treatments-xml/data/03/A8/A4/03A8A444FF8CC466FF08FBB0FAF7FD87.xml

<document id="1990774D49BFAEE5D2E4ED43C78111B3" ID-DOI="10.11646/phytotaxa.252.1.2" ID-ISSN="1179-3163" ID-Zenodo-Dep="13674915" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725405123220" checkinUser="felipe" docAuthor="Sang, Julia &amp; Kiew, Ruth" docDate="2016" docId="03A8A444FF8CC466FF08FBB0FAF7FD87" docLanguage="en" docName="phytotaxa.252.1.2.pdf" docOrigin="Phytotaxa 252 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.252.1.2" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Begonia compacta S. Julia &amp; Kiew 2016, spec. nov." docType="treatment" docVersion="2" lastPageNumber="25" masterDocId="FF91DC3CFF8AC46EFF80FFDBFFE9FFA6" masterDocTitle="Begonia (Begoniaceae) from Batang Ai National Park and vicinity, Sarawak, Borneo, including six new species" masterLastPageNumber="30" masterPageNumber="17" pageNumber="23" updateTime="1725426159086" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="FE04E47FD83AA66FEC13A90E2459B108">Begonia (Begoniaceae) from Batang Ai National Park and vicinity, Sarawak, Borneo, including six new species</mods:title>
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<mods:namePart id="296F5095DC16CF64DBC2399F448E4BC4">Sang, Julia</mods:namePart>
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<mods:namePart id="F69BE4F5615CE344E3AAB6E5848E0021">Kiew, Ruth</mods:namePart>
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<mods:title id="5622F6649952245DED29748300899AF9">Phytotaxa</mods:title>
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<mods:date id="05DE9E6B43286D9A4C9019D8BE72E015">2016</mods:date>
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<paragraph id="8BBE1552FF8CC468FF08FBB0FCF7FB23" blockId="6.[136,798,1130,1157]" box="[136,798,1130,1157]" pageId="6" pageNumber="23">
<heading id="D0F6A23EFF8CC468FF08FBB0FCF7FB23" box="[136,798,1130,1157]" fontSize="11" level="2" pageId="6" pageNumber="23" reason="3">
<emphasis id="B975C940FF8CC468FF08FBB0FF74FB23" bold="true" box="[136,157,1131,1157]" italics="true" pageId="6" pageNumber="23">5.</emphasis>
<taxonomicName id="4C016ED1FF8CC468FF24FBB1FDC7FB23" ID-CoL="7SF74" authority="S.Julia &amp; Kiew" authorityName="S. Julia &amp; Kiew" authorityYear="2016" box="[164,558,1130,1157]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="6" pageNumber="23" phylum="Tracheophyta" rank="species" species="compacta" status="sp. nov.">
<emphasis id="B975C940FF8CC468FF24FBB1FE93FB23" bold="true" box="[164,378,1130,1157]" italics="true" pageId="6" pageNumber="23">Begonia compacta</emphasis>
S.Julia &amp; Kiew
</taxonomicName>
, 
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<taxonomicNameLabel id="A246743BFF8CC468FDB9FBB7FD4EFB23" box="[569,679,1132,1157]" pageId="6" pageNumber="23" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="133A09D7FF8CC468FD35FBB7FCFFFB23" box="[693,790,1132,1157]" captionStart="FIGURE 4" captionStartId="7.[136,229,1680,1701]" captionTargetBox="[292,1300,398,1648]" captionTargetId="figure-101@7.[284,1304,389,1654]" captionTargetPageId="7" captionText="FIGURE 4. Begonia compacta. A. Habit; B. Stipule; C. Male inflorescence surrounded by persistent bracteoles; D. Female inflorescence surrounded by persistent bracteole; E. Female flower showing tepals and pistil; F. Fruits surrounded by persistent bracteole; G. Cross section of ovary. All photos from SFC 2751 by Julia Sang (A–F) and Ling Chea Yiing (G)." figureDoi="http://doi.org/10.5281/zenodo.13674927" httpUri="https://zenodo.org/record/13674927/files/figure.png" pageId="6" pageNumber="23">Figure 4</figureCitation>
)
</heading>
</paragraph>
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<subSubSection id="C31B46D9FF8CC468FF08FB4BFE9EFB0F" box="[136,375,1168,1193]" pageId="6" pageNumber="23" type="description">
