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<document id="B5AF7725110124F85F541F9A3B23B3A3" ID-DOI="10.5281/zenodo.277308" ID-GBIF-Dataset="61dd45a6-918a-4b12-8157-6090f7514613" ID-ISSN="1175-5326" ID-Zenodo-Dep="277308" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460286829391" checkinUser="plazi" docAuthor="Rosa, Simone Policena" docDate="2011" docId="03C58797BE01292DFF57FA9CFA2CF65E" docLanguage="en" docName="zt02831p022.pdf" docOrigin="Zootaxa 2831" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Globothorax" docType="key" docVersion="8" lastPageNumber="3" masterDocId="FFFCFFEFBE00292FFFC0FF89FFB7F724" masterDocTitle="New species of Triplonychus Candèze and Globothorax Fleutiaux from Brazil (Coleoptera, Elateridae, Cardiophorinae)" masterLastPageNumber="22" masterPageNumber="1" pageNumber="2" updateTime="1698250676213" updateUser="plazi">
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<mods:titleInfo id="1FCCB43BBEDE7BD595FF5E952EA4C6C7">
<mods:title id="90D3757F8965E0BCF0AC9F1DAC3E9D36">New species of Triplonychus Candèze and Globothorax Fleutiaux from Brazil (Coleoptera, Elateridae, Cardiophorinae)</mods:title>
</mods:titleInfo>
<mods:name id="8774D2497DE57DA852F3AB4992460AF8" type="personal">
<mods:role id="A83F4FBCF88CABB4981A84B5209C461F">
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<mods:namePart id="062FA970E6DC02693369539CB0B3AA5C">Rosa, Simone Policena</mods:namePart>
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<mods:typeOfResource id="433CDE5C6A087640F269809EDF7C856E">text</mods:typeOfResource>
<mods:relatedItem id="2831927C50B45569977899EF3CB51877" type="host">
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<mods:title id="44D995B6E44D0DDE64AAE30812EA8A44">Zootaxa</mods:title>
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<mods:part id="9E04516FB4CBE071CE049F79B1CF21FB">
<mods:date id="CFFB36414B8A962A2790240D4ABF05DF">2011</mods:date>
<mods:detail id="93BA40F5085C1F9DB04B17C6798CD199" type="volume">
<mods:number id="B5B1BD2FBAAEF65B49B3B90D40076CC2">2831</mods:number>
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<mods:classification id="24988BC8D8CA71439FDFBBF2D53770D4">journal article</mods:classification>
<mods:identifier id="365B97ED7D66FA8322B8EA0E928FC49A" type="DOI">10.5281/zenodo.277308</mods:identifier>
<mods:identifier id="F64395A0123B1B5613DCA59A7EF57BD0" type="GBIF-Dataset">61dd45a6-918a-4b12-8157-6090f7514613</mods:identifier>
<mods:identifier id="C0097CD49FCB35389D52887B2EE23560" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="794FE75238AE903D4919D88FA1C37EE2" type="Zenodo-Dep">277308</mods:identifier>
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<treatment id="03C58797BE01292DFF57FA9CFA2CF65E" ID-DOI="http://doi.org/10.5281/zenodo.6188942" ID-GBIF-Taxon="119397853" ID-Zenodo-Dep="6188942" LSID="urn:lsid:plazi:treatment:03C58797BE01292DFF57FA9CFA2CF65E" httpUri="http://treatment.plazi.org/id/03C58797BE01292DFF57FA9CFA2CF65E" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
<subSubSection id="C376650ABE01292EFF57FA9CFCE8F20B" box="[151,863,1301,1327]" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="8BD33681BE01292EFF57FA9CFCE8F20B" blockId="1.[151,863,1301,1327]" box="[151,863,1301,1327]" pageId="1" pageNumber="2">
<heading id="D09B81EDBE01292EFF57FA9CFCE8F20B" bold="true" box="[151,863,1301,1327]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
<emphasis id="B918EA93BE01292EFF57FA9CFCE8F20B" bold="true" box="[151,863,1301,1327]" pageId="1" pageNumber="2">
Key to Brazilian species of
<taxonomicName id="4C6C4D02BE01292EFE28FA9CFD36F20B" box="[488,641,1301,1327]" class="Insecta" family="Elateridae" genus="Globothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B918EA93BE01292EFE28FA9CFD36F20B" bold="true" box="[488,641,1301,1327]" italics="true" pageId="1" pageNumber="2">Globothorax</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6C4D02BE01292EFD7FFA9CFCE8F20B" box="[703,863,1301,1327]" class="Insecta" family="Elateridae" genus="Triplonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B918EA93BE01292EFD7FFA9CFCE8F20B" bold="true" box="[703,863,1301,1327]" italics="true" pageId="1" pageNumber="2">Triplonychus</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C376650ABE01292DFF57FAD1FA2CF65E" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="key">
<keyStep id="30982E24BE01292EFF57FAD1FA2BF2E1" pageId="1" pageNumber="2">
<paragraph id="8BD33681BE01292EFF57FAD1FA2CF2AF" blockId="1.[151,1436,1366,2035]" pageId="1" pageNumber="2">
<keyLead id="309D95B4BE01292EFF57FAD1FA2CF2AF" pageId="1" pageNumber="2">
1. Prothorax with lateral sides strongly rounded (
<figureCitation id="13572A04BE01292EFD5DFAD1FD51F249" box="[669,742,1368,1389]" captionStart="FIGURES 49 – 60" captionStartId="13.[151,264,1958,1981]" captionTargetBox="[205,1354,198,1910]" captionTargetId="figure@13.[194,1363,193,1911]" captionTargetPageId="13" captionText="FIGURES 49 – 60. Globothorax latidens sp. nov. 49, antenna; 50, labrum; 51, mandible; 52, labium; 53, maxilla; 54 – 56, prothorax (dorsal, lateral, ventral); 57, 58, pterothorax (ventral, lateroventral); 59, wing; 60, abdomen." httpUri="https://zenodo.org/record/277315/files/figure.png" pageId="1" pageNumber="2">Fig. 54</figureCitation>
