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<document id="D7DA57FFD86B590C6284DE8EF4B55F01" ID-CLB-Dataset="2598" ID-DOI="10.5852/ejt.2022.788.1625" ID-GBIF-Dataset="d6e6119f-4100-4cfc-9cc3-0d6951ee2c28" ID-ISSN="2118-9773" ID-Zenodo-Dep="5846636" ID-ZooBank="46C3CFCA-0CED-4432-AFD8-F4CFC1E0E1E7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1642092502222" checkinUser="felipe" docAuthor="Castillo, Stephanie, Rédei, Dávid &amp; Weirauch, Christiane" docDate="2022" docId="015D8E70FFC5FFD4FD6C02A0FCD1BF76" docLanguage="en" docName="ejt-788_1-95.pdf" docOrigin="European Journal of Taxonomy 788 (1)" docSource="http://dx.doi.org/10.5852/ejt.2022.788.1625" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Voconia tridens Castillo &amp; Rédei &amp; Weirauch 2022, sp. nov." docType="treatment" docUuid="82E67D5A-E019-40EE-A019-E3BCD5E781DA" docUuidSource="ZooBank" docVersion="10" lastPageNumber="81" masterDocId="FD64F608FF8BFF84FFE7031FFFACBA3D" masterDocTitle="Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs" masterLastPageNumber="95" masterPageNumber="1" pageNumber="79" updateTime="1699302197121" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="9F90B509F6533311675C36B27F551A44">Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs</mods:title>
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<mods:namePart id="3B0664EE49B48BA501F04C63494A3CFD">Castillo, Stephanie</mods:namePart>
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<mods:affiliation id="FCFD21230FDD71976AD400A8C0CEB07D">University of California, Riverside, 900 University Avenue, Riverside, CA 92521, USA.</mods:affiliation>
<mods:nameIdentifier id="F9F4EB3708C546E2DA3C506E21E17A86" type="email">scast032@ucr.edu</mods:nameIdentifier>
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<mods:namePart id="0B8C43AA32BD72783BE64A01669B9386">Rédei, Dávid</mods:namePart>
<mods:nameIdentifier id="24E7C0B6EB4C6E75F7CCAB3ECB799B3A" type="LSID">74DB4DB4-E632-4CA9-B820-2070E7EB18D4</mods:nameIdentifier>
<mods:affiliation id="1D1DFBF9CD663FB730970D13DBB2D9BD">National Chung Hsing University, 145 Xingda Rd., 40227 Taichung, Taiwan.</mods:affiliation>
<mods:nameIdentifier id="1D20A7265307917F4E528739C55C1603" type="email">david.redei@gmail.com</mods:nameIdentifier>
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<mods:namePart id="AB824ABF19EBFCDA17443641ABCBA39A">Weirauch, Christiane</mods:namePart>
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<mods:affiliation id="2B6DD9DEBE6004B1B8B5405C5FDD884B">University of California, Riverside, 900 University Avenue, Riverside, CA 92521, USA.</mods:affiliation>
<mods:nameIdentifier id="CE476561A5B0BAF7A2D3F9253B09979E" type="email">christiane.weirauch@ucr.edu</mods:nameIdentifier>
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<mods:title id="76A8272803AF0E8CF1CEFE050BA95A18">European Journal of Taxonomy</mods:title>
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<mods:date id="1A8144DA092FBD2E0BC3751A3DB4A653">2022</mods:date>
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<mods:number id="DE0272C60A091043D0D54EA878C01260">2022-01-11</mods:number>
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<treatment id="015D8E70FFC5FFD4FD6C02A0FCD1BF76" ID-DOI="http://doi.org/10.5281/zenodo.5849751" ID-GBIF-Taxon="192123835" ID-Zenodo-Dep="5849751" LSID="urn:lsid:zoobank.org:act:82E67D5A-E019-40EE-A019-E3BCD5E781DA" httpUri="http://treatment.plazi.org/id/015D8E70FFC5FFD4FD6C02A0FCD1BF76" lastPageId="80" lastPageNumber="81" pageId="78" pageNumber="79">
<subSubSection id="C1EE6CEDFFC5FFCAFD6C02A0FC05BBE4" box="[651,937,446,473]" pageId="78" pageNumber="79" type="nomenclature">
<paragraph id="894B3F66FFC5FFCAFD6C02A0FC05BBE4" blockId="78.[359,1230,446,543]" box="[651,937,446,473]" pageId="78" pageNumber="79">
<heading id="D203880AFFC5FFCAFD6C02A0FC05BBE4" box="[651,937,446,473]" centered="true" fontSize="11" level="2" pageId="78" pageNumber="79" reason="2">
<taxonomicName id="4EF444E5FFC5FFCAFD6C02A0FCEBBBE4" authority="Castillo &amp; Rédei &amp; Weirauch, 2022" authorityName="Castillo &amp; Rédei &amp; Weirauch" authorityYear="2022" box="[651,839,446,473]" class="Insecta" family="Reduviidae" genus="Voconia" kingdom="Animalia" order="Hemiptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="tridens" status="sp. nov.">
<emphasis id="BB80E374FFC5FFCAFD6C02A0FCEBBBE4" bold="true" box="[651,839,446,473]" italics="true" pageId="78" pageNumber="79">Voconia tridens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0B35E0FFFC5FFCAFCA902A0FC05BBE4" box="[846,937,447,473]" pageId="78" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C1EE6CEDFFC5FFCAFE8002FDFC1FB822" pageId="78" pageNumber="79" type="description">
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<uri id="FD653364FFC5FFCAFE8002FDFB62BBC0" box="[359,1230,482,509]" pageId="78" pageNumber="79">
urn:lsid:zoobank.org:act:
<uuid id="FD5205B3FFC5FFCAFD6302FDFB62BBC0" box="[644,1230,482,509]" pageId="78" pageNumber="79">82E67D5A-E019-40EE-A019-E3BCD5E781DA</uuid>
</uri>
</paragraph>
<paragraph id="894B3F66FFC5FFCAFD67011AFC1FB822" blockId="78.[359,1230,446,543]" box="[640,947,517,543]" pageId="78" pageNumber="79">
<figureCitation id="11CF23E3FFC5FFCAFD67011AFD44B822" box="[640,744,517,543]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="8.[190,233,1950,1976]" captionStartId-1="9.[189,232,1883,1909]" captionTargetBox-0="[204,1371,808,1898]" captionTargetBox-1="[199,1395,651,1845]" captionTargetId-0="figure-168@8.[189,1398,793,1908]" captionTargetId-1="figure-137@9.[189,1398,644,1845]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="Fig. 1. Strict consensus tree resulting from equal weighting analysis. Jