="[151,701,1715,1739]" pageId="17" pageNumber="278">
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<taxonomicName id="4C4D4D29FFA2FF81FF744DF1FE3FF964" box="[151,452,1715,1739]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="278" phylum="Chordata" rank="species" species="intermedius">
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<emphasis id="B939EAB8FFA2FF81FF744DF1FEDEF964" box="[151,293,1715,1739]" italics="true" pageId="17" pageNumber="278">Cyrtodactylus</emphasis>
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cf.
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<emphasis id="B939EAB8FFA2FF81FEAF4DF1FE3FF964" box="[332,452,1715,1739]" italics="true" pageId="17" pageNumber="278">intermedius</emphasis>
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</taxonomicName>
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.
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<bibRefCitation id="EFDC4B5BFFA2FF81FE334DF1FD43F964" author="Geissler, P. & Hartmann, T. & Ihlow, F. & Neang, T. & Seng, R. & Wagner, P. & Bohme, W." box="[464,696,1715,1739]" pageId="17" pageNumber="278" pagination="40 - 63" refId="ref12170" refString="Geissler, P., Hartmann, T., Ihlow, F., Neang, T., Seng, R., Wagner, P. & Bohme, W. (2019) Herpetofauna of the Phnom Kulen National Park, northern Cambodia - an annotated checklist. Cambodian Journal of Natural History, 2019, 40 - 63. https: // doi. org / 10.7717 / peerj. 7841 / fig- 1" type="journal article" year="2019">
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Geissler
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<emphasis id="B939EAB8FFA2FF81FDC94DF6FDA6F964" box="[554,605,1715,1739]" italics="true" pageId="17" pageNumber="278">et al.</emphasis>
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2019:49
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</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3576521FFA2FF82FF744DBBFEB3FF79" lastPageId="18" lastPageNumber="279" pageId="17" pageNumber="278" type="materials_examined">
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<paragraph id="8BF236AAFFA2FF81FF744DBBFEDEF8F4" blockId="17.[151,1437,1785,2028]" pageId="17" pageNumber="278">
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<materialsCitation id="3B253CF7FFA2FF81FF744DBBFEDEF8F4" ID-GBIF-Occurrence="3068459301" collectingDate="2011-06-02" collectionCode="ZFMK" country="Cambodia" county="Banteay Srei District" elevation="490" latitude="13.689167" location="Adult" longLatPrecision="21" longitude="104.015274" municipality="Phnom Kbal" pageId="17" pageNumber="278" specimenCode="ZFMK 92573" specimenCount="1" stateProvince="Siem Reap" typeStatus="holotype">
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<emphasis id="B939EAB8FFA2FF81FF744DBBFEF0F8BC" bold="true" box="[151,267,1785,1811]" pageId="17" pageNumber="278">
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<typeStatus id="54F68808FFA2FF81FF744DBBFEFDF8BC" box="[151,262,1785,1811]" pageId="17" pageNumber="278" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<location id="8E926071FFA2FF81FEF14DBBFEA9F8BC" LSID="urn:lsid:plazi:treatment:03E487BCFFA2FF84FF744D64FD1EFCBA:8E926071FFA2FF81FEF14DBBFEA9F8BC" box="[274,338,1785,1811]" country="Cambodia" county="Banteay Srei District" latitude="13.689167" longLatPrecision="21" longitude="104.015274" municipality="Phnom Kbal" name="Adult" pageId="17" pageNumber="278" stateProvince="Siem Reap">Adult</location>
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male
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<specimenCode id="DBEB9ED1FFA2FF81FE7B4DB8FDCFF8BB" box="[408,564,1785,1812]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="17" pageNumber="278" type="Museum">ZFMK 92573</specimenCode>
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from
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<location id="8E926071FFA2FF81FD984DBBFC46F8BC" LSID="urn:lsid:plazi:treatment:03E487BCFFA2FF84FF744D64FD1EFCBA:8E926071FFA2FF81FD984DBBFC46F8BC" box="[635,957,1785,1811]" country="Cambodia" county="Banteay Srei District" latitude="13.689167" longLatPrecision="21" longitude="104.015274" municipality="Phnom Kbal" name="Phnom Kulen National Park" pageId="17" pageNumber="278" stateProvince="Siem Reap">Phnom Kulen National Park</location>
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,
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<collectingMunicipality id="6B96ACD0FFA2FF81FC2A4DBBFBA1F8BC" box="[969,1114,1785,1811]" pageId="17" pageNumber="278">Phnom Kbal</collectingMunicipality>
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,
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<collectingCounty id="62934E26FFA2FF81FB854DB8FAADF8BC" box="[1126,1366,1785,1812]" pageId="17" pageNumber="278">Banteay Srei District</collectingCounty>
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,
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<collectingRegion id="4989F848FFA2FF81FA804DBBFEC5F898" country="Cambodia" name="Siem Reap" pageId="17" pageNumber="278">Siem Reap Province</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="F35A763AFFA2FF81FEA94C5FFE46F898" box="[330,445,1821,1847]" name="Cambodia" pageId="17" pageNumber="278">Cambodia</collectingCountry>
|
|||
|
|
(
|
|||
|
|
<geoCoordinate id="EE79506DFFA2FF81FE2E4C5FFDA6F898" box="[461,605,1821,1847]" degrees="13" direction="north" minutes="41" orientation="latitude" pageId="17" pageNumber="278" precision="15" seconds="21" value="13.689167">13°41’21’’N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="EE79506DFFA2FF81FD8A4C5FFD04F897" box="[617,767,1821,1848]" degrees="104" direction="east" minutes="00" orientation="longitude" pageId="17" pageNumber="278" precision="15" seconds="55" value="104.015274">104°00’55’’E</geoCoordinate>
|
|||
|
|
°N at about
|
|||
|
|
<quantity id="4CB59B4FFFA2FF81FC6B4C5FFC34F897" box="[904,975,1821,1848]" metricMagnitude="2" metricUnit="m" metricValue="4.9" pageId="17" pageNumber="278" unit="m" value="490.0">
|
|||
|
|
<elevation id="0060D199FFA2FF81FC6B4C5FFC34F897" box="[904,975,1821,1848]" metricMagnitude="2" metricUnit="m" metricValue="4.9" pageId="17" pageNumber="278" unit="m" value="490.0">490 m</elevation>
|
|||
|
|
</quantity>
|
|||
|
|
in elevation) collected by TH and PG on
|
|||
|
|
<date id="FFF3106AFFA2FF81FF744C03FEDAF8F4" box="[151,289,1857,1884]" pageId="17" pageNumber="278" value="2011-06-02">
|
|||
|
|
<collectingDate id="EFB7E982FFA2FF81FF744C03FEDAF8F4" box="[151,289,1857,1884]" pageId="17" pageNumber="278" value="2011-06-02">2 June 2011</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
.