), mandibles with lateroventral margin curved 90o to apex (
<figureCitation id="13572A04BE01292EFAD7FAD1FAEAF249" box="[1303,1373,1368,1389]" captionStart="FIGURES 68 – 76" captionStartId="15.[151,264,1443,1466]" captionTargetBox="[151,1436,193,1421]" captionTargetId="figure@15.[151,1436,193,1422]" captionTargetPageId="15" captionText="FIGURES 68 – 76. Triplonychus cruspinosus sp. nov. 68, maxilla; 69, detail of maxilla; 70, labium. T. tibialatus sp. nov. 71, maxilla; 72, detail of maxilla; 73, labium. Head: 74, T. cruspinosus sp. nov.; 75, T. tibialatus sp. nov.; 76, Globothorax latidens sp. nov." httpUri="https://zenodo.org/record/277317/files/figure.png" pageId="1" pageNumber="2">Fig. 76</figureCitation>
)................................................................................ 2
<taxonomicName id="4C6C4D02BE01292EFB45FAFFFA2CF2AF" authority="Fleutiaux, 1891" authorityName="Fleutiaux" authorityYear="1891" box="[1157,1435,1398,1419]" class="Insecta" family="Elateridae" genus="Globothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B918EA93BE01292EFB45FAFFFB48F2AF" box="[1157,1279,1398,1419]" italics="true" pageId="1" pageNumber="2">Globothorax</emphasis>
<bibRefCitation id="EFFD4B70BE01292EFAC5FAFFFA2CF2AF" author="Fleutiaux" box="[1285,1435,1398,1419]" pageId="1" pageNumber="2" refString="Fleutiaux, E. (1891) Description d'un genre nouveau d'Elaterides. Annales de la Societe Entomologique de Belgique, 35, 232 - 233." type="journal article" year="1891">Fleutiaux, 1891</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD33681BE01292EFF57FA1AFA2BF2E1" blockId="1.[151,1436,1366,2035]" pageId="1" pageNumber="2">
<keyLead id="309D95B4BE01292EFF57FA1AFA2BF2E1" pageId="1" pageNumber="2">
- Prothorax with lateral sides subparallel to rounded (
<figureCitation id="13572A04BE01292EFD1EFA1AFC93F28C" box="[734,804,1427,1448]" captionStart="FIGURES 1 – 8" captionStartId="3.[151,264,1819,1842]" captionTargetBox="[154,1426,218,1746]" captionTargetId="figure@3.[152,1437,211,1765]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Triplonychus cruspinosus sp. nov. 1, antenna; 2, mandible (dorsal); 3 – 5, prothorax (dorsal, lateral, ventral); 6, abdomen; 7, 8, pterothorax (ventral, lateroventral)." httpUri="https://zenodo.org/record/277309/files/figure.png" pageId="1" pageNumber="2">Figs. 3</figureCitation>
,
<figureCitation id="13572A04BE01292EFCF1FA1AFCFCF28C" box="[817,843,1427,1448]" captionStart="FIGURES 15 – 25" captionStartId="6.[151,264,1978,2001]" captionTargetBox="[215,1386,193,1941]" captionTargetId="figure@6.[195,1400,193,1965]" captionTargetPageId="6" captionText="FIGURES 15 – 25. Triplonychus crassifemoris sp. nov. 15, antenna; 16, labrum; 17, mandible; 18, maxilla; 19, labium; 20, head; 21 – 23, prothorax (dorsal, lateral, ventral); 24, 25, pterothorax (ventral, lateroventral)." httpUri="https://zenodo.org/record/277311/files/figure.png" pageId="1" pageNumber="2">21</figureCitation>
), mandibles with lateroventral margin gradually (
<figureCitation id="13572A04BE01292EFAF1FA1AFACEF28C" box="[1329,1401,1427,1448]" captionStart="FIGURES 68 – 76" captionStartId="15.[151,264,1443,1466]" captionTargetBox="[151,1436,193,1421]" captionTargetId="figure@15.[151,1436,193,1422]" captionTargetPageId="15" captionText="FIGURES 68 – 76. Triplonychus cruspinosus sp. nov. 68, maxilla; 69, detail of maxilla; 70, labium. T. tibialatus sp. nov. 71, maxilla; 72, detail of maxilla; 73, labium. Head: 74, T. cruspinosus sp. nov.; 75, T. tibialatus sp. nov.; 76, Globothorax latidens sp. nov." httpUri="https://zenodo.org/record/277317/files/figure.png" pageId="1" pageNumber="2">Fig. 74</figureCitation>
) or abruptly (
<figureCitation id="13572A04BE01292EFEFEFA39FE33F2E1" box="[318,388,1456,1477]" captionStart="FIGURES 68 – 76" captionStartId="15.[151,264,1443,1466]" captionTargetBox="[151,1436,193,1421]" captionTargetId="figure@15.[151,1436,193,1422]" captionTargetPageId="15" captionText="FIGURES 68 – 76. Triplonychus cruspinosus sp. nov. 68, maxilla; 69, detail of maxilla; 70, labium. T. tibialatus sp. nov. 71, maxilla; 72, detail of maxilla; 73, labium. Head: 74, T. cruspinosus sp. nov.; 75, T. tibialatus sp. nov.; 76, Globothorax latidens sp. nov." httpUri="https://zenodo.org/record/277317/files/figure.png" pageId="1" pageNumber="2">Fig. 75</figureCitation>
) curved to apex................................................. 3
<taxonomicName id="4C6C4D02BE01292EFB4AFA38FA2BF2E1" authority="Candeze, 1860" authorityName="Candeze" authorityYear="1860" box="[1162,1436,1456,1478]" class="Insecta" family="Elateridae" genus="Triplonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B918EA93BE01292EFB4AFA38FAB0F2E2" box="[1162,1287,1457,1478]" italics="true" pageId="1" pageNumber="2">Triplonychus</emphasis>