<paragraph id="8BBE1552FF8CC468FF08FB4BFE9EFB0F" blockId="6.[136,375,1168,1193]" box="[136,375,1168,1193]" pageId="6" pageNumber="23">
<emphasis id="B975C940FF8CC468FF08FB4BFF36FB0F" bold="true" box="[136,223,1168,1193]" pageId="6" pageNumber="23">Section</emphasis>
<taxonomicName id="4C016ED1FF8CC468FF66FB4BFE9EFB0F" box="[230,375,1168,1193]" class="Magnoliopsida" family="Begoniaceae" genus="Petermannia" kingdom="Plantae" order="Cucurbitales" pageId="6" pageNumber="23" phylum="Tracheophyta" rank="genus">
<emphasis id="B975C940FF8CC468FF66FB4BFE9EFB0F" box="[230,375,1168,1193]" italics="true" pageId="6" pageNumber="23">Petermannia</emphasis>
</taxonomicName>
</paragraph>
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<subSubSection id="C31B46D9FF8CC468FF08FB01FCDEFAFD" pageId="6" pageNumber="23" type="diagnosis">
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<emphasis id="B975C940FF8CC468FF08FB01FF1AFB49" bold="true" box="[136,243,1242,1263]" pageId="6" pageNumber="23">Diagnosis:</emphasis>
—Similar to 
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<emphasis id="B975C940FF8CC468FEF1FB01FDF9FB49" box="[369,528,1242,1263]" italics="true" pageId="6" pageNumber="23">Begonia vulgare</emphasis>
S. 
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in habit, leaf shape and inflorescence that has a basal pair of female flowers and male flowers in a dense cluster above but it is different in its shorter stem to 
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tall (vs. to 
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tall in 
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<emphasis id="B975C940FF8CC468FAB9FB25FA74FAB5" box="[1337,1437,1278,1299]" italics="true" pageId="6" pageNumber="23">B. vulgare</emphasis>
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), longer petioles 
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long (vs. 
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), shorter inflorescences, 
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(vs. 
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) and the shorter fruit, 15–25 × 
<quantity id="4CF9B8B7FF8CC468FAEFFAF9FF0BFAFD" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="2.4" metricValueMin="1.2" pageId="6" pageNumber="23" unit="mm" value="18.0" valueMax="24.0" valueMin="12.0">12–24 mm</quantity>
(vs. the more oblong fruit, 35–45 × 
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in 
<taxonomicName id="4C016ED1FF8CC468FD46FA9DFCC0FAFD" authorityName="S. Julia &amp; Kiew" authorityYear="2013" box="[710,809,1350,1371]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="6" pageNumber="23" phylum="Tracheophyta" rank="species" species="vulgare">
<emphasis id="B975C940FF8CC468FD46FA9DFCC0FAFD" box="[710,809,1350,1371]" italics="true" pageId="6" pageNumber="23">B. vulgare</emphasis>
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).
</paragraph>
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<emphasis id="B975C940FF8CC468FF08FAB1FF2BFAD9" bold="true" box="[136,194,1386,1407]" pageId="6" pageNumber="23">Type:</emphasis>
— 
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. Borneo. 
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. 
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: 
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, 
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, 
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and 
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, 
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, 
<date id="FFBF3392FF8CC468FAD5FAB1FF06FA05" pageId="6" pageNumber="23" value="2015-08-01">
<collectingDate id="EFFBCA7AFF8CC468FAD5FAB1FF06FA05" pageId="6" pageNumber="23" value="2015-08-01">1 August 2015</collectingDate>
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, 
<emphasis id="B975C940FF8CC468FF79FA55FE25FA05" box="[249,460,1422,1443]" italics="true" pageId="6" pageNumber="23">
Julia et al. 
