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<document ID-DOI="10.11646/zootaxa.5264.1.8" ID-ISSN="1175-5326" ID-Zenodo-Dep="7836278" ID-ZooBank="49FCE66C-0EEE-46A1-B170-F1DDA1837107" approvalRequired="3" approvalRequired_for_taxonomicNames="2" approvalRequired_for_textStreams="1" checkinTime="1681726520946" checkinUser="plazi" docAuthor="Anju, K., Thejass, P., Binoy, C. &amp; Kumar, P. Girish" docDate="2023" docId="039287F7FF828507FF2C84C1FE60FE7E" docLanguage="en" docName="zootaxa.5264.1.8.pdf" docOrigin="Zootaxa 5264 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5264.1.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ceropales (Ceropales) keralaensis Anju, Binoy &amp; Thejass 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="127" masterDocId="FFABFF8FFF87850FFFBB856FFFC5FFAF" masterDocTitle="Taxonomic study on the spider wasp genus Ceropales Latreille, 1796 (Pompilidae: Ceropalinae) with description of two new species from India" masterLastPageNumber="128" masterPageNumber="119" pageNumber="124" updateTime="1681726933130" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomic study on the spider wasp genus Ceropales Latreille, 1796 (Pompilidae: Ceropalinae) with description of two new species from India</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Anju, K.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-3312-7721</mods:nameIdentifier>
<mods:affiliation>P. G. &amp; Research Department of Zoology, Government College Madappally, Vatakara, Kozhikode, Kerala- 673102, India, Affiliated to University of Calicut. &amp; anjukappu 91 @ gmail. com; https: // orcid. org / 0000 - 0003 - 3312 - 7721</mods:affiliation>
<mods:nameIdentifier type="email">anjukappu91@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Thejass, P.</mods:namePart>
<mods:affiliation>P. G. &amp; Research Department of Zoology, Government College Madappally, Vatakara, Kozhikode, Kerala- 673102, India, Affiliated to University of Calicut.</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Binoy, C.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-4309-5460</mods:nameIdentifier>
<mods:affiliation>Department of Zoology, Sree Narayana Guru College, Chelannur, Kozhikode, Kerala- 673616, India, Affiliated to University of Calicut. binoy _ doz @ uoc. ac. in; https: // orcid. org / 0000 - 0003 - 4309 - 5460</mods:affiliation>
<mods:nameIdentifier type="email">binoy_doz@uoc.ac.in</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kumar, P. Girish</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-2121-0165</mods:nameIdentifier>
<mods:affiliation>Western Ghats Regional Centre, Zoological Survey of India, Eranhipalam, Kozhikode, Kerala- 673006, India. kpgiris @ gmail. com; https: // orcid. org / 0000 - 0003 - 2121 - 0165</mods:affiliation>
<mods:nameIdentifier type="email">kpgiris@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-04-12</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5264</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>119</mods:start>
<mods:end>128</mods:end>
</mods:extent>
</mods:part>
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<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5264.1.8</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5264.1.8</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7836278</mods:identifier>
<mods:identifier type="ZooBank">49FCE66C-0EEE-46A1-B170-F1DDA1837107</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7836286" ID-Zenodo-Dep="7836286" LSID="urn:lsid:plazi:treatment:039287F7FF828507FF2C84C1FE60FE7E" httpUri="http://treatment.plazi.org/id/039287F7FF828507FF2C84C1FE60FE7E" lastPageId="8" lastPageNumber="127" pageId="5" pageNumber="124">
<subSubSection box="[151,964,430,457]" pageId="5" pageNumber="124" type="nomenclature">
<paragraph blockId="5.[151,964,430,523]" box="[151,964,430,457]" pageId="5" pageNumber="124">
<heading bold="true" box="[151,964,430,457]" fontSize="11" level="1" pageId="5" pageNumber="124" reason="1">
<emphasis bold="true" box="[151,964,430,457]" pageId="5" pageNumber="124">
<taxonomicName authority="Anju, Binoy &amp; Thejass" authorityName="Anju, Binoy &amp; Thejass" authorityYear="2023" box="[151,857,430,457]" class="Insecta" family="Pompilidae" genus="Ceropales" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="keralaensis" status="sp. nov." subGenus="Ceropales">
<emphasis bold="true" box="[151,272,430,457]" italics="true" pageId="5" pageNumber="124">Ceropales</emphasis>
(
<emphasis bold="true" box="[289,410,430,457]" italics="true" pageId="5" pageNumber="124">Ceropales</emphasis>
)
<emphasis bold="true" box="[427,564,430,457]" italics="true" pageId="5" pageNumber="124">keralaensis</emphasis>
Anju, Binoy &amp; Thejass
</taxonomicName>
,
<taxonomicNameLabel box="[870,964,431,457]" pageId="5" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="124" type="description">
<paragraph blockId="5.[151,964,430,523]" box="[151,275,467,493]" pageId="5" pageNumber="124">
<figureCitation box="[151,275,467,493]" captionStart-0="FIGURES 11–20" captionStart-1="FIGURES 21–23" captionStartId-0="6.[151,264,1899,1923]" captionStartId-1="7.[151,264,1969,1993]" captionTargetBox-0="[219,1368,181,1875]" captionTargetBox-1="[158,1430,1317,1946]" captionTargetId-0="figure-19@6.[219,1368,181,1875]" captionTargetId-1="figure-486@7.[158,1430,1317,1946]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURES 11–20. Ceropales (Ceropales) keralaensis Anju, Binoy &amp; Thejass, sp. nov., holotype, ♀. 11, habitus, lateral view; 12, habitus, dorsal view; 13, head, lateral view; 14, head, frontal view; 15, head, dorsal view; 16, mesosoma, dorsal view; 17, mesosoma, lateral view; 18, wings; 19, metasoma, lateral view; 20, metasoma, dorsal view. Scale bar: 11–2 mm; 12, 13, 17, 18, 20–1 mm; 14, 15, 16, 19–0.5 mm." captionText-1="FIGURES 21–23. Ceropales (Ceropales) keralaensis Anju, Binoy &amp; Thejass, sp. nov., paratype, J. 21, habitus, lateral view; 22, hypopygium; 23, genitalia, dorsal view. Scale bar: 21–2 mm; 22, 23–0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7836284" figureDoi-1="http://doi.org/10.5281/zenodo.7836288" httpUri-0="https://zenodo.org/record/7836284/files/figure.png" httpUri-1="https://zenodo.org/record/7836288/files/figure.png" pageId="5" pageNumber="124">Figs 11–23</figureCitation>
</paragraph>
<paragraph blockId="5.[151,964,430,523]" box="[151,884,499,523]" pageId="5" pageNumber="124">
<uri box="[151,884,499,523]" pageId="5" pageNumber="124">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="124" type="materials_examined">
<paragraph blockId="5.[151,1437,565,2032]" pageId="5" pageNumber="124">
<emphasis bold="true" box="[151,325,565,591]" pageId="5" pageNumber="124">Type material.</emphasis>
<materialsCitation collectingDate="2020-01-16" collectionCode="ZSIK, WGRC" collectorName="Coll. C. Binoy &amp; Regd. No." country="India" latitude="11.482499" location="Viyyur" longLatPrecision="21" longitude="75.685" municipality="Kozhikode district" pageId="5" pageNumber="124" specimenCount="1" specimenCount-female="1" stateProvince="Kerala" typeStatus="holotype">
<typeStatus box="[332,439,565,591]" pageId="5" pageNumber="124">Holotype</typeStatus>
,
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,
<emphasis bold="true" box="[484,548,565,591]" pageId="5" pageNumber="124">
<collectingCountry box="[484,548,565,591]" name="India" pageId="5" pageNumber="124">India</collectingCountry>
</emphasis>
:
<collectingRegion box="[563,640,565,591]" country="India" name="Kerala" pageId="5" pageNumber="124">Kerala</collectingRegion>
,
<collectingMunicipality box="[651,862,565,591]" pageId="5" pageNumber="124">Kozhikode district</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:039287F7FF828507FF2C84C1FE60FE7E:8EE4603AFF82850AFCD3875AFC73FDE0" box="[872,950,565,591]" country="India" latitude="11.482499" longLatPrecision="21" longitude="75.685" municipality="Kozhikode district" name="Viyyur" pageId="5" pageNumber="124" stateProvince="Kerala">Viyyur</location>
(
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,
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),
<date box="[1261,1370,565,591]" pageId="5" pageNumber="124" value="2020-01-16">
<collectingDate box="[1261,1370,565,591]" pageId="5" pageNumber="124" value="2020-01-16">16.i.2020</collectingDate>
</date>
,
<collectorName pageId="5" pageNumber="124">Coll. C. Binoy</collectorName>
, [
<emphasis bold="true" box="[274,350,601,627]" pageId="5" pageNumber="124">
<collectionCode box="[274,343,601,627]" pageId="5" pageNumber="124">ZSIK</collectionCode>
]
</emphasis>
<collectionCode box="[357,419,601,627]" pageId="5" pageNumber="124">ZSIK</collectionCode>
<collectorName box="[426,541,601,628]" pageId="5" pageNumber="124">Regd. No.</collectorName>
ZSI/
<collectionCode box="[597,681,601,627]" pageId="5" pageNumber="124">WGRC</collectionCode>
/IR/INV. 22059.