<bibRefCitation id="EFFD4B70BE01292EFACDFA39FA2BF2E1" author="Candeze" box="[1293,1436,1456,1477]" pageId="1" pageNumber="2" refString="Candeze, E. (1860) Monographie des Elaterides. Vol. 3. Memoires de la Societe Royale des Sciences de Liege, 15, 512 pp." type="book" year="1860">Candèze, 1860</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30982E24BE01292EFF57FA47FA2CF11F" pageId="1" pageNumber="2">
<paragraph id="8BD33681BE01292EFF57FA47FA2CF124" blockId="1.[151,1436,1366,2035]" pageId="1" pageNumber="2">
<keyLead id="309D95B4BE01292EFF57FA47FA2CF124" pageId="1" pageNumber="2">
2. Frontal carina present and produced, labrum semielliptical, posterior leg with trochanter convex, tibia slightly widened apicad with long and fine spiniform setae (
<figureCitation id="13572A04BE01292EFDF6FA62FD0FF124" box="[566,696,1515,1536]" captionStart="FIGURES 105 – 108" captionStartId="20.[151,264,932,955]" captionTargetBox="[206,1379,196,903]" captionTargetId="figure@20.[191,1395,193,911]" captionTargetPageId="20" captionText="FIGURES 105 – 108. Holotype of Globothorax chevrolati Fleutiaux, 1891 (EC 1939). 105, dorsal; 106, ventral; 107, lateral; 108, labels. (Courtesy of MNHN Collection, Paris)." httpUri="https://zenodo.org/record/277322/files/figure.png" pageId="1" pageNumber="2">Fig. 105–107</figureCitation>
).....................................
<taxonomicName id="4C6C4D02BE01292EFB41FA65FA2CF124" authority="Fleutiaux, 1891" authorityName="Fleutiaux" authorityYear="1891" box="[1153,1435,1515,1537]" class="Insecta" family="Elateridae" genus="Globothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="chevrolati">
<emphasis id="B918EA93BE01292EFB41FA65FB48F125" box="[1153,1279,1516,1537]" italics="true" pageId="1" pageNumber="2">G. chevrolati</emphasis>
<bibRefCitation id="EFFD4B70BE01292EFAC5FA62FA2CF124" author="Fleutiaux" box="[1285,1435,1515,1536]" pageId="1" pageNumber="2" refString="Fleutiaux, E. (1891) Description d'un genre nouveau d'Elaterides. Annales de la Societe Entomologique de Belgique, 35, 232 - 233." type="journal article" year="1891">Fleutiaux, 1891</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD33681BE01292EFF57F981FA2CF11F" blockId="1.[151,1436,1366,2035]" pageId="1" pageNumber="2">
<keyLead id="309D95B4BE01292EFF57F981FA2CF11F" pageId="1" pageNumber="2">
- Frontal carina absent, labrum trapezoidal with apex acute, posterior leg with trochanter and femur strongly convex, tibia wide with short and stout spiniform setae (
<figureCitation id="13572A04BE01292EFD85F9AFFD3CF11F" box="[581,651,1574,1595]" captionStart="FIGURES 77 – 90" captionStartId="16.[151,264,1935,1958]" captionTargetBox="[184,1388,193,1898]" captionTargetId="figure@16.[184,1389,193,1898]" captionTargetPageId="16" captionText="FIGURES 77 – 90. Anterior, median and posterior legs (outer surface): 77 – 79, Triplonychus cruspinosus sp. nov.; 80 – 82, T. crassifemoris sp. nov.; 83 – 85, T. tibialatus sp. nov.; 86 – 88, Globothorax latidens sp. nov. Claws: 89, T. cruspinosus sp. nov.; 90, G. latidens sp. nov." httpUri="https://zenodo.org/record/277318/files/figure.png" pageId="1" pageNumber="2">Fig. 88</figureCitation>
) (MT) (
<figureCitation id="13572A04BE01292EFD19F9AFFC9EF11F" box="[729,809,1574,1595]" captionStart="FIGURES 49 – 60" captionStartId="13.[151,264,1958,1981]" captionTargetBox="[205,1354,198,1910]" captionTargetId="figure@13.[194,1363,193,1911]" captionTargetPageId="13" captionText="FIGURES 49 – 60. Globothorax latidens sp. nov. 49, antenna; 50, labrum; 51, mandible; 52, labium; 53, maxilla; 54 – 56, prothorax (dorsal, lateral, ventral); 57, 58, pterothorax (ventral, lateroventral); 59, wing; 60, abdomen." httpUri="https://zenodo.org/record/277315/files/figure.png" pageId="1" pageNumber="2">Figs. 50</figureCitation>
,
<figureCitation id="13572A04BE01292EFCF3F9AFFCFAF11F" box="[819,845,1574,1595]" captionStart="FIGURES 68 – 76" captionStartId="15.[151,264,1443,1466]" captionTargetBox="[151,1436,193,1421]" captionTargetId="figure@15.[151,1436,193,1422]" captionTargetPageId="15" captionText="FIGURES 68 – 76. Triplonychus cruspinosus sp. nov. 68, maxilla; 69, detail of maxilla; 70, labium. T. tibialatus sp. nov. 71, maxilla; 72, detail of maxilla; 73, labium. Head: 74, T. cruspinosus sp. nov.; 75, T. tibialatus sp. nov.; 76, Globothorax latidens sp. nov." httpUri="https://zenodo.org/record/277317/files/figure.png" pageId="1" pageNumber="2">76</figureCitation>
,
<figureCitation id="13572A04BE01292EFC97F9AFFCCBF11F" box="[855,892,1574,1595]" captionStart="FIGURES 101 – 104" captionStartId="19.[151,264,1959,1982]" captionTargetBox="[222,1358,193,1919]" captionTargetId="figure@19.[222,1370,193,1925]" captionTargetPageId="19" captionText="FIGURES 101 – 104. Dorsal habitus of holotypes: 101, Triplonychus cruspinosus sp. nov. (4,5 mm length); 102, T. crassifemoris sp. nov. (7 mm length); 103, T. tibialatus sp. nov. (8 mm length); 104, Globothorax latidens sp. nov. (6 mm length)." httpUri="https://zenodo.org/record/277321/files/figure.png" pageId="1" pageNumber="2">104</figureCitation>
)............................