<specimenCode id="DBA7BD29FF8CC468FEEBFA55FE25FA05" box="[363,460,1422,1443]" collectionCode="SFC" country="China" httpUri="http://grbio.org/cool/myie-bfyd" name="Laboratory of Fishes" pageId="6" pageNumber="23">SFC 2751</specimenCode>
</emphasis>
(
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<collectionCode id="ED108D97FF8CC468FDB4FA55FD83FA05" box="[564,618,1422,1443]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" lsid="urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="6" pageNumber="23" type="Herbarium">SAR</collectionCode>
!; 
<typeStatus id="54BAABF0FF8CC468FDF7FA55FD2FFA05" box="[631,710,1422,1443]" pageId="6" pageNumber="23" type="isotype">isotypes</typeStatus>
<collectionCode id="ED108D97FF8CC468FD4CFA55FCE9FA05" box="[716,768,1422,1443]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15093" lsid="urn:lsid:biocol.org:col:15093" name="Forest Research Institute Malaysia" pageId="6" pageNumber="23" type="Herbarium">KEP</collectionCode>
!, 
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!)
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.
</paragraph>
<paragraph id="8BBE1552FF8CC469FF08FA0FFA7CFE83" blockId="6.[136,1452,1492,2093]" lastBlockId="7.[136,1452,160,365]" lastPageId="7" lastPageNumber="24" pageId="6" pageNumber="23">
Cane-like begonia 
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tall. Stems erect, reddish, 
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thick, little-branched, succulent, minutely pubescent; internodes 3–8(–16) cm long, thicker at nodes. Stipules brownish, lanceolate, 11–19(–28) × 5–8(–12) mm, margin entire, keeled, apex acuminate, caducous. Leaves alternate, distant, oblique, held horizontally; petioles reddish, (4–) 
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long, grooved, minutely pubescent; lamina plain green on both surfaces, in life succulent, drying papery, matt, ovate to obovate, 15–22 × 
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, broad side 
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wide, often markedly concave on the narrower side, base slightly cordate, sometimes slightly overlapping, basal lobes 
<quantity id="4CF9B8B7FF8CC468FCC9F953FC33F907" box="[841,986,1672,1697]" metricMagnitude="-3" metricUnit="m" metricValue="9.25" metricValueMax="11.0" metricValueMin="7.5" pageId="6" pageNumber="23" unit="cm" value="0.925" valueMax="1.1" valueMin="0.75">0.75–1.1 cm</quantity>
, margin shallowly dentate, apex sharply acuminate, 
<quantity id="4CF9B8B7FF8CC468FE88F977FE86F963" box="[264,367,1708,1733]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="6" pageNumber="23" unit="cm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 cm</quantity>
long; venation palmate-pinnate, veins pale green on both surfaces, minutely pubescent, 3–5 veins on either side of the midrib, 3–5 veins in the basal lobe, prominent on both surfaces. Inflorescences protogynous, axillary in the upper leaf axils, rachis 
<quantity id="4CF9B8B7FF8CC468FDC6F92FFD58F8AB" box="[582,689,1780,1805]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="5.0" metricValueMin="1.5" pageId="6" pageNumber="23" unit="cm" value="3.25" valueMax="5.0" valueMin="1.5">1.5–5 cm</quantity>
long, with a pair of female flowers below and a dense cluster of male flowers above, each cymule enveloped in upto 2 pairs of bracteoles. Bracts reddish, lanceolate, 12–20 × 
<quantity id="4CF9B8B7FF8CC468FAF4F8C3FF5FF8F3" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="6" pageNumber="23" unit="mm" value="10.0" valueMax="12.0" valueMin="8.0">8–12 mm</quantity>
, margin entire, caducous; bracteoles broadly lanceolate, setose at apex, increasing in size to 11–13 × 
<quantity id="4CF9B8B7FF8CC468FAB9F8E7FA4EF8F3" box="[1337,1447,1852,1877]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="6" pageNumber="23" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8–10 mm</quantity>
, then abruptly diminishing in size towards the developing apex. Male flowers with pale pink pedicels, 
<quantity id="4CF9B8B7FF8CC468FA8CF8BBFA83F8DF" box="[1292,1386,1888,1913]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="6" pageNumber="23" unit="mm" value="6.0" valueMax="8.0" valueMin="4.0">4–8 mm</quantity>
