<documentid="CA47A8BD2AC1B5CD7CDD8CF1D8B04E8B"ID-DOI="10.11646/zootaxa.5560.1.1"ID-ISSN="1175-5334"ID-Zenodo-Dep="14595436"ID-ZooBank="0F942970-010E-4775-856E-31CA016DAD50"IM.bibliography_approvedBy="felipe"IM.illustrations_approvedBy="felipe"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="felipe"IM.tables_requiresApprovalFor="GgImagineBatch"IM.taxonomicNames_approvedBy="felipe"IM.treatmentCitations_approvedBy="felipe"IM.treatments_approvedBy="felipe"checkinTime="1735931741530"checkinUser="plazi"docAuthor="Haddad, Charles Richard, Wiśniewski, Konrad & Wesołowska, Wanda"docDate="2024"docId="03FF87E70C350077DAE3857BB2FDF8EA"docLanguage="en"docName="zootaxa.5560.1.1.pdf"docOrigin="Zootaxa 5560 (1)"docSource="https://doi.org/10.11646/zootaxa.5560.1.1"docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph"docStyleId="5EBBA59367AD13919D70D935FA04F6A3"docStyleName="Zootaxa.2013-.monograph"docStyleVersion="19"docTitle="Hyllus ornatus Haddad, Wiśniewski & Wesołowska, 2024, sp. nov."docType="treatment"docVersion="2"lastPageNumber="30"masterDocId="FFC6FF9F0C29006ADA748055B129FF90"masterDocTitle="The jumping spiders of Mozambique (Araneae: Salticidae)"masterLastPageNumber="92"masterPageNumber="1"pageNumber="29"updateTime="1735934017513"updateUser="ExternalLinkService">
<mods:affiliationid="3D9C80CC513A96EC0A37DBB856B42A3A">Department of Zoology & Entomology, University of the Free State, P. O. Box 339, Bloemfontein 9300, South Africa & Department of Biodiversity and Evolutionary Taxonomy, University of Wrocław, Przybyszewskiego 65, 51 - 148 Wrocław, Poland & Department of Zoology & Entomology, University of the Free State, P. O. Box 339, Bloemfontein 9300, South Africa & wanda. h. wesolowska @ gmail. com; https: // orcid. org / 0000 - 0002 - 4411 - 1058 * Corresponding author</mods:affiliation>
<taxonomicNameid="4C564D720C350076DAE3857BB062FAD9"authorityName="Haddad & Wiśniewski & Wesołowska"authorityYear="2024"box="[151,331,1326,1353]"class="Arachnida"family="Salticidae"genus="Hyllus"kingdom="Animalia"order="Araneae"pageId="28"pageNumber="29"phylum="Arthropoda"rank="species"species="ornatus"status="sp. nov.">
<figureCitationid="136D2A740C350076DAE38506B036FAFD"box="[151,287,1363,1389]"captionStart="FIGURE 18"captionStartId="29.[151,250,1529,1553]"captionTargetBox="[227,1361,274,1503]"captionTargetId="figure-42@29.[227,1361,270,1503]"captionTargetPageId="29"captionText="FIGURE 18. Hyllus lwoffi Berland & Millot, 1941, female (A), and H. ornatus sp. nov., male holotype and female paratype (B–E). A, B. General appearance, dorsal view; C. Palpal organ, ventral view; D. Same, lateral view; E. Epigyne."figureDoi="http://doi.org/10.5281/zenodo.14595476"httpUri="https://zenodo.org/record/14595476/files/figure.png"pageId="28"pageNumber="29">Figs 18B–E</figureCitation>
<figureCitationid="136D2A740C350076DB5E8506B06FFAFD"box="[298,326,1363,1389]"captionStart="FIGURE 19"captionStartId="30.[151,250,1440,1464]"captionTargetBox="[177,1392,186,1400]"captionTargetId="figure-21@30.[151,1436,176,1414]"captionTargetPageId="30"captionText="FIGURE 19. Hyllus ornatus sp. nov., male holotype (A–C) and female paratype (D, E). A. Chelicera of male; B. Palpal organ, ventral view; C. Same, lateral view; D. Epigyne; E. Internal structure of epigyne."figureDoi="http://doi.org/10.5281/zenodo.14595480"httpUri="https://zenodo.org/record/14595480/files/figure.png"pageId="28"pageNumber="29">19</figureCitation>