<taxonomicName id="4C6C4D02BE01292EFB23F9AFFAFEF11F" box="[1251,1353,1574,1595]" class="Insecta" family="Elateridae" genus="Globothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="latidens" status="sp. nov.">
<emphasis id="B918EA93BE01292EFB23F9AFFAFEF11F" box="[1251,1353,1574,1595]" italics="true" pageId="1" pageNumber="2">G. latidens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57E8BE01292EFA8FF9AFFA2CF11F" box="[1359,1435,1574,1595]" pageId="1" pageNumber="2" rank="species">
<emphasis id="B918EA93BE01292EFA8FF9AFFA2CF11F" bold="true" box="[1359,1435,1574,1595]" pageId="1" pageNumber="2">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30982E24BE01292EFF57F9CAFA2BF151" pageId="1" pageNumber="2">
<paragraph id="8BD33681BE01292EFF57F9CAFA2BF17C" blockId="1.[151,1436,1366,2035]" box="[151,1436,1603,1624]" pageId="1" pageNumber="2">
<keyLead id="309D95B4BE01292EFF57F9CAFA2BF17C" box="[151,1436,1603,1624]" pageId="1" pageNumber="2">3 (1). Prothorax longer than wide............................................................................. 4</keyLead>
</paragraph>
<paragraph id="8BD33681BE01292EFF57F9E9FA2BF151" blockId="1.[151,1436,1366,2035]" box="[151,1436,1632,1653]" pageId="1" pageNumber="2">
<keyLead id="309D95B4BE01292EFF57F9E9FA2BF151" box="[151,1436,1632,1653]" pageId="1" pageNumber="2">- Prothorax as wide as or wider than long.................................................................. 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="30982E24BE01292EFF57F9F7FA2CF194" pageId="1" pageNumber="2">
<paragraph id="8BD33681BE01292EFF57F9F7FA2BF1B7" blockId="1.[151,1436,1366,2035]" box="[151,1436,1662,1683]" pageId="1" pageNumber="2">
<keyLead id="309D95B4BE01292EFF57F9F7FA2BF1B7" box="[151,1436,1662,1683]" pageId="1" pageNumber="2">4 (3). Frontal carina strongly produced, mandibles tridentate........................................................ 5</keyLead>
</paragraph>
<paragraph id="8BD33681BE01292EFF57F912FA2CF194" blockId="1.[151,1436,1366,2035]" box="[151,1435,1691,1713]" pageId="1" pageNumber="2">
<keyLead id="309D95B4BE01292EFF57F912FA2CF194" box="[151,1435,1691,1713]" pageId="1" pageNumber="2">
- Frontal carina not produced, mandibles unidentate (MS) (
<figureCitation id="13572A04BE01292EFCC8F912FCEEF194" box="[776,857,1691,1712]" captionStart="FIGURES 34 – 42" captionStartId="9.[151,264,1984,2007]" captionTargetBox="[219,1376,198,1945]" captionTargetId="figure@9.[194,1388,193,1949]" captionTargetPageId="9" captionText="FIGURES 34 – 42. Triplonychus tibialatus sp. nov. 34, antenna; 35, mandible; 36 – 38, prothorax (dorsal, lateral, ventral); 39, abdomen; 40, 41 pterothorax (ventral, lateroventral); 42, wing." httpUri="https://zenodo.org/record/277313/files/figure.png" pageId="1" pageNumber="2">Figs. 35</figureCitation>
,
<figureCitation id="13572A04BE01292EFCA3F912FCCAF194" box="[867,893,1691,1712]" captionStart="FIGURES 68 – 76" captionStartId="15.[151,264,1443,1466]" captionTargetBox="[151,1436,193,1421]" captionTargetId="figure@15.[151,1436,193,1422]" captionTargetPageId="15" captionText="FIGURES 68 – 76. Triplonychus cruspinosus sp. nov. 68, maxilla; 69, detail of maxilla; 70, labium. T. tibialatus sp. nov. 71, maxilla; 72, detail of maxilla; 73, labium. Head: 74, T. cruspinosus sp. nov.; 75, T. tibialatus sp. nov.; 76, Globothorax latidens sp. nov." httpUri="https://zenodo.org/record/277317/files/figure.png" pageId="1" pageNumber="2">75</figureCitation>
,
<figureCitation id="13572A04BE01292EFC47F912FC1BF194" box="[903,940,1691,1712]" captionStart="FIGURES 101 – 104" captionStartId="19.[151,264,1959,1982]" captionTargetBox="[222,1358,193,1919]" captionTargetId="figure@19.[222,1370,193,1925]" captionTargetPageId="19" captionText="FIGURES 101 – 104. Dorsal habitus of holotypes: 101, Triplonychus cruspinosus sp. nov. (4,5 mm length); 102, T. crassifemoris sp. nov. (7 mm length); 103, T. tibialatus sp. nov. (8 mm length); 104, Globothorax latidens sp. nov. (6 mm length)." httpUri="https://zenodo.org/record/277321/files/figure.png" pageId="1" pageNumber="2">103</figureCitation>
).......................
