treatments-xml/data/03/F8/87/03F887C28765FFBEFF05FC1898E8FE10.xml

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<document id="9FE3CD94970BD5AAEB1AB5EE244E0844" ID="10.11646/zootaxa.4247.2.5SLASH10635" ID-CLB-Dataset="36194" ID-DOI="10.11646/zootaxa.4247.2.5" ID-GBIF-Dataset="fbd48589-66ea-48b1-a4c7-87843a599066" ID-ISSN="1175-5326" ID-Zenodo-Dep="438232" ID-ZooBank="C736CA96-99A5-40D2-B944-A50410552E37" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1490600583814" checkinUser="plazi" docAuthor="Kosterin, Oleg E., Akimbekova, Nazymgul, Dubatolov, Vladimir V. &amp; Sivec, Ignac" docDate="2017" docId="03F887C28765FFBEFF05FC1898E8FE10" docLanguage="en" docName="zootaxa.4247.2.5SLASH10635.pdf" docOrigin="Zootaxa 4247 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Taeniopteryx araneoides Klapalek 1902" docType="treatment" docVersion="8" lastPageNumber="148" masterDocId="FFC1FFBA8764FFB9FF92FFCA9C63FFDE" masterDocTitle="A stonefly species extinct in Europe (Taeniopteryx araneoides Klapalek, 1902, Taeniopterygidae, Plecoptera) is thriving in the Irtysh River in West Siberia and North Kazakhstan" masterLastPageNumber="148" masterPageNumber="141" pageNumber="142" updateTime="1698438416906" updateUser="ExternalLinkService">
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<mods:title id="2CE720C4063F65F9C486AACC71EF233C">A stonefly species extinct in Europe (Taeniopteryx araneoides Klapalek, 1902, Taeniopterygidae, Plecoptera) is thriving in the Irtysh River in West Siberia and North Kazakhstan</mods:title>
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<mods:namePart id="4B6ECB919A6F5ADE2D9D52767D015678">Kosterin, Oleg E.</mods:namePart>
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<mods:namePart id="33BE66DA5916971A0BF19DB320AA640B">Akimbekova, Nazymgul</mods:namePart>
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<mods:namePart id="D1976D097ED55C2264905A5EAD4917C1">Dubatolov, Vladimir V.</mods:namePart>
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<mods:namePart id="30A5BEBBF66959096BBB27C9387B4A1C">Sivec, Ignac</mods:namePart>
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<treatment id="03F887C28765FFBEFF05FC1898E8FE10" ID-DOI="http://doi.org/10.5281/zenodo.6053065" ID-GBIF-Taxon="127852517" ID-Zenodo-Dep="6053065" LSID="urn:lsid:plazi:treatment:03F887C28765FFBEFF05FC1898E8FE10" httpUri="http://treatment.plazi.org/id/03F887C28765FFBEFF05FC1898E8FE10" lastPageId="7" lastPageNumber="148" pageId="1" pageNumber="142">
<subSubSection id="C34B655F8765FFB8FF05FC189D63FBD0" pageId="1" pageNumber="142" type="nomenclature">
<paragraph id="8BEE36D48765FFB8FF05FC189EE1FC32" blockId="1.[151,642,978,1038]" box="[151,642,978,1004]" pageId="1" pageNumber="142">
<heading id="D0A681B88765FFB8FF05FC189EE1FC32" bold="true" box="[151,642,978,1004]" fontSize="11" level="1" pageId="1" pageNumber="142" reason="1">
<taxonomicName id="4C514D578765FFB8FF05FC189EE1FC32" ID-CoL="54HPS" authority="Klapalek, 1902" authorityName="Klapalek" authorityYear="1902" box="[151,642,978,1004]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="araneoides">
<emphasis id="B925EAC68765FFB8FF05FC189EE1FC32" bold="true" box="[151,642,978,1004]" pageId="1" pageNumber="142">
<emphasis id="B925EAC68765FFB8FF05FC189DDCFC32" bold="true" box="[151,447,978,1004]" italics="true" pageId="1" pageNumber="142">Taeniopteryx araneoides</emphasis>
Klápalek, 1902
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BEE36D48765FFB8FF05FC3F9D63FBD0" blockId="1.[151,642,978,1038]" box="[151,256,1013,1038]" pageId="1" pageNumber="142">
<figureCitation id="136A2A518765FFB8FF05FC3F9D63FBD0" box="[151,256,1013,1038]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1601,1623]" captionStartId-1="3.[151,250,1949,1971]" captionStartId-2="4.[151,250,1974,1996]" captionTargetBox-0="[340,1247,630,1577]" captionTargetBox-1="[340,1245,951,1912]" captionTargetBox-2="[418,1144,193,1886]" captionTargetId-0="figure@2.[340,1247,630,1577]" captionTargetId-1="figure@3.[340,1247,951,1924]" captionTargetId-2="figure@4.[418,1144,193,1905]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Taeniopteryx araneoides photographed at the Irtysh River in the Omsk centre: a—males on driftwood (two not fully mature with grey legs and a mature one with black legs), 29 iv 2006; b—female, 4 v 1979; c—attempt at mating of teneral male and a female with crumpled wings, 4 v 1979." captionText-1="FIGURE 2. Details of Taeniopteryx araneoides males from Omsk, 3 v 2003: a e—mature male, f—male with a not yet fully expanded epiproct; a—general habitus; b—wings; c—end of abdomen in lateral view; d—the same, dorsal view; e—the same, ventral view; f—laterodorsal view; scale bar 1 mm." captionText-2="FIGURE 3. Detail of a female of Taeniopteryx araneoides from Omsk, 3 v 2003: a—wings; b—end of abdomen in dorsoposterior view; d—the same, ventral view; scale bar 1 mm." httpUri-0="https://zenodo.org/record/438233/files/figure.png" httpUri-1="https://zenodo.org/record/438234/files/figure.png" httpUri-2="https://zenodo.org/record/438235/files/figure.png" pageId="1" pageNumber="142">Figs. 13</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34B655F8765FFB8FF05FBF49FBDF8B6" pageId="1" pageNumber="142" type="description">
<paragraph id="8BEE36D48765FFB8FF05FBF49910FB8D" blockId="1.[151,1395,1085,1107]" box="[151,1395,1085,1107]" pageId="1" pageNumber="142">
Reports for Siberia:
<bibRefCitation id="EFC04B258765FFB8FEF7FBF79E38FB8D" author="Zapekina-Dulkeit" box="[357,603,1085,1107]" pageId="1" pageNumber="142" refString="Zapekina-Dulkeit, Y. I. (1957) Vesnyanki Altae-Sayanskikh gornotaezhnykh vodoemov. In: Abstracts of a conference of zoologists of Siberia. AN SSSR, Novosibirsk, pp. 27 - 28. [in Russian]" type="book chapter" year="1957">Zapekina-Dulkeit, 1957</bibRefCitation>
: 2728;
<bibRefCitation id="EFC04B258765FFB8FD23FBF79FD2FB8D" author="Zapekina-Dulkeit" box="[689,945,1085,1107]" pageId="1" pageNumber="142" refString="Zapekina-Dulkeit, Y. I. (1971 a) Dopolnenie k faune vesnyanok (Plecoptera) zapovednika &quot; Stolby i smezhnykh territoriy. Trudy gosudarstvennogo zapovednika &quot; Stolby, 8, 88 - 90 [in Russian]" type="journal article" year="1971" yearSuffix="a">Zapekina-Dulkeit, 1971a</bibRefCitation>
: 8890;
<bibRefCitation id="EFC04B258765FFB8FB9AFBF79969FB8D" author="Zapekina-Dulkeit" box="[1032,1290,1085,1107]" pageId="1" pageNumber="142" refString="Zapekina-Dulkeit, Y. I. (1971 b) Vesnyanki (Plecoptera) Sibiri. TXIII International Congress of Entomology, Moscow, 2 - 9 August, Abstracts, 1, 223 - 224. [in Russian]" type="journal article" year="1971" yearSuffix="b">Zapekina-Dulkeit, 1971b</bibRefCitation>
: 223224.
</paragraph>
<paragraph id="8BEE36D48765FFB8FF05FBB59939FADB" blockId="1.[151,1437,1151,2005]" pageId="1" pageNumber="142">
<materialsCitation id="3B393C898765FFB8FF05FBB59F9DFB3E" ID-GBIF-Occurrence="1449840846" collectingDate="1995-05-02" collectorName="O. Kosterin" country="Russia" county="Om' River" elevation="68" location="Specimens" municipality="Irtysh River" pageId="1" pageNumber="142" specimenCount="58" specimenCount-female="5" specimenCount-male="53" stateProvince="Omsk">
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<location id="8E8E600F8765FFB8FF05FBB59D76FB46" LSID="urn:lsid:plazi:treatment:03F887C28765FFBEFF05FC1898E8FE10:8E8E600F8765FFB8FF05FBB59D76FB46" box="[151,277,1151,1176]" country="Russia" county="Om' River" municipality="Irtysh River" name="Specimens" pageId="1" pageNumber="142" stateProvince="Omsk">Specimens</location>
studied.
</emphasis>
<collectingCountry id="F34676448765FFB8FE17FBB59DB1FB46" box="[389,466,1151,1176]" name="Russia" pageId="1" pageNumber="142">Russia</collectingCountry>
,
<location id="8E8E600F8765FFB8FE4DFB4A9E11FB46" LSID="urn:lsid:plazi:treatment:03F887C28765FFBEFF05FC1898E8FE10:8E8E600F8765FFB8FE4DFB4A9E11FB46" box="[479,626,1151,1176]" country="Russia" county="Om' River" municipality="Irtysh River" name="West Siberia" pageId="1" pageNumber="142" stateProvince="Omsk">West Siberia</location>
,
<collectingRegion id="4995F8368765FFB8FDEDFBB59EA1FB46" box="[639,706,1151,1176]" country="Russia" name="Omsk" pageId="1" pageNumber="142">Omsk</collectingRegion>
city centre, the
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right bank downstream of the
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mouth, 5459' N, 732122'' E,
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a.s.l.