long; tepals 2, pale yellowish green, oval, 5–6 × 
<quantity id="4CF9B8B7FF8CC468FDFCF85FFD34F83B" box="[636,733,1924,1949]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="6" pageNumber="23" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 mm</quantity>
, margin entire, apex rounded; stamens 29–32, cluster globose, subsessile; filaments pale yellow, 
<quantity id="4CF9B8B7FF8CC468FD8AF873FD94F867" box="[522,637,1960,1985]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="6" pageNumber="23" unit="mm" value="0.9" valueMax="1.0" valueMin="0.8">0.8–1 mm</quantity>
long; anthers pale yellow, obovate, ca. 1.5 × 
<quantity id="4CF9B8B7FF8CC468FB05F873FB37F867" box="[1157,1246,1960,1985]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="23" unit="mm" value="0.8">0.8 mm</quantity>
, apex emarginate. Female flowers with greenish pedicels, 
<quantity id="4CF9B8B7FF8CC468FDC7F817FD4DF843" box="[583,676,1996,2021]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="6" pageNumber="23" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
long; ovary pale green, wings tips pale green, obovoid, 9–10 × 
<quantity id="4CF9B8B7FF8CC468FAF4F817FF5FF7AF" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="6" pageNumber="23" unit="mm" value="10.0" valueMax="12.0" valueMin="8.0">8–12 mm</quantity>
, wings 3, equal, locules 3, placentas 2 per locule; tepals 5, pale green, outer 4 tepals ovate, 7–8 × 
<quantity id="4CF9B8B7FF8CC468FB73F82BFABBF7AF" box="[1267,1362,2032,2057]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="6" pageNumber="23" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 mm</quantity>
, margin entire, apex broadly acute, inner tepals ovate, ca. 6 × 
<quantity id="4CF9B8B7FF8CC468FD52F7CFFCFDF78B" box="[722,788,2068,2093]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="23" unit="mm" value="5.0">5 mm</quantity>
, margin entire, apex rounded; styles 3, pale yellow, 
<quantity id="4CF9B8B7FF8CC468FAD0F7CFFA45F78B" box="[1360,1452,2068,2093]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="6" pageNumber="23" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
long, divided to base, deeply Y-shaped; stigma pale yellow with reddish stripe, papillose forming a continuous twisted band. Capsules single or a pair, pale green, triangular or sometimes to slightly oblong-triangular in outline, 15–25 × 
<quantity id="4CF9B8B7FF8DC469FF08FF33FEEBFEA7" box="[136,258,232,257]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="2.4" metricValueMin="1.2" pageId="7" pageNumber="24" unit="mm" value="18.0" valueMax="24.0" valueMin="12.0">12–24 mm</quantity>
, glabrous, locules 3, wings 3, equal, rounded proximally and expanded and pointed distally, wings 
<quantity id="4CF9B8B7FF8DC469FAD0FF33FA45FEA7" box="[1360,1452,232,257]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="7" pageNumber="24" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6–7 mm</quantity>
wide, thinly fibrous, dehiscing between locule and wing; pedicel pendent, 
<quantity id="4CF9B8B7FF8DC469FC49FED7FBDAFE83" box="[969,1075,268,293]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="11.0" metricValueMin="7.0" pageId="7" pageNumber="24" unit="mm" value="9.0" valueMax="11.0" valueMin="7.0">7–11 mm</quantity>
long, reddish towards the base.
</paragraph>
</subSubSection>
<subSubSection id="C31B46D9FF8DC469FF3DFEEBFEB4FECB" pageId="7" pageNumber="24" type="etymology">
<paragraph id="8BBE1552FF8DC469FF3DFEEBFEB4FECB" blockId="7.[136,1452,160,365]" pageId="7" pageNumber="24">
<emphasis id="B975C940FF8DC469FF3DFEEBFEA1FEEF" bold="true" box="[189,328,304,329]" pageId="7" pageNumber="24">Etymology:</emphasis>
—Latin, 
<taxonomicName id="4C016ED1FF8DC469FE2EFEEBFDF2FEEF" authorityName="S. Julia &amp; Kiew" authorityYear="2016" box="[430,539,304,329]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="7" pageNumber="24" phylum="Tracheophyta" rank="species" species="compacta">
<emphasis id="B975C940FF8DC469FE2EFEEBFDF2FEEF" box="[430,539,304,329]" italics="true" pageId="7" pageNumber="24">compacta</emphasis>
</taxonomicName>
= close-packed, referring to the compact male inflorescence subtended by many pairs of bracteoles.