|
|||
|
|
</materialsCitation>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BF236AAFFA2FF82FF244C27FEB3FF79" blockId="17.[151,1437,1785,2028]" lastBlockId="18.[151,1437,150,2013]" lastPageId="18" lastPageNumber="279" pageId="17" pageNumber="278">
|
|||
|
|
<materialsCitation id="3B253CF7FFA2FF82FF244C27FEBEFF79" ID-GBIF-Occurrence="3068459303" collectingDate="2008-09" collectingDateMax="2011-06" collectingDateMin="2008-09" collectionCode="ZFMK" country="Cambodia" county="Banteay Srei District" elevation="496" lastPageId="18" lastPageNumber="279" latitude="13.689167" location="Adult" longLatPrecision="21" longitude="104.015274" municipality="Phnom Kbal" pageId="17" pageNumber="278" specimenCode="ZFMK 88356, ZFMK 88357, ZFMK 88359, ZFMK 88358, ZFMK 90311, ZFMK 92572, ZFMK 92574, ZFMK 92575" specimenCount="2" stateProvince="Siem Reap" typeStatus="paratype">
|
|||
|
|
<emphasis id="B939EAB8FFA2FF81FF244C27FEBDF8D0" bold="true" box="[199,326,1893,1919]" pageId="17" pageNumber="278">
|
|||
|
|
<typeStatus id="54F68808FFA2FF81FF244C27FEBAF8D0" box="[199,321,1893,1919]" pageId="17" pageNumber="278" type="paratype">Paratypes</typeStatus>
|
|||
|
|
.
|
|||
|
|
</emphasis>
|
|||
|
|
Adult females
|
|||
|
|
<specimenCode id="DBEB9ED1FFA2FF81FE164C24FD6EF82F" box="[501,661,1893,1920]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="17" pageNumber="278" type="Museum">ZFMK 88356</specimenCode>
|
|||
|
|
,
|
|||
|
|
<specimenCode id="DBEB9ED1FFA2FF81FD414C24FCBAF82F" box="[674,833,1893,1920]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="17" pageNumber="278" type="Museum">ZFMK 88357</specimenCode>
|
|||
|
|
,
|
|||
|
|
<specimenCode id="DBEB9ED1FFA2FF81FCAD4C24FC10F82F" box="[846,1003,1893,1920]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="17" pageNumber="278" type="Museum">ZFMK 88359</specimenCode>
|
|||
|
|
and subadult male
|
|||
|
|
<specimenCode id="DBEB9ED1FFA2FF81FB284C24FA93F82F" box="[1227,1384,1893,1920]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="17" pageNumber="278" type="Museum">ZFMK 88358</specimenCode>
|
|||
|
|
collected by TH in
|
|||
|
|
<date id="FFF3106AFFA2FF81FEBF4CCBFDE1F80C" box="[348,538,1929,1955]" pageId="17" pageNumber="278" value="2008-09">
|
|||
|
|
<collectingDate id="EFB7E982FFA2FF81FEBF4CCBFDE1F80C" box="[348,538,1929,1955]" pageId="17" pageNumber="278" value="2008-09">September 2008</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
.
|
|||
|
|
<location id="8E926071FFA2FF81FDC44CCBFD9CF80C" LSID="urn:lsid:plazi:treatment:03E487BCFFA2FF84FF744D64FD1EFCBA:8E926071FFA2FF81FDC44CCBFD9CF80C" box="[551,615,1929,1955]" country="Cambodia" county="Banteay Srei District" latitude="13.689167" longLatPrecision="21" longitude="104.015274" municipality="Phnom Kbal" name="Adult" pageId="17" pageNumber="278" stateProvince="Siem Reap">Adult</location>
|
|||
|
|
female CBC (Centre for Biological Conservation) 03030 and juvenile
|
|||
|
|
<specimenCode id="DBEB9ED1FFA2FF81FF744CECFECBF868" box="[151,304,1965,1992]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="17" pageNumber="278" type="Museum">ZFMK 90311</specimenCode>
|
|||
|
|
collected by TH in
|
|||
|
|
<date id="FFF3106AFFA2FF81FDEA4CECFD85F868" box="[521,638,1965,1992]" pageId="17" pageNumber="278" value="2009-06">
|
|||
|
|
<collectingDate id="EFB7E982FFA2FF81FDEA4CECFD85F868" box="[521,638,1965,1992]" pageId="17" pageNumber="278" value="2009-06">June 2009</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
.
|
|||
|
|
<location id="8E926071FFA2FF81FD654CEFFD3DF868" LSID="urn:lsid:plazi:treatment:03E487BCFFA2FF84FF744D64FD1EFCBA:8E926071FFA2FF81FD654CEFFD3DF868" box="[646,710,1965,1991]" country="Cambodia" county="Banteay Srei District" latitude="13.689167" longLatPrecision="21" longitude="104.015274" municipality="Phnom Kbal" name="Adult" pageId="17" pageNumber="278" stateProvince="Siem Reap">Adult</location>
|
|||
|
|
females
|
|||
|
|
<specimenCode id="DBEB9ED1FFA2FF81FCCB4CECFC3EF867" box="[808,965,1965,1992]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="17" pageNumber="278" type="Museum">ZFMK 92572</specimenCode>
|
|||
|
|
,
|
|||
|
|
<specimenCode id="DBEB9ED1FFA2FF81FC2C4CECFB97F867" box="[975,1132,1965,1992]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="17" pageNumber="278" type="Museum">ZFMK 92574</specimenCode>
|
|||
|
|
,
|
|||
|
|
<specimenCode id="DBEB9ED1FFA2FF81FB954CECFAF4F867" box="[1142,1295,1965,1992]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="17" pageNumber="278" type="Museum">ZFMK 92575</specimenCode>
|
|||
|
|
collected by TH and PG in
|
|||
|
|
<date id="FFF3106AFFA2FF81FEDA4C90FE56F844" box="[313,429,2001,2028]" pageId="17" pageNumber="278" value="2011-06">
|
|||
|
|
<collectingDate id="EFB7E982FFA2FF81FEDA4C90FE56F844" box="[313,429,2001,2028]" pageId="17" pageNumber="278" value="2011-06">June 2011</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
.