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,
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leg.:
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(of them 8 had non-expanded and 3 partly expanded anal appendages),
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in alcohol,
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,
<specimenCount id="9D57FD5D8765FFB8FD40FB029E9EFB01" box="[722,765,1224,1248]" pageId="1" pageNumber="142" type="female">4 ♀</specimenCount>
dry on pins,
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<collectingDate id="EFABE9FC8765FFB8FC04FB029F9DFB3E" box="[918,1022,1223,1248]" pageId="1" pageNumber="142" value="1995-05-02">2 v 1995</collectingDate>
</date>
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;
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<specimenCount id="9D57FD5D8765FFB8FB99FB0D9826FB01" box="[1035,1093,1223,1248]" pageId="1" pageNumber="142" type="male">15 ♂</specimenCount>
<specimenCount id="9D57FD5D8765FFB8FBDFFB029813FB3E" box="[1101,1136,1224,1248]" pageId="1" pageNumber="142" type="female">1♀</specimenCount>
, the same place,
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<collectingDate id="EFABE9FC8765FFB8FAA5FB0299FFFB3E" box="[1335,1436,1224,1248]" pageId="1" pageNumber="142" value="2003-05-03">3 v 2003</collectingDate>
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(specimens deposited in the
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of
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and
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of
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<collectionCode id="ED40AE118765FFB8FBB1FB26983FFADA" box="[1059,1116,1260,1284]" country="Myanmar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14256" name="Union of Burma Applied Research Institute" pageId="1" pageNumber="142">RAS</collectionCode>
,
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,
<collectingCountry id="F34676448765FFB8FB6DFB26992CFADB" box="[1279,1359,1260,1285]" name="Russia" pageId="1" pageNumber="142">Russia</collectingCountry>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEE36D48765FFB8FF05FAC59FBDF8B6" blockId="1.[151,1437,1151,2005]" pageId="1" pageNumber="142">
<materialsCitation id="3B393C898765FFB8FF05FAC59872FA93" ID-GBIF-Occurrence="1449840857" collectingDate="2014-04-20" collectorName="Pavlodar &amp; Akku &amp; I. Sivec &amp; N. Akimbekova" country="Kazakhstan" location="Pavlodar Province" municipality="Irtysh River" pageId="1" pageNumber="142" specimenCount="28" specimenCount-female="25" specimenCount-male="3">
<collectingCountry id="F34676448765FFB8FF05FAC59D7EFAF6" box="[151,285,1295,1320]" name="Kazakhstan" pageId="1" pageNumber="142">Kazakhstan</collectingCountry>
,
<location id="8E8E600F8765FFB8FEB8FAC59D9CFAF6" LSID="urn:lsid:plazi:treatment:03F887C28765FFBEFF05FC1898E8FE10:8E8E600F8765FFB8FEB8FAC59D9CFAF6" box="[298,511,1295,1320]" country="Kazakhstan" municipality="Irtysh River" name="Pavlodar Province" pageId="1" pageNumber="142">Pavlodar Province</location>
:
<specimenCount id="9D57FD5D8765FFB8FD9EFADA9E58FAF9" box="[524,571,1296,1320]" pageId="1" pageNumber="142" type="male">3 ♂</specimenCount>
,
<specimenCount id="9D57FD5D8765FFB8FDDBFAC59EE7FAF9" box="[585,644,1295,1320]" pageId="1" pageNumber="142" type="female">25 ♀</specimenCount>
,
<date id="FFEF10148765FFB8FD07FADA9F76FAF6" box="[661,789,1295,1320]" pageId="1" pageNumber="142" value="2014-04-20">
<collectingDate id="EFABE9FC8765FFB8FD07FADA9F76FAF6" box="[661,789,1295,1320]" pageId="1" pageNumber="142" value="2014-04-20">20 iv 2014</collectingDate>
</date>
, 5128 N, 7747 E, the
<collectingMunicipality id="6B8AACAE8765FFB8FBB6FAC598CEFAF6" box="[1060,1197,1295,1320]" pageId="1" pageNumber="142">Irtysh River</collectingMunicipality>
right bank upstream
<collectorName id="26A453028765FFB8FF05FAFE9C98FA93" box="[151,251,1332,1357]" pageId="1" pageNumber="142">Pavlodar</collectorName>
at
<collectorName id="26A453028765FFB8FE8DFAFE9D3EFA93" box="[287,349,1332,1357]" pageId="1" pageNumber="142">Akku</collectorName>
(formerly Lebyazhye) village,
<collectorName id="26A453028765FFB8FD50FAFF9F7FFA93" box="[706,796,1332,1357]" pageId="1" pageNumber="142">I. Sivec</collectorName>
,
<collectorName id="26A453028765FFB8FCBAFAFF9FBEFA93" box="[808,989,1332,1357]" pageId="1" pageNumber="142">N. Akimbekova</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B393C898765FFB8FB8CFAFE9DAAFAAE" ID-GBIF-Occurrence="1449840850" collectingDate="2015-04-20" collectorName="N. Akimbekova" country="Kazakhstan" location="Pavlodar Province" municipality="Irtysh River" pageId="1" pageNumber="142" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
<specimenCount id="9D57FD5D8765FFB8FB8CFAFE982FFA92" box="[1054,1100,1332,1356]" pageId="1" pageNumber="142" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D57FD5D8765FFB8FBCBFAFE98E7FA92" box="[1113,1156,1332,1356]" pageId="1" pageNumber="142" type="female">2 ♀</specimenCount>
, the same locality,
<date id="FFEF10148765FFB8FAF0FAFE9CB1FAAE" pageId="1" pageNumber="142" value="2015-04-20">
<collectingDate id="EFABE9FC8765FFB8FAF0FAFE9CB1FAAE" pageId="1" pageNumber="142" value="2015-04-20">20 iv 2015</collectingDate>
</date>
,
<collectorName id="26A453028765FFB8FF4DFA929DF7FAAE" box="[223,404,1367,1392]" pageId="1" pageNumber="142">N. Akimbekova</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B393C898765FFB8FE45FA92980BFAAE" ID-GBIF-Occurrence="1449840847" box="[471,1128,1367,1392]" collectingDate="2016-04-15" collectorName="N. Akimbekova" country="Kazakhstan" location="Pavlodar Province" municipality="Irtysh River" pageId="1" pageNumber="142" specimenCount="19" specimenCount-male="19">
<specimenCount id="9D57FD5D8765FFB8FE45FA929E70FAB1" box="[471,531,1368,1392]" pageId="1" pageNumber="142" type="male">19 ♂</specimenCount>
, the same locality,
<date id="FFEF10148765FFB8FD60FA9D9F13FAAE" box="[754,880,1367,1392]" pageId="1" pageNumber="142" value="2016-04-15">
<collectingDate id="EFABE9FC8765FFB8FD60FA9D9F13FAAE" box="[754,880,1367,1392]" pageId="1" pageNumber="142" value="2016-04-15">15 iv 2016</collectingDate>
</date>
,
<collectorName id="26A453028765FFB8FCEFFA929850FAAE" box="[893,1075,1367,1392]" pageId="1" pageNumber="142">N. Akimbekova</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B393C898765FFB8FBE4FA929ECAFA4B" ID-GBIF-Occurrence="1449840854" collectingDate="2016-04-23" collectorName="N. Akimbekova" country="Kazakhstan" location="Pavlodar Province" municipality="Irtysh River" pageId="1" pageNumber="142" specimenCount="105" specimenCount-female="97" specimenCount-male="8">
<specimenCount id="9D57FD5D8765FFB8FBE4FA9298C7FAB1" box="[1142,1188,1368,1392]" pageId="1" pageNumber="142" type="male">8 ♂</specimenCount>
,
<specimenCount id="9D57FD5D8765FFB8FB20FA92988FFAB1" box="[1202,1260,1368,1392]" pageId="1" pageNumber="142" type="female">97 ♀</specimenCount>
, 4 exuviae, the same locality,
<date id="FFEF10148765FFB8FEABFAB69DD1FA4B" box="[313,434,1404,1429]" pageId="1" pageNumber="142" value="2016-04-23">
<collectingDate id="EFABE9FC8765FFB8FEABFAB69DD1FA4B" box="[313,434,1404,1429]" pageId="1" pageNumber="142" value="2016-04-23">23 iv 2016</collectingDate>
</date>
,
<collectorName id="26A453028765FFB8FE53FAB79E16FA4B" box="[449,629,1404,1429]" pageId="1" pageNumber="142">N. Akimbekova</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B393C898765FFB8FD24FAB69F51FA66" ID-GBIF-Occurrence="1449840861" collectingDate="2016-04-23" collectorName="N. Akimbekova" country="Kazakhstan" county="Dzhambul village" location="Irtysh River" municipality="Pavlodar" pageId="1" pageNumber="142" specimenCount="23" specimenCount-female="23">
<specimenCount id="9D57FD5D8765FFB8FD24FAB69E8CFA4A" box="[694,751,1404,1429]" pageId="1" pageNumber="142" type="female">23 ♀</specimenCount>
, the
<location id="8E8E600F8765FFB8FCB5FAB69FCDFA4B" LSID="urn:lsid:plazi:treatment:03F887C28765FFBEFF05FC1898E8FE10:8E8E600F8765FFB8FCB5FAB69FCDFA4B" box="[807,942,1404,1429]" country="Kazakhstan" county="Dzhambul village" municipality="Pavlodar" name="Irtysh River" pageId="1" pageNumber="142">Irtysh River</location>
right bank upstream
<collectingMunicipality id="6B8AACAE8765FFB8FB32FAB69967FA4B" box="[1184,1284,1404,1429]" pageId="1" pageNumber="142">Pavlodar</collectingMunicipality>
at
<collectingCounty id="628F4E588765FFB8FAB5FAB69C85FA66" pageId="1" pageNumber="142">Dzhambul village</collectingCounty>
, 5130 N, 7741 E,
<date id="FFEF10148765FFB8FE53FA6A9E5DFA66" box="[449,574,1439,1464]" pageId="1" pageNumber="142" value="2016-04-23">
<collectingDate id="EFABE9FC8765FFB8FE53FA6A9E5DFA66" box="[449,574,1439,1464]" pageId="1" pageNumber="142" value="2016-04-23">23 iv 2016</collectingDate>
</date>
,
<collectorName id="26A453028765FFB8FDD8FA6A9E9DFA66" box="[586,766,1439,1464]" pageId="1" pageNumber="142">N. Akimbekova</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B393C898765FFB8FCADFA6A9877FA03" ID-GBIF-Occurrence="1449840858" collectingDate="2015-04-24" collectorName="N. Akimbekova" country="Kazakhstan" county="Zhanatan village" location="Irtysh River" municipality="Pavlodar" pageId="1" pageNumber="142" specimenCount="8" specimenCount-female="7" specimenCount-male="1">
<specimenCount id="9D57FD5D8765FFB8FCADFA6A9F0FFA69" box="[831,876,1440,1464]" pageId="1" pageNumber="142" type="male">1 ♂</specimenCount>
, the
<location id="8E8E600F8765FFB8FC31FA559849FA66" LSID="urn:lsid:plazi:treatment:03F887C28765FFBEFF05FC1898E8FE10:8E8E600F8765FFB8FC31FA559849FA66" box="[931,1066,1439,1464]" country="Kazakhstan" county="Zhanatan village" municipality="Pavlodar" name="Irtysh River" pageId="1" pageNumber="142">Irtysh River</location>
right bank upstream
<collectingMunicipality id="6B8AACAE8765FFB8FA8EFA5599E3FA66" box="[1308,1408,1439,1464]" pageId="1" pageNumber="142">Pavlodar</collectingMunicipality>
at
<collectingCounty id="628F4E588765FFB8FF05FA0E9D30FA03" box="[151,339,1476,1501]" pageId="1" pageNumber="142">Zhanatan village</collectingCounty>
env.,
<specimenCount id="9D57FD5D8765FFB8FE0AFA0F9DA0FA02" box="[408,451,1477,1500]" pageId="1" pageNumber="142" type="female">7 ♀</specimenCount>
, 5132 N, 7738 E,
<date id="FFEF10148765FFB8FD30FA0E9F7FFA03" box="[674,796,1476,1501]" pageId="1" pageNumber="142" value="2015-04-24">
<collectingDate id="EFABE9FC8765FFB8FD30FA0E9F7FFA03" box="[674,796,1476,1501]" pageId="1" pageNumber="142" value="2015-04-24">24 iv 2015</collectingDate>
</date>
,