<figureCitationid="136D2A740C350076D91085CEB2E2FA25"box="[868,971,1435,1461]"captionStart="FIGURE 18"captionStartId="29.[151,250,1529,1553]"captionTargetBox="[227,1361,274,1503]"captionTargetId="figure-42@29.[227,1361,270,1503]"captionTargetPageId="29"captionText="FIGURE 18. Hyllus lwoffi Berland & Millot, 1941, female (A), and H. ornatus sp. nov., male holotype and female paratype (B–E). A, B. General appearance, dorsal view; C. Palpal organ, ventral view; D. Same, lateral view; E. Epigyne."figureDoi="http://doi.org/10.5281/zenodo.14595476"httpUri="https://zenodo.org/record/14595476/files/figure.png"pageId="28"pageNumber="29">Fig. 18B</figureCitation>
<taxonomicNameid="4C564D720C350076DA9C85EAB3FAFA49"authority="Wanless & Clark, 1975"authorityName="Wanless & Clark"authorityYear="1975"box="[232,723,1470,1497]"class="Arachnida"family="Salticidae"genus="Hyllus"kingdom="Animalia"order="Araneae"pageId="28"pageNumber="29"phylum="Arthropoda"rank="species"species="tuberculatus">
<bibRefCitationid="EFC74B000C350076DBB885EAB3FAFA49"author="Wanless, F. R. & Clark, D. J."box="[460,723,1470,1497]"pageId="28"pageNumber="29"pagination="273 - 296"refId="ref48329"refString="Wanless, F. R. & Clark, D. J. (1975) On a collection of spiders of the family Salticidae from the Ivory Coast (Araneae). Revue Zoologique Africaine, 89, 273 - 296."type="journal article"year="1975">Wanless & Clark, 1975</bibRefCitation>
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but can be recognized by the clearly longer embolus (compare
<figureCitationid="136D2A740C350076DAE385B6B1D0FA6D"box="[151,249,1507,1533]"captionStart="FIGURE 18"captionStartId="29.[151,250,1529,1553]"captionTargetBox="[227,1361,274,1503]"captionTargetId="figure-42@29.[227,1361,270,1503]"captionTargetPageId="29"captionText="FIGURE 18. Hyllus lwoffi Berland & Millot, 1941, female (A), and H. ornatus sp. nov., male holotype and female paratype (B–E). A, B. General appearance, dorsal view; C. Palpal organ, ventral view; D. Same, lateral view; E. Epigyne."figureDoi="http://doi.org/10.5281/zenodo.14595476"httpUri="https://zenodo.org/record/14595476/files/figure.png"pageId="28"pageNumber="29">Fig. 18C</figureCitation>
<figureCitationid="136D2A740C350076DB5985B6B075FA6D"box="[301,348,1507,1533]"captionStart="FIGURE 19"captionStartId="30.[151,250,1440,1464]"captionTargetBox="[177,1392,186,1400]"captionTargetId="figure-21@30.[151,1436,176,1414]"captionTargetPageId="30"captionText="FIGURE 19. Hyllus ornatus sp. nov., male holotype (A–C) and female paratype (D, E). A. Chelicera of male; B. Palpal organ, ventral view; C. Same, lateral view; D. Epigyne; E. Internal structure of epigyne."figureDoi="http://doi.org/10.5281/zenodo.14595480"httpUri="https://zenodo.org/record/14595480/files/figure.png"pageId="28"pageNumber="29">19B</figureCitation>
<bibRefCitationid="EFC74B000C350076D86285B6B28EFA6D"author="Wesolowska, W. & Russell-Smith, A."box="[534,935,1507,1533]"pageId="28"pageNumber="29"pagination="1 - 143"refId="ref49851"refString="Wesolowska, W. & Russell-Smith, A. (2022) Jumping spiders from Ivory Coast collected by J. - C. Ledoux (Araneae, Salticidae). European Journal of Taxonomy, 841, 1 - 143. https: // doi. org / 10.5852 / ejt. 2022.841.1943"type="journal article"year="2022">Wesołowska & Russell-Smith 2022</bibRefCitation>