<taxonomicName id="4C6C4D02BE01292EFB17F915FAFEF195" box="[1239,1353,1692,1713]" class="Insecta" family="Elateridae" genus="Triplonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="tibialatus" status="sp. nov.">
<emphasis id="B918EA93BE01292EFB17F915FAFEF195" box="[1239,1353,1692,1713]" italics="true" pageId="1" pageNumber="2">T. tibialatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57E8BE01292EFA8FF912FA2CF194" box="[1359,1435,1691,1712]" pageId="1" pageNumber="2" rank="species">
<emphasis id="B918EA93BE01292EFA8FF912FA2CF194" bold="true" box="[1359,1435,1691,1712]" pageId="1" pageNumber="2">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30982E24BE01292EFF57F931FA2BF1CF" pageId="1" pageNumber="2">
<paragraph id="8BD33681BE01292EFF57F931FA2BF1E9" blockId="1.[151,1436,1366,2035]" box="[151,1436,1720,1741]" pageId="1" pageNumber="2">
<keyLead id="309D95B4BE01292EFF57F931FA2BF1E9" box="[151,1436,1720,1741]" pageId="1" pageNumber="2">5 (4). Pronotum with double punctation......................................................................... 7</keyLead>
</paragraph>
<paragraph id="8BD33681BE01292EFF57F95FFA2BF1CF" blockId="1.[151,1436,1366,2035]" box="[151,1436,1750,1771]" pageId="1" pageNumber="2">
<keyLead id="309D95B4BE01292EFF57F95FFA2BF1CF" box="[151,1436,1750,1771]" pageId="1" pageNumber="2">
- Pronotum with single punctation (PA, BA).........................................
<taxonomicName id="4C6C4D02BE01292EFBB4F95FFA2BF1CF" authority="Eschscholtz, 1829" authorityName="Eschscholtz" authorityYear="1829" box="[1140,1436,1750,1771]" class="Insecta" family="Elateridae" genus="Triplonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="ephipiger">
<emphasis id="B918EA93BE01292EFBB4F95FFB50F1CF" box="[1140,1255,1750,1771]" italics="true" pageId="1" pageNumber="2">T. ephipiger</emphasis>
Eschscholtz, 1829
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30982E24BE01292EFF57F97AFA2CF001" pageId="1" pageNumber="2">
<paragraph id="8BD33681BE01292EFF57F97AFA2CF02C" blockId="1.[151,1436,1366,2035]" box="[151,1435,1779,1800]" pageId="1" pageNumber="2">
<keyLead id="309D95B4BE01292EFF57F97AFA2CF02C" box="[151,1435,1779,1800]" pageId="1" pageNumber="2">6 (3). Pronotum with double punctation....................................................................... 10</keyLead>
</paragraph>
<paragraph id="8BD33681BE01292EFF57F899FA2CF001" blockId="1.[151,1436,1366,2035]" box="[151,1435,1808,1829]" pageId="1" pageNumber="2">
<keyLead id="309D95B4BE01292EFF57F899FA2CF001" box="[151,1435,1808,1829]" pageId="1" pageNumber="2">- Pronotum with single punctation......................................................................... 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="30982E24BE01292EFF57F8A7FA2BF044" pageId="1" pageNumber="2">
<paragraph id="8BD33681BE01292EFF57F8A7FA2BF067" blockId="1.[151,1436,1366,2035]" box="[151,1436,1838,1859]" pageId="1" pageNumber="2">
<keyLead id="309D95B4BE01292EFF57F8A7FA2BF067" box="[151,1436,1838,1859]" pageId="1" pageNumber="2">
7 (5). Pronotum dark brown to black with a pair of longitudinal orange stripes (AM, PA, MT)........
<taxonomicName id="4C6C4D02BE01292EFB54F8A7FA2BF067" authority="Erichson, 1846" authorityName="Erichson" authorityYear="1846" box="[1172,1436,1838,1859]" class="Insecta" family="Elateridae" genus="Triplonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="plagiatus">
<emphasis id="B918EA93BE01292EFB54F8A7FAB3F067" box="[1172,1284,1838,1859]" italics="true" pageId="1" pageNumber="2">T. plagiatus</emphasis>
Erichson, 1846
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD33681BE01292EFF57F8C2FA2BF044" blockId="1.[151,1436,1366,2035]" box="[151,1436,1867,1888]" pageId="1" pageNumber="2">
<keyLead id="309D95B4BE01292EFF57F8C2FA2BF044" box="[151,1436,1867,1888]" pageId="1" pageNumber="2">- Prothorax brown to black without orange stripes............................................................ 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30982E24BE01292EFF57F8E1FA2CF0BF" pageId="1" pageNumber="2">
<paragraph id="8BD33681BE01292EFF57F8E1FA2BF059" blockId="1.[151,1436,1366,2035]" box="[151,1436,1896,1917]" pageId="1" pageNumber="2">
<keyLead id="309D95B4BE01292EFF57F8E1FA2BF059" box="[151,1436,1896,1917]" pageId="1" pageNumber="2">8 (7). Elytra evenly brown or brown with anterior margin reddish-brown.............................................. 9</keyLead>
</paragraph>
<paragraph id="8BD33681BE01292EFF57F80FFA2CF0BF" blockId="1.[151,1436,1366,2035]" box="[151,1435,1926,1947]" pageId="1" pageNumber="2">
<keyLead id="309D95B4BE01292EFF57F80FFA2CF0BF" box="[151,1435,1926,1947]" pageId="1" pageNumber="2">
- Elytra brown with lateral and sutural edges dark brown to black (PA).......................