</materialsCitation>
</paragraph>
<paragraph blockId="5.[151,1437,565,2032]" pageId="5" pageNumber="124">
<materialsCitation collectingDate="2022-06-18" collectionCode="V, ZSIK, WGRC" collectorName="Coll. &amp; D. Hegde &amp; Party &amp; Regd. No." county="Udupi district" latitude="13.86389" location="Areshiroor" longLatPrecision="21" longitude="74.80945" municipality="Mookambika" pageId="5" pageNumber="124" specimenCount="1" specimenCount-female="1" stateProvince="Karnataka" typeStatus="paratype">
<emphasis bold="true" box="[199,326,637,663]" pageId="5" pageNumber="124">
<typeStatus box="[199,321,637,663]" pageId="5" pageNumber="124">Paratypes</typeStatus>
.
</emphasis>
<collectingRegion box="[336,451,637,663]" country="India" name="Karnataka" pageId="5" pageNumber="124">Karnataka</collectingRegion>
:
<specimenCount box="[469,504,637,663]" pageId="5" pageNumber="124" type="female">1♀</specimenCount>
,
<collectingCounty box="[521,682,637,663]" pageId="5" pageNumber="124">Udupi district</collectingCounty>
,
<collectingMunicipality box="[696,845,637,663]" pageId="5" pageNumber="124">Mookambika</collectingMunicipality>
WLS,
<location LSID="urn:lsid:plazi:treatment:039287F7FF828507FF2C84C1FE60FE7E:8EE4603AFF82850AFC1A8712FBDFFD38" box="[929,1050,637,663]" county="Udupi district" latitude="13.86389" longLatPrecision="21" longitude="74.80945" municipality="Mookambika" name="Areshiroor" pageId="5" pageNumber="124" stateProvince="Karnataka">Areshiroor</location>
(
<geoCoordinate box="[1068,1204,639,666]" degrees="13" direction="north" minutes="51" orientation="latitude" pageId="5" pageNumber="124" precision="15" seconds="50" value="13.86389">13º51′50′′N</geoCoordinate>
,
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),
<date pageId="5" pageNumber="124" value="2022-06-18">
<collectingDate pageId="5" pageNumber="124" value="2022-06-18">18.vi. 2022</collectingDate>
</date>
,
<collectorName box="[225,281,673,699]" pageId="5" pageNumber="124">Coll.</collectorName>
<collectionCode box="[291,313,674,700]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="124" type="Museum">V</collectionCode>
.
<collectorName box="[317,427,673,700]" pageId="5" pageNumber="124">D. Hegde</collectorName>
&amp;
<collectorName box="[471,531,674,700]" pageId="5" pageNumber="124">Party</collectorName>
, [
<emphasis bold="true" box="[556,632,673,699]" pageId="5" pageNumber="124">
<collectionCode box="[556,625,673,699]" pageId="5" pageNumber="124">ZSIK</collectionCode>
]
</emphasis>
<collectionCode box="[643,705,673,699]" pageId="5" pageNumber="124">ZSIK</collectionCode>
<collectorName box="[717,836,673,700]" pageId="5" pageNumber="124">Regd. No.</collectorName>
ZSI/
<collectionCode box="[897,983,673,699]" pageId="5" pageNumber="124">WGRC</collectionCode>
/IR/INV.22064
</materialsCitation>
;
<materialsCitation collectingDate="2019-06-30" collectionCode="ZSIK, WGRC" collectorName="Coll. K. P. Hanima Raveendran &amp; Regd. No." country="India" latitude="11.6866665" location="Payanthong" longLatPrecision="21" longitude="75.67389" pageId="5" pageNumber="124" specimenCount="1" stateProvince="Kerala" typeStatus="paratype">
<collectingRegion box="[1166,1241,673,699]" country="India" name="Kerala" pageId="5" pageNumber="124">Kerala</collectingRegion>
: 3J,
<collectingCountry box="[1313,1436,673,699]" name="India" pageId="5" pageNumber="124">Kozhikode</collectingCountry>
district,
<location LSID="urn:lsid:plazi:treatment:039287F7FF828507FF2C84C1FE60FE7E:8EE4603AFF82850AFF4E87AAFEBCFD70" box="[245,377,709,735]" country="India" latitude="11.6866665" longLatPrecision="21" longitude="75.67389" name="Payanthong" pageId="5" pageNumber="124" stateProvince="Kerala">Payanthong</location>
(
<geoCoordinate box="[395,529,711,738]" degrees="11" direction="north" minutes="41" orientation="latitude" pageId="5" pageNumber="124" precision="15" seconds="12" value="11.6866665">11º41′12′′N</geoCoordinate>
,
<geoCoordinate box="[543,676,711,738]" degrees="75" direction="east" minutes="40" orientation="longitude" pageId="5" pageNumber="124" precision="15" seconds="26" value="75.67389">75º40′26′′E</geoCoordinate>
),
<date box="[697,820,709,735]" pageId="5" pageNumber="124" value="2019-06-30">
<collectingDate box="[697,820,709,735]" pageId="5" pageNumber="124" value="2019-06-30">30.vi.2019</collectingDate>
</date>
,
<collectorName box="[834,1190,709,736]" pageId="5" pageNumber="124">Coll. K.P. Hanima Raveendran</collectorName>
, [
<emphasis bold="true" box="[1213,1289,709,735]" pageId="5" pageNumber="124">
<collectionCode box="[1213,1282,709,735]" pageId="5" pageNumber="124">ZSIK</collectionCode>
]
</emphasis>
<collectionCode box="[1298,1360,709,735]" pageId="5" pageNumber="124">ZSIK</collectionCode>
<collectorName pageId="5" pageNumber="124">Regd. No.</collectorName>
ZSI/
<collectionCode box="[249,333,745,771]" pageId="5" pageNumber="124">WGRC</collectionCode>
/IR/INV.22060
<emphasis bold="true" box="[500,514,745,771]" pageId="5" pageNumber="124">–</emphasis>
22062
</materialsCitation>
;
<materialsCitation collectingDate="2019-01-04" collectionCode="ZSIK, WGRC" collectorName="Coll. P. Girish Kumar &amp; Regd. No." country="India" latitude="11.1075" location="Anaikatti" longLatPrecision="21" longitude="76.76611" municipality="Coimbatore district" pageId="5" pageNumber="124" specimenCount="1" specimenCount-female="1" stateProvince="Tamil Nadu" typeStatus="paratype">
<collectingRegion box="[599,732,745,771]" country="India" name="Tamil Nadu" pageId="5" pageNumber="124">Tamil Nadu</collectingRegion>
:
<specimenCount box="[747,782,745,771]" pageId="5" pageNumber="124" type="female">1♀</specimenCount>
,
<collectingMunicipality box="[796,1017,745,771]" pageId="5" pageNumber="124">Coimbatore district</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:039287F7FF828507FF2C84C1FE60FE7E:8EE4603AFF82850AFBB98786FBAFFCAC" box="[1026,1130,745,771]" country="India" latitude="11.1075" longLatPrecision="21" longitude="76.76611" municipality="Coimbatore district" name="Anaikatti" pageId="5" pageNumber="124" stateProvince="Tamil Nadu">Anaikatti</location>
(
<geoCoordinate box="[1146,1280,747,774]" degrees="11" direction="north" minutes="06" orientation="latitude" pageId="5" pageNumber="124" precision="15" seconds="27" value="11.1075">11º06′27′′N</geoCoordinate>
,
<geoCoordinate box="[1292,1424,747,774]" degrees="76" direction="east" minutes="45" orientation="longitude" pageId="5" pageNumber="124" precision="15" seconds="58" value="76.76611">76º45′58′′E</geoCoordinate>
),
<date box="[151,246,781,807]" pageId="5" pageNumber="124" value="2019-01-04">