). The female is distinctive in having a unique form of the epigyne, which is triangular with two strongly sclerotized flaps in the centre (
<figureCitationid="136D2A740C350076DEF48652B5CCF9B1"box="[1152,1253,1543,1569]"captionStart="FIGURE 18"captionStartId="29.[151,250,1529,1553]"captionTargetBox="[227,1361,274,1503]"captionTargetId="figure-42@29.[227,1361,270,1503]"captionTargetPageId="29"captionText="FIGURE 18. Hyllus lwoffi Berland & Millot, 1941, female (A), and H. ornatus sp. nov., male holotype and female paratype (B–E). A, B. General appearance, dorsal view; C. Palpal organ, ventral view; D. Same, lateral view; E. Epigyne."figureDoi="http://doi.org/10.5281/zenodo.14595476"httpUri="https://zenodo.org/record/14595476/files/figure.png"pageId="28"pageNumber="29">Fig. 18E</figureCitation>
<materialsCitationid="3B3E3CAC0C350076DB08861AB5AFF9F9"box="[380,1158,1614,1641]"collectionCode="SMF"collectorName="C. F. Roewer"country="Mozambique"location="Tete"pageId="28"pageNumber="29"specimenCode="SMF 9699"specimenCount="1"specimenCount-male="1"stateProvince="Tete"typeStatus="holotype">
<specimenCodeid="DBF09E8A0C350076D988861AB553F9F9"box="[1020,1146,1615,1641]"collectionCode="SMF"country="Germany"httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838"lsid="urn:lsid:biocol.org:col:34838"name="Forschungsinstitut und Natur-Museum Senckenberg"pageId="28"pageNumber="29">SMF 9699</specimenCode>
<specimenCodeid="DBF09E8A0C350076D9268626B2F9F91D"box="[850,976,1651,1677]"collectionCode="SMF"country="Germany"httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838"lsid="urn:lsid:biocol.org:col:34838"name="Forschungsinstitut und Natur-Museum Senckenberg"pageId="28"pageNumber="29">SMF 9682</specimenCode>
<figureCitationid="136D2A740C350076DEB086EEB400F945"box="[1220,1321,1723,1749]"captionStart="FIGURE 18"captionStartId="29.[151,250,1529,1553]"captionTargetBox="[227,1361,274,1503]"captionTargetId="figure-42@29.[227,1361,270,1503]"captionTargetPageId="29"captionText="FIGURE 18. Hyllus lwoffi Berland & Millot, 1941, female (A), and H. ornatus sp. nov., male holotype and female paratype (B–E). A, B. General appearance, dorsal view; C. Palpal organ, ventral view; D. Same, lateral view; E. Epigyne."figureDoi="http://doi.org/10.5281/zenodo.14595476"httpUri="https://zenodo.org/record/14595476/files/figure.png"pageId="28"pageNumber="29">Fig. 18B</figureCitation>
. Carapace oval, brown, lighter in centre and darkening to margins, vicinity of eyes black. White patch formed by white hairs at fovea, broad white streaks along lateral margins of carapace. Very long white hairs on lower edge of clypeus. Chelicerae dark brown, with single large tooth on retromargin (
<figureCitationid="136D2A740C350076D9238772B296F8D1"box="[855,959,1831,1857]"captionStart="FIGURE 19"captionStartId="30.[151,250,1440,1464]"captionTargetBox="[177,1392,186,1400]"captionTargetId="figure-21@30.[151,1436,176,1414]"captionTargetPageId="30"captionText="FIGURE 19. Hyllus ornatus sp. nov., male holotype (A–C) and female paratype (D, E). A. Chelicera of male; B. Palpal organ, ventral view; C. Same, lateral view; D. Epigyne; E. Internal structure of epigyne."figureDoi="http://doi.org/10.5281/zenodo.14595480"httpUri="https://zenodo.org/record/14595480/files/figure.png"pageId="28"pageNumber="29">Fig. 19A</figureCitation>