<collectorName id="26A453028765FFB8FCB9FA0F9F83FA03" box="[811,992,1476,1501]" pageId="1" pageNumber="142">N. Akimbekova</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B393C898765FFB8FBB2FA0E9DA7F9DE" ID-GBIF-Occurrence="1449840855" collectingDate="2015-04-25" collectorName="N. Akimbekova" country="Kazakhstan" county="Zhanatan village" location="Irtysh River" municipality="Pavlodar" pageId="1" pageNumber="142" specimenCount="10" specimenCount-female="8" specimenCount-male="2">
<specimenCount id="9D57FD5D8765FFB8FBB2FA0E982FFA02" box="[1056,1100,1476,1501]" pageId="1" pageNumber="142" type="male">2 ♂</specimenCount>
,
<specimenCount id="9D57FD5D8765FFB8FBC9FA0E98E5FA02" box="[1115,1158,1476,1500]" pageId="1" pageNumber="142" type="female">8 ♀</specimenCount>
, the same locality,
<date id="FFEF10148765FFB8FAF0FA0E9CACF9DE" pageId="1" pageNumber="142" value="2015-04-25">
<collectingDate id="EFABE9FC8765FFB8FAF0FA0E9CACF9DE" pageId="1" pageNumber="142" value="2015-04-25">25 iv 2015</collectingDate>
</date>
,
<collectorName id="26A453028765FFB8FF4FFA229DF2F9DE" box="[221,401,1511,1536]" pageId="1" pageNumber="142">N. Akimbekova</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B393C898765FFB8FE42FA2D9E8AF9FB" ID-GBIF-Occurrence="1449840856" collectingDate="2016-04-23" collectorName="N. Akimbekova" country="Kazakhstan" county="Tlektes village" location="Irtysh River" municipality="Pavlodar" pageId="1" pageNumber="142" specimenCount="26" specimenCount-female="22" specimenCount-male="4">
<specimenCount id="9D57FD5D8765FFB8FE42FA2D9D99FA21" box="[464,506,1511,1536]" pageId="1" pageNumber="142" type="male">4 ♂</specimenCount>
,
<specimenCount id="9D57FD5D8765FFB8FD9AFA229E22FA21" box="[520,577,1512,1536]" pageId="1" pageNumber="142" type="female">22 ♀</specimenCount>
, 3 exuviae, the
<location id="8E8E600F8765FFB8FD61FA2D9F1AF9DE" LSID="urn:lsid:plazi:treatment:03F887C28765FFBEFF05FC1898E8FE10:8E8E600F8765FFB8FD61FA2D9F1AF9DE" box="[755,889,1511,1536]" country="Kazakhstan" county="Tlektes village" municipality="Pavlodar" name="Irtysh River" pageId="1" pageNumber="142">Irtysh River</location>
right bank upstream
<collectingMunicipality id="6B8AACAE8765FFB8FBFAFA2D98AFF9DE" box="[1128,1228,1511,1536]" pageId="1" pageNumber="142">Pavlodar</collectingMunicipality>
at
<collectingCounty id="628F4E588765FFB8FB7DFA2D99FBF9DE" box="[1263,1432,1511,1536]" pageId="1" pageNumber="142">Tlektes village</collectingCounty>
, 5149 N, 7727 E,
<date id="FFEF10148765FFB8FEFCF9C69D8FF9FB" box="[366,492,1548,1573]" pageId="1" pageNumber="142" value="2016-04-23">
<collectingDate id="EFABE9FC8765FFB8FEFCF9C69D8FF9FB" box="[366,492,1548,1573]" pageId="1" pageNumber="142" value="2016-04-23">23 iv 2016</collectingDate>
</date>
,
<collectorName id="26A453028765FFB8FE6FF9C79ED7F9FB" box="[509,692,1548,1573]" pageId="1" pageNumber="142">N. Akimbekova</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B393C898765FFB8FD6AF9C698E2F996" ID-GBIF-Occurrence="1449840852" collectingDate="2016-04-19" collectorName="N. Akimbekova" country="Kazakhstan" county="Novoyamyshevo village" location="Irtysh River" municipality="Pavlodar" pageId="1" pageNumber="142" specimenCount="45" specimenCount-exuvia="1" specimenCount-female="7" specimenCount-male="37">
<specimenCount id="9D57FD5D8765FFB8FD6AF9C69F57F9FA" box="[760,820,1548,1573]" pageId="1" pageNumber="142" type="male">37 ♂</specimenCount>
,
<specimenCount id="9D57FD5D8765FFB8FCD6F9C79F12F9FA" box="[836,881,1549,1572]" pageId="1" pageNumber="142" type="female">7 ♀</specimenCount>
,
<specimenCount id="9D57FD5D8765FFB8FC10F9C69F8BF9FB" box="[898,1000,1548,1573]" pageId="1" pageNumber="142" type="exuvia">1 exuvia</specimenCount>
, the
<location id="8E8E600F8765FFB8FBB0F9C698CFF9FB" LSID="urn:lsid:plazi:treatment:03F887C28765FFBEFF05FC1898E8FE10:8E8E600F8765FFB8FBB0F9C698CFF9FB" box="[1058,1196,1548,1573]" country="Kazakhstan" county="Novoyamyshevo village" municipality="Pavlodar" name="Irtysh River" pageId="1" pageNumber="142">Irtysh River</location>
right bank upstream
<collectingMunicipality id="6B8AACAE8765FFB8FF05F9E59C98F996" box="[151,251,1583,1608]" pageId="1" pageNumber="142">Pavlodar</collectingMunicipality>
at
<collectingCounty id="628F4E588765FFB8FE8DF9E59E57F996" box="[287,564,1583,1608]" pageId="1" pageNumber="142">Novoyamyshevo village</collectingCounty>
, 5155 N, 7719 E,
<date id="FFEF10148765FFB8FC82F9FA9FE9F996" box="[784,906,1583,1608]" pageId="1" pageNumber="142" value="2016-04-19">
<collectingDate id="EFABE9FC8765FFB8FC82F9FA9FE9F996" box="[784,906,1583,1608]" pageId="1" pageNumber="142" value="2016-04-19">19 iv 2016</collectingDate>
</date>
,
<collectorName id="26A453028765FFB8FC0AF9FA982EF996" box="[920,1101,1583,1608]" pageId="1" pageNumber="142">N. Akimbekova</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B393C898765FFB8FB1FF9E59EC6F9B3" ID-GBIF-Occurrence="1449840849" collectingDate="2016-04-23" collectorName="N. Akimbekova" country="Kazakhstan" county="Novoyamyshevo village" location="Irtysh River" municipality="Pavlodar" pageId="1" pageNumber="142" specimenCount="7" specimenCount-exuvia="1" specimenCount-female="5" specimenCount-male="1">
<specimenCount id="9D57FD5D8765FFB8FB1FF9E598DBF999" box="[1165,1208,1583,1608]" pageId="1" pageNumber="142" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D57FD5D8765FFB8FB5AF9E59890F999" box="[1224,1267,1583,1608]" pageId="1" pageNumber="142" type="female">5 ♀</specimenCount>
,
<specimenCount id="9D57FD5D8765FFB8FA98F9FA990EF996" box="[1290,1389,1583,1608]" pageId="1" pageNumber="142" type="exuvia">1 exuvia</specimenCount>
, the same locality,
<date id="FFEF10148765FFB8FEAAF99E9DD3F9B3" box="[312,432,1620,1645]" pageId="1" pageNumber="142" value="2016-04-23">
<collectingDate id="EFABE9FC8765FFB8FEAAF99E9DD3F9B3" box="[312,432,1620,1645]" pageId="1" pageNumber="142" value="2016-04-23">23 iv 2016</collectingDate>
</date>
,
<collectorName id="26A453028765FFB8FE2CF99F9E11F9B3" box="[446,626,1620,1645]" pageId="1" pageNumber="142">N. Akimbekova</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B393C898765FFB8FD23F99E997BF9B3" ID-GBIF-Occurrence="1449840848" box="[689,1304,1620,1645]" collectingDate="2016-05-06" collectorName="N. Akimbekova" country="Kazakhstan" county="Novoyamyshevo village" location="Irtysh River" municipality="Pavlodar" pageId="1" pageNumber="142" specimenCount="19" specimenCount-female="19">
<specimenCount id="9D57FD5D8765FFB8FD23F99E9E8AF9B2" box="[689,745,1620,1645]" pageId="1" pageNumber="142" type="female">19 ♀</specimenCount>
, the same locality,
<date id="FFEF10148765FFB8FC53F99E9847F9B3" box="[961,1060,1620,1645]" pageId="1" pageNumber="142" value="2016-05-06">
<collectingDate id="EFABE9FC8765FFB8FC53F99E9847F9B3" box="[961,1060,1620,1645]" pageId="1" pageNumber="142" value="2016-05-06">6 v 2016</collectingDate>
</date>
,
<collectorName id="26A453028765FFB8FBA0F99F9886F9B3" box="[1074,1253,1620,1645]" pageId="1" pageNumber="142">N. Akimbekova</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B393C898765FFB8FAB7F99E9D38F96B" ID-GBIF-Occurrence="1449840851" collectingDate="2016-04-21" collectorName="N. Akimbekova" country="Kazakhstan" county="Sychovka village" location="Irtysh River" municipality="Pavlodar" pageId="1" pageNumber="142" specimenCount="33" specimenCount-female="3" specimenCount-male="30">
<specimenCount id="9D57FD5D8765FFB8FAB7F99E993FF9B2" box="[1317,1372,1620,1645]" pageId="1" pageNumber="142" type="male">30 ♂</specimenCount>
,
<specimenCount id="9D57FD5D8765FFB8FAF9F99E99F6F9B2" box="[1387,1429,1620,1644]" pageId="1" pageNumber="142" type="female">3 ♀</specimenCount>
, 1 skin, the
<location id="8E8E600F8765FFB8FE89F9BD9DC7F94E" LSID="urn:lsid:plazi:treatment:03F887C28765FFBEFF05FC1898E8FE10:8E8E600F8765FFB8FE89F9BD9DC7F94E" box="[283,420,1655,1680]" country="Kazakhstan" county="Sychovka village" municipality="Pavlodar" name="Irtysh River" pageId="1" pageNumber="142">Irtysh River</location>
right bank downstream
<collectingMunicipality id="6B8AACAE8765FFB8FD52F9BD9F47F94E" box="[704,804,1655,1680]" pageId="1" pageNumber="142">Pavlodar</collectingMunicipality>
at
<collectingCounty id="628F4E588765FFB8FCDEF9BD9870F94E" box="[844,1043,1655,1680]" pageId="1" pageNumber="142">Sychovka village</collectingCounty>
, 5236 N 7645 E,
<date id="FFEF10148765FFB8FB60F9B29910F94E" box="[1266,1395,1655,1680]" pageId="1" pageNumber="142" value="2016-04-21">
<collectingDate id="EFABE9FC8765FFB8FB60F9B29910F94E" box="[1266,1395,1655,1680]" pageId="1" pageNumber="142" value="2016-04-21">21 iv 2016</collectingDate>
</date>
,
<collectorName id="26A453028765FFB8FA13F9B29D4BF96B" pageId="1" pageNumber="142">N. Akimbekova</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B393C898765FFB8FEF5F9569E0BF906" ID-GBIF-Occurrence="1449840853" collectingDate="2015-04-22" collectorName="N. Akimbekova" country="Kazakhstan" county="Naberezhnoe village" elevation="10" location="Irtysh River" municipality="Pavlodar" pageId="1" pageNumber="142" specimenCount="82" specimenCount-female="82">
<quantity id="4CA99B318765FFB8FEF5F9569DFBF96A" box="[359,408,1692,1716]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="142" unit="m" value="10.0">
<elevation id="007CD1E78765FFB8FEF5F9569DFBF96A" box="[359,408,1692,1716]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="142" unit="m" value="10.0">10m</elevation>
</quantity>
#,
<specimenCount id="9D57FD5D8765FFB8FE29F9569D90F96A" box="[443,499,1692,1716]" pageId="1" pageNumber="142" type="female">82 ♀</specimenCount>
, the
<location id="8E8E600F8765FFB8FDB8F9569ED2F96B" LSID="urn:lsid:plazi:treatment:03F887C28765FFBEFF05FC1898E8FE10:8E8E600F8765FFB8FDB8F9569ED2F96B" box="[554,689,1692,1717]" country="Kazakhstan" county="Naberezhnoe village" municipality="Pavlodar" name="Irtysh River" pageId="1" pageNumber="142">Irtysh River</location>
right bank downstream
<collectingMunicipality id="6B8AACAE8765FFB8FC53F9569846F96B" box="[961,1061,1692,1717]" pageId="1" pageNumber="142">Pavlodar</collectingMunicipality>
at
<collectingCounty id="628F4E588765FFB8FBD5F9569952F96B" box="[1095,1329,1692,1717]" pageId="1" pageNumber="142">Naberezhnoe village</collectingCounty>
, 5241 N, 7643 E,
<date id="FFEF10148765FFB8FF69F90A9D10F906" box="[251,371,1727,1752]" pageId="1" pageNumber="142" value="2015-04-22">
<collectingDate id="EFABE9FC8765FFB8FF69F90A9D10F906" box="[251,371,1727,1752]" pageId="1" pageNumber="142" value="2015-04-22">22 iv 2015</collectingDate>
</date>
,
<collectorName id="26A453028765FFB8FE13F90A9E56F906" box="[385,565,1727,1752]" pageId="1" pageNumber="142">N. Akimbekova</collectorName>
leg.