<collectingDate box="[151,246,781,807]" pageId="5" pageNumber="124" value="2019-01-04">4.i.2019</collectingDate>
</date>
,
<collectorName box="[257,503,781,808]" pageId="5" pageNumber="124">Coll. P. Girish Kumar</collectorName>
, [
<emphasis bold="true" box="[523,599,781,807]" pageId="5" pageNumber="124">
<collectionCode box="[523,592,781,807]" pageId="5" pageNumber="124">ZSIK</collectionCode>
]
</emphasis>
<collectionCode box="[606,668,781,807]" pageId="5" pageNumber="124">ZSIK</collectionCode>
<collectorName box="[675,790,781,808]" pageId="5" pageNumber="124">Regd. No.</collectorName>
ZSI/
<collectionCode box="[847,933,781,807]" pageId="5" pageNumber="124">WGRC</collectionCode>
/IR/INV.22063
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="124" type="diagnosis">
<paragraph blockId="5.[151,1437,565,2032]" pageId="5" pageNumber="124">
<emphasis bold="true" box="[199,322,817,843]" pageId="5" pageNumber="124">Diagnosis.</emphasis>
This new species closely resembles the Chinese species
<taxonomicName authorityName="Latreille" authorityYear="1796" box="[957,1071,817,843]" class="Insecta" family="Pompilidae" genus="Ceropales" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis box="[957,1071,817,843]" italics="true" pageId="5" pageNumber="124">Ceropales</emphasis>
</taxonomicName>
<taxonomicName authority="Lu &amp; Li, 2019" authorityName="Lu &amp; Li" authorityYear="2019" box="[1078,1436,817,844]" class="Insecta" family="Pompilidae" genus="Ceropales" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="longisulcata" subGenus="Ceropales">
(
<emphasis box="[1088,1107,818,844]" italics="true" pageId="5" pageNumber="124">C</emphasis>
.)
<emphasis box="[1129,1267,817,843]" italics="true" pageId="5" pageNumber="124">longisulcata</emphasis>
<bibRefCitation author="Lu, H. &amp; Bashir, N. H. &amp; Li, Q. &amp; Ma, L." box="[1275,1436,817,844]" pageId="5" pageNumber="124" pagination="211 - 220" refId="ref7077" refString="Lu, H., Bashir, N. H., Li, Q. &amp; Ma, L. (2019) Two new species of Ceropales (Ceropales) Latreille (Hymenoptera, Pompilidae), with a key to species from China. Zootaxa, 4551 (2), 211 - 220. https: // doi. org / 10.11646 / zootaxa. 4551.2.5." type="journal article" year="2019">
Lu &amp; Li,
<quantity box="[1380,1436,817,843]" metricMagnitude="1" metricUnit="m" metricValue="5.12826" pageId="5" pageNumber="124" unit="in" value="2019.0">2019</quantity>
</bibRefCitation>
</taxonomicName>
in the key to the species of the subgenus
<taxonomicName authority="Latreille" authorityName="Latreille" authorityYear="1796" box="[598,813,853,879]" class="Insecta" family="Pompilidae" genus="Ceropales" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis box="[598,712,853,879]" italics="true" pageId="5" pageNumber="124">Ceropales</emphasis>
Latreille
</taxonomicName>
from
<collectingCountry box="[878,945,853,879]" name="China" pageId="5" pageNumber="124">China</collectingCountry>
(
<bibRefCitation author="Lu, H. &amp; Bashir, N. H. &amp; Li, Q. &amp; Ma, L." box="[959,1118,853,879]" pageId="5" pageNumber="124" pagination="211 - 220" refId="ref7077" refString="Lu, H., Bashir, N. H., Li, Q. &amp; Ma, L. (2019) Two new species of Ceropales (Ceropales) Latreille (Hymenoptera, Pompilidae), with a key to species from China. Zootaxa, 4551 (2), 211 - 220. https: // doi. org / 10.11646 / zootaxa. 4551.2.5." type="journal article" year="2019">
Lu
<emphasis box="[997,1052,853,879]" italics="true" pageId="5" pageNumber="124">et al.</emphasis>
2019
</bibRefCitation>
). The two species are similar in having flat propodeum with lateral side rather shallowly impressed and with deep medial longitudinal groove; frons remarkably flat anterior to antennal sockets and bent in obtuse-angle on 2/3 away from antennal sockets; ocelli forming acute triangle; and the scutellum and postscutellum remarkably raised, higher than mesonotum in lateral view. However, the new species differs from
<taxonomicName authorityName="Lu &amp; Li" authorityYear="2019" box="[762,937,997,1024]" class="Insecta" family="Pompilidae" genus="Ceropales" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="longisulcata">
<emphasis box="[762,937,997,1024]" italics="true" pageId="5" pageNumber="124">C. longisulcata</emphasis>
</taxonomicName>
in having fore wing with upper margin of SMC2 and SMC3 equal in length (in
<taxonomicName authorityName="Lu &amp; Li" authorityYear="2019" box="[582,720,1033,1059]" class="Insecta" family="Pompilidae" genus="Ceropales" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="longisulcata">
<emphasis box="[582,720,1033,1059]" italics="true" pageId="5" pageNumber="124">longisulcata</emphasis>
</taxonomicName>
, upper margin of SMC3 shorter than SMC2); frons and vertex with combination of small punctures mixed with large punctures (in
<taxonomicName authorityName="Lu &amp; Li" authorityYear="2019" box="[916,1054,1069,1095]" class="Insecta" family="Pompilidae" genus="Ceropales" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="longisulcata">
<emphasis box="[916,1054,1069,1095]" italics="true" pageId="5" pageNumber="124">longisulcata</emphasis>
</taxonomicName>
, deeper and densely set punctures on frons and vertex); frons black without any marking in the middle (in
<taxonomicName authorityName="Lu &amp; Li" authorityYear="2019" box="[998,1136,1105,1131]" class="Insecta" family="Pompilidae" genus="Ceropales" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="longisulcata">
<emphasis box="[998,1136,1105,1131]" italics="true" pageId="5" pageNumber="124">longisulcata</emphasis>
</taxonomicName>
, frons with a yellow spot medially); numerous long yellowish brown semi erect setae on frons and vertex (in
<taxonomicName authorityName="Lu &amp; Li" authorityYear="2019" box="[1077,1215,1141,1167]" class="Insecta" family="Pompilidae" genus="Ceropales" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="longisulcata">
<emphasis box="[1077,1215,1141,1167]" italics="true" pageId="5" pageNumber="124">longisulcata</emphasis>
</taxonomicName>
, such setae absent); and non-confluent ivory patch on face with completely pale yellow clypeus and labrum (in
<taxonomicName authorityName="Lu &amp; Li" authorityYear="2019" box="[1159,1297,1177,1203]" class="Insecta" family="Pompilidae" genus="Ceropales" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="longisulcata">
<emphasis box="[1159,1297,1177,1203]" italics="true" pageId="5" pageNumber="124">longisulcata</emphasis>
</taxonomicName>
, ivory patch confluent on face with clypeus and labrum having median dark patch).