</paragraph>
</subSubSection>
<caption id="DF7E45DAFF8DC469FF08F94BFB5FF94B" ID-DOI="http://doi.org/10.5281/zenodo.13674927" ID-Zenodo-Dep="13674927" httpUri="https://zenodo.org/record/13674927/files/figure.png" pageId="7" pageNumber="24" startId="7.[136,229,1680,1701]" targetBox="[292,1300,398,1648]" targetPageId="7" targetType="figure">
<paragraph id="8BBE1552FF8DC469FF08F94BFB5FF94B" blockId="7.[136,1452,1680,1773]" pageId="7" pageNumber="24">
<emphasis id="B975C940FF8DC469FF08F94BFEE8F903" bold="true" box="[136,257,1680,1701]" pageId="7" pageNumber="24">FIGURE 4.</emphasis>
<emphasis id="B975C940FF8DC469FE8BF94BFE26F903" box="[267,463,1680,1701]" italics="true" pageId="7" pageNumber="24">
<taxonomicName id="4C016ED1FF8DC469FE8BF94BFE23F903" authorityName="S. Julia &amp; Kiew" authorityYear="2016" box="[267,458,1680,1701]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="7" pageNumber="24" phylum="Tracheophyta" rank="species" species="compacta">Begonia compacta</taxonomicName>
.
</emphasis>
A. Habit; B. Stipule; C. Male inflorescence surrounded by persistent bracteoles; D. Female inflorescence surrounded by persistent bracteole; E. Female flower showing tepals and pistil; F. Fruits surrounded by persistent bracteole; G. Cross section of ovary. All photos from SFC 2751 by Julia Sang (A–F) and Ling Chea Yiing (G).
</paragraph>
</caption>
<subSubSection id="C31B46D9FF8DC466FF3DF89AFAF7FD87" lastPageId="8" lastPageNumber="25" pageId="7" pageNumber="24" type="distribution">
<paragraph id="8BBE1552FF8DC469FF3DF89AFD5DF7A8" blockId="7.[136,1451,1857,2062]" pageId="7" pageNumber="24">
<emphasis id="B975C940FF8DC469FF3DF89AFEB0F8FC" bold="true" box="[189,345,1857,1882]" pageId="7" pageNumber="24">Distribution:</emphasis>
— 
<collectingCountry id="F31655C2FF8DC469FEF6F89AFDE3F8FC" box="[374,522,1857,1882]" name="Malaysia" pageId="7" pageNumber="24">MALAYSIA</collectingCountry>
. Borneo. 
<collectingCountry id="F31655C2FF8DC469FDF5F89AFD30F8FC" box="[629,729,1857,1882]" name="Malaysia" pageId="7" pageNumber="24">Sarawak</collectingCountry>
. Endemic in 
<collectingCountry id="F31655C2FF8DC469FCEAF89AFC24F8FC" box="[874,973,1857,1882]" name="Malaysia" pageId="7" pageNumber="24">Sarawak</collectingCountry>
, known only from the Batang Ai. 
<emphasis id="B975C940FF8DC469FF3DF8BEFECAF8D8" bold="true" box="[189,291,1893,1918]" pageId="7" pageNumber="24">Habitat:</emphasis>
—Riparian forest to 
<quantity id="4CF9B8B7FF8DC469FD8AF8BEFDB8F8D8" box="[522,593,1893,1918]" metricMagnitude="2" metricUnit="m" metricValue="1.27" pageId="7" pageNumber="24" unit="m" value="127.0">127 m</quantity>
elevation on steep earth or rocky slopes above the river bank in light shade. 
<emphasis id="B975C940FF8DC469FF3DF852FDD4F804" bold="true" box="[189,573,1929,1954]" pageId="7" pageNumber="24">Additional specimen examined:</emphasis>
— 
<collectingCountry id="F31655C2FF8DC469FDD9F852FD04F804" box="[601,749,1929,1954]" name="Malaysia" pageId="7" pageNumber="24">MALAYSIA</collectingCountry>
. Borneo. 