</materialsCitation>
; 70 ♂, 14 ♀, the same place,
<date id="FFEF10148765FFB8FC21F90A9848F906" box="[947,1067,1727,1752]" pageId="1" pageNumber="142" value="2016-04-30">30 iv 2016</date>
, N. Akimbekova leg.; 170 ♂, 75 ♀, 7 exuviae, the same place,
<date id="FFEF10148765FFB8FD9AF92E9EE8F923" box="[520,651,1764,1789]" pageId="1" pageNumber="142" value="2016-04-16">16 iv 2016</date>
, N. Akimbekova leg.; 1 ♂, 5 ♀, the same place,
<date id="FFEF10148765FFB8FB78F92E9913F923" box="[1258,1392,1764,1789]" pageId="1" pageNumber="142" value="2016-04-24">24 iv 2016</date>
, N. Akimbekova leg.;
<materialsCitation id="3B393C898765FFB8FEF8F8CD9EBDF89B" ID-GBIF-Occurrence="1449840845" collectingDate="2016-04-24" collectorName="N. Akimbekova" country="Kazakhstan" county="Grigoryevka village" location="Irtysh River" municipality="Pavlodar" pageId="1" pageNumber="142" specimenCount="19" specimenCount-exuvia="1" specimenCount-female="10" specimenCount-male="8">
<specimenCount id="9D57FD5D8765FFB8FEF8F8CD9DF6F8C1" box="[362,405,1799,1824]" pageId="1" pageNumber="142" type="male">8 ♂</specimenCount>
,
<specimenCount id="9D57FD5D8765FFB8FE37F8C29DBCF8C1" box="[421,479,1800,1824]" pageId="1" pageNumber="142" type="female">10 ♀</specimenCount>
,
<specimenCount id="9D57FD5D8765FFB8FE7CF8C29E31F8FE" box="[494,594,1799,1824]" pageId="1" pageNumber="142" type="exuvia">1 exuvia</specimenCount>
, the
<location id="8E8E600F8765FFB8FD18F8CD9F71F8FE" LSID="urn:lsid:plazi:treatment:03F887C28765FFBEFF05FC1898E8FE10:8E8E600F8765FFB8FD18F8CD9F71F8FE" box="[650,786,1799,1824]" country="Kazakhstan" county="Grigoryevka village" municipality="Pavlodar" name="Irtysh River" pageId="1" pageNumber="142">Irtysh River</location>
right bank downstream
<collectingMunicipality id="6B8AACAE8765FFB8FBBBF8CD98EEF8FE" box="[1065,1165,1799,1824]" pageId="1" pageNumber="142">Pavlodar</collectingMunicipality>
at
<collectingCounty id="628F4E588765FFB8FB20F8CD99FBF8FE" box="[1202,1432,1799,1824]" pageId="1" pageNumber="142">Grigoryevka village</collectingCounty>
, 5238 N 7644 E,
<date id="FFEF10148765FFB8FEF7F8E69D81F89B" box="[357,482,1836,1861]" pageId="1" pageNumber="142" value="2016-04-24">
<collectingDate id="EFABE9FC8765FFB8FEF7F8E69D81F89B" box="[357,482,1836,1861]" pageId="1" pageNumber="142" value="2016-04-24">24 iv 2016</collectingDate>
</date>
,
<collectorName id="26A453028765FFB8FE61F8E79ECBF89B" box="[499,680,1836,1861]" pageId="1" pageNumber="142">N. Akimbekova</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B393C898765FFB8FD7EF8E69FBDF8B6" ID-GBIF-Occurrence="1449840860" collectingDate="2014-04-24" collectionCode="PSU" collectorName="I. Sivec &amp; N. Akimbekova" country="Kazakhstan" location="Irtysh River" municipality="Terenkol" pageId="1" pageNumber="142" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D57FD5D8765FFB8FD7EF8E69F7BF89A" box="[748,792,1836,1861]" pageId="1" pageNumber="142" type="female">1 ♀</specimenCount>
,
<location id="8E8E600F8765FFB8FCBBF8E69FD1F89B" LSID="urn:lsid:plazi:treatment:03F887C28765FFBEFF05FC1898E8FE10:8E8E600F8765FFB8FCBBF8E69FD1F89B" box="[809,946,1836,1861]" country="Kazakhstan" municipality="Terenkol" name="Irtysh River" pageId="1" pageNumber="142">Irtysh River</location>
right bank at
<collectingMunicipality id="6B8AACAE8765FFB8FBC5F8E698D9F89B" box="[1111,1210,1836,1861]" pageId="1" pageNumber="142">Terenkol</collectingMunicipality>
(formerly Kachiry) village, 5304 N, 7606 E,
<date id="FFEF10148765FFB8FE2FF89A9E5BF8B6" box="[445,568,1871,1896]" pageId="1" pageNumber="142" value="2014-04-24">
<collectingDate id="EFABE9FC8765FFB8FE2FF89A9E5BF8B6" box="[445,568,1871,1896]" pageId="1" pageNumber="142" value="2014-04-24">24 iv 2014</collectingDate>
</date>
,
<collectorName id="26A453028765FFB8FDD0F89A9EF8F8B6" box="[578,667,1871,1896]" pageId="1" pageNumber="142">I. Sivec</collectorName>
,
<collectorName id="26A453028765FFB8FD34F89A9F3AF8B6" box="[678,857,1871,1896]" pageId="1" pageNumber="142">N. Akimbekova</collectorName>
leg. (
<collectionCode id="ED40AE118765FFB8FC0AF89A9FB2F8B6" box="[920,977,1872,1896]" country="USA" httpUri="http://grbio.org/cool/5mg9-yyc1" name="Portland State University, Vertebrate Biology Museum" pageId="1" pageNumber="142">PSU</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34B655F8765FFBEFF55F8BE98E8FE10" lastPageId="7" lastPageNumber="148" pageId="1" pageNumber="142" type="discussion">
<paragraph id="8BEE36D48765FFBBFF55F8BE9E66FE9D" blockId="1.[151,1437,1151,2005]" lastBlockId="2.[151,1437,151,576]" lastPageId="2" lastPageNumber="143" pageId="1" pageNumber="142">
<emphasis id="B925EAC68765FFB8FF55F8BE9D58F853" bold="true" box="[199,315,1908,1933]" pageId="1" pageNumber="142">Remarks.</emphasis>
The above males are in agreement with the detailed descriptions and figures of
<taxonomicName id="4C514D578765FFB8FB57F8BF993DF853" box="[1221,1374,1908,1933]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="araneoides">
<emphasis id="B925EAC68765FFB8FB57F8BF993DF853" box="[1221,1374,1908,1933]" italics="true" pageId="1" pageNumber="142">T. araneoides</emphasis>
</taxonomicName>
from the Danube River (Klápalek 1902, 1905;
<bibRefCitation id="EFC04B258765FFB8FD11F8529F42F86F" author="Bianchi" box="[643,801,1944,1969]" pageId="1" pageNumber="142" refString="Bianchi, Y. L. (1905) Vesnyanki Plecoptera. In: Jacobson, G. G. &amp; Bianchi, Y. L. (Ed.), Orthoptera and Pseudoneuroptera from the Russian Empire and neighbouring lands. A. F. De Vriena Publ., St-Petersburg, pp. 502 - 634. [in Russian]" type="book chapter" year="1905">Bianchi 1905</bibRefCitation>
;
<bibRefCitation id="EFC04B258765FFB8FCA3F8529F80F86E" author="Zhiltzova" box="[817,995,1944,1969]" pageId="1" pageNumber="142" refString="Zhiltzova, L. A. (2003) Insecta Plecoptera. Plecoptera, Gruppe Euholognatha. Fauna of Russia and Neighbouring Countries, New Series, 145, 1 - 538. [in Russian, English / German title]" type="journal article" year="2003">Zhiltzova 2003</bibRefCitation>
) exhibiting diagnostic features of the species, the vesicle situated in the apical half of sternite IX, tergite X not incised apically (
<figureCitation id="136A2A518765FFB8FB57F876996FF80B" box="[1221,1292,1980,2005]" captionStart="FIGURE 2" captionStartId="3.[151,250,1949,1971]" captionTargetBox="[340,1245,951,1912]" captionTargetId="figure@3.[340,1247,951,1924]" captionTargetPageId="3" captionText="FIGURE 2. Details of Taeniopteryx araneoides males from Omsk, 3 v 2003: a e—mature male, f—male with a not yet fully expanded epiproct; a—general habitus; b—wings; c—end of abdomen in lateral view; d—the same, dorsal view; e—the same, ventral view; f—laterodorsal view; scale bar 1 mm." httpUri="https://zenodo.org/record/438234/files/figure.png" pageId="1" pageNumber="142">Fig. 2</figureCitation>
d); the outer portion of the paraproct expanded basally and acute apically, curved at the tip (
<figureCitation id="136A2A518766FFBBFB8DFF5D9806FF6E" box="[1055,1125,151,176]" captionStart="FIGURE 2" captionStartId="3.[151,250,1949,1971]" captionTargetBox="[340,1245,951,1912]" captionTargetId="figure@3.[340,1247,951,1924]" captionTargetPageId="3" captionText="FIGURE 2. Details of Taeniopteryx araneoides males from Omsk, 3 v 2003: a e—mature male, f—male with a not yet fully expanded epiproct; a—general habitus; b—wings; c—end of abdomen in lateral view; d—the same, dorsal view; e—the same, ventral view; f—laterodorsal view; scale bar 1 mm." httpUri="https://zenodo.org/record/438234/files/figure.png" pageId="2" pageNumber="143">Fig. 2</figureCitation>
c), the cerci produced to an inner tooth (
<figureCitation id="136A2A518766FFBBFEB2FF769D01FF0B" box="[288,354,188,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1949,1971]" captionTargetBox="[340,1245,951,1912]" captionTargetId="figure@3.[340,1247,951,1924]" captionTargetPageId="3" captionText="FIGURE 2. Details of Taeniopteryx araneoides males from Omsk, 3 v 2003: a e—mature male, f—male with a not yet fully expanded epiproct; a—general habitus; b—wings; c—end of abdomen in lateral view; d—the same, dorsal view; e—the same, ventral view; f—laterodorsal view; scale bar 1 mm." httpUri="https://zenodo.org/record/438234/files/figure.png" pageId="2" pageNumber="143">Fig. 2</figureCitation>
cf), in front of which there is a tiny projection (the vestigial second segment of the cercus) (
<figureCitation id="136A2A518766FFBBFAFDFF769CC6FF26" captionStart="FIGURE 2" captionStartId="3.[151,250,1949,1971]" captionTargetBox="[340,1245,951,1912]" captionTargetId="figure@3.[340,1247,951,1924]" captionTargetPageId="3" captionText="FIGURE 2. Details of Taeniopteryx araneoides males from Omsk, 3 v 2003: a e—mature male, f—male with a not yet fully expanded epiproct; a—general habitus; b—wings; c—end of abdomen in lateral view; d—the same, dorsal view; e—the same, ventral view; f—laterodorsal view; scale bar 1 mm." httpUri="https://zenodo.org/record/438234/files/figure.png" pageId="2" pageNumber="143">Fig. 2</figureCitation>
cd), a large upright epiproct (
<figureCitation id="136A2A518766FFBBFE65FF159E58FF26" box="[503,571,223,248]" captionStart="FIGURE 2" captionStartId="3.[151,250,1949,1971]" captionTargetBox="[340,1245,951,1912]" captionTargetId="figure@3.[340,1247,951,1924]" captionTargetPageId="3" captionText="FIGURE 2. Details of Taeniopteryx araneoides males from Omsk, 3 v 2003: a e—mature male, f—male with a not yet fully expanded epiproct; a—general habitus; b—wings; c—end of abdomen in lateral view; d—the same, dorsal view; e—the same, ventral view; f—laterodorsal view; scale bar 1 mm." httpUri="https://zenodo.org/record/438234/files/figure.png" pageId="2" pageNumber="143">Fig. 2</figureCitation>
c). The male habitus (
<figureCitation id="136A2A518766FFBBFCB8FF159F0AFF26" box="[810,873,223,248]" captionStart="FIGURE 1" captionStartId="2.[151,250,1601,1623]" captionTargetBox="[340,1247,630,1577]" captionTargetId="figure@2.[340,1247,630,1577]" captionTargetPageId="2" captionText="FIGURE 1. Taeniopteryx araneoides photographed at the Irtysh River in the Omsk centre: a—males on driftwood (two not fully mature with grey legs and a mature one with black legs), 29 iv 2006; b—female, 4 v 1979; c—attempt at mating of teneral male and a female with crumpled wings, 4 v 1979." httpUri="https://zenodo.org/record/438233/files/figure.png" pageId="2" pageNumber="143">Fig. 1</figureCitation>
a, 2a) is identical to a photograph of a specimen in the Hungarian Natural History Museum,
<collectingRegion id="4995F8368766FFBBFDE9FECC9E85FEC1" box="[635,742,262,287]" country="Hungary" name="Budapest" pageId="2" pageNumber="143">Budapest</collectingRegion>
, collected at the beginning of the 20th century at
<collectingRegion id="4995F8368766FFBBFAA6FECC99FFFEC1" box="[1332,1436,262,287]" country="Hungary" name="Budapest" pageId="2" pageNumber="143">Budapest</collectingRegion>
(
<bibRefCitation id="EFC04B258766FFBBFF0CFEE19D97FE9D" author="Kovcs" box="[158,500,298,323]" pageId="2" pageNumber="143" refString="Kovcs, T. &amp; Muranyi, D. (2008) Taeniopterygidae Klapalek, 1905 species in Hungary (Plecoptera). Folio historico naturalia musei matraensis, 32, 103 - 113." type="journal article" year="2008">Kovcs &amp; Murányi 2008: fig. 1</bibRefCitation>
).