), white hairs covering promarginal surface of chelicerae. Sternum light brown, labium and endites dark brown with white tips.Abdomen ovoid, with contrasting pattern; background dark brown, with transverse streak composed of white hairs near anterior edge, large yellow patch on posterior half, posterior tip white (
<figureCitationid="136D2A740C350076D8E487C6B3D1F83D"box="[656,760,1939,1965]"captionStart="FIGURE 18"captionStartId="29.[151,250,1529,1553]"captionTargetBox="[227,1361,274,1503]"captionTargetId="figure-42@29.[227,1361,270,1503]"captionTargetPageId="29"captionText="FIGURE 18. Hyllus lwoffi Berland & Millot, 1941, female (A), and H. ornatus sp. nov., male holotype and female paratype (B–E). A, B. General appearance, dorsal view; C. Palpal organ, ventral view; D. Same, lateral view; E. Epigyne."figureDoi="http://doi.org/10.5281/zenodo.14595476"httpUri="https://zenodo.org/record/14595476/files/figure.png"pageId="28"pageNumber="29">Fig. 18B</figureCitation>
). Sparse long brown bristles on abdominal dorsum. Venter with wide grey streak. Spinnerets light brown. Legs brownish, two anterior pairs dark brown, larger than others. Spines long, leg hairs brown and colourless. Distal ends of tibiae blackish. Palp light brown, clothed in long brown and white hairs. Structure of palpal organ in
<figureCitationid="136D2A740C340077D8DD80C2B215FF21"box="[681,828,151,177]"captionStart="FIGURE 18"captionStartId="29.[151,250,1529,1553]"captionTargetBox="[227,1361,274,1503]"captionTargetId="figure-42@29.[227,1361,270,1503]"captionTargetPageId="29"captionText="FIGURE 18. Hyllus lwoffi Berland & Millot, 1941, female (A), and H. ornatus sp. nov., male holotype and female paratype (B–E). A, B. General appearance, dorsal view; C. Palpal organ, ventral view; D. Same, lateral view; E. Epigyne."figureDoi="http://doi.org/10.5281/zenodo.14595476"httpUri="https://zenodo.org/record/14595476/files/figure.png"pageId="29"pageNumber="30">Figs 18C, D</figureCitation>
<figureCitationid="136D2A740C340077D93F80C2B288FF21"box="[843,929,151,177]"captionStart="FIGURE 19"captionStartId="30.[151,250,1440,1464]"captionTargetBox="[177,1392,186,1400]"captionTargetId="figure-21@30.[151,1436,176,1414]"captionTargetPageId="30"captionText="FIGURE 19. Hyllus ornatus sp. nov., male holotype (A–C) and female paratype (D, E). A. Chelicera of male; B. Palpal organ, ventral view; C. Same, lateral view; D. Epigyne; E. Internal structure of epigyne."figureDoi="http://doi.org/10.5281/zenodo.14595480"httpUri="https://zenodo.org/record/14595480/files/figure.png"pageId="29"pageNumber="30">19B, C</figureCitation>
<figureCitationid="136D2A740C340077DB4480EEB0BEFF45"box="[304,407,187,213]"captionStart="FIGURE 18"captionStartId="29.[151,250,1529,1553]"captionTargetBox="[227,1361,274,1503]"captionTargetId="figure-42@29.[227,1361,270,1503]"captionTargetPageId="29"captionText="FIGURE 18. Hyllus lwoffi Berland & Millot, 1941, female (A), and H. ornatus sp. nov., male holotype and female paratype (B–E). A, B. General appearance, dorsal view; C. Palpal organ, ventral view; D. Same, lateral view; E. Epigyne."figureDoi="http://doi.org/10.5281/zenodo.14595476"httpUri="https://zenodo.org/record/14595476/files/figure.png"pageId="29"pageNumber="30">Fig. 18C</figureCitation>