</paragraph>
<paragraph blockId="5.[151,1437,565,2032]" pageId="5" pageNumber="124">
<taxonomicName authority="Anju &amp; Thejass &amp; Binoy &amp; Kumar, 2023" authorityName="Anju &amp; Thejass &amp; Binoy &amp; Kumar" authorityYear="2023" box="[199,448,1249,1275]" class="Insecta" family="Pompilidae" genus="Ceropales" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="keralaensis" status="sp. nov.">
<emphasis box="[199,448,1249,1275]" italics="true" pageId="5" pageNumber="124">Ceropales keralaensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[453,542,1249,1275]" pageId="5" pageNumber="124">
<taxonomicNameLabel box="[453,542,1249,1275]" pageId="5" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is also similar to the other species described here
<taxonomicName authority="Anju &amp; Thejass &amp; Binoy &amp; Kumar, 2023" authorityName="Anju &amp; Thejass &amp; Binoy &amp; Kumar" authorityYear="2023" box="[1096,1313,1249,1275]" class="Insecta" family="Pompilidae" genus="Ceropales" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="anaghae" status="sp. nov.">
<emphasis box="[1096,1313,1249,1275]" italics="true" pageId="5" pageNumber="124">Ceropales anaghae</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1319,1408,1249,1275]" pageId="5" pageNumber="124">
<taxonomicNameLabel box="[1319,1408,1249,1275]" pageId="5" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in general appearance and coloration. However, it differs from
<taxonomicName authority="Anju &amp; Thejass &amp; Binoy &amp; Kumar, 2023" authorityName="Anju &amp; Thejass &amp; Binoy &amp; Kumar" authorityYear="2023" box="[818,946,1285,1312]" class="Insecta" family="Pompilidae" genus="Ceropales" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="anaghae" status="sp. nov.">
<emphasis box="[818,946,1285,1312]" italics="true" pageId="5" pageNumber="124">C. anaghae</emphasis>
</taxonomicName>
<emphasis bold="true" box="[952,1040,1285,1311]" pageId="5" pageNumber="124">
<taxonomicNameLabel box="[952,1040,1285,1311]" pageId="5" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in having numerous long yellowish brown semi erect setae on frons and vertex (in
<taxonomicName authorityName="Anju &amp; Thejass &amp; Binoy &amp; Kumar" authorityYear="2023" box="[687,783,1321,1347]" class="Insecta" family="Pompilidae" genus="Ceropales" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="anaghae">
<emphasis box="[687,783,1321,1347]" italics="true" pageId="5" pageNumber="124">anaghae</emphasis>
</taxonomicName>
such setae absent); non-confluent ivory patch on face (in
<taxonomicName authorityName="Anju &amp; Thejass &amp; Binoy &amp; Kumar" authorityYear="2023" box="[151,247,1357,1383]" class="Insecta" family="Pompilidae" genus="Ceropales" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="anaghae">
<emphasis box="[151,247,1357,1383]" italics="true" pageId="5" pageNumber="124">anaghae</emphasis>
</taxonomicName>
, ivory patch confluent on face); clypeus completely pale yellow without any black marking (in
<taxonomicName authorityName="Anju &amp; Thejass &amp; Binoy &amp; Kumar" authorityYear="2023" box="[1333,1429,1357,1383]" class="Insecta" family="Pompilidae" genus="Ceropales" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="anaghae">
<emphasis box="[1333,1429,1357,1383]" italics="true" pageId="5" pageNumber="124">anaghae</emphasis>
</taxonomicName>
, median black spot on ivory coloured clypeus); medial emargination of compound eye not deep (in
<taxonomicName authorityName="Anju &amp; Thejass &amp; Binoy &amp; Kumar" authorityYear="2023" box="[1333,1429,1393,1419]" class="Insecta" family="Pompilidae" genus="Ceropales" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="anaghae">
<emphasis box="[1333,1429,1393,1419]" italics="true" pageId="5" pageNumber="124">anaghae</emphasis>
</taxonomicName>
, compound eye with deep medial emargination); head coarsely pitted with numerous large punctures (in
<taxonomicName authorityName="Anju &amp; Thejass &amp; Binoy &amp; Kumar" authorityYear="2023" box="[1333,1429,1429,1455]" class="Insecta" family="Pompilidae" genus="Ceropales" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="anaghae">
<emphasis box="[1333,1429,1429,1455]" italics="true" pageId="5" pageNumber="124">anaghae</emphasis>
</taxonomicName>
, head coarsely pitted with very few large punctures and not as prominent); antenna pale yellowish ventrally, brownish black dorsally (in
<taxonomicName authorityName="Anju &amp; Thejass &amp; Binoy &amp; Kumar" authorityYear="2023" box="[364,460,1501,1527]" class="Insecta" family="Pompilidae" genus="Ceropales" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="anaghae">
<emphasis box="[364,460,1501,1527]" italics="true" pageId="5" pageNumber="124">anaghae</emphasis>
</taxonomicName>
, antenna entirely brownish black); and mandible black with apex reddish brown (in
<taxonomicName authorityName="Anju &amp; Thejass &amp; Binoy &amp; Kumar" authorityYear="2023" box="[151,247,1537,1563]" class="Insecta" family="Pompilidae" genus="Ceropales" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="anaghae">
<emphasis box="[151,247,1537,1563]" italics="true" pageId="5" pageNumber="124">anaghae</emphasis>
</taxonomicName>
, basal 2/3
<superScript attach="left" box="[362,376,1537,1551]" fontSize="6" pageId="5" pageNumber="124">rd</superScript>
of mandible pale yellowish with apex reddish brown).
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="126" pageId="5" pageNumber="124" type="description">
<paragraph blockId="5.[151,1437,565,2032]" box="[199,1142,1573,1600]" pageId="5" pageNumber="124">
<emphasis bold="true" box="[199,446,1573,1599]" pageId="5" pageNumber="124">Description. Female.</emphasis>
(
<emphasis bold="true" box="[462,530,1573,1599]" pageId="5" pageNumber="124">11–20</emphasis>
). Body length
<quantity box="[698,801,1573,1600]" metricMagnitude="-3" metricUnit="m" metricValue="4.83" pageId="5" pageNumber="124" unit="mm" value="4.83">4.83 mm</quantity>
; length of fore wing
<quantity box="[1035,1138,1573,1600]" metricMagnitude="-3" metricUnit="m" metricValue="4.07" pageId="5" pageNumber="124" unit="mm" value="4.07">4.07 mm</quantity>
.
</paragraph>
<paragraph blockId="5.[151,1437,565,2032]" pageId="5" pageNumber="124">
<emphasis bold="true" box="[199,287,1609,1636]" pageId="5" pageNumber="124">
<emphasis bold="true" box="[199,280,1609,1636]" italics="true" pageId="5" pageNumber="124">Colour</emphasis>
.
</emphasis>
Overall colour black with antenna brownish black dorsally, pale yellowish ventrally; frons between antennal sockets and adjacent to the medial emargination of compound eyes pale yellow; lower frons below antenna black; bands on outer margin of compound eyes pale yellow; clypeus and labrum pale yellow; mandible black with apex reddish brown; pronotum black with band on posterior margin and posterolateral angle pale yellowish; mesonotum entirely black; metanotum black with medial patch pale yellow; tegula pale yellow; propodeum black with posterolateral corner pale yellow; large spot on outer face of all coxae pale yellow, remainder black; trochanter of fore and mid leg brownish black, that of hind leg black; all femora and tibia honey brown; fore and mid tarsus light honey brown, hind tarsus dark brown; hind tibial spur pale yellow; posterior yellowish white band on T1–T5; T6 with large yellowish white spot; wings hyaline; apical region of fore wing with light brown infumation; veins and pterostigma dark brown.
</paragraph>
<paragraph blockId="5.[151,1437,565,2032]" lastBlockId="7.[151,1437,151,1293]" lastPageId="7" lastPageNumber="126" pageId="5" pageNumber="124">
<emphasis bold="true" box="[199,341,1969,1996]" pageId="5" pageNumber="124">
<emphasis bold="true" box="[199,334,1969,1996]" italics="true" pageId="5" pageNumber="124">Pubescence</emphasis>
.