<taxonomicName id="4C6C4D02BE01292EFB5CF80FFA2CF0BF" authority="Candeze, 1860" authorityName="Candeze" authorityYear="1860" box="[1180,1435,1926,1947]" class="Insecta" family="Elateridae" genus="Triplonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="ventralis">
<emphasis id="B918EA93BE01292EFB5CF80FFAB0F0BF" box="[1180,1287,1926,1947]" italics="true" pageId="1" pageNumber="2">T. ventralis</emphasis>
<bibRefCitation id="EFFD4B70BE01292EFACDF80FFA2CF0BF" author="Candeze" box="[1293,1435,1926,1947]" pageId="1" pageNumber="2" refString="Candeze, E. (1860) Monographie des Elaterides. Vol. 3. Memoires de la Societe Royale des Sciences de Liege, 15, 512 pp." type="book" year="1860">Candèze, 1860</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30982E24BE01292EFF57F82AFA2CF0D7" pageId="1" pageNumber="2">
<paragraph id="8BD33681BE01292EFF57F82AFA2BF0F2" blockId="1.[151,1436,1366,2035]" pageId="1" pageNumber="2">
<keyLead id="309D95B4BE01292EFF57F82AFA2BF0F2" pageId="1" pageNumber="2">
9 (8). Elytra brown with humeral margin reddish-brown, interstices 3, 5, and 7 carinate at apex, legs yellowish with apex of femur, dorsal margin of tibiae and tarsi darker (BA, PE)......................................
<taxonomicName id="4C6C4D02BE01292EFB48F848FA2BF0F2" authority="Erichson, 1840" authorityName="Erichson" authorityYear="1840" box="[1160,1436,1985,2006]" class="Insecta" family="Elateridae" genus="Triplonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="longicollis">
<emphasis id="B918EA93BE01292EFB48F848FAB3F0F2" box="[1160,1284,1985,2006]" italics="true" pageId="1" pageNumber="2">T. longicollis</emphasis>
Erichson, 1840
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD33681BE01292EFF57F857FA2CF0D7" blockId="1.[151,1436,1366,2035]" box="[151,1435,2014,2035]" pageId="1" pageNumber="2">
<keyLead id="309D95B4BE01292EFF57F857FA2CF0D7" box="[151,1435,2014,2035]" pageId="1" pageNumber="2">
- Elytra evenly colored, interstices 3, 5, 7 not carinate, legs evenly yellowish (PA).................
<taxonomicName id="4C6C4D02BE01292EFB76F857FA2CF0D7" authority="Candeze, 1865" authorityName="Candeze" authorityYear="1865" box="[1206,1435,2014,2035]" class="Insecta" family="Elateridae" genus="Triplonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="fulvus">
<emphasis id="B918EA93BE01292EFB76F857FAB0F0D7" box="[1206,1287,2014,2035]" italics="true" pageId="1" pageNumber="2">T. fulvus</emphasis>
<bibRefCitation id="EFFD4B70BE01292EFACDF857FA2CF0D7" author="Candeze" box="[1293,1435,2014,2035]" pageId="1" pageNumber="2" refString="Candeze, E. (1865) Elaterides nouveaux. Fascicule Premier. Memoires Couronnes par l'Academie Royale des Sciences, Belgique, 17, 1 - 63." type="journal article" year="1865">Candèze, 1865</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30982E24BE02292DFF57FF1EFA2CF7C3" pageId="2" pageNumber="3">
<paragraph id="8BD33681BE02292DFF57FF1EFA2CF7EE" blockId="2.[151,1436,151,378]" pageId="2" pageNumber="3">
<keyLead id="309D95B4BE02292DFF57FF1EFA2CF7EE" pageId="2" pageNumber="3">
10 (6). Mandibles unidentate, meso- and metatibiae tibiae broadened, metafemur convex (MS) (
<figureCitation id="13572A04BE02292DFBF3FF1EFB34F788" box="[1075,1155,151,172]" captionStart="FIGURES 34 – 42" captionStartId="9.[151,264,1984,2007]" captionTargetBox="[219,1376,198,1945]" captionTargetId="figure@9.[194,1388,193,1949]" captionTargetPageId="9" captionText="FIGURES 34 – 42. Triplonychus tibialatus sp. nov. 34, antenna; 35, mandible; 36 – 38, prothorax (dorsal, lateral, ventral); 39, abdomen; 40, 41 pterothorax (ventral, lateroventral); 42, wing." httpUri="https://zenodo.org/record/277313/files/figure.png" pageId="2" pageNumber="3">Figs. 35</figureCitation>
,
<figureCitation id="13572A04BE02292DFB4DFF1EFB10F788" box="[1165,1191,151,172]" captionStart="FIGURES 68 – 76" captionStartId="15.[151,264,1443,1466]" captionTargetBox="[151,1436,193,1421]" captionTargetId="figure@15.[151,1436,193,1422]" captionTargetPageId="15" captionText="FIGURES 68 – 76. Triplonychus cruspinosus sp. nov. 68, maxilla; 69, detail of maxilla; 70, labium. T. tibialatus sp. nov. 71, maxilla; 72, detail of maxilla; 73, labium. Head: 74, T. cruspinosus sp. nov.; 75, T. tibialatus sp. nov.; 76, Globothorax latidens sp. nov." httpUri="https://zenodo.org/record/277317/files/figure.png" pageId="2" pageNumber="3">75</figureCitation>
,
<figureCitation id="13572A04BE02292DFB71FF1EFB58F788" box="[1201,1263,151,172]" captionStart="FIGURES 77 – 90" captionStartId="16.[151,264,1935,1958]" captionTargetBox="[184,1388,193,1898]" captionTargetId="figure@16.[184,1389,193,1898]" captionTargetPageId="16" captionText="FIGURES 77 – 90. Anterior, median and posterior legs (outer surface): 77 – 79, Triplonychus cruspinosus sp. nov.; 80 – 82, T. crassifemoris sp. nov.; 83 – 85, T. tibialatus sp. nov.; 86 – 88, Globothorax latidens sp. nov. Claws: 89, T. cruspinosus sp. nov.; 90, G. latidens sp. nov." httpUri="https://zenodo.org/record/277318/files/figure.png" pageId="2" pageNumber="3">84, 85</figureCitation>
,