<taxonomicNameid="4C564D720C340077DB5D85AFB380F982"authority="Berland & Millot, 1941"authorityName="Berland & Millot"authorityYear="1941"box="[297,681,1530,1554]"class="Arachnida"family="Salticidae"genus="Hyllus"kingdom="Animalia"order="Araneae"pageId="29"pageNumber="30"phylum="Arthropoda"rank="species"species="lwoffi">
<bibRefCitationid="EFC74B000C340077DBC085AFB380F982"author="Berland, L. & Millot, J."box="[436,681,1530,1554]"pageId="29"pageNumber="30"pagination="297 - 423"refId="ref44514"refString="Berland, L. & Millot, J. (1941) Les araignees de l'Afrique Occidentale Francaise I. - Les salticides. Memoires du Museum National d'Histoire Naturelle de Paris (N. S.), 12, 297 - 423."type="journal article"year="1941">Berland & Millot, 1941</bibRefCitation>
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, female (A), and
<taxonomicNameid="4C564D720C340077D92A85AFB2E2F981"authorityName="Haddad & Wiśniewski & Wesołowska"authorityYear="2024"box="[862,971,1530,1553]"class="Arachnida"family="Salticidae"genus="Hyllus"kingdom="Animalia"order="Araneae"pageId="29"pageNumber="30"phylum="Arthropoda"rank="species"species="ornatus"status="sp. nov.">
Measurements: Cephalothorax: length 5.3, width 4.4, height 2.3. Eye field: length 2.3, anterior and posterior width 3.5. Abdomen: length 5.6, width 3.9. Shape of body and colouration as in male. Some white hairs between anterior eyes and on frontal surface of chelicerae. Epigyne triangular, two small semicircular depressions in centre with strongly sclerotized flaps above (
<figureCitationid="136D2A740C340077D8FB874DB3D1F8A2"box="[655,760,1816,1842]"captionStart="FIGURE 18"captionStartId="29.[151,250,1529,1553]"captionTargetBox="[227,1361,274,1503]"captionTargetId="figure-42@29.[227,1361,270,1503]"captionTargetPageId="29"captionText="FIGURE 18. Hyllus lwoffi Berland & Millot, 1941, female (A), and H. ornatus sp. nov., male holotype and female paratype (B–E). A, B. General appearance, dorsal view; C. Palpal organ, ventral view; D. Same, lateral view; E. Epigyne."figureDoi="http://doi.org/10.5281/zenodo.14595476"httpUri="https://zenodo.org/record/14595476/files/figure.png"pageId="29"pageNumber="30">Figs 18E</figureCitation>
<figureCitationid="136D2A740C340077D970874DB211F8A2"box="[772,824,1816,1842]"captionStart="FIGURE 19"captionStartId="30.[151,250,1440,1464]"captionTargetBox="[177,1392,186,1400]"captionTargetId="figure-21@30.[151,1436,176,1414]"captionTargetPageId="30"captionText="FIGURE 19. Hyllus ornatus sp. nov., male holotype (A–C) and female paratype (D, E). A. Chelicera of male; B. Palpal organ, ventral view; C. Same, lateral view; D. Epigyne; E. Internal structure of epigyne."figureDoi="http://doi.org/10.5281/zenodo.14595480"httpUri="https://zenodo.org/record/14595480/files/figure.png"pageId="29"pageNumber="30">19D</figureCitation>
<figureCitationid="136D2A740C340077DE3B874DB59BF8A2"box="[1103,1202,1816,1842]"captionStart="FIGURE 19"captionStartId="30.[151,250,1440,1464]"captionTargetBox="[177,1392,186,1400]"captionTargetId="figure-21@30.[151,1436,176,1414]"captionTargetPageId="30"captionText="FIGURE 19. Hyllus ornatus sp. nov., male holotype (A–C) and female paratype (D, E). A. Chelicera of male; B. Palpal organ, ventral view; C. Same, lateral view; D. Epigyne; E. Internal structure of epigyne."figureDoi="http://doi.org/10.5281/zenodo.14595480"httpUri="https://zenodo.org/record/14595480/files/figure.png"pageId="29"pageNumber="30">Fig. 19E</figureCitation>