</emphasis>
Body covered with white pubescence on head and mesosoma, metasoma bare; lower frons densely pubescent; head with numerous yellowish brown setae concentrated on upper frons and vertex (
<figureCitation box="[1337,1421,2005,2031]" captionStart="FIGURES 11–20" captionStartId="6.[151,264,1899,1923]" captionTargetBox="[219,1368,181,1875]" captionTargetId="figure-19@6.[219,1368,181,1875]" captionTargetPageId="6" captionText="FIGURES 11–20. Ceropales (Ceropales) keralaensis Anju, Binoy &amp; Thejass, sp. nov., holotype, ♀. 11, habitus, lateral view; 12, habitus, dorsal view; 13, head, lateral view; 14, head, frontal view; 15, head, dorsal view; 16, mesosoma, dorsal view; 17, mesosoma, lateral view; 18, wings; 19, metasoma, lateral view; 20, metasoma, dorsal view. Scale bar: 11–2 mm; 12, 13, 17, 18, 20–1 mm; 14, 15, 16, 19–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836284" httpUri="https://zenodo.org/record/7836284/files/figure.png" pageId="5" pageNumber="124">Fig. 13</figureCitation>
); clypeus and labrum having pubescence moderately; small sensillae present mainly along the ventral side of antennal segments (
<figureCitation box="[273,357,187,213]" captionStart="FIGURES 11–20" captionStartId="6.[151,264,1899,1923]" captionTargetBox="[219,1368,181,1875]" captionTargetId="figure-19@6.[219,1368,181,1875]" captionTargetPageId="6" captionText="FIGURES 11–20. Ceropales (Ceropales) keralaensis Anju, Binoy &amp; Thejass, sp. nov., holotype, ♀. 11, habitus, lateral view; 12, habitus, dorsal view; 13, head, lateral view; 14, head, frontal view; 15, head, dorsal view; 16, mesosoma, dorsal view; 17, mesosoma, lateral view; 18, wings; 19, metasoma, lateral view; 20, metasoma, dorsal view. Scale bar: 11–2 mm; 12, 13, 17, 18, 20–1 mm; 14, 15, 16, 19–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836284" httpUri="https://zenodo.org/record/7836284/files/figure.png" pageId="7" pageNumber="126">Fig. 13</figureCitation>
); pronotum with moderately dense yellowish brown setae (
<figureCitation box="[1035,1120,187,213]" captionStart="FIGURES 11–20" captionStartId="6.[151,264,1899,1923]" captionTargetBox="[219,1368,181,1875]" captionTargetId="figure-19@6.[219,1368,181,1875]" captionTargetPageId="6" captionText="FIGURES 11–20. Ceropales (Ceropales) keralaensis Anju, Binoy &amp; Thejass, sp. nov., holotype, ♀. 11, habitus, lateral view; 12, habitus, dorsal view; 13, head, lateral view; 14, head, frontal view; 15, head, dorsal view; 16, mesosoma, dorsal view; 17, mesosoma, lateral view; 18, wings; 19, metasoma, lateral view; 20, metasoma, dorsal view. Scale bar: 11–2 mm; 12, 13, 17, 18, 20–1 mm; 14, 15, 16, 19–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836284" httpUri="https://zenodo.org/record/7836284/files/figure.png" pageId="7" pageNumber="126">Fig. 17</figureCitation>
); pubescence on mesosoma concentrated along the lateral margins; pubescence on propodeum moderate, midline somewhat bare.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7836284" ID-Zenodo-Dep="7836284" httpUri="https://zenodo.org/record/7836284/files/figure.png" pageId="6" pageNumber="125" startId="6.[151,264,1899,1923]" targetBox="[219,1368,181,1875]" targetPageId="6">
<paragraph blockId="6.[151,1437,1899,2032]" pageId="6" pageNumber="125">
<emphasis bold="true" box="[151,340,1899,1923]" pageId="6" pageNumber="125">FIGURES 11–20.</emphasis>
<taxonomicName authority="Anju, Binoy &amp; Thejass" authorityName="Anju, Binoy &amp; Thejass" authorityYear="2023" box="[346,944,1899,1924]" class="Insecta" family="Pompilidae" genus="Ceropales" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="keralaensis" status="sp. nov." subGenus="Ceropales">
<emphasis box="[346,449,1899,1923]" italics="true" pageId="6" pageNumber="125">Ceropales</emphasis>
(
<emphasis box="[465,568,1899,1923]" italics="true" pageId="6" pageNumber="125">Ceropales</emphasis>
)
<emphasis box="[584,699,1899,1923]" italics="true" pageId="6" pageNumber="125">keralaensis</emphasis>
Anju, Binoy &amp; Thejass
</taxonomicName>
,
<emphasis bold="true" box="[955,1037,1900,1924]" pageId="6" pageNumber="125">
<taxonomicNameLabel box="[955,1037,1900,1924]" pageId="6" pageNumber="125" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, ♀. 11, habitus, lateral view; 12, habitus, dorsal view; 13, head, lateral view; 14, head, frontal view; 15, head, dorsal view; 16, mesosoma, dorsal view; 17, mesosoma, lateral view; 18, wings; 19, metasoma, lateral view; 20, metasoma, dorsal view. Scale bar: 11–2 mm; 12, 13, 17, 18, 20–1 mm; 14, 15, 16, 19–0.5 mm.
</paragraph>
</caption>
<paragraph blockId="7.[151,1437,151,1293]" pageId="7" pageNumber="126">
<emphasis bold="true" box="[199,268,258,285]" pageId="7" pageNumber="126">
<emphasis bold="true" box="[199,261,258,285]" italics="true" pageId="7" pageNumber="126">Head</emphasis>
.
</emphasis>
Head oblong in frontal view, 1.2× as wide as long (
<figureCitation box="[883,970,259,285]" captionStart="FIGURES 11–20" captionStartId="6.[151,264,1899,1923]" captionTargetBox="[219,1368,181,1875]" captionTargetId="figure-19@6.[219,1368,181,1875]" captionTargetPageId="6" captionText="FIGURES 11–20. Ceropales (Ceropales) keralaensis Anju, Binoy &amp; Thejass, sp. nov., holotype, ♀. 11, habitus, lateral view; 12, habitus, dorsal view; 13, head, lateral view; 14, head, frontal view; 15, head, dorsal view; 16, mesosoma, dorsal view; 17, mesosoma, lateral view; 18, wings; 19, metasoma, lateral view; 20, metasoma, dorsal view. Scale bar: 11–2 mm; 12, 13, 17, 18, 20–1 mm; 14, 15, 16, 19–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836284" httpUri="https://zenodo.org/record/7836284/files/figure.png" pageId="7" pageNumber="126">Fig. 14</figureCitation>
); frons occupying the interantennal area conspicuously elevated; a shallow median frontal line seen from anterior ocellus to interantennal space (
<figureCitation box="[1337,1422,295,321]" captionStart="FIGURES 11–20" captionStartId="6.[151,264,1899,1923]" captionTargetBox="[219,1368,181,1875]" captionTargetId="figure-19@6.[219,1368,181,1875]" captionTargetPageId="6" captionText="FIGURES 11–20. Ceropales (Ceropales) keralaensis Anju, Binoy &amp; Thejass, sp. nov., holotype, ♀. 11, habitus, lateral view; 12, habitus, dorsal view; 13, head, lateral view; 14, head, frontal view; 15, head, dorsal view; 16, mesosoma, dorsal view; 17, mesosoma, lateral view; 18, wings; 19, metasoma, lateral view; 20, metasoma, dorsal view. Scale bar: 11–2 mm; 12, 13, 17, 18, 20–1 mm; 14, 15, 16, 19–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836284" httpUri="https://zenodo.org/record/7836284/files/figure.png" pageId="7" pageNumber="126">Fig. 14</figureCitation>
); frons and vertex coarsely pitted throughout, large punctures more prominent between pitted surface; compound eyes reniform, inner margins distinctly sinuate, strongly convergent below; UID as long as MID, LID 0.5× UID and MID (