<collectingCountry id="F31655C2FF8DC469FCDFF852FC2AF804" box="[863,963,1929,1954]" name="Malaysia" pageId="7" pageNumber="24">Sarawak</collectingCountry>
. Lubok Antu District: Batang Ai National Park, Sungai Bebiyong Besai, 
<date id="FFBF3392FF8DC469FE5EF876FD6CF860" box="[478,645,1965,1990]" pageId="7" pageNumber="24" value="2015-08-02">2 August 2015</date>
, 
<emphasis id="B975C940FF8DC469FD10F876FC97F860" box="[656,894,1965,1990]" italics="true" pageId="7" pageNumber="24">Julia et al. SFC 2760</emphasis>
(KEP, SAR, SNP). 
<emphasis id="B975C940FF8DC469FF3DF80AFDCDF84C" bold="true" box="[189,548,2001,2026]" pageId="7" pageNumber="24">Proposed conservation status:</emphasis>
—Least Concern as the species occurs within a Totally Protected Area and their habitat is not threatened by any local disturbance.
</paragraph>
<paragraph id="8BBE1552FF82C466FF08FF45FBA5FF7B" blockId="8.[136,1453,158,221]" pageId="8" pageNumber="25">
<emphasis id="B975C940FF82C466FF08FF45FF74FF1E" bold="true" box="[136,157,158,184]" italics="true" pageId="8" pageNumber="25">6.</emphasis>
<taxonomicName id="4C016ED1FF82C466FF21FF45FDE4FF1F" authority="C. W. Lin" authorityName="C. W. Lin" box="[161,525,158,185]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="8" pageNumber="25" phylum="Tracheophyta" rank="species" species="darthvaderiana">
<emphasis id="B975C940FF82C466FF21FF45FE44FF1F" bold="true" box="[161,429,158,185]" italics="true" pageId="8" pageNumber="25">Begonia darthvaderiana</emphasis>
<materialsCitation id="3B691F0FFF82C466FE31FF7BFDE4FF1F" box="[433,525,160,185]" collectingDate="2013-07-05" collectionCode="C, I, W, SAR, E, HAST, KEP, TAIF" country="Malaysia" county="Peng" location="Borneo" municipality="Type" pageId="8" pageNumber="25" specimenCount="2" stateProvince="Sarawak" typeStatus="holotype">C.W.Lin</materialsCitation>
</taxonomicName>
&amp; C.-I 
<collectingCounty id="62DF6DDEFF82C466FDDBFF7BFD78FF1F" box="[603,657,160,185]" pageId="8" pageNumber="25">Peng</collectingCounty>
(2014: 132). 
<collectingMunicipality id="6BDA8F28FF82C466FC9DFF7BFCBCFF1F" box="[797,853,160,185]" pageId="8" pageNumber="25">Type</collectingMunicipality>
:— 
<collectingCountry id="F31655C2FF82C466FCF4FF7BFBEFFF1F" box="[884,1030,160,185]" name="Malaysia" pageId="8" pageNumber="25">MALAYSIA</collectingCountry>
. 
<location id="8EDE4389FF82C466FB8DFF7BFBB6FF1F" LSID="urn:lsid:plazi:treatment:03A8A444FF8CC466FF08FBB0FAF7FD87:8EDE4389FF82C466FB8DFF7BFBB6FF1F" box="[1037,1119,160,185]" country="Malaysia" county="Peng" municipality="Type" name="Borneo" pageId="8" pageNumber="25" stateProvince="Sarawak">Borneo</location>
, 
<collectingRegion id="49C5DBB0FF82C466FBE7FF7BFB2EFF1F" box="[1127,1223,160,185]" country="Malaysia" name="Sarawak" pageId="8" pageNumber="25">Sarawak</collectingRegion>
. 