</paragraph>
<paragraph id="8BEE36D48766FFBBFF55FE84985EFE0D" blockId="2.[151,1437,151,576]" pageId="2" pageNumber="143">
The female has no transverse vein in the subcostal field (
<figureCitation id="136A2A518766FFBBFCD3FE849FE1FEB9" box="[833,898,334,359]" captionStart="FIGURE 3" captionStartId="4.[151,250,1974,1996]" captionTargetBox="[418,1144,193,1886]" captionTargetId="figure@4.[418,1144,193,1905]" captionTargetPageId="4" captionText="FIGURE 3. Detail of a female of Taeniopteryx araneoides from Omsk, 3 v 2003: a—wings; b—end of abdomen in dorsoposterior view; d—the same, ventral view; scale bar 1 mm." httpUri="https://zenodo.org/record/438235/files/figure.png" pageId="2" pageNumber="143">Fig. 3</figureCitation>
a) as found in
<taxonomicName id="4C514D578766FFBBFBB0FE9A98D9FEB9" box="[1058,1210,334,359]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="143" phylum="Arthropoda" rank="species" species="araneoides">
<emphasis id="B925EAC68766FFBBFBB0FE9A98D9FEB9" box="[1058,1210,334,359]" italics="true" pageId="2" pageNumber="143">T. araneoides</emphasis>
</taxonomicName>
and in contrast to
<taxonomicName id="4C514D578766FFBBFA15FE9A9C9FFE55" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="143" phylum="Arthropoda" rank="species" species="nebulosa">
<emphasis id="B925EAC68766FFBBFA15FE9A9C9FFE55" italics="true" pageId="2" pageNumber="143">T. nebulosa</emphasis>
</taxonomicName>
with a crossvein (Klápalek 1902,
<bibRefCitation id="EFC04B258766FFBBFD12FEB89F79FE55" author="Bianchi" box="[640,794,370,395]" pageId="2" pageNumber="143" refString="Bianchi, Y. L. (1905) Vesnyanki Plecoptera. In: Jacobson, G. G. &amp; Bianchi, Y. L. (Ed.), Orthoptera and Pseudoneuroptera from the Russian Empire and neighbouring lands. A. F. De Vriena Publ., St-Petersburg, pp. 502 - 634. [in Russian]" type="book chapter" year="1905">Bianchi 1905</bibRefCitation>
,
<bibRefCitation id="EFC04B258766FFBBFCB5FEB89FB4FE55" author="Zhiltzova" box="[807,983,370,395]" pageId="2" pageNumber="143" refString="Zhiltzova, L. A. (2003) Insecta Plecoptera. Plecoptera, Gruppe Euholognatha. Fauna of Russia and Neighbouring Countries, New Series, 145, 1 - 538. [in Russian, English / German title]" type="journal article" year="2003">Zhiltzova 2003</bibRefCitation>
). The above diagnostic features confirm our species identification. Even if there are some subtle differences from the presently extinct European populations, they cannot be compared without available fresh European specimens.
</paragraph>
<paragraph id="8BEE36D48766FFBBFF55FE2A98E8FD9E" blockId="2.[151,1437,151,576]" pageId="2" pageNumber="143">
It should be noted that the series from
<collectingRegion id="4995F8368766FFBBFDE6FE149ED4FE29" box="[628,695,478,503]" country="Russia" name="Omsk" pageId="2" pageNumber="143">Omsk</collectingRegion>
includes eight males with exposed cerci not fully exposed and the epiproct still concealed inside a nearly flatly truncated tip of abdomen. At least three males had the epiproct small and rounded (
<figureCitation id="136A2A518766FFBBFEA3FDEC9D16FDE1" box="[305,373,550,575]" captionStart="FIGURE 2" captionStartId="3.[151,250,1949,1971]" captionTargetBox="[340,1245,951,1912]" captionTargetId="figure@3.[340,1247,951,1924]" captionTargetPageId="3" captionText="FIGURE 2. Details of Taeniopteryx araneoides males from Omsk, 3 v 2003: a e—mature male, f—male with a not yet fully expanded epiproct; a—general habitus; b—wings; c—end of abdomen in lateral view; d—the same, dorsal view; e—the same, ventral view; f—laterodorsal view; scale bar 1 mm." httpUri="https://zenodo.org/record/438234/files/figure.png" pageId="2" pageNumber="143">Fig. 2</figureCitation>
f). Presumingly, these specimens were collected soon after emergence.
</paragraph>
<caption id="DF2E665C8766FFBBFF05F98B9EFBF94A" httpUri="https://zenodo.org/record/438233/files/figure.png" pageId="2" pageNumber="143" targetBox="[340,1247,630,1577]" targetPageId="2">
<paragraph id="8BEE36D48766FFBBFF05F98B9EFBF94A" blockId="2.[151,1436,1601,1684]" pageId="2" pageNumber="143">
<emphasis id="B925EAC68766FFBBFF05F98B9D76F988" bold="true" box="[151,277,1601,1623]" pageId="2" pageNumber="143">FIGURE 1.</emphasis>
<taxonomicName id="4C514D578766FFBBFE8FF9889E7AF989" box="[285,537,1601,1623]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="143" phylum="Arthropoda" rank="species" species="araneoides">
<emphasis id="B925EAC68766FFBBFE8FF9889E7AF989" box="[285,537,1601,1623]" italics="true" pageId="2" pageNumber="143">Taeniopteryx araneoides</emphasis>
</taxonomicName>
photographed at the Irtysh River in the Omsk centre: a—males on driftwood (two not fully mature with grey legs and a mature one with black legs), 29 iv 2006; b—female, 4 v 1979; c—attempt at mating of teneral male and a female with crumpled wings, 4 v 1979.
</paragraph>
</caption>
<paragraph id="8BEE36D48766FFBAFF55F9769CB0FE2B" blockId="2.[151,1437,1724,2037]" lastBlockId="3.[151,1437,151,933]" lastPageId="3" lastPageNumber="144" pageId="2" pageNumber="143">
<emphasis id="B925EAC68766FFBBFF55F9769DB1F90B" bold="true" box="[199,466,1724,1749]" pageId="2" pageNumber="143">Habitat and behavior.</emphasis>
At
<collectingRegion id="4995F8368766FFBBFD90F9769E2BF90B" box="[514,584,1724,1749]" country="Russia" name="Omsk" pageId="2" pageNumber="143">Omsk</collectingRegion>
, the Irtysh River has a strong current but not fast flowing, approximately
<quantity id="4CA99B318766FFBBFF05F92A9D78F926" box="[151,283,1760,1785]" metricMagnitude="2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="2" pageNumber="143" unit="m" value="400.0" valueMax="500.0" valueMin="300.0">300500 m</quantity>
wide, with sandy loam (a mixture of clay and sand) bottom and banks and rather turbid water. According to the limited observations by OK at
<collectingRegion id="4995F8368766FFBBFD26F8CE9E99F8C3" box="[692,762,1796,1821]" country="Russia" name="Omsk" pageId="2" pageNumber="143">Omsk</collectingRegion>
,
<taxonomicName id="4C514D578766FFBBFC97F8CF9FFEF8C3" box="[773,925,1796,1821]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="143" phylum="Arthropoda" rank="species" species="araneoides">
<emphasis id="B925EAC68766FFBBFC97F8CF9FFEF8C3" box="[773,925,1796,1821]" italics="true" pageId="2" pageNumber="143">T. araneoides</emphasis>
</taxonomicName>
inhabits the main stem of the river, including at the city center, as well as large side channels. Adults of
<taxonomicName id="4C514D578766FFBBFCD6F8E39F83F89F" box="[836,992,1832,1857]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="143" phylum="Arthropoda" rank="species" species="araneoides">
<emphasis id="B925EAC68766FFBBFCD6F8E39F83F89F" box="[836,992,1832,1857]" italics="true" pageId="2" pageNumber="143">T. araneoides</emphasis>
</taxonomicName>
began emerging immediately after ice break in late April, caused by a sudden increase in discharge and followed by massive drifting of ice (Fig. 4). This sudden release was initiated several days before natural ice breakup by use of explosives upstream from the city to break up the ice to reduce flooding. In the city centre, the ice does not cover the river completely throughout the winter. Usually a large open water area along the right bank occurs because of inflow of warmer municipal waste water discharge. After ice break, or occasionally before at open water areas, larvae could be observed swimming at the water surface towards the bank by intensively wiggling their bodies, often in large numbers until early May. The following dates of the larval observations were:
<date id="FFEF10148767FFBAFD65FF769F18FF0B" box="[759,891,188,213]" pageId="3" pageNumber="144" value="1975-04-13">13 IV 1975</date>
(one individual) (ice break on
<date id="FFEF10148767FFBAFB70FF769909FF0B" box="[1250,1386,188,213]" pageId="3" pageNumber="144" value="1975-04-17">17 IV 1975</date>
); 22 (one) and 24 (several)
<date id="FFEF10148767FFBAFE0DFF2A9D9CFF26" box="[415,511,224,248]" pageId="3" pageNumber="144" value="1977-04">IV 1977</date>
(ice break on
<date id="FFEF10148767FFBAFD38FF2A9F50FF26" box="[682,819,224,248]" pageId="3" pageNumber="144" value="1977-04-18">18 IV 1977</date>
);
<date id="FFEF10148767FFBAFCD8FF2A9F9AFF26" box="[842,1017,224,248]" pageId="3" pageNumber="144" value="1978-04-28" valueMax="1978-04-29" valueMin="1978-04-28">2829 IV 1978</date>
(one each day) (ice break on
<date id="FFEF10148767FFBAFACBFF2A9CB0FEC2" pageId="3" pageNumber="144" value="1978-04-27">27 IV 1978</date>
); 23 (one) and 29 (numerous) IV, 6 (extremely numerous) and 8 (few)
<date id="FFEF10148767FFBAFC6AFECF982CFEC2" box="[1016,1103,260,284]" pageId="3" pageNumber="144" value="1979-05">V 1979</date>
. Upon reaching the bank, the larvae move to concrete structures for 1020 minutes until finding shelter in cracks or crevices in the concrete or among driftwood or gravel, where adult eclosion takes place. Apparently, female larvae eclose only into adults in deeper, shaded-sheltered sites, whereas male larvae eclose among gravel or even in more exposed areas. Three larvae were observed after reaching the concreted river embankment on
<date id="FFEF10148767FFBAFC2DFE5E98FFFE72" box="[959,1180,404,429]" pageId="3" pageNumber="144" value="1979-04-29" valueMax="1979-05-06" valueMin="1979-04-29">29 IV and 6 V 1979</date>
crawling
<quantity id="4CA99B318767FFBAFA9EFE5E9930FE72" box="[1292,1363,404,428]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="3" pageNumber="144" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">12 m</quantity>
above the water, then turning and approaching the water again and eventually eclosing close to proximity of the waters edge.