<figureCitation id="13572A04BE02292DFB39FF1EFA96F788" box="[1273,1313,151,172]" captionStart="FIGURES 101 – 104" captionStartId="19.[151,264,1959,1982]" captionTargetBox="[222,1358,193,1919]" captionTargetId="figure@19.[222,1370,193,1925]" captionTargetPageId="19" captionText="FIGURES 101 – 104. Dorsal habitus of holotypes: 101, Triplonychus cruspinosus sp. nov. (4,5 mm length); 102, T. crassifemoris sp. nov. (7 mm length); 103, T. tibialatus sp. nov. (8 mm length); 104, Globothorax latidens sp. nov. (6 mm length)." httpUri="https://zenodo.org/record/277321/files/figure.png" pageId="2" pageNumber="3">103</figureCitation>
)..............................................................................................
<taxonomicName id="4C6C4D02BE02292DFB17FF3CFAFEF7EE" box="[1239,1353,181,202]" class="Insecta" family="Elateridae" genus="Triplonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="tibialatus" status="sp. nov.">
<emphasis id="B918EA93BE02292DFB17FF3CFAFEF7EE" box="[1239,1353,181,202]" italics="true" pageId="2" pageNumber="3">T. tibialatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57E8BE02292DFA8FFF3CFA2CF7EE" box="[1359,1435,181,202]" pageId="2" pageNumber="3" rank="species">
<emphasis id="B918EA93BE02292DFA8FFF3CFA2CF7EE" bold="true" box="[1359,1435,181,202]" pageId="2" pageNumber="3">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BD33681BE02292DFF57FF5BFA2CF7C3" blockId="2.[151,1436,151,378]" box="[151,1435,210,231]" pageId="2" pageNumber="3">
<keyLead id="309D95B4BE02292DFF57FF5BFA2CF7C3" box="[151,1435,210,231]" pageId="2" pageNumber="3">
- Mandibles tridentate, legs slender (PA)...................................................
<taxonomicName id="4C6C4D02BE02292DFB7EFF5BFA2CF7C3" authority="Candeze, 1860" authorityName="Candeze" authorityYear="1860" box="[1214,1435,210,231]" class="Insecta" family="Elateridae" genus="Triplonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="rufus">
<emphasis id="B918EA93BE02292DFB7EFF5BFAB2F7C3" box="[1214,1285,210,231]" italics="true" pageId="2" pageNumber="3">T. rufus</emphasis>
<bibRefCitation id="EFFD4B70BE02292DFACDFF5BFA2CF7C3" author="Candeze" box="[1293,1435,210,231]" pageId="2" pageNumber="3" refString="Candeze, E. (1860) Monographie des Elaterides. Vol. 3. Memoires de la Societe Royale des Sciences de Liege, 15, 512 pp." type="book" year="1860">Candèze, 1860</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30982E24BE02292DFF57FF66FA2CF606" pageId="2" pageNumber="3">
<paragraph id="8BD33681BE02292DFF57FF66FA2BF620" blockId="2.[151,1436,151,378]" box="[151,1436,239,260]" pageId="2" pageNumber="3">
<keyLead id="309D95B4BE02292DFF57FF66FA2BF620" box="[151,1436,239,260]" pageId="2" pageNumber="3">11 (6). Mandibles small, legs slender......................................................................... 12</keyLead>
</paragraph>
<paragraph id="8BD33681BE02292DFF57FE84FA2CF606" blockId="2.[151,1436,151,378]" box="[151,1435,269,290]" pageId="2" pageNumber="3">
<keyLead id="309D95B4BE02292DFF57FE84FA2CF606" box="[151,1435,269,290]" pageId="2" pageNumber="3">
- Mandibles large, metafemur broadened and convex, tibiae widened apicad (MA) (
<figureCitation id="13572A04BE02292DFC33FE84FBF4F606" box="[1011,1091,269,290]" captionStart="FIGURES 15 – 25" captionStartId="6.[151,264,1978,2001]" captionTargetBox="[215,1386,193,1941]" captionTargetId="figure@6.[195,1400,193,1965]" captionTargetPageId="6" captionText="FIGURES 15 – 25. Triplonychus crassifemoris sp. nov. 15, antenna; 16, labrum; 17, mandible; 18, maxilla; 19, labium; 20, head; 21 – 23, prothorax (dorsal, lateral, ventral); 24, 25, pterothorax (ventral, lateroventral)." httpUri="https://zenodo.org/record/277311/files/figure.png" pageId="2" pageNumber="3">Figs. 20</figureCitation>
,
<figureCitation id="13572A04BE02292DFB8DFE84FBD0F606" box="[1101,1127,269,290]" captionStart="FIGURES 77 – 90" captionStartId="16.[151,264,1935,1958]" captionTargetBox="[184,1388,193,1898]" captionTargetId="figure@16.[184,1389,193,1898]" captionTargetPageId="16" captionText="FIGURES 77 – 90. Anterior, median and posterior legs (outer surface): 77 – 79, Triplonychus cruspinosus sp. nov.; 80 – 82, T. crassifemoris sp. nov.; 83 – 85, T. tibialatus sp. nov.; 86 – 88, Globothorax latidens sp. nov. Claws: 89, T. cruspinosus sp. nov.; 90, G. latidens sp. nov." httpUri="https://zenodo.org/record/277318/files/figure.png" pageId="2" pageNumber="3">82</figureCitation>
,
<figureCitation id="13572A04BE02292DFBB1FE84FB21F606" box="[1137,1174,269,290]" captionStart="FIGURES 101 – 104" captionStartId="19.[151,264,1959,1982]" captionTargetBox="[222,1358,193,1919]" captionTargetId="figure@19.[222,1370,193,1925]" captionTargetPageId="19" captionText="FIGURES 101 – 104. Dorsal habitus of holotypes: 101, Triplonychus cruspinosus sp. nov. (4,5 mm length); 102, T. crassifemoris sp. nov. (7 mm length); 103, T. tibialatus sp. nov. (8 mm length); 104, Globothorax latidens sp. nov. (6 mm length)." httpUri="https://zenodo.org/record/277321/files/figure.png" pageId="2" pageNumber="3">102</figureCitation>
).