<figureCitation box="[159,244,403,429]" captionStart="FIGURES 11–20" captionStartId="6.[151,264,1899,1923]" captionTargetBox="[219,1368,181,1875]" captionTargetId="figure-19@6.[219,1368,181,1875]" captionTargetPageId="6" captionText="FIGURES 11–20. Ceropales (Ceropales) keralaensis Anju, Binoy &amp; Thejass, sp. nov., holotype, ♀. 11, habitus, lateral view; 12, habitus, dorsal view; 13, head, lateral view; 14, head, frontal view; 15, head, dorsal view; 16, mesosoma, dorsal view; 17, mesosoma, lateral view; 18, wings; 19, metasoma, lateral view; 20, metasoma, dorsal view. Scale bar: 11–2 mm; 12, 13, 17, 18, 20–1 mm; 14, 15, 16, 19–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836284" httpUri="https://zenodo.org/record/7836284/files/figure.png" pageId="7" pageNumber="126">Fig. 14</figureCitation>
); ocellar area slightly raised, ocelli in acute triangle, OOD 1.95× POD (
<figureCitation box="[1063,1147,402,429]" captionStart="FIGURES 11–20" captionStartId="6.[151,264,1899,1923]" captionTargetBox="[219,1368,181,1875]" captionTargetId="figure-19@6.[219,1368,181,1875]" captionTargetPageId="6" captionText="FIGURES 11–20. Ceropales (Ceropales) keralaensis Anju, Binoy &amp; Thejass, sp. nov., holotype, ♀. 11, habitus, lateral view; 12, habitus, dorsal view; 13, head, lateral view; 14, head, frontal view; 15, head, dorsal view; 16, mesosoma, dorsal view; 17, mesosoma, lateral view; 18, wings; 19, metasoma, lateral view; 20, metasoma, dorsal view. Scale bar: 11–2 mm; 12, 13, 17, 18, 20–1 mm; 14, 15, 16, 19–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836284" httpUri="https://zenodo.org/record/7836284/files/figure.png" pageId="7" pageNumber="126">Fig. 15</figureCitation>
); clypeus slightly convex in lateral view, more than twice as broad as long, anterior margin slightly emarginated medially (
<figureCitation box="[1250,1335,439,465]" captionStart="FIGURES 11–20" captionStartId="6.[151,264,1899,1923]" captionTargetBox="[219,1368,181,1875]" captionTargetId="figure-19@6.[219,1368,181,1875]" captionTargetPageId="6" captionText="FIGURES 11–20. Ceropales (Ceropales) keralaensis Anju, Binoy &amp; Thejass, sp. nov., holotype, ♀. 11, habitus, lateral view; 12, habitus, dorsal view; 13, head, lateral view; 14, head, frontal view; 15, head, dorsal view; 16, mesosoma, dorsal view; 17, mesosoma, lateral view; 18, wings; 19, metasoma, lateral view; 20, metasoma, dorsal view. Scale bar: 11–2 mm; 12, 13, 17, 18, 20–1 mm; 14, 15, 16, 19–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836284" httpUri="https://zenodo.org/record/7836284/files/figure.png" pageId="7" pageNumber="126">Fig. 14</figureCitation>
); labrum highly exposed beneath the clypeus (
<figureCitation box="[569,653,475,501]" captionStart="FIGURES 11–20" captionStartId="6.[151,264,1899,1923]" captionTargetBox="[219,1368,181,1875]" captionTargetId="figure-19@6.[219,1368,181,1875]" captionTargetPageId="6" captionText="FIGURES 11–20. Ceropales (Ceropales) keralaensis Anju, Binoy &amp; Thejass, sp. nov., holotype, ♀. 11, habitus, lateral view; 12, habitus, dorsal view; 13, head, lateral view; 14, head, frontal view; 15, head, dorsal view; 16, mesosoma, dorsal view; 17, mesosoma, lateral view; 18, wings; 19, metasoma, lateral view; 20, metasoma, dorsal view. Scale bar: 11–2 mm; 12, 13, 17, 18, 20–1 mm; 14, 15, 16, 19–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836284" httpUri="https://zenodo.org/record/7836284/files/figure.png" pageId="7" pageNumber="126">Fig. 14</figureCitation>
); mandible bidentate, malar space reduced; antenna thick, scape 1.4× as long as wide and 2.3× pedicel in profile, flagellomeres almost twice as long as broad, F1–F5 almost of equal length (
<figureCitation box="[236,320,547,573]" captionStart="FIGURES 11–20" captionStartId="6.[151,264,1899,1923]" captionTargetBox="[219,1368,181,1875]" captionTargetId="figure-19@6.[219,1368,181,1875]" captionTargetPageId="6" captionText="FIGURES 11–20. Ceropales (Ceropales) keralaensis Anju, Binoy &amp; Thejass, sp. nov., holotype, ♀. 11, habitus, lateral view; 12, habitus, dorsal view; 13, head, lateral view; 14, head, frontal view; 15, head, dorsal view; 16, mesosoma, dorsal view; 17, mesosoma, lateral view; 18, wings; 19, metasoma, lateral view; 20, metasoma, dorsal view. Scale bar: 11–2 mm; 12, 13, 17, 18, 20–1 mm; 14, 15, 16, 19–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836284" httpUri="https://zenodo.org/record/7836284/files/figure.png" pageId="7" pageNumber="126">Fig. 13</figureCitation>
).
</paragraph>
<paragraph blockId="7.[151,1437,151,1293]" pageId="7" pageNumber="126">
<emphasis bold="true" box="[199,329,583,609]" pageId="7" pageNumber="126">
<emphasis bold="true" box="[199,322,583,609]" italics="true" pageId="7" pageNumber="126">Mesosoma</emphasis>
.
</emphasis>
Pronotum 0.5× as long as wide, densely pitted with large punctures scattered irregularly in between; anterolateral angle acute, posterolateral angle rounded (
<figureCitation box="[770,852,619,645]" captionStart="FIGURES 11–20" captionStartId="6.[151,264,1899,1923]" captionTargetBox="[219,1368,181,1875]" captionTargetId="figure-19@6.[219,1368,181,1875]" captionTargetPageId="6" captionText="FIGURES 11–20. Ceropales (Ceropales) keralaensis Anju, Binoy &amp; Thejass, sp. nov., holotype, ♀. 11, habitus, lateral view; 12, habitus, dorsal view; 13, head, lateral view; 14, head, frontal view; 15, head, dorsal view; 16, mesosoma, dorsal view; 17, mesosoma, lateral view; 18, wings; 19, metasoma, lateral view; 20, metasoma, dorsal view. Scale bar: 11–2 mm; 12, 13, 17, 18, 20–1 mm; 14, 15, 16, 19–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836284" httpUri="https://zenodo.org/record/7836284/files/figure.png" pageId="7" pageNumber="126">Fig. 17</figureCitation>
); posterior margin nearly straight; mesoscutum 1.2× as wide as long, densely pitted along with irregularly scattered large punctures (
<figureCitation box="[1045,1129,655,681]" captionStart="FIGURES 11–20" captionStartId="6.[151,264,1899,1923]" captionTargetBox="[219,1368,181,1875]" captionTargetId="figure-19@6.[219,1368,181,1875]" captionTargetPageId="6" captionText="FIGURES 11–20. Ceropales (Ceropales) keralaensis Anju, Binoy &amp; Thejass, sp. nov., holotype, ♀. 11, habitus, lateral view; 12, habitus, dorsal view; 13, head, lateral view; 14, head, frontal view; 15, head, dorsal view; 16, mesosoma, dorsal view; 17, mesosoma, lateral view; 18, wings; 19, metasoma, lateral view; 20, metasoma, dorsal view. Scale bar: 11–2 mm; 12, 13, 17, 18, 20–1 mm; 14, 15, 16, 19–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836284" httpUri="https://zenodo.org/record/7836284/files/figure.png" pageId="7" pageNumber="126">Fig. 16</figureCitation>
); parapsidal furrow present through the lateral sides of mesoscutum longitudinally, reaching close to anterior margin (
<figureCitation box="[1163,1247,691,717]" captionStart="FIGURES 11–20" captionStartId="6.[151,264,1899,1923]" captionTargetBox="[219,1368,181,1875]" captionTargetId="figure-19@6.[219,1368,181,1875]" captionTargetPageId="6" captionText="FIGURES 11–20. Ceropales (Ceropales) keralaensis Anju, Binoy &amp; Thejass, sp. nov., holotype, ♀. 11, habitus, lateral view; 12, habitus, dorsal view; 13, head, lateral view; 14, head, frontal view; 15, head, dorsal view; 16, mesosoma, dorsal view; 17, mesosoma, lateral view; 18, wings; 19, metasoma, lateral view; 20, metasoma, dorsal view. Scale bar: 11–2 mm; 12, 13, 17, 18, 20–1 mm; 14, 15, 16, 19–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836284" httpUri="https://zenodo.org/record/7836284/files/figure.png" pageId="7" pageNumber="126">Fig. 16</figureCitation>