</paragraph>
<paragraph id="8BEE36D48767FFBAFF55FE359EFFFC7B" blockId="3.[151,1437,151,933]" pageId="3" pageNumber="144">
The dates of observation of adults were as follows:
<date id="FFEF10148767FFBAFC80FDCA9FADFDC6" box="[786,974,511,536]" pageId="3" pageNumber="144" value="1975-04-24" valueMax="1975-05-05" valueMin="1975-04-24">24 IV5 V 1975</date>
;
<date id="FFEF10148767FFBAFC49FE359803FDC6" box="[987,1120,511,536]" pageId="3" pageNumber="144" value="1977-04-26">26 IV 1977</date>
;
<date id="FFEF10148767FFBAFBFFFDCA9978FDC6" box="[1133,1307,512,536]" pageId="3" pageNumber="144" value="1978-04-28" valueMax="1978-04-29" valueMin="1978-04-28">2829 IV 1978</date>
;
<date id="FFEF10148767FFBAFABAFE359CB0FDE2" pageId="3" pageNumber="144" value="1979-04-26" valueMax="1979-05-08" valueMin="1979-04-26">26 IV8 v 1979</date>
;
<date id="FFEF10148767FFBAFF73FDEE9D0DFDE3" box="[225,366,548,573]" pageId="3" pageNumber="144" value="1995-05-02" valueMax="1995-05-04" valueMin="1995-05-02">24 V 1995</date>
;
<date id="FFEF10148767FFBAFEEFFDEE9E68FDE2" box="[381,523,548,572]" pageId="3" pageNumber="144" value="2003-05-01" valueMax="2003-05-03" valueMin="2003-05-01">13 V 2003</date>
;
<date id="FFEF10148767FFBAFD8BFDEE9EB4FDE3" box="[537,727,548,573]" pageId="3" pageNumber="144" value="2006-04-29" valueMax="2006-05-02" valueMin="2006-04-29">29 IV2 v 2006</date>
;
<date id="FFEF10148767FFBAFD77FDEE9F17FDE2" box="[741,884,548,572]" pageId="3" pageNumber="144" value="2007-05-01" valueMax="2007-05-04" valueMin="2007-05-01">14 V 2007</date>
. On
<date id="FFEF10148767FFBAFC3FFDEE9855FDE3" box="[941,1078,548,573]" pageId="3" pageNumber="144" value="1975-04-24">24 IV 1975</date>
, a male was found active on a concrete parapet when the air temperature was 0˚ C. In the first days of emergence, the males predominated, walking over the sandy river bank or on concrete embankments (e.g. Fig. 4) a dozen meters from the water edge, and congregating up to 3040 individuals under driftwood or in cracks or crevices in the concrete embankments near the waters edge. In these aggregations, a portion of males had the epiproct still unfolded and a few were teneral (e.g. on
<date id="FFEF10148767FFBAFEDFFD129DDEFD2E" box="[333,445,727,752]" pageId="3" pageNumber="144" value="1995-05-02">2 V 1995</date>
,
<date id="FFEF10148767FFBAFE59FD129E5EFD2E" box="[459,573,728,752]" pageId="3" pageNumber="144" value="2003-05-03">3 V 2003</date>
). On two separate occasions (on
<date id="FFEF10148767FFBAFC29FD1298C8FD2E" box="[955,1195,727,752]" pageId="3" pageNumber="144" value="1979-04-24" valueMax="1979-05-04" valueMin="1979-04-24">24 IV and 4 V 1979</date>
), captive males were observed to stop walking from time to time for a brief period to produce several quick drumming beats with the tip of abdomen upon substrate, as has been reported in this genus (Stewart et al. 1998). Mating was observed once on
<date id="FFEF10148767FFBAFF05FC8E9D67FC82" box="[151,260,836,860]" pageId="3" pageNumber="144" value="1979-05-04">4 V 1979</date>
. Males appeared uninterested in all females, but one with crumpled wings and yellowish liquid under it (just eclosed or injured?). Several males mounted her one by one for a short period of time, whereas one was observed mounted for a longer period of time.
</paragraph>
<caption id="DF2E665C8767FFBAFF05F8579EC2F82F" httpUri="https://zenodo.org/record/438234/files/figure.png" pageId="3" pageNumber="144" targetBox="[340,1245,951,1912]" targetPageId="3">
<paragraph id="8BEE36D48767FFBAFF05F8579EC2F82F" blockId="3.[151,1436,1949,2033]" pageId="3" pageNumber="144">
<emphasis id="B925EAC68767FFBAFF05F8579D70F86C" bold="true" box="[151,275,1949,1971]" pageId="3" pageNumber="144">FIGURE 2.</emphasis>
Details of
<taxonomicName id="4C514D578767FFBAFE14F8549EE3F86D" box="[390,640,1949,1971]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="144" phylum="Arthropoda" rank="species" species="araneoides">
<emphasis id="B925EAC68767FFBAFE14F8549EE3F86D" box="[390,640,1949,1971]" italics="true" pageId="3" pageNumber="144">Taeniopteryx araneoides</emphasis>
</taxonomicName>
males from Omsk, 3 v 2003: ae—mature male, f—male with a not yet fully expanded epiproct; a—general habitus; b—wings; c—end of abdomen in lateral view; d—the same, dorsal view; e—the same, ventral view; f—laterodorsal view; scale bar 1 mm.
</paragraph>
</caption>
<caption id="DF2E665C8760FFBDFF05F87C9F45F835" httpUri="https://zenodo.org/record/438235/files/figure.png" pageId="4" pageNumber="145" targetBox="[418,1144,193,1886]" targetPageId="4">
<paragraph id="8BEE36D48760FFBDFF05F87C9F45F835" blockId="4.[151,1436,1974,2027]" pageId="4" pageNumber="145">
<emphasis id="B925EAC68760FFBDFF05F87C9D7AF815" bold="true" box="[151,281,1974,1996]" pageId="4" pageNumber="145">FIGURE 3.</emphasis>
Detail of a female of
<taxonomicName id="4C514D578760FFBDFD8FF87D9F7DF812" box="[541,798,1974,1996]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="145" phylum="Arthropoda" rank="species" species="araneoides">
<emphasis id="B925EAC68760FFBDFD8FF87D9F7DF812" box="[541,798,1974,1996]" italics="true" pageId="4" pageNumber="145">Taeniopteryx araneoides</emphasis>
</taxonomicName>
from Omsk, 3 v 2003: a—wings; b—end of abdomen in dorsoposterior view; d—the same, ventral view; scale bar 1 mm.
</paragraph>
</caption>
<paragraph id="8BEE36D48761FFBCFF05FA3B9DA8F99A" pageId="5" pageNumber="146">
<table id="F951C47487610046FF05FA3B99FFF99A" box="[151,1436,1521,1604]" gridcols="3" gridrows="3" pageId="5" pageNumber="146">
<tr id="3561349687610046FF05FA3B99FFF9D9" box="[151,1436,1521,1543]" gridrow="0" pageId="5" pageNumber="146">
<th id="76B05DEA87610046FF05FA3B9DBDF9D9" box="[151,478,1521,1543]" gridcol="0" gridrow="0" pageId="5" pageNumber="146">
<emphasis id="B925EAC68761FFBCFF05FA3B9D7BF9D8" bold="true" box="[151,280,1521,1543]" pageId="5" pageNumber="146">FIGURE 4.</emphasis>
Ice break up and
</th>
<th id="76B05DEA87610046FE75FA3B9F19F9D9" box="[487,890,1521,1543]" gridcol="1" gridrow="0" pageId="5" pageNumber="146">drift at the Irtysh River in the Omsk</th>
<th id="76B05DEA87610046FC13FA3B99FFF9D9" box="[897,1436,1521,1543]" gridcol="2" gridrow="0" pageId="5" pageNumber="146">city centre (12 IV 2008) triggering emergence of</th>
</tr>
<tr id="3561349687610046FF05F9DA99FFF9F8" box="[151,1436,1552,1574]" gridrow="1" pageId="5" pageNumber="146">
<th id="76B05DEA87610046FF05F9DA9DBDF9F8" box="[151,478,1552,1574]" gridcol="0" gridrow="1" pageId="5" pageNumber="146">
<taxonomicName id="4C514D578761FFBCFF05F9DB9DF0F9F8" box="[151,403,1552,1574]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="146" phylum="Arthropoda" rank="species" species="araneoides">
<emphasis id="B925EAC68761FFBCFF05F9DB9DF0F9F8" box="[151,403,1552,1574]" italics="true" pageId="5" pageNumber="146">Taeniopteryx araneoides</emphasis>
</taxonomicName>
, adults
</th>
<td id="76B05DEA87610046FE75F9DA9F19F9F8" box="[487,890,1552,1574]" gridcol="1" gridrow="1" pageId="5" pageNumber="146">appearing in the following days on the</td>
<td id="76B05DEA87610046FC13F9DA99FFF9F8" box="[897,1436,1552,1574]" gridcol="2" gridrow="1" pageId="5" pageNumber="146">concrete embankments of the right bank seen in the</td>
</tr>
<tr id="3561349687610046FF05F9E499FFF99A" box="[151,1436,1582,1604]" gridrow="2" pageId="5" pageNumber="146" rowspan-1="1" rowspan-2="1">
<th id="76B05DEA87610046FF05F9E49DBDF99A" box="[151,478,1582,1604]" gridcol="0" gridrow="2" pageId="5" pageNumber="146">central part of the photograph.</th>
</tr>
</table>
</paragraph>
<paragraph id="8BEE36D48761FFBCFF55F9B89F0FF8C4" blockId="5.[151,1436,1649,2034]" pageId="5" pageNumber="146">
At later dates of adult occurrence, females became more abundant. In sunny weather they could be observed gliding towards the river bank fluttering their wings over the water surface. Several females would be observed resting on the water surface for periods of time. Females would fold their wings when reaching the bank (observations of
<date id="FFEF10148761FFBCFECBF9149DA4F928" box="[345,455,1758,1782]" pageId="5" pageNumber="146" value="2003-05-01">1 V 2003</date>
and
<date id="FFEF10148761FFBCFD96F9149E14F928" box="[516,631,1758,1782]" pageId="5" pageNumber="146" value="2007-05-04">4 V 2007</date>
). Only once (
<date id="FFEF10148761FFBCFC84F9149FE4F928" box="[790,903,1758,1782]" pageId="5" pageNumber="146" value="2003-05-03">3 V 2003</date>
) a female was observed flying over the water directly to the river bank, most likely returning after oviposition.
</paragraph>
<paragraph id="8BEE36D48761FFBFFF55F8EF9F4FFF26" blockId="5.[151,1436,1649,2034]" lastBlockId="6.[151,1437,151,2028]" lastPageId="6" lastPageNumber="147" pageId="5" pageNumber="146">
According to observations by NA,
<taxonomicName id="4C514D578761FFBCFDF3F8EC9E9EF8E0" box="[609,765,1829,1854]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="146" phylum="Arthropoda" rank="species" species="araneoides">
<emphasis id="B925EAC68761FFBCFDF3F8EC9E9EF8E0" box="[609,765,1829,1854]" italics="true" pageId="5" pageNumber="146">T. araneoides</emphasis>
</taxonomicName>
was abundant at the Irtysh River upstream of the city of Pavlodar on
<date id="FFEF10148761FFBCFEB4F8809DB0F8BC" box="[294,467,1865,1890]" pageId="5" pageNumber="146" value="2014-06-19" valueMax="2014-06-30" valueMin="2014-06-19">1930 VI 2014</date>
2016, but has not been collected within the city limits.
<taxonomicName id="4C514D578761FFBCFBC3F8809904F8BC" box="[1105,1383,1865,1890]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="146" phylum="Arthropoda" rank="species" species="araneoides">
<emphasis id="B925EAC68761FFBCFBC3F8809904F8BC" box="[1105,1383,1865,1890]" italics="true" pageId="5" pageNumber="146">Taeniopteryx araneoides</emphasis>
</taxonomicName>
also occurred downstream of the city but surrounding steep bluffs made observations difficult including attempts to estimate the relative abundance of this species. Females were found on grasses (mostly
<taxonomicName id="4C514D578761FFBCFB59F858994AF874" box="[1227,1321,1938,1962]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="5" pageNumber="146" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
) growing immediately at the river edge. During the first days of adult emergence, many males would hide in loose sand of a small bluff, later wandering in search of females and periodically drum. Adults of
<taxonomicName id="4C514D578761FFBCFBBEF81098A6F82C" box="[1068,1221,2009,2034]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="146" phylum="Arthropoda" rank="species" species="araneoides">
<emphasis id="B925EAC68761FFBCFBBEF81098A6F82C" box="[1068,1221,2009,2034]" italics="true" pageId="5" pageNumber="146">T. araneoides</emphasis>
</taxonomicName>
were found mostly in the morning. No larvae were found at depths up to 0.