|
|||
|
|
<location id="8E926071FFA2FF81FE554C93FE21F844" LSID="urn:lsid:plazi:treatment:03E487BCFFA2FF84FF744D64FD1EFCBA:8E926071FFA2FF81FE554C93FE21F844" box="[438,474,2001,2027]" country="Cambodia" county="Banteay Srei District" latitude="13.689167" longLatPrecision="21" longitude="104.015274" municipality="Phnom Kbal" name="All" pageId="17" pageNumber="278" stateProvince="Siem Reap">All</location>
|
|||
|
|
<typeStatus id="54F68808FFA2FF81FE024C90FDB7F843" box="[481,588,2002,2028]" pageId="17" pageNumber="278" type="paratype">paratypes</typeStatus>
|
|||
|
|
were collected on
|
|||
|
|
<location id="8E926071FFA2FF81FCFD4C93FC02F844" LSID="urn:lsid:plazi:treatment:03E487BCFFA2FF84FF744D64FD1EFCBA:8E926071FFA2FF81FCFD4C93FC02F844" box="[798,1017,2001,2027]" country="Cambodia" county="Banteay Srei District" latitude="13.689167" longLatPrecision="21" longitude="104.015274" municipality="Phnom Kbal" name="Phnom Kbal Spean" pageId="17" pageNumber="278" stateProvince="Siem Reap">Phnom Kbal Spean</location>
|
|||
|
|
,
|
|||
|
|
<location id="8E926071FFA2FF81FBE74C93FAB9F844" LSID="urn:lsid:plazi:treatment:03E487BCFFA2FF84FF744D64FD1EFCBA:8E926071FFA2FF81FBE74C93FAB9F844" box="[1028,1346,2001,2027]" country="Cambodia" county="Banteay Srei District" latitude="13.689167" longLatPrecision="21" longitude="104.015274" municipality="Phnom Kbal" name="Phnom Kulen National Park" pageId="17" pageNumber="278" stateProvince="Siem Reap">Phnom Kulen National Park</location>
|
|||
|
|
,
|
|||
|
|
<collectingMunicipality id="6B96ACD0FFA2FF82FAAE4C93FF2BFF1E" lastPageId="18" lastPageNumber="279" pageId="17" pageNumber="278">Phnom Kbal</collectingMunicipality>
|
|||
|
|
,
|
|||
|
|
<collectingCounty id="62934E26FFA1FF82FF3F4BD5FE37FF1E" box="[220,460,151,177]" pageId="18" pageNumber="279">Banteay Srei District</collectingCounty>
|
|||
|
|
,
|
|||
|
|
<collectingRegion id="4989F848FFA1FF82FE344BD5FD3BFF1E" box="[471,704,151,177]" country="Cambodia" name="Siem Reap" pageId="18" pageNumber="279">Siem Reap Province</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="F35A763AFFA1FF82FD2F4BD5FCC4FF1E" box="[716,831,151,177]" name="Cambodia" pageId="18" pageNumber="279">Cambodia</collectingCountry>
|
|||
|
|
(
|
|||
|
|
<geoCoordinate id="EE79506DFFA1FF82FCB34BD5FC1BFF1E" box="[848,992,151,177]" degrees="13" direction="north" minutes="41" orientation="latitude" pageId="18" pageNumber="279" precision="15" seconds="21" value="13.689167">13°41’21’’N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="EE79506DFFA1FF82FC0F4BD4FB78FF1E" box="[1004,1155,150,177]" degrees="104" direction="east" minutes="00" orientation="longitude" pageId="18" pageNumber="279" precision="15" seconds="55" value="104.015274">104°00’55’’E</geoCoordinate>
|
|||
|
|
°N between 480 and
|
|||
|
|
<quantity id="4CB59B4FFFA1FF82FA914BD5FF56FF7A" metricMagnitude="2" metricUnit="m" metricValue="4.96" pageId="18" pageNumber="279" unit="m" value="496.0">
|
|||
|
|
<elevation id="0060D199FFA1FF82FA914BD5FF56FF7A" metricMagnitude="2" metricUnit="m" metricValue="4.96" pageId="18" pageNumber="279" unit="m" value="496.0">496 m</elevation>
|
|||
|
|
</quantity>
|
|||
|
|
in elevation)
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3576521FFA1FF82FF244B9CFD45FDB6" pageId="18" pageNumber="279" type="diagnosis">
|
|||
|
|
<paragraph id="8BF236AAFFA1FF82FF244B9CFD45FDB6" blockId="18.[151,1437,150,2013]" pageId="18" pageNumber="279">
|
|||
|
|
<emphasis id="B939EAB8FFA1FF82FF244B9CFEB9FF57" bold="true" box="[199,322,222,248]" pageId="18" pageNumber="279">Diagnosis.</emphasis>
|
|||
|
|
<taxonomicName id="4C4D4D29FFA1FF82FEA84B9DFD93FF56" authority="Grismer & Geissler & Neang & Hartmann & Wagner & Poyarkov, 2021" authorityName="Grismer & Geissler & Neang & Hartmann & Wagner & Poyarkov" authorityYear="2021" box="[331,616,223,249]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="279" phylum="Chordata" rank="species" species="kulenensis" status="sp. nov.">
|
|||
|
|
<emphasis id="B939EAB8FFA1FF82FEA84B9DFD93FF56" box="[331,616,223,249]" italics="true" pageId="18" pageNumber="279">Cyrtodactylus kulenensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B939EAB8FFA1FF82FD924B9DFD35FF56" bold="true" box="[625,718,223,249]" pageId="18" pageNumber="279">
|
|||
|
|
<taxonomicNameLabel id="A20A57C3FFA1FF82FD924B9DFD35FF56" box="[625,718,223,249]" pageId="18" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
can be separated from all other species of the
|
|||
|
|
<taxonomicName id="4C4D4D29FFA1FF82FB174B9DFA67FF56" box="[1268,1436,223,249]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="279" phylum="Chordata" rank="species" species="intermedius">
|
|||
|
|
<emphasis id="B939EAB8FFA1FF82FB174B9DFA67FF56" box="[1268,1436,223,249]" italics="true" pageId="18" pageNumber="279">C. intermedius</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
group by having 8–11 supralabials, 8–11 infralabials, 33–38 paravertebral tubercles, 17–19 rows of longitudinally arranged tubercles, 38–44 ventrals, seven or eight expanded subdigital lamellae on the fourth toe, 11–13 unexpanded subdigital lamellae on the fourth toe, 18–21 total subdigital lamellae on the fourth toe, 12–21 enlarged femorals, nine or 10 enlarged precloacals, two or three rows of enlarged post-precloacals, two or three postcloacal tubercles, nine or 10 precloacal pores in males (N=2), enlarged femorals and enlarged precloacals continuous, proximal femorals less than one-half the size of the distal femorals, no digital pockets, maximum SVL
|
|||
|
|
<quantity id="4CB59B4FFFA1FF82FB734AF5FB03FE7E" box="[1168,1272,439,465]" metricMagnitude="-2" metricUnit="m" metricValue="8.959999999999999" pageId="18" pageNumber="279" unit="mm" value="89.6">89.6 mm</quantity>
|
|||
|
|
, no large dark blotches on top of head, three or four well-defined brown body bands edged with white tubercles, and body bands narrower than the beige interspaces (
|
|||
|
|
<tableCitation id="C6CF0311FFA1FF82FDD14ABDFD69FDB6" box="[562,658,511,537]" captionStart="TABLE 1" captionStartId="5.[151,239,461,487]" captionText="TABLE 1. Summary statistics of diagnostic meristic characters and morphological and color pattern characters of the species of the Cyrtodactylus intermedius group. Highlighted character states are those that differ discretely or significantly (p <0.05) from C. kulenensis sp. nov. Data for C. intermedius come from Smith (1917) and Taylor (1963)." pageId="18" pageNumber="279">Tables 1</tableCitation>
|
|||
|
|
,
|
|||
|
|
<tableCitation id="C6CF0311FFA1FF82FD434ABDFD4AFDB6" box="[672,689,511,537]" captionStart="TABLE 4" captionStartId="22.[151,239,474,500]" captionText="TABLE 4. Meristic, mensural (mm), and color pattern data from the type series of Cyrtodactylus kulenensis sp. nov. Abbreviations are listed in the Materials and methods. R = regenerated. Paired values are listed right/left. / = data unobtainable." pageId="18" pageNumber="279">4</tableCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3576521FFA1FF83FF244960FBB9F948" lastPageId="19" lastPageNumber="280" pageId="18" pageNumber="279" type="description">
|
|||
|
|
<paragraph id="8BF236AAFFA1FF82FF244960FAEAFB9A" blockId="18.[151,1437,150,2013]" pageId="18" pageNumber="279">
|
|||
|
|
<emphasis id="B939EAB8FFA1FF82FF244960FDBCFD93" bold="true" box="[199,583,546,573]" pageId="18" pageNumber="279">
|
|||
|
|
Description of
|
|||
|
|
<typeStatus id="54F68808FFA1FF82FE994961FE1AFD92" box="[378,481,547,573]" pageId="18" pageNumber="279" type="holotype">holotype</typeStatus>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="13762A2FFFA1FF82FE124960FDC3FD93" box="[497,568,546,572]" captionStart="FIGURE 7" captionStartId="19.[151,250,1279,1303]" captionTargetBox="[173,1421,189,1250]" captionTargetId="figure-17@19.[166,1421,181,1256]" captionTargetPageId="19" captionText="FIGURE 7. Holotype of adult male Cyrtodactylus kulenensis sp. nov. (ZFMK 92573. A. Dorsal view. B. Ventral view of femoral and precloacal regions. C. Ventral view of left foot. D. Lateral view of head. E. Dorsal view of head. F. Ventral view of gular region." figureDoi="http://doi.org/10.5281/zenodo.4636385" httpUri="https://zenodo.org/record/4636385/files/figure.png" pageId="18" pageNumber="279">Fig. 7</figureCitation>
|
|||
|
|
).