); mesoscutellum with conspicuous prescutellar sulcus (
<figureCitation box="[568,649,727,753]" captionStart="FIGURES 11–20" captionStartId="6.[151,264,1899,1923]" captionTargetBox="[219,1368,181,1875]" captionTargetId="figure-19@6.[219,1368,181,1875]" captionTargetPageId="6" captionText="FIGURES 11–20. Ceropales (Ceropales) keralaensis Anju, Binoy &amp; Thejass, sp. nov., holotype, ♀. 11, habitus, lateral view; 12, habitus, dorsal view; 13, head, lateral view; 14, head, frontal view; 15, head, dorsal view; 16, mesosoma, dorsal view; 17, mesosoma, lateral view; 18, wings; 19, metasoma, lateral view; 20, metasoma, dorsal view. Scale bar: 11–2 mm; 12, 13, 17, 18, 20–1 mm; 14, 15, 16, 19–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836284" httpUri="https://zenodo.org/record/7836284/files/figure.png" pageId="7" pageNumber="126">Fig. 16</figureCitation>
); middle portion raised in profile; punctuations clearly visible in middle portion than on lateral regions (
<figureCitation box="[501,583,763,789]" captionStart="FIGURES 11–20" captionStartId="6.[151,264,1899,1923]" captionTargetBox="[219,1368,181,1875]" captionTargetId="figure-19@6.[219,1368,181,1875]" captionTargetPageId="6" captionText="FIGURES 11–20. Ceropales (Ceropales) keralaensis Anju, Binoy &amp; Thejass, sp. nov., holotype, ♀. 11, habitus, lateral view; 12, habitus, dorsal view; 13, head, lateral view; 14, head, frontal view; 15, head, dorsal view; 16, mesosoma, dorsal view; 17, mesosoma, lateral view; 18, wings; 19, metasoma, lateral view; 20, metasoma, dorsal view. Scale bar: 11–2 mm; 12, 13, 17, 18, 20–1 mm; 14, 15, 16, 19–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836284" httpUri="https://zenodo.org/record/7836284/files/figure.png" pageId="7" pageNumber="126">Fig. 16</figureCitation>
); mesopleuron with scattered punctures (
<figureCitation box="[1040,1123,763,789]" captionStart="FIGURES 11–20" captionStartId="6.[151,264,1899,1923]" captionTargetBox="[219,1368,181,1875]" captionTargetId="figure-19@6.[219,1368,181,1875]" captionTargetPageId="6" captionText="FIGURES 11–20. Ceropales (Ceropales) keralaensis Anju, Binoy &amp; Thejass, sp. nov., holotype, ♀. 11, habitus, lateral view; 12, habitus, dorsal view; 13, head, lateral view; 14, head, frontal view; 15, head, dorsal view; 16, mesosoma, dorsal view; 17, mesosoma, lateral view; 18, wings; 19, metasoma, lateral view; 20, metasoma, dorsal view. Scale bar: 11–2 mm; 12, 13, 17, 18, 20–1 mm; 14, 15, 16, 19–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836284" httpUri="https://zenodo.org/record/7836284/files/figure.png" pageId="7" pageNumber="126">Fig. 17</figureCitation>
); metanotum conspicuously elevated and convex in profile (
<figureCitation box="[513,598,799,825]" captionStart="FIGURES 11–20" captionStartId="6.[151,264,1899,1923]" captionTargetBox="[219,1368,181,1875]" captionTargetId="figure-19@6.[219,1368,181,1875]" captionTargetPageId="6" captionText="FIGURES 11–20. Ceropales (Ceropales) keralaensis Anju, Binoy &amp; Thejass, sp. nov., holotype, ♀. 11, habitus, lateral view; 12, habitus, dorsal view; 13, head, lateral view; 14, head, frontal view; 15, head, dorsal view; 16, mesosoma, dorsal view; 17, mesosoma, lateral view; 18, wings; 19, metasoma, lateral view; 20, metasoma, dorsal view. Scale bar: 11–2 mm; 12, 13, 17, 18, 20–1 mm; 14, 15, 16, 19–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836284" httpUri="https://zenodo.org/record/7836284/files/figure.png" pageId="7" pageNumber="126">Fig. 17</figureCitation>
); surface moderately micropunctured; metapostnotum very much reduced and narrow with few transverse rugae; metapleuron with fine, sparse punctures; propodeum medium sized, 1.21× as wide as long along the middle, conspicuously flat (
<figureCitation box="[724,808,871,897]" captionStart="FIGURES 11–20" captionStartId="6.[151,264,1899,1923]" captionTargetBox="[219,1368,181,1875]" captionTargetId="figure-19@6.[219,1368,181,1875]" captionTargetPageId="6" captionText="FIGURES 11–20. Ceropales (Ceropales) keralaensis Anju, Binoy &amp; Thejass, sp. nov., holotype, ♀. 11, habitus, lateral view; 12, habitus, dorsal view; 13, head, lateral view; 14, head, frontal view; 15, head, dorsal view; 16, mesosoma, dorsal view; 17, mesosoma, lateral view; 18, wings; 19, metasoma, lateral view; 20, metasoma, dorsal view. Scale bar: 11–2 mm; 12, 13, 17, 18, 20–1 mm; 14, 15, 16, 19–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836284" httpUri="https://zenodo.org/record/7836284/files/figure.png" pageId="7" pageNumber="126">Fig. 17</figureCitation>
); surface with microscopically fine, transverse rugulose sculpturing (
<figureCitation box="[295,380,907,933]" captionStart="FIGURES 11–20" captionStartId="6.[151,264,1899,1923]" captionTargetBox="[219,1368,181,1875]" captionTargetId="figure-19@6.[219,1368,181,1875]" captionTargetPageId="6" captionText="FIGURES 11–20. Ceropales (Ceropales) keralaensis Anju, Binoy &amp; Thejass, sp. nov., holotype, ♀. 11, habitus, lateral view; 12, habitus, dorsal view; 13, head, lateral view; 14, head, frontal view; 15, head, dorsal view; 16, mesosoma, dorsal view; 17, mesosoma, lateral view; 18, wings; 19, metasoma, lateral view; 20, metasoma, dorsal view. Scale bar: 11–2 mm; 12, 13, 17, 18, 20–1 mm; 14, 15, 16, 19–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836284" httpUri="https://zenodo.org/record/7836284/files/figure.png" pageId="7" pageNumber="126">Fig. 16</figureCitation>
); median sulcus reaching nearly half length of propodeum; posterior margin nearly straight, posterolateral angle rounded; lateral sides with shallow impression medially.
</paragraph>
<paragraph blockId="7.[151,1437,151,1293]" pageId="7" pageNumber="126">
<emphasis bold="true" box="[199,261,979,1005]" pageId="7" pageNumber="126">
<emphasis bold="true" box="[199,253,979,1005]" italics="true" pageId="7" pageNumber="126">Legs</emphasis>
.
</emphasis>
Tarsal claws normal, bent apically; short, erect and acute subapical tooth present; hind tibial spur long, 0.7× hind basitarsus.
</paragraph>
<paragraph blockId="7.[151,1437,151,1293]" pageId="7" pageNumber="126">
<emphasis bold="true" box="[199,278,1051,1077]" pageId="7" pageNumber="126">
<emphasis bold="true" box="[199,271,1051,1077]" italics="true" pageId="7" pageNumber="126">Wings</emphasis>
.
</emphasis>
Fore wing with upper margin of SMC2 and SMC3 equal in length, lower margin of SMC3 longer than that of SMC2; SMC2 receiving crossvein
<emphasis box="[636,707,1087,1113]" italics="true" pageId="7" pageNumber="126">1m-cu</emphasis>
medially, SMC3 receiving crossvein
<emphasis box="[1139,1209,1087,1113]" italics="true" pageId="7" pageNumber="126">2m-cu</emphasis>
medially; length of SMC3 less than its distance from the wing apex; vein
<emphasis box="[756,773,1123,1149]" italics="true" pageId="7" pageNumber="126">A</emphasis>
and crossvein
<emphasis box="[940,990,1123,1148]" italics="true" pageId="7" pageNumber="126">cu-a</emphasis>
of hind wing forming obtuse angle.