<location id="8EDE4389FF82C466FB4FFF7BFA40FF1F" LSID="urn:lsid:plazi:treatment:03A8A444FF8CC466FF08FBB0FAF7FD87:8EDE4389FF82C466FB4FFF7BFA40FF1F" box="[1231,1449,160,185]" country="Malaysia" county="Peng" municipality="Type" name="Lubok Antu District" pageId="8" pageNumber="25" stateProvince="Sarawak">Lubok Antu District</location>
: 
<location id="8EDE4389FF82C466FF08FF1FFF1EFF7B" LSID="urn:lsid:plazi:treatment:03A8A444FF8CC466FF08FBB0FAF7FD87:8EDE4389FF82C466FF08FF1FFF1EFF7B" box="[136,247,196,221]" country="Malaysia" county="Peng" municipality="Type" name="Batang Ai" pageId="8" pageNumber="25" stateProvince="Sarawak">Batang Ai</location>
, 
<date id="FFBF3392FF82C466FE81FF1FFE69FF7B" box="[257,384,196,221]" pageId="8" pageNumber="25" value="2013-07-05">
<collectingDate id="EFFBCA7AFF82C466FE81FF1FFE69FF7B" box="[257,384,196,221]" pageId="8" pageNumber="25" value="2013-07-05">5 July 2013</collectingDate>
</date>
, 
<emphasis id="B975C940FF82C466FE0AFF1FFDFEFF7B" box="[394,535,196,221]" italics="true" pageId="8" pageNumber="25">C.W. Lin 549</emphasis>
(
<typeStatus id="54BAABF0FF82C466FDA4FF1FFD6DFF7B" box="[548,644,196,221]" pageId="8" pageNumber="25" type="holotype">holotype</typeStatus>
SAR n.v.; 
<typeStatus id="54BAABF0FF82C466FD78FF1FFCB9FF7B" box="[760,848,196,221]" pageId="8" pageNumber="25" type="isotype">isotypes</typeStatus>
E, HAST, KEP, TAIF).
</paragraph>
<paragraph id="8BBE1552FF82C466FF08FED7FB94FE83" blockId="8.[136,1452,268,545]" box="[136,1149,268,293]" pageId="8" pageNumber="25">
<emphasis id="B975C940FF82C466FF08FED7FECDFE83" bold="true" box="[136,292,268,293]" pageId="8" pageNumber="25">Distribution:</emphasis>
—Endemic in 
<collectingCountry id="F31655C2FF82C466FE47FED7FDC2FE83" box="[455,555,268,293]" name="Malaysia" pageId="8" pageNumber="25">Sarawak</collectingCountry>
; known only from Batang Ai in Lubok Antu District.
</paragraph>
<paragraph id="8BBE1552FF82C466FF3DFEEBFDC1FECB" blockId="8.[136,1452,268,545]" pageId="8" pageNumber="25">
<emphasis id="B975C940FF82C466FF3DFEEBFECAFEEF" bold="true" box="[189,291,304,329]" pageId="8" pageNumber="25">Habitat:</emphasis>
—In lowland mixed dipterocarp forest at 
<quantity id="4CF9B8B7FF82C466FD76FEEBFC9FFEEF" box="[758,886,304,329]" metricMagnitude="2" metricUnit="m" metricValue="1.65" metricValueMax="2.0" metricValueMin="1.3" pageId="8" pageNumber="25" unit="m" value="165.0" valueMax="200.0" valueMin="130.0">130–200 m</quantity>
elevation in valleys, on lower to upper soil slopes or soil at the cliff base in deep shade.
</paragraph>
<paragraph id="8BBE1552FF82C466FF3DFEA3FB9CFE13" blockId="8.[136,1452,268,545]" pageId="8" pageNumber="25">
<emphasis id="B975C940FF82C466FF3DFEA3FEE0FE37" bold="true" box="[189,265,376,401]" pageId="8" pageNumber="25">Notes:</emphasis>
—The species was collected from forest areas outside the Batang Ai National Park boundary (Lin, C.W. 
<emphasis id="B975C940FF82C466FF08FE47FEEAFE13" box="[136,259,412,437]" italics="true" pageId="8" pageNumber="25">pers.comm</emphasis>
.). So far no record of this species has been recorded within the Park boundary.
</paragraph>
<paragraph id="8BBE1552FF82C466FF3DFE1BFAF7FD87" blockId="8.[136,1452,268,545]" pageId="8" pageNumber="25">
<emphasis id="B975C940FF82C466FF3DFE1BFDCCFE7F" bold="true" box="[189,549,448,473]" pageId="8" pageNumber="25">Proposed conservation status:</emphasis>
—Data Deficient. The species was previously recorded from forest area outside the park boundary and the species occurrence within the park boundary is yet to be ascertained. It may also be under pressure from commercial collecting because it is currently the most sought after begonia from 
<collectingCountry id="F31655C2FF82C466FB36FDD3FAF3FD87" box="[1206,1306,520,545]" name="Malaysia" pageId="8" pageNumber="25">Sarawak</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>