|
|||
|
|
</emphasis>
|
|||
|
|
Adult male SVL
|
|||
|
|
<quantity id="4CB59B4FFFA1FF82FCF24961FC82FD92" box="[785,889,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="8.77" pageId="18" pageNumber="279" unit="mm" value="87.7">87.7 mm</quantity>
|
|||
|
|
; head moderate in length (HL/SVL 0.28), width (HW/HL 0.67), somewhat flattened (HD/HL 0.36), distinct from neck, triangular in dorsal profile; lores concave slightly anteriorly, weakly inflated posteriorly; prefrontal region concave; canthus rostralis rounded; snout elongate (ES/HL 0.37), rounded in dorsal profile; eye large (ED/HL 0.30); ear opening elliptical, obliquely oriented, moderate in size; eye to ear distance slightly greater than diameter of eye; rostral rectangular, partially divided dorsally by inverted Y-shaped furrow, bordered posteriorly by large left and right supranasals and one small azygous internasal, bordered laterally by first supralabials; external nares bordered anteriorly by rostral, dorsally by large supranasal, posteriorly by two smaller postnasals, bordered ventrally by first supralabial; 11,10(R,L) rectangular supralabials extending to below midpoint of eye, second supralabial slightly larger than first; 11,10(R,L) infralabials tapering smoothly to just below and slightly past posterior margin of eye; scales of rostrum and lores flat to domed, larger than granular scales on top of head and occiput; scales of occiput intermixed with distinct, small tubercles; superciliaries elongate, largest anteriorly; mental triangular, bordered laterally by first infralabials and posteriorly by large left and right trapezoidal postmentals contacting medially for 40% of their length posterior to mental; one row of slightly enlarged, elongate sublabials extending posteriorly to fourth infralabial; gular and throat scales small, granular, grading posteriorly into slightly larger, flatter, smooth, imbricate, pectoral and ventral scales.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BF236AAFFA1FF82FF244F7DFD16FA9E" blockId="18.[151,1437,150,2013]" pageId="18" pageNumber="279">Body relatively short (AG/SVL 0.44) with well-defined ventrolateral folds; dorsal scales small, granular interspersed with larger, conical, semi-regularly arranged, moderately keeled tubercles; tubercles extend from top of head onto base of tail forming transverse rows, terminating at regenerated portion of tail; smaller tubercles extend anteriorly onto nape and occiput, diminishing in size and distinction on top of head; approximately 19 longitudinal rows of tubercles at midbody; approximately 38 paravertebral tubercles; 39 flat, imbricate, ventral scales much larger than dorsal scales; 10 large, pore-bearing, precloacal scales; no deep precloacal groove or depression; and three rows of large post-precloacal scales on midline.</paragraph>
|
|||
|
|
<paragraph id="8BF236AAFFA1FF82FF244E79FCE0F886" blockId="18.[151,1437,150,2013]" pageId="18" pageNumber="279">Forelimbs moderate in stature, relatively short (FL/SVL 0.14); granular scales of forearm slightly larger than those on body, interspersed with small tubercles; palmar scales rounded, slightly raised; interdigital pocketing absent; digits well-developed, inflected at basal interphalangeal joints; digits narrower distal to inflections; subdigital lamellae wide, transversely expanded proximal to joint inflections, narrower transverse lamellae distal to joint inflections; claws well-developed, claw base sheathed by a dorsal and ventral scale; hind limbs more robust than forelimbs, moderate in length (TBL/SVL 0.18), covered dorsally by granular scales interspersed with large, conical tubercles and anteriorly by flat, slightly larger scales; ventral scales of thigh flat, imbricate, larger than dorsals; subtibial scales flat, imbricate; one row of 9,10(R,L) enlarged femoral scales not continuous with enlarged precloacal scales, terminating distally before knee; proximal femoral scales smaller than distal femorals, the latter forming an abrupt union with smaller, rounded, ventral scales of posteroventral margin of thigh; femoral pores absent; plantar scales flat; interdigital pocketing absent; digits relatively long, well-developed, inflected at basal interphalangeal joints; 8(R,L) wide, transversely expanded subdigital lamellae on fourth toe proximal to joint inflection that extend onto sole; 12,12(R,L) narrower, transverse lamellae distal to inflection; 20 total subdigital lamellae; and claws welldeveloped, sheathed by a dorsal and ventral scale at base.</paragraph>
|
|||
|
|
<paragraph id="8BF236AAFFA1FF82FF244C71FE7DF872" blockId="18.[151,1437,150,2013]" pageId="18" pageNumber="279">
|
|||
|
|
Tail
|
|||
|
|
<quantity id="4CB59B4FFFA1FF82FF184C71FE95F8E2" box="[251,366,1843,1869]" metricMagnitude="-1" metricUnit="m" metricValue="1.084" pageId="18" pageNumber="279" unit="mm" value="108.4">108.4 mm</quantity>
|
|||
|
|
in length, first
|
|||
|
|
<quantity id="4CB59B4FFFA1FF82FDC34C71FD83F8E2" box="[544,632,1843,1869]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="18" pageNumber="279" unit="mm" value="8.1">8.1 mm</quantity>
|
|||
|
|
original,
|
|||
|
|
<quantity id="4CB59B4FFFA1FF82FD054C71FCC5F8E2" box="[742,830,1843,1869]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="18" pageNumber="279" unit="mm" value="6.2">6.2 mm</quantity>
|
|||
|
|