<specimenCount id="9D57FD5D8762FFBFFCA9FF5D9F11FF71" box="[827,882,151,176]" pageId="6" pageNumber="147" type="generic">5 m.</specimenCount>
It is noteworthy that at Pavlodar, not a single individual was found along the left bank of the Irtysh River, possibly because the main flow of the river is close to the right bank, whereas at the left bank, the current is slow.
</paragraph>
<paragraph id="8BEE36D48762FFBFFF55FECE9FB1FD7D" blockId="6.[151,1437,151,2028]" pageId="6" pageNumber="147">
<emphasis id="B925EAC68762FFBFFF55FECE9D2FFEC3" bold="true" box="[199,332,260,285]" pageId="6" pageNumber="147">Discussion.</emphasis>
The
<quantity id="4CA99B318762FFBFFE14FECE9D93FEC3" box="[390,496,260,285]" metricMagnitude="3" metricUnit="m" metricValue="4.248" pageId="6" pageNumber="147" unit="km" value="4.248">4,248 km</quantity>
long Irtysh River flows from its source at the western slope of the Mongolian Altai Mountains at 4725' N, 9013'' in the Xinjiang Autonomous Region of China, traversing through the territory of Kazakhstan to Russia, passing the large Lake Zaisan and a series of three adjacent Bukhtarma, UstKamenogorsk and Shulbinsk reservoirs, and enters the Ob River at 6105' N, 6850''. There are five large cities along the river, Oskemen (Ust-Kamenogorsk), Semey (Semipalatinsk) and Pavlodar in Kazakhstan and Omsk and Khanty- Mansiysk (at the mouth) in Russia.
<taxonomicName id="4C514D578762FFBFFDB0FE7C9F54FE10" box="[546,823,437,462]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="araneoides">
<emphasis id="B925EAC68762FFBFFDB0FE7C9F54FE10" box="[546,823,437,462]" italics="true" pageId="6" pageNumber="147">Taeniopteryx araneoides</emphasis>
</taxonomicName>
is not known from the Irtysh River further upstream of Pavlodar, e.g. at the city of Oskemen, where the river becomes more rapid, as well as downstream of Omsk.
<bibRefCitation id="EFC04B258762FFBFFF05FE369D08FDCB" author="Devyatkov" box="[151,363,508,533]" pageId="6" pageNumber="147" refString="Devyatkov, V. I. (2002) Pervye svedeniya po faune (Insecta: Plecoptera) basseyna verkhnego Irtysha. Zoologicheskie issledovaniya v Kazakhstane: sostoyaniye i perspektivy (proceedings of international conference), Almaty, 2002, 225 - 227. [in Russian]" type="book chapter" year="2002">Devyatkov (2002)</bibRefCitation>
did not include
<taxonomicName id="4C514D578762FFBFFDD3FE379E81FDCB" box="[577,738,508,533]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="araneoides">
<emphasis id="B925EAC68762FFBFFDD3FE379E81FDCB" box="[577,738,508,533]" italics="true" pageId="6" pageNumber="147">T. araneoides</emphasis>
</taxonomicName>
in his checklist of stoneflies of the Upper Irtysh Basin. Additionally, occurrence of this species has not been confirmed from the large Yenisei River in Central Siberia, as suggested by a record by
<bibRefCitation id="EFC04B258762FFBFFE2FFD899EADFD82" author="Zapekina-Dulkeit" box="[445,718,579,604]" pageId="6" pageNumber="147" refString="Zapekina-Dulkeit, Y. I. (1957) Vesnyanki Altae-Sayanskikh gornotaezhnykh vodoemov. In: Abstracts of a conference of zoologists of Siberia. AN SSSR, Novosibirsk, pp. 27 - 28. [in Russian]" type="book chapter" year="1957">Zapekina-Dulkeit (1957</bibRefCitation>
,
<bibRefCitation id="EFC04B258762FFBFFD4EFD899F42FD85" author="Zapekina-Dulkeit" box="[732,801,579,603]" pageId="6" pageNumber="147" refString="Zapekina-Dulkeit, Y. I. (1971 a) Dopolnenie k faune vesnyanok (Plecoptera) zapovednika &quot; Stolby i smezhnykh territoriy. Trudy gosudarstvennogo zapovednika &quot; Stolby, 8, 88 - 90 [in Russian]" type="journal article" year="1971" yearSuffix="a">1971a</bibRefCitation>
) from Krasnoyarsk, without specifying the name of the river, half a century earlier. Recent studies of stoneflies at the origin of the Yenisei River at the junction of the Biy Khem and Kaa Khem rivers in Tuva (
<bibRefCitation id="EFC04B258762FFBFFDAEFD409EDEFD7C" author="Zaika" box="[572,701,650,675]" pageId="6" pageNumber="147" refString="Zaika, V. V. (2011) The stoneflies (Insecta, Ectognatha, Plecoptera) of Maliy Enisey River (Tyva). Euroasian Entomological Journal, 10 (1), 10 - 14. [in Russian, English summary]" type="journal article" year="2011">Zaika 2011</bibRefCitation>
) did not list this species.
</paragraph>
<paragraph id="8BEE36D48762FFBFFF55FD649D8BFC40" blockId="6.[151,1437,151,2028]" pageId="6" pageNumber="147">
<taxonomicName id="4C514D578762FFBFFF55FD649DBEFD18" box="[199,477,685,710]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="araneoides">
<emphasis id="B925EAC68762FFBFFF55FD649DBEFD18" box="[199,477,685,710]" italics="true" pageId="6" pageNumber="147">Taeniopteryx araneoides</emphasis>
</taxonomicName>
is obviously a species of large rivers (a potamon species), as the Danube, Dniester, Elbe rivers in Europe, the Irtysh River in West Siberia and
<collectingCountry id="F34676448762FFBFFCAEFD1B9FDCFD34" box="[828,959,721,746]" name="Kazakhstan" pageId="6" pageNumber="147">Kazakhstan</collectingCountry>
and perhaps, the Yenisei River in Central Siberia. These rivers historically have been impacted by water pollution likely causing the extinction of of
<taxonomicName id="4C514D578762FFBFFA15FD3C9D6AFCED" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="araneoide">
<emphasis id="B925EAC68762FFBFFA15FD3C9D6AFCED" italics="true" pageId="6" pageNumber="147">T. araneoide</emphasis>
</taxonomicName>
s in Central Europe during the 20th century.
<collectingRegion id="4995F8368762FFBFFD65FCD09F59FCED" box="[759,826,794,819]" country="Russia" name="Omsk" pageId="6" pageNumber="147">Omsk</collectingRegion>
is a city with a population of 1.18 million people and includes a large industrial centre. Nevertheless,
<taxonomicName id="4C514D578762FFBFFD29FCF59F36FC89" box="[699,853,830,855]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="araneoides">
<emphasis id="B925EAC68762FFBFFD29FCF59F36FC89" box="[699,853,830,855]" italics="true" pageId="6" pageNumber="147">T. araneoides</emphasis>
</taxonomicName>
appears to be surviving well, even at the center of city. This species, however, has not been collected within the city limits of Pavlodar, a city of third the size, with a population of 335,000 people.
</paragraph>
<paragraph id="8BEE36D48762FFBFFF55FC629E79FB03" blockId="6.[151,1437,151,2028]" pageId="6" pageNumber="147">
The geographical distribution of
<taxonomicName id="4C514D578762FFBFFDDCFC639E8EFC1F" box="[590,749,936,961]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="araneoides">
<emphasis id="B925EAC68762FFBFFDDCFC639E8EFC1F" box="[590,749,936,961]" italics="true" pageId="6" pageNumber="147">T. araneoides</emphasis>
</taxonomicName>
appears to be enigmatic, with a disjunction of
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between the Dniester and Irtysh rivers, and the seeming absence of this species from the Ob River at the city of Novosibirsk in West Siberia situated at the same latitude. The Ob River is similar to Irtysh River and is just
<quantity id="4CA99B318762FFBFFADAFC2599FFFBD6" box="[1352,1436,1007,1032]" metricMagnitude="5" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="147" unit="km" value="600.0">600 km</quantity>
east of Omsk. The reports of
<taxonomicName id="4C514D578762FFBFFE75FBDE9EE2FBF2" box="[487,641,1043,1068]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="araneoides">
<emphasis id="B925EAC68762FFBFFE75FBDE9EE2FBF2" box="[487,641,1043,1068]" italics="true" pageId="6" pageNumber="147">T. araneoides</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFC04B258762FFBFFD3FFBD99FDCFBF2" author="Zapekina-Dulkeit" box="[685,959,1043,1068]" pageId="6" pageNumber="147" refString="Zapekina-Dulkeit, Y. I. (1957) Vesnyanki Altae-Sayanskikh gornotaezhnykh vodoemov. In: Abstracts of a conference of zoologists of Siberia. AN SSSR, Novosibirsk, pp. 27 - 28. [in Russian]" type="book chapter" year="1957">Zapekina-Dulkeit (1957</bibRefCitation>
,
<bibRefCitation id="EFC04B258762FFBFFC5EFBD99871FBF5" author="Zapekina-Dulkeit" box="[972,1042,1043,1067]" pageId="6" pageNumber="147" refString="Zapekina-Dulkeit, Y. I. (1971 a) Dopolnenie k faune vesnyanok (Plecoptera) zapovednika &quot; Stolby i smezhnykh territoriy. Trudy gosudarstvennogo zapovednika &quot; Stolby, 8, 88 - 90 [in Russian]" type="journal article" year="1971" yearSuffix="a">1971a</bibRefCitation>
,
<bibRefCitation id="EFC04B258762FFBFFB8DFBD9984DFBF2" author="Zapekina-Dulkeit" box="[1055,1070,1043,1068]" pageId="6" pageNumber="147" refString="Zapekina-Dulkeit, Y. I. (1971 b) Vesnyanki (Plecoptera) Sibiri. TXIII International Congress of Entomology, Moscow, 2 - 9 August, Abstracts, 1, 223 - 224. [in Russian]" type="journal article" year="1971" yearSuffix="b">b</bibRefCitation>
) from Krasnoyarsk (further
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NEE of Novosibirsk), however, suggests that
<taxonomicName id="4C514D578762FFBFFD5EFBFD9F05FB91" box="[716,870,1078,1103]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="araneoides">
<emphasis id="B925EAC68762FFBFFD5EFBFD9F05FB91" box="[716,870,1078,1103]" italics="true" pageId="6" pageNumber="147">T. araneoides</emphasis>
</taxonomicName>
may occur at the next major river to the east, the Yenisei. The common Palearctic
<taxonomicName id="4C514D578762FFBFFD9BFB919EEAFBAD" box="[521,649,1114,1139]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="nebulosa">
<emphasis id="B925EAC68762FFBFFD9BFB919EEAFBAD" box="[521,649,1114,1139]" italics="true" pageId="6" pageNumber="147">T. nebulosa</emphasis>
</taxonomicName>
is distributed from the Atlantic to Pacific coasts of Eurasia (
<bibRefCitation id="EFC04B258762FFBFFABDFB909CB0FB48" author="Zhiltzova" pageId="6" pageNumber="147" refString="Zhiltzova, L. A. (2003) Insecta Plecoptera. Plecoptera, Gruppe Euholognatha. Fauna of Russia and Neighbouring Countries, New Series, 145, 1 - 538. [in Russian, English / German title]" type="journal article" year="2003">Zhiltzova 2003</bibRefCitation>
), but has not been recorded from the Irtysh River either at Pavlodar or Omsk, but is common at the Ob River and its ttributaries, the Inya River and Berd River. There is also a female specimen in the ISEA collected at the junction of the Inya and Ob rivers.