<taxonomicName id="4C6C4D02BE02292DFB70FE84FAFEF606" box="[1200,1353,269,290]" class="Insecta" family="Elateridae" genus="Triplonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="crassifemoris" status="sp. nov.">
<emphasis id="B918EA93BE02292DFB70FE84FAFEF606" box="[1200,1353,269,290]" italics="true" pageId="2" pageNumber="3">T. crassifemoris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57E8BE02292DFA8FFE84FA2CF606" box="[1359,1435,269,290]" pageId="2" pageNumber="3" rank="species">
<emphasis id="B918EA93BE02292DFA8FFE84FA2CF606" bold="true" box="[1359,1435,269,290]" pageId="2" pageNumber="3">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30982E24BE02292DFF57FEA3FA2CF65E" pageId="2" pageNumber="3">
<paragraph id="8BD33681BE02292DFF57FEA3FA2CF61B" blockId="2.[151,1436,151,378]" box="[151,1435,298,319]" pageId="2" pageNumber="3">
<keyLead id="309D95B4BE02292DFF57FEA3FA2CF61B" box="[151,1435,298,319]" pageId="2" pageNumber="3">
12 (11). Antenna reaching hind angles of prothorax, elytral interstices 3, 5 and 7 carinate at apex (PA)....
<taxonomicName id="4C6C4D02BE02292DFB61FEA3FA2CF61B" authority="Candeze, 1860" authorityName="Candeze" authorityYear="1860" box="[1185,1435,298,319]" class="Insecta" family="Elateridae" genus="Triplonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="costatus">
<emphasis id="B918EA93BE02292DFB61FEA3FAB0F61B" box="[1185,1287,298,319]" italics="true" pageId="2" pageNumber="3">T. costatus</emphasis>
<bibRefCitation id="EFFD4B70BE02292DFACDFEA3FA2CF61B" author="Candeze" box="[1293,1435,298,319]" pageId="2" pageNumber="3" refString="Candeze, E. (1860) Monographie des Elaterides. Vol. 3. Memoires de la Societe Royale des Sciences de Liege, 15, 512 pp." type="book" year="1860">Candèze, 1860</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD33681BE02292DFF57FECEFA2CF65E" blockId="2.[151,1436,151,378]" pageId="2" pageNumber="3">
<keyLead id="309D95B4BE02292DFF57FECEFA2CF65E" pageId="2" pageNumber="3">
- Antenna surpassing the hind angles of prothorax by two or three antennomeres, elytral interstices 3, 5 and 7 not carinate at apex (AM) (
<figureCitation id="13572A04BE02292DFE98FEECFE04F65E" box="[344,435,357,378]" captionStart="FIGURES 101 – 104" captionStartId="19.[151,264,1959,1982]" captionTargetBox="[222,1358,193,1919]" captionTargetId="figure@19.[222,1370,193,1925]" captionTargetPageId="19" captionText="FIGURES 101 – 104. Dorsal habitus of holotypes: 101, Triplonychus cruspinosus sp. nov. (4,5 mm length); 102, T. crassifemoris sp. nov. (7 mm length); 103, T. tibialatus sp. nov. (8 mm length); 104, Globothorax latidens sp. nov. (6 mm length)." httpUri="https://zenodo.org/record/277321/files/figure.png" pageId="2" pageNumber="3">Figs. 101</figureCitation>
)................................................................
<taxonomicName id="4C6C4D02BE02292DFB7FFEECFAFFF65E" box="[1215,1352,357,378]" class="Insecta" family="Elateridae" genus="Triplonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="cruspinosus" status="sp. nov.">
<emphasis id="B918EA93BE02292DFB7FFEECFAFFF65E" box="[1215,1352,357,378]" italics="true" pageId="2" pageNumber="3">T. cruspinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57E8BE02292DFA8FFEECFA2CF65E" box="[1359,1435,357,378]" pageId="2" pageNumber="3" rank="species">
<emphasis id="B918EA93BE02292DFA8FFEECFA2CF65E" bold="true" box="[1359,1435,357,378]" pageId="2" pageNumber="3">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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