in width at base, tapering to a point; dorsal scales of original portion, flat, square; regenerated portion covered with small, smooth subcircular scales; median row of transversely expanded subcaudal scales, significantly larger than dorsal caudal scales on original portion; base of tail bearing hemipenal swellings; three conical postcloacal tubercles at base of hemipenal swellings; and postcloacal scales flat, imbricate.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF326622FFA0FF83FF744FBDFF24FACF" ID-DOI="http://doi.org/10.5281/zenodo.4636385" ID-Zenodo-Dep="4636385" httpUri="https://zenodo.org/record/4636385/files/figure.png" pageId="19" pageNumber="280" startId="19.[151,250,1279,1303]" targetBox="[173,1421,189,1250]" targetPageId="19">
|
|||
|
|
<paragraph id="8BF236AAFFA0FF83FF744FBDFF24FACF" blockId="19.[151,1437,1279,1376]" pageId="19" pageNumber="280">
|
|||
|
|
<emphasis id="B939EAB8FFA0FF83FF744FBDFEE9FAB7" bold="true" box="[151,274,1279,1304]" pageId="19" pageNumber="280">FIGURE 7.</emphasis>
|
|||
|
|
Holotype of adult male
|
|||
|
|
<taxonomicName id="4C4D4D29FFA0FF83FDE44E42FCFEFAB7" authority="Grismer & Geissler & Neang & Hartmann & Wagner & Poyarkov, 2021" authorityName="Grismer & Geissler & Neang & Hartmann & Wagner & Poyarkov" authorityYear="2021" box="[519,773,1280,1304]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="280" phylum="Chordata" rank="species" species="kulenensis" status="sp. nov.">
|
|||
|
|
<emphasis id="B939EAB8FFA0FF83FDE44E42FCFEFAB7" box="[519,773,1280,1304]" italics="true" pageId="19" pageNumber="280">Cyrtodactylus kulenensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B939EAB8FFA0FF83FCE94E42FCA1FAB7" bold="true" box="[778,858,1280,1304]" pageId="19" pageNumber="280">
|
|||
|
|
<taxonomicNameLabel id="A20A57C3FFA0FF83FCE94E42FCA1FAB7" box="[778,858,1280,1304]" pageId="19" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
(ZFMK 92573. A. Dorsal view. B. Ventral view of femoral and precloacal regions. C. Ventral view of left foot. D. Lateral view of head. E. Dorsal view of head. F. Ventral view of gular region.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BF236AAFFA0FF83FF244ECAFAC3F99C" blockId="19.[151,1437,1416,2019]" pageId="19" pageNumber="280">
|
|||
|
|
<emphasis id="B939EAB8FFA0FF83FF244ECAFD9FFA0D" bold="true" box="[199,612,1416,1442]" pageId="19" pageNumber="280">
|
|||
|
|
Coloration in preservation (
|
|||
|
|
<figureCitation id="13762A2FFFA0FF83FDF34ECAFDADFA0D" box="[528,598,1416,1442]" captionStart="FIGURE 7" captionStartId="19.[151,250,1279,1303]" captionTargetBox="[173,1421,189,1250]" captionTargetId="figure-17@19.[166,1421,181,1256]" captionTargetPageId="19" captionText="FIGURE 7. Holotype of adult male Cyrtodactylus kulenensis sp. nov. (ZFMK 92573. A. Dorsal view. B. Ventral view of femoral and precloacal regions. C. Ventral view of left foot. D. Lateral view of head. E. Dorsal view of head. F. Ventral view of gular region." figureDoi="http://doi.org/10.5281/zenodo.4636385" httpUri="https://zenodo.org/record/4636385/files/figure.png" pageId="19" pageNumber="280">Fig. 7</figureCitation>
|
|||
|
|
).
|
|||
|
|
</emphasis>
|
|||
|
|
Ground color of top of head, limbs, and dorsum brown, immaculate; wide dark-brown nuchal loop edged with white tubercles extends from posterior margin of one orbit across nape to posterior margin of opposing orbit; four dark-brown, straight-edged, bands bordered with white tubercles and slightly wider beige interspaces extend from forelimb insertion to groin, no caudal bands on unicolor brown, regenerated tail; and ventral surfaces of gular region, limbs and body dull-white to beige, bearing fine, dark, stippling.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BF236AAFFA0FF83FF244D7EFBB9F948" blockId="19.[151,1437,1416,2019]" pageId="19" pageNumber="280">
|
|||
|
|
<emphasis id="B939EAB8FFA0FF83FF244D7EFEC4F9F9" bold="true" box="[199,319,1596,1622]" pageId="19" pageNumber="280">Variation.</emphasis>
|
|||
|
|
<materialsCitation id="3B253CF7FFA0FF83FEA54D7FFBC6F948" ID-GBIF-Occurrence="3068459302" collectionCode="ZMFK" pageId="19" pageNumber="280" specimenCode="ZMFK 88356, ZMFK 88356-57" specimenCount="2" typeStatus="paratype">
|
|||
|
|
The
|
|||
|
|
<typeStatus id="54F68808FFA0FF83FE994D7FFE1EF9F8" box="[378,485,1597,1623]" pageId="19" pageNumber="280" type="paratype">paratypes</typeStatus>
|
|||
|
|
generally resemble the coloration and pattern of the
|
|||
|
|
<typeStatus id="54F68808FFA0FF83FBA24D7FFB58F9F8" box="[1089,1187,1597,1623]" pageId="19" pageNumber="280" type="holotype">holotype</typeStatus>
|
|||
|
|
with the exception of
|
|||
|
|
<collectionCode id="ED5CAE6FFFA0FF83FF744D23FF1EF9D4" box="[151,229,1633,1659]" pageId="19" pageNumber="280">ZMFK</collectionCode>
|
|||
|
|
<quantity id="4CB59B4FFFA0FF83FF0F4D22FEB5F9D4" box="[236,334,1632,1659]" metricMagnitude="3" metricUnit="m" metricValue="2.2442932" pageId="19" pageNumber="280" unit="in" value="88358.0">88358 in</quantity>
|
|||
|
|
which the first band is divided into three, oval-shaped parts, and the second band is nearly divided on the midline of the body and in
|
|||
|
|
<specimenCode id="DBEB9ED1FFA0FF83FDFE4DC7FD47F930" box="[541,700,1668,1695]" collectionCode="ZMFK" pageId="19" pageNumber="280">ZMFK 88356</specimenCode>
|
|||
|
|
, where the first band is divided on the midline (
|
|||
|
|
<figureCitation id="13762A2FFFA0FF83FB3C4DC7FADEF930" box="[1247,1317,1669,1695]" captionStart="FIGURE 8" captionStartId="20.[151,250,1955,1979]" captionTargetBox="[151,1436,813,1932]" captionTargetId="figure-317@20.[151,1436,813,1932]" captionTargetPageId="20" captionText="FIGURE 8. In situ view of Cyrtodactlus kulenensis sp. nov. A. ZFMK 92575 adult female. B. ZFMK 92572 adult female. C. ZFMK 88357 adult female. D. ZFMK 88358 subadult male." figureDoi="http://doi.org/10.5281/zenodo.4636387" httpUri="https://zenodo.org/record/4636387/files/figure.png" pageId="19" pageNumber="280">Fig. 8</figureCitation>
|
|||
|
|
). With the exception of
|
|||
|
|
<specimenCode id="DBEB9ED1FFA0FF83FECE4DEBFE0DF96C" box="[301,502,1704,1731]" collectionCode="ZMFK" pageId="19" pageNumber="280">ZMFK 88356–57</specimenCode>
|
|||
|
|
, the dark caudal bands are wider than the light caudal bands and encircle the tail. Meristic variation is provided in
|
|||
|
|
<tableCitation id="C6CF0311FFA0FF83FDE54D8FFDA7F948" box="[518,604,1741,1767]" captionStart="TABLE 4" captionStartId="22.[151,239,474,500]" captionText="TABLE 4. Meristic, mensural (mm), and color pattern data from the type series of Cyrtodactylus kulenensis sp. nov. Abbreviations are listed in the Materials and methods. R = regenerated. Paired values are listed right/left. / = data unobtainable." pageId="19" pageNumber="280">Table 4</tableCitation>
|
|||
|
|
. Coloration in life is illustrated in
|
|||
|
|
<figureCitation id="13762A2FFFA0FF83FC3E4D8FFBC6F948" box="[989,1085,1741,1767]" captionStart="FIGURE 8" captionStartId="20.[151,250,1955,1979]" captionTargetBox="[151,1436,813,1932]" captionTargetId="figure-317@20.[151,1436,813,1932]" captionTargetPageId="20" captionText="FIGURE 8. In situ view of Cyrtodactlus kulenensis sp. nov. A. ZFMK 92575 adult female. B. ZFMK 92572 adult female. C. ZFMK 88357 adult female. D. ZFMK 88358 subadult male." figureDoi="http://doi.org/10.5281/zenodo.4636387" httpUri="https://zenodo.org/record/4636387/files/figure.png" pageId="19" pageNumber="280">Figure 8</figureCitation>
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3576521FFA0FF83FF244DB2FC1BF880" pageId="19" pageNumber="280" type="distribution">
|
|||
|
|
<paragraph id="8BF236AAFFA0FF83FF244DB2FC1BF880" blockId="19.[151,1437,1416,2019]" pageId="19" pageNumber="280">
|
|||
|
|
<emphasis id="B939EAB8FFA0FF83FF244DB2FE9AF8A5" bold="true" box="[199,353,1776,1802]" pageId="19" pageNumber="280">Distribution.</emphasis>
|
|||
|
|
<taxonomicName id="4C4D4D29FFA0FF83FE8B4DB3FD7AF8A4" authority="Grismer & Geissler & Neang & Hartmann & Wagner & Poyarkov, 2021" authorityName="Grismer & Geissler & Neang & Hartmann & Wagner & Poyarkov" authorityYear="2021" box="[360,641,1777,1803]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="280" phylum="Chordata" rank="species" species="kulenensis" status="sp. nov.">
|
|||
|
|
<emphasis id="B939EAB8FFA0FF83FE8B4DB3FD7AF8A4" box="[360,641,1777,1803]" italics="true" pageId="19" pageNumber="280">Cyrtodactylus kulenensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B939EAB8FFA0FF83FD6B4DB3FD19F8A4" bold="true" box="[648,738,1777,1803]" pageId="19" pageNumber="280">
|
|||
|
|
<taxonomicNameLabel id="A20A57C3FFA0FF83FD6B4DB3FD19F8A4" box="[648,738,1777,1803]" pageId="19" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
is known only from the
|
|||
|
|
<typeStatus id="54F68808FFA0FF83FC144DB3FBDCF8A4" box="[1015,1063,1777,1803]" pageId="19" pageNumber="280">type</typeStatus>
|
|||
|
|
locality in the Phnom Kulen National Park, Banteay Srei District,
|
|||
|
|
<collectingRegion id="4989F848FFA0FF83FDFB4C57FD04F880" box="[536,767,1813,1839]" country="Cambodia" name="Siem Reap" pageId="19" pageNumber="280">Siem Reap Province</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="F35A763AFFA0FF83FCE84C57FC85F880" box="[779,894,1813,1839]" name="Cambodia" pageId="19" pageNumber="280">Cambodia</collectingCountry>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="13762A2FFFA0FF83FC6E4C57FC34F880" box="[909,975,1813,1839]" captionStart="FIGURE 1" captionStartId="1.[151,250,1793,1817]" captionTargetBox="[151,1436,309,1770]" captionTargetId="figure-81@1.[151,1436,309,1770]" captionTargetPageId="1" captionText="FIGURE 1. Distribution of the species of the Cyrtodactylus intermedius group." figureDoi="http://doi.org/10.5281/zenodo.4636373" httpUri="https://zenodo.org/record/4636373/files/figure.png" pageId="19" pageNumber="280">Fig. 1</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3576521FFA0FF84FF244C7BFD1EFCBA" lastPageId="20" lastPageNumber="281" pageId="19" pageNumber="280" type="etymology">
|
|||
|
|
<paragraph id="8BF236AAFFA0FF83FF244C7BFDF8F834" blockId="19.[151,1437,1416,2019]" pageId="19" pageNumber="280">
|
|||
|
|
<emphasis id="B939EAB8FFA0FF83FF244C7BFEB5F8FC" bold="true" box="[199,334,1849,1875]" pageId="19" pageNumber="280">Etymology.</emphasis>
|
|||
|
|
The specific epithet
|
|||
|
|
<taxonomicName id="4C4D4D29FFA0FF83FDDE4C7BFD49F8FC" authorityName="Grismer & Geissler & Neang & Hartmann & Wagner & Poyarkov" authorityYear="2021" box="[573,690,1849,1875]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="280" phylum="Chordata" rank="species" species="kulenensis">
|
|||
|
|
<emphasis id="B939EAB8FFA0FF83FDDE4C7BFD49F8FC" box="[573,690,1849,1875]" italics="true" pageId="19" pageNumber="280">kulenensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is a Latinized toponymic adjective named after Phnom (=Mount) Kulen National Park, Banteay Srei District,
|
|||
|
|
<collectingRegion id="4989F848FFA0FF83FD624C1FFC9CF8D8" box="[641,871,1885,1911]" country="Cambodia" name="Siem Reap" pageId="19" pageNumber="280">Siem Reap Province</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="F35A763AFFA0FF83FC914C1FFC1EF8D8" box="[882,997,1885,1911]" name="Cambodia" pageId="19" pageNumber="280">Cambodia</collectingCountry>
|
|||
|
|
. “Phnom Kulen” means “the Mountain of Lychees” in Khmer language.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BF236AAFFA0FF84FF244CE6FE59FD92" blockId="19.[151,1437,1416,2019]" lastBlockId="20.[151,1437,151,789]" lastPageId="20" lastPageNumber="281" pageId="19" pageNumber="280">
|
|||
|
|
<emphasis id="B939EAB8FFA0FF83FF244CE6FE9DF811" bold="true" box="[199,358,1956,1982]" pageId="19" pageNumber="280">Comparisons</emphasis>
|
|||
|
|
.
|
|||
|
|
<taxonomicName id="4C4D4D29FFA0FF83FE954CE7FD6AF810" authority="Grismer & Geissler & Neang & Hartmann & Wagner & Poyarkov, 2021" authorityName="Grismer & Geissler & Neang & Hartmann & Wagner & Poyarkov" authorityYear="2021" box="[374,657,1957,1983]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="280" phylum="Chordata" rank="species" species="kulenensis" status="sp. nov.">
|
|||
|
|
<emphasis id="B939EAB8FFA0FF83FE954CE7FD6AF810" box="[374,657,1957,1983]" italics="true" pageId="19" pageNumber="280">Cyrtodactylus kulenensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B939EAB8FFA0FF83FD794CE7FD0DF810" bold="true" box="[666,758,1957,1983]" pageId="19" pageNumber="280">
|
|||
|
|
<taxonomicNameLabel id="A20A57C3FFA0FF83FD794CE7FD0DF810" box="[666,758,1957,1983]" pageId="19" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
is most closely related to
|
|||
|
|
<taxonomicName id="4C4D4D29FFA0FF83FBC44CE7FB3FF810" authorityName="Gray" authorityYear="1827" box="[1063,1220,1957,1983]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="280" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis id="B939EAB8FFA0FF83FBC44CE7FB3FF810" box="[1063,1220,1957,1983]" italics="true" pageId="19" pageNumber="280">Cyrtodactylus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
sp. 2 but is differentiated from it by having a significantly higher mean number of supralabial scales (SL), paravertebral tubercles (PVT), enlarged femoral scales (FS), and enlarged precloacal scales (PS). It is differentiated from
|
|||
|
|
<taxonomicName id="4C4D4D29FFA7FF84FB154BD5FA67FF1E" box="[1270,1436,151,177]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="281" phylum="Chordata" rank="species" sensu="stricto" species="intermedius">
|
|||
|
|
<emphasis id="B939EAB8FFA7FF84FB154BD5FA67FF1E" box="[1270,1436,151,177]" italics="true" pageId="20" pageNumber="281">C. intermedius</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A20A57C3FFA7FF84FF744BF9FF40FF7A" box="[151,187,187,213]" pageId="20" pageNumber="281" sensu="stricto">s.s.</taxonomicNameLabel>
|
|||
|
|
by having continuous as opposed to discontinuous enlarged femoral and precloacal scales (
|
|||
|
|
<bibRefCitation id="EFDC4B5BFFA7FF84FB224BF9FAB2FF7A" author="Smith, M. A." box="[1217,1353,187,213]" pageId="20" pageNumber="281" pagination="221 - 225" refId="ref14811" refString="Smith, M. A. (1917) Descriptions of new reptiles and a new batrachian from Siam. The Journal of the Natural History Society of Siam, 2, 221 - 225." type="journal article" year="1917">Smith 1917</bibRefCitation>
|
|||
|
|
). It can be further differentiated from all other members of the
|
|||
|
|
<taxonomicName id="4C4D4D29FFA7FF84FD1F4B9DFC5AFF56" box="[764,929,223,249]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="281" phylum="Chordata" rank="species" species="intermedius">
|
|||
|
|
<emphasis id="B939EAB8FFA7FF84FD1F4B9DFC5AFF56" box="[764,929,223,249]" italics="true" pageId="20" pageNumber="281">C. intermedius</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
group by having a significantly greater mean number of supralabials and paravertebral tubercles. It differs further from
|
|||
|
|
<emphasis id="B939EAB8FFA7FF84FC024A41FD5DFEEE" italics="true" pageId="20" pageNumber="281">
|
|||
|
|
<taxonomicName id="4C4D4D29FFA7FF84FC024A41FB81FEB2" box="[993,1146,259,285]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="281" phylum="Chordata" rank="species" species="auralensis">C. auralensis</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C4D4D29FFA7FF84FB644A41FADFFEB2" box="[1159,1316,259,285]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="281" phylum="Chordata" rank="species" species="bokorensis">C. bokorensis</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C4D4D29FFA7FF84FAD24A41FEF3FEEF" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="281" phylum="Chordata" rank="species" species="cardamomensis">C. cardamomensis</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C4D4D29FFA7FF84FEF74A65FE2BFEEE" box="[276,464,295,321]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="281" phylum="Chordata" rank="species" species="phuquocensis">C. phuquocensis</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C4D4D29FFA7FF84FE3F4A65FD59FEEE" box="[476,674,295,321]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="281" phylum="Chordata" rank="species" species="thylcodactylus">C. thylcodactylus</taxonomicName>
|
|||
|
|
,
|
|||
|
|
</emphasis>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C4D4D29FFA7FF84FD3E4A65FC6DFEEF" box="[733,918,295,321]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="281" phylum="Chordata" rank="species" species="septimontium">
|
|||
|
|
<emphasis id="B939EAB8FFA7FF84FD3E4A65FC6DFEEF" box="[733,918,295,321]" italics="true" pageId="20" pageNumber="281">C. septimontium</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by having a higher mean number of proximal expanded subdigital lamellae (TLE) and is further differentiated from
|
|||
|
|
<emphasis id="B939EAB8FFA7FF84FC474A09FEACFE26" italics="true" pageId="20" pageNumber="281">
|
|||
|
|
<taxonomicName id="4C4D4D29FFA7FF84FC474A09FBC4FECA" box="[932,1087,331,357]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="281" phylum="Chordata" rank="species" species="bokorensis">C. bokorensis</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C4D4D29FFA7FF84FBAA4A09FB2DFECA" box="[1097,1238,331,357]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="281" phylum="Chordata" rank="species" species="laagensis">C. laagensis</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C4D4D29FFA7FF84FB024A09FA63FECA" box="[1249,1432,331,357]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="281" phylum="Chordata" rank="species" species="kohrongensis">C. kohrongensis</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C4D4D29FFA7FF84FF744A2DFEA8FE26" box="[151,339,367,393]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="281" phylum="Chordata" rank="species" species="phuquocensis">C. phuquocensis</taxonomicName>
|
|||
|
|
,
|
|||
|
|
</emphasis>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C4D4D29FFA7FF84FE6D4A2DFDBDFE27" box="[398,582,367,393]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="281" phylum="Chordata" rank="species" species="septimontium">
|
|||
|
|
<emphasis id="B939EAB8FFA7FF84FE6D4A2DFDBDFE27" box="[398,582,367,393]" italics="true" pageId="20" pageNumber="281">C. septimontium</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in having a higher mean number of enlarged precloacal scales. Other differences between
|
|||
|
|
<taxonomicName id="4C4D4D29FFA7FF84FEDC4AD1FE2FFE02" authority="Grismer & Geissler & Neang & Hartmann & Wagner & Poyarkov, 2021" authorityName="Grismer & Geissler & Neang & Hartmann & Wagner & Poyarkov" authorityYear="2021" box="[319,468,403,429]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="281" phylum="Chordata" rank="species" species="kulenensis" status="sp. nov.">
|
|||
|
|
<emphasis id="B939EAB8FFA7FF84FEDC4AD1FE2FFE02" box="[319,468,403,429]" italics="true" pageId="20" pageNumber="281">C. kulenensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B939EAB8FFA7FF84FE3A4AD1FDC9FE02" bold="true" box="[473,562,403,429]" pageId="20" pageNumber="281">
|
|||
|
|
<taxonomicNameLabel id="A20A57C3FFA7FF84FE3A4AD1FDC9FE02" box="[473,562,403,429]" pageId="20" pageNumber="281" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
and various combinations of species are illustrated in
|
|||
|
|
<figureCitation id="13762A2FFFA7FF84FB694AD1FB13FE02" box="[1162,1256,402,429]" captionStart="FIGURE 5" captionStartId="16.[151,250,1717,1741]" captionTargetBox="[156,1430,842,1687]" captionTargetId="figure-139@16.[151,1436,840,1692]" captionTargetPageId="16" captionText="FIGURE 5. Boxplot comparisons of meristic characters among the species of the Cyrtodactylus intermedius group. Light-blue circles are means and the black horizontal bars are medians. Asterisks denote species bearing statistically significant mean differences from C. kulenensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4636379" httpUri="https://zenodo.org/record/4636379/files/figure.png" pageId="20" pageNumber="281">Figure 5</figureCitation>
|
|||
|
|
and listed
|
|||
|
|
<tableCitation id="C6CF0311FFA7FF84FABD4AD1FF5DFE7E" captionStart="TABLE 1" captionStartId="5.[151,239,461,487]" captionText="TABLE 1. Summary statistics of diagnostic meristic characters and morphological and color pattern characters of the species of the Cyrtodactylus intermedius group. Highlighted character states are those that differ discretely or significantly (p <0.05) from C. kulenensis sp. nov. Data for C. intermedius come from Smith (1917) and Taylor (1963)." pageId="20" pageNumber="281">Table 1</tableCitation>
|
|||
|
|
.
|
|||
|
|
<taxonomicName id="4C4D4D29FFA7FF84FF504AF5FE30FE7E" authority="Grismer & Geissler & Neang & Hartmann & Wagner & Poyarkov, 2021" authorityName="Grismer & Geissler & Neang & Hartmann & Wagner & Poyarkov" authorityYear="2021" box="[179,459,439,465]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="281" phylum="Chordata" rank="species" species="kulenensis" status="sp. nov.">
|
|||
|
|
<emphasis id="B939EAB8FFA7FF84FF504AF5FE30FE7E" box="[179,459,439,465]" italics="true" pageId="20" pageNumber="281">Cyrtodactylus kulenensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B939EAB8FFA7FF84FE314AF5FDD0FE7E" bold="true" box="[466,555,439,465]" pageId="20" pageNumber="281">
|
|||
|
|
<taxonomicNameLabel id="A20A57C3FFA7FF84FE314AF5FDD0FE7E" box="[466,555,439,465]" pageId="20" pageNumber="281" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
is well-separated from all other species in all three multivariate analyses (
|
|||
|
|
<figureCitation id="13762A2FFFA7FF84FA874AF5FF5EFE5A" captionStart="FIGURE 4" captionStartId="14.[151,250,1460,1484]" captionTargetBox="[153,1433,188,1427]" captionTargetId="figure-25@14.[151,1436,181,1436]" captionTargetPageId="14" captionText="FIGURE 4. A. PCA of the species the Cyrtodactylus intermedius group based on meristic characters. Plot points with white borders are centroids. B. DAPC of same. C. Histograms of the factor loadings of the characters contributing most to the variation along PC1, PC2, PC3, and PC4. Abbreviations are in the materials and methods." figureDoi="http://doi.org/10.5281/zenodo.4636377" httpUri="https://zenodo.org/record/4636377/files/figure.png" pageId="20" pageNumber="281">Figs. 4</figureCitation>
|
|||
|
|
and
|
|||
|
|
<figureCitation id="13762A2FFFA7FF84FF3D4A99FF15FE59" box="[222,238,475,502]" captionStart="FIGURE 6" captionStartId="17.[151,250,1108,1132]" captionTargetBox="[153,1436,187,1079]" captionTargetId="figure-17@17.[151,1436,181,1084]" captionTargetPageId="17" captionText="FIGURE 6. MFA scatter plot showing the morphospatial relationships among the species of the Cyrtodactylus intermedius group and bar graphs showing the percent contribution of each data type to the overall variation in the data set. The dashed red line in the bar graphs indicates the expected average value if the contributions of each data type were equal." figureDoi="http://doi.org/10.5281/zenodo.4636381" httpUri="https://zenodo.org/record/4636381/files/figure.png" pageId="20" pageNumber="281">6</figureCitation>
|
|||
|
|
). Additionally,
|
|||
|
|
<taxonomicName id="4C4D4D29FFA7FF84FE7D4A99FDCEFE5A" authority="Grismer & Geissler & Neang & Hartmann & Wagner & Poyarkov, 2021" authorityName="Grismer & Geissler & Neang & Hartmann & Wagner & Poyarkov" authorityYear="2021" box="[414,565,475,501]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="281" phylum="Chordata" rank="species" species="kulenensis" status="sp. nov.">
|
|||
|
|
<emphasis id="B939EAB8FFA7FF84FE7D4A99FDCEFE5A" box="[414,565,475,501]" italics="true" pageId="20" pageNumber="281">C. kulenensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B939EAB8FFA7FF84FDDE4A99FD6CFE5A" bold="true" box="[573,663,475,501]" pageId="20" pageNumber="281">
|
|||
|
|
<taxonomicNameLabel id="A20A57C3FFA7FF84FDDE4A99FD6CFE5A" box="[573,663,475,501]" pageId="20" pageNumber="281" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
differs from its sister species
|
|||
|
|
<taxonomicName id="4C4D4D29FFA7FF84FC0D4A99FB70FE5A" authorityName="Gray" authorityYear="1827" box="[1006,1163,475,501]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="281" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis id="B939EAB8FFA7FF84FC0D4A99FB70FE5A" box="[1006,1163,475,501]" italics="true" pageId="20" pageNumber="281">Cyrtodactylus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
sp. 2 by an uncorrected pairwise sequence divergence of 4.4%. It is likely this percentage would be much different using a full read of ND2 and its flanking tRNAs.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BF236AAFFA7FF84FF244905FD1EFCBA" blockId="20.[151,1437,151,789]" pageId="20" pageNumber="281">
|
|||
|
|
<emphasis id="B939EAB8FFA7FF84FF244905FE7FFDCF" bold="true" box="[199,388,582,609]" pageId="20" pageNumber="281">Natural history.</emphasis>
|
|||
|
|
The Phnom Kulen National Park encompasses a small (
|
|||
|
|
<quantity id="4CB59B4FFFA7FF84FC0F4905FBBAFDCE" box="[1004,1089,583,609]" metricMagnitude="5" metricUnit="m" metricValue="3.7399999999999998" pageId="20" pageNumber="281" unit="km" value="374.0">374 km</quantity>
|
|||
|
|
<superScript id="7C389BE2FFA7FF84FBA24904FBB2FDFB" attach="left" box="[1089,1097,582,596]" fontSize="6" pageId="20" pageNumber="281">2</superScript>
|
|||
|
|
), forested, hilly, habitat-island in the Phnom Kulen mountain massif surrounded by the lowlands of northern
|
|||
|
|
<collectingCountry id="F35A763AFFA7FF84FBF54929FB72FD2A" box="[1046,1161,619,645]" name="Cambodia" pageId="20" pageNumber="281">Cambodia</collectingCountry>
|
|||
|
|
in the Tonle Sap Basin. The massif reaches
|
|||
|
|
<quantity id="4CB59B4FFFA7FF84FE9049CCFE41FD06" box="[371,442,654,681]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="20" pageNumber="281" unit="m" value="500.0">500 m</quantity>
|
|||
|
|
in elevation and supports semi-evergreen forest on the hillsides and upper elevations and degraded dry deciduous dipterocarp forest at lower elevations. According to
|
|||
|
|
<bibRefCitation id="EFDC4B5BFFA7FF84FBE149F1FB0DFD62" author="Geissler, P. & Hartmann, T. & Ihlow, F. & Neang, T. & Seng, R. & Wagner, P. & Bohme, W." box="[1026,1270,691,717]" pageId="20" pageNumber="281" pagination="40 - 63" refId="ref12170" refString="Geissler, P., Hartmann, T., Ihlow, F., Neang, T., Seng, R., Wagner, P. & Bohme, W. (2019) Herpetofauna of the Phnom Kulen National Park, northern Cambodia - an annotated checklist. Cambodian Journal of Natural History, 2019, 40 - 63. https: // doi. org / 10.7717 / peerj. 7841 / fig- 1" type="journal article" year="2019">
|
|||
|
|
Geissler
|
|||
|
|
<emphasis id="B939EAB8FFA7FF84FB8549F1FB5BFD62" box="[1126,1184,691,717]" italics="true" pageId="20" pageNumber="281">et al.</emphasis>
|
|||
|
|
(2019)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
, all specimens were collected from 17:30–23:00 hrs in the month of June and September hiding in rock crevices or while foraging on sandstone rocks in semi-evergreen forest (
|
|||
|
|
<figureCitation id="13762A2FFFA7FF84FD7149B9FD2CFCBA" box="[658,727,763,789]" captionStart="FIGURE 9" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[212,1375,181,1919]" captionTargetId="figure-17@21.[212,1375,181,1919]" captionTargetPageId="21" captionText="FIGURE 9. Habitat of Cyrtodactylus kulenensis sp. nov. at Phnom Kulen National Park, Phnom Kbal Spean, Banteay Srei District, Siem Reap Province, Cambodia." figureDoi="http://doi.org/10.5281/zenodo.4636389" httpUri="https://zenodo.org/record/4636389/files/figure.png" pageId="20" pageNumber="281">Fig. 9</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|