</paragraph>
<paragraph id="8BEE36D48762FFBFFF55FB2D9C9CF9C1" blockId="6.[151,1437,151,2028]" pageId="6" pageNumber="147">
Another possible cause of the extinction of
<taxonomicName id="4C514D578762FFBFFD23FB239F2AFADE" box="[689,841,1255,1280]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="araneoides">
<emphasis id="B925EAC68762FFBFFD23FB239F2AFADE" box="[689,841,1255,1280]" italics="true" pageId="6" pageNumber="147">T. araneoides</emphasis>
</taxonomicName>
in Europe may be directly related to river regulation by dams and reservoirs preventing natural spring flooding that may serve as a trigger for adult emergence as supported by observations at Omsk. In this respect it is important to note that middle reach of the Irtysh River is still free flowing, with Pavlodar located
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and Omsk
<quantity id="4CA99B318762FFBFFC13FA999FBCFAB5" box="[897,991,1362,1387]" metricMagnitude="5" metricUnit="m" metricValue="7.3" pageId="6" pageNumber="147" unit="km" value="730.0">730 km</quantity>
(air km) from the nearest upstream impoundment, the Shulbinsk Dam and reservoir. In contrast, the Ob River is dammed within the city limits of Novosibirsk, forming the large Obskoe Reservoir. It is also noteworthy that the reports of
<taxonomicName id="4C514D578762FFBFFB2CFA509938FA6C" box="[1214,1371,1433,1458]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="araneoides">
<emphasis id="B925EAC68762FFBFFB2CFA509938FA6C" box="[1214,1371,1433,1458]" italics="true" pageId="6" pageNumber="147">T. araneoides</emphasis>
</taxonomicName>
from Krasnoyarsk are from
<date id="FFEF10148762FFBFFE07FA759E70FA06" box="[405,531,1471,1496]" pageId="6" pageNumber="147" value="1957" valueMax="1971">19571971</date>
(
<bibRefCitation id="EFC04B258762FFBFFDB0FA759F4FFA06" author="Zapekina-Dulkeit" box="[546,812,1471,1496]" pageId="6" pageNumber="147" refString="Zapekina-Dulkeit, Y. I. (1957) Vesnyanki Altae-Sayanskikh gornotaezhnykh vodoemov. In: Abstracts of a conference of zoologists of Siberia. AN SSSR, Novosibirsk, pp. 27 - 28. [in Russian]" type="book chapter" year="1957">Zapekina-Dulkeit 1957</bibRefCitation>
,
<bibRefCitation id="EFC04B258762FFBFFCAAFA0A9FE3FA06" author="Zapekina-Dulkeit" box="[824,896,1472,1496]" pageId="6" pageNumber="147" refString="Zapekina-Dulkeit, Y. I. (1971 a) Dopolnenie k faune vesnyanok (Plecoptera) zapovednika &quot; Stolby i smezhnykh territoriy. Trudy gosudarstvennogo zapovednika &quot; Stolby, 8, 88 - 90 [in Russian]" type="journal article" year="1971" yearSuffix="a">1971a</bibRefCitation>
), the Krasnoyarsk Hydropower Station (the 10th largest in the world) was constructed in
<date id="FFEF10148762FFBFFDCAFA299EBAFA22" box="[600,729,1507,1532]" pageId="6" pageNumber="147" value="1956" valueMax="1972">19561972</date>
, with
<taxonomicName id="4C514D578762FFBFFC8FFA2E9FD5FA22" box="[797,950,1507,1532]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="araneoides">
<emphasis id="B925EAC68762FFBFFC8FFA2E9FD5FA22" box="[797,950,1507,1532]" italics="true" pageId="6" pageNumber="147">T. araneoides</emphasis>
</taxonomicName>
collected just before the Yenisei River was dammed.
</paragraph>
<paragraph id="8BEE36D48762FFBEFF55F9E19FC1FEC5" blockId="6.[151,1437,151,2028]" lastBlockId="7.[151,1436,151,462]" lastPageId="7" lastPageNumber="148" pageId="6" pageNumber="147">
<taxonomicName id="4C514D578762FFBFFF55F9E19DB8F99C" box="[199,475,1577,1602]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="araneoides">
<emphasis id="B925EAC68762FFBFFF55F9E19DB8F99C" box="[199,475,1577,1602]" italics="true" pageId="6" pageNumber="147">Taeniopteryx araneoides</emphasis>
</taxonomicName>
and
<taxonomicName id="4C514D578762FFBFFD83F9E19EF2F99C" box="[529,657,1577,1602]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="nebulosa">
<emphasis id="B925EAC68762FFBFFD83F9E19EF2F99C" box="[529,657,1577,1602]" italics="true" pageId="6" pageNumber="147">T. nebulosa</emphasis>
</taxonomicName>
may be competing with and excluding each other in large rivers. Two ecogeographical scenarios may posited: 1) Two closely related and ecologically similar species most likely did not disperse over the Palaearctic simultaneously, whenever this dispersal occurred. It could be hypothesized that
<taxonomicName id="4C514D578762FFBFFA15F9B89D77F973" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="araneoides">
<emphasis id="B925EAC68762FFBFFA15F9B89D77F973" italics="true" pageId="6" pageNumber="147">T. araneoides</emphasis>
</taxonomicName>
dispersed initially over its former range. Subsequently,
<taxonomicName id="4C514D578762FFBFFC02F95F986CF973" box="[912,1039,1684,1709]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="nebulosa">
<emphasis id="B925EAC68762FFBFFC02F95F986CF973" box="[912,1039,1684,1709]" italics="true" pageId="6" pageNumber="147">T. nebulosa</emphasis>
</taxonomicName>
dispersed over the same extensive area and displaced
<taxonomicName id="4C514D578762FFBFFE12F9739E7DF90E" box="[384,542,1719,1744]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="araneoides">
<emphasis id="B925EAC68762FFBFFE12F9739E7DF90E" box="[384,542,1719,1744]" italics="true" pageId="6" pageNumber="147">T. araneoides</emphasis>
</taxonomicName>
from most large rivers.
<taxonomicName id="4C514D578762FFBFFCDEF9739805F90E" box="[844,1126,1719,1744]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="araneoides">
<emphasis id="B925EAC68762FFBFFCDEF9739805F90E" box="[844,1126,1719,1744]" italics="true" pageId="6" pageNumber="147">Taeniopteryx araneoides</emphasis>
</taxonomicName>
apparently maintained its stronghold in the Irtysh River (and probably Yenisei River) as well as in Central Europe, but later disappeared from the latter, at least in part, because of anthropogenic impacts. The fact that
<taxonomicName id="4C514D578762FFBFFB9BF93598EDF8C9" box="[1033,1166,1790,1815]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="nebulosa">
<emphasis id="B925EAC68762FFBFFB9BF93598EDF8C9" box="[1033,1166,1790,1815]" italics="true" pageId="6" pageNumber="147">T. nebulosa</emphasis>
</taxonomicName>
still occurs in Europe, whereas
<taxonomicName id="4C514D578762FFBFFF6EF8E99DFBF8E5" box="[252,408,1826,1851]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="araneoides">
<emphasis id="B925EAC68762FFBFFF6EF8E99DFBF8E5" box="[252,408,1826,1851]" italics="true" pageId="6" pageNumber="147">T. araneoides</emphasis>
</taxonomicName>
became extinct, indirectly supports the assumption that
<taxonomicName id="4C514D578762FFBFFBB7F8E998CAF8E5" box="[1061,1193,1826,1851]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="nebulosa">
<emphasis id="B925EAC68762FFBFFBB7F8E998CAF8E5" box="[1061,1193,1826,1851]" italics="true" pageId="6" pageNumber="147">T. nebulosa</emphasis>
</taxonomicName>
is more ecologically adaptive or tolerate, inhabiting a wide range of streams, from slow large rivers to smaller fast flowing rivulets and brooks (
<bibRefCitation id="EFC04B258762FFBFFF63F8A39DC1F85F" author="Zhiltzova" box="[241,418,1897,1922]" pageId="6" pageNumber="147" refString="Zhiltzova, L. A. (2003) Insecta Plecoptera. Plecoptera, Gruppe Euholognatha. Fauna of Russia and Neighbouring Countries, New Series, 145, 1 - 538. [in Russian, English / German title]" type="journal article" year="2003">Zhiltzova 2003</bibRefCitation>
). Such smaller streams may serve as refugia for
<taxonomicName id="4C514D578762FFBFFC2EF8A09859F85C" box="[956,1082,1897,1922]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="nebulosa">
<emphasis id="B925EAC68762FFBFFC2EF8A09859F85C" box="[956,1082,1897,1922]" italics="true" pageId="6" pageNumber="147">T. nebulosa</emphasis>
</taxonomicName>
as water quality deteriorated in the larger streams and rivers due to anthropogenic perturbations. It is not surprising that
<taxonomicName id="4C514D578762FFBFFBE5F844996DF87B" box="[1143,1294,1932,1957]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="araneoides">
<emphasis id="B925EAC68762FFBFFBE5F844996DF87B" box="[1143,1294,1932,1957]" italics="true" pageId="6" pageNumber="147">T. araneoides</emphasis>
</taxonomicName>
, confined to large rivers, has persisted only in relatively pristine and unimpacted large lotic systems of Siberia and
<collectingCountry id="F34676448762FFBFFA80F86599FBF816" box="[1298,1432,1967,1992]" name="Kazakhstan" pageId="6" pageNumber="147">Kazakhstan</collectingCountry>
, but disappeared from Europe. This interpretation assumes that
<taxonomicName id="4C514D578762FFBFFCCEF81E9F95F832" box="[860,1014,2003,2028]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="araneoides">
<emphasis id="B925EAC68762FFBFFCCEF81E9F95F832" box="[860,1014,2003,2028]" italics="true" pageId="6" pageNumber="147">T. araneoides</emphasis>
</taxonomicName>
is a relict species in currently known localities. 2) Both species could have dispersed over the Palaearctic at the same time or not, but
<taxonomicName id="4C514D578763FFBEFB47FF53990DFF6E" box="[1237,1390,151,176]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="148" phylum="Arthropoda" rank="species" species="araneoides">
<emphasis id="B925EAC68763FFBEFB47FF53990DFF6E" box="[1237,1390,151,176]" italics="true" pageId="7" pageNumber="148">T. araneoides</emphasis>
</taxonomicName>
is a species of large rivers and was more successful in these habitats than
<emphasis id="B925EAC68763FFBEFC75FF7699FFFF0A" box="[999,1436,187,212]" italics="true" pageId="7" pageNumber="148">
<taxonomicName id="4C514D578763FFBEFC75FF769812FF0A" box="[999,1137,187,212]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="148" phylum="Arthropoda" rank="species" species="nebulosa">T. nebulosa</taxonomicName>
.
<taxonomicName id="4C514D578763FFBEFB13FF7699FFFF0A" box="[1153,1436,187,212]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="148" phylum="Arthropoda" rank="species" species="araneoides">Taeniopteryx araneoides</taxonomicName>
</emphasis>
however was intolerant to river regulation and has been extirpated from these large rivers and far downstream from these impoundments, allowing
<taxonomicName id="4C514D578763FFBEFE67FEC99E16FEC5" box="[501,629,258,283]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="148" phylum="Arthropoda" rank="species" species="nebulosa">
<emphasis id="B925EAC68763FFBEFE67FEC99E16FEC5" box="[501,629,258,283]" italics="true" pageId="7" pageNumber="148">T. nebulosa</emphasis>
</taxonomicName>
to occupy this open niche.
</paragraph>
<paragraph id="8BEE36D48763FFBEFF55FEED98E8FE10" blockId="7.[151,1436,151,462]" pageId="7" pageNumber="148">
To test these hypotheses, more data of occurrence for both species in Siberia is required. One of the ways to test the above hypotheses would be reintroducing
<taxonomicName id="4C514D578763FFBEFD58FE819F00FEBC" box="[714,867,329,354]" class="Insecta" family="Taeniopterygidae" genus="Taeniopteryx" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="148" phylum="Arthropoda" rank="species" species="araneoides">
<emphasis id="B925EAC68763FFBEFD58FE819F00FEBC" box="[714,867,329,354]" italics="true" pageId="7" pageNumber="148">T. araneoides</emphasis>
</taxonomicName>
in the Danube River, now much less polluted than a century ago, using the Irtysh River population as a possible source. Hopefully, the Danube countries and European Union would be interested in such a restoration project attempting to recover their lost biodiversity. This attempt could reveal if ecological limiting factors for the species still occur in the Danube.
</paragraph>
</subSubSection>
</treatment>
</document>