</paragraph>
<paragraph blockId="7.[151,1437,151,1293]" pageId="7" pageNumber="126">
<emphasis bold="true" box="[199,326,1159,1185]" pageId="7" pageNumber="126">
<emphasis bold="true" box="[199,319,1159,1185]" italics="true" pageId="7" pageNumber="126">Metasoma</emphasis>
.
</emphasis>
Fusiform; as broad and long as mesosoma; smooth (
<figureCitation box="[954,1041,1159,1185]" captionStart="FIGURES 11–20" captionStartId="6.[151,264,1899,1923]" captionTargetBox="[219,1368,181,1875]" captionTargetId="figure-19@6.[219,1368,181,1875]" captionTargetPageId="6" captionText="FIGURES 11–20. Ceropales (Ceropales) keralaensis Anju, Binoy &amp; Thejass, sp. nov., holotype, ♀. 11, habitus, lateral view; 12, habitus, dorsal view; 13, head, lateral view; 14, head, frontal view; 15, head, dorsal view; 16, mesosoma, dorsal view; 17, mesosoma, lateral view; 18, wings; 19, metasoma, lateral view; 20, metasoma, dorsal view. Scale bar: 11–2 mm; 12, 13, 17, 18, 20–1 mm; 14, 15, 16, 19–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836284" httpUri="https://zenodo.org/record/7836284/files/figure.png" pageId="7" pageNumber="126">Fig. 19</figureCitation>
); T1 sessile, parallel sided; lower margin of S6 flat in lateral view, pointed apically.
</paragraph>
<paragraph blockId="7.[151,1437,151,1293]" box="[199,419,1230,1257]" pageId="7" pageNumber="126">
<emphasis bold="true" box="[199,419,1230,1257]" pageId="7" pageNumber="126">
Male. (
<figureCitation box="[281,412,1230,1256]" captionStart="FIGURES 21–23" captionStartId="7.[151,264,1969,1993]" captionTargetBox="[158,1430,1317,1946]" captionTargetId="figure-486@7.[158,1430,1317,1946]" captionTargetPageId="7" captionText="FIGURES 21–23. Ceropales (Ceropales) keralaensis Anju, Binoy &amp; Thejass, sp. nov., paratype, J. 21, habitus, lateral view; 22, hypopygium; 23, genitalia, dorsal view. Scale bar: 21–2 mm; 22, 23–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836288" httpUri="https://zenodo.org/record/7836288/files/figure.png" pageId="7" pageNumber="126">Figs 21–23</figureCitation>
)
</emphasis>
</paragraph>
<paragraph blockId="7.[151,1437,151,1293]" box="[199,1147,1267,1293]" pageId="7" pageNumber="126">
Resembles female in most characters. Hypopygium and genitalia as in
<figureCitation box="[993,1143,1267,1293]" captionStart="FIGURES 21–23" captionStartId="7.[151,264,1969,1993]" captionTargetBox="[158,1430,1317,1946]" captionTargetId="figure-486@7.[158,1430,1317,1946]" captionTargetPageId="7" captionText="FIGURES 21–23. Ceropales (Ceropales) keralaensis Anju, Binoy &amp; Thejass, sp. nov., paratype, J. 21, habitus, lateral view; 22, hypopygium; 23, genitalia, dorsal view. Scale bar: 21–2 mm; 22, 23–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836288" httpUri="https://zenodo.org/record/7836288/files/figure.png" pageId="7" pageNumber="126">Figs 22 &amp; 23</figureCitation>
.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7836288" ID-Zenodo-Dep="7836288" httpUri="https://zenodo.org/record/7836288/files/figure.png" pageId="7" pageNumber="126" startId="7.[151,264,1969,1993]" targetBox="[158,1430,1317,1946]" targetPageId="7">
<paragraph blockId="7.[151,1436,1969,2030]" pageId="7" pageNumber="126">
<emphasis bold="true" box="[151,341,1969,1993]" pageId="7" pageNumber="126">FIGURES 21–23.</emphasis>
<taxonomicName authority="Anju, Binoy &amp; Thejass" authorityName="Anju, Binoy &amp; Thejass" authorityYear="2023" box="[348,945,1970,1994]" class="Insecta" family="Pompilidae" genus="Ceropales" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="126" phylum="Arthropoda" rank="species" species="keralaensis" status="sp. nov." subGenus="Ceropales">
<emphasis box="[348,451,1970,1994]" italics="true" pageId="7" pageNumber="126">Ceropales</emphasis>
(
<emphasis box="[466,569,1970,1994]" italics="true" pageId="7" pageNumber="126">Ceropales</emphasis>
)
<emphasis box="[585,700,1970,1994]" italics="true" pageId="7" pageNumber="126">keralaensis</emphasis>
Anju, Binoy &amp; Thejass
</taxonomicName>
,
<emphasis bold="true" box="[956,1037,1970,1994]" pageId="7" pageNumber="126">
<taxonomicNameLabel box="[956,1037,1970,1994]" pageId="7" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype, J. 21, habitus, lateral view; 22, hypopygium; 23, genitalia, dorsal view. Scale bar: 21–2 mm; 22, 23–0.5 mm.
</paragraph>
</caption>
<subSubSection box="[199,1324,151,177]" pageId="8" pageNumber="127" type="etymology">
<paragraph blockId="8.[151,1437,151,465]" box="[199,1324,151,177]" pageId="8" pageNumber="127">
<emphasis bold="true" box="[199,334,151,177]" pageId="8" pageNumber="127">Etymology.</emphasis>
The species name refers to the Indian state of
<collectingRegion box="[855,932,151,177]" country="India" name="Kerala" pageId="8" pageNumber="127">Kerala</collectingRegion>
, where the
<typeStatus box="[1060,1158,151,177]" pageId="8" pageNumber="127">holotype</typeStatus>
was collected.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="127" type="description">
<paragraph blockId="8.[151,1437,151,465]" pageId="8" pageNumber="127">
<emphasis bold="true" box="[199,319,186,212]" pageId="8" pageNumber="127">Variation.</emphasis>
Relative density of pubescence varies from moderately to densely setose in
<typeStatus box="[1199,1306,187,213]" pageId="8" pageNumber="127">paratypes</typeStatus>
along with variations in intensity of body coloration.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="127" type="distribution">
<paragraph blockId="8.[151,1437,151,465]" box="[199,800,258,285]" pageId="8" pageNumber="127">
<emphasis bold="true" box="[199,353,258,284]" pageId="8" pageNumber="127">Distribution.</emphasis>
<collectingCountry box="[359,417,259,285]" name="India" pageId="8" pageNumber="127">India</collectingCountry>
(
<collectingRegion box="[432,511,259,285]" country="India" name="Kerala" pageId="8" pageNumber="127">Kerala</collectingRegion>
,
<collectingRegion box="[522,640,259,285]" country="India" name="Karnataka" pageId="8" pageNumber="127">Karnataka</collectingRegion>
,
<collectingRegion box="[650,788,259,285]" country="India" name="Tamil Nadu" pageId="8" pageNumber="127">Tamil Nadu</collectingRegion>
).
</paragraph>
<paragraph blockId="8.[151,1437,151,465]" pageId="8" pageNumber="127">
<emphasis bold="true" box="[199,431,294,320]" pageId="8" pageNumber="127">Sex differentiation.</emphasis>
Females of
<taxonomicName authority="Anju &amp; Thejass &amp; Binoy &amp; Kumar, 2023" authorityName="Anju &amp; Thejass &amp; Binoy &amp; Kumar" authorityYear="2023" box="[578,742,295,321]" class="Insecta" family="Pompilidae" genus="Ceropales" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="species" species="keralaensis" status="sp. nov.">
<emphasis box="[578,742,295,321]" italics="true" pageId="8" pageNumber="127">C. keralaensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[752,844,295,321]" pageId="8" pageNumber="127">
<taxonomicNameLabel box="[752,844,295,321]" pageId="8" pageNumber="127" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resemble males in almost all characters except for the typical genital and antennal sex differentiation. Additionally, females are distinguishable by the presence of bicoloured frons (medial area black flanked by yellow on either sides) whereas, in males, the medial part of the frons is pale yellow; in females, the mesoscutellum is completely black, while in males, the mesoscutellum is black with a pale-yellow spot.
</paragraph>
</subSubSection>
</treatment>
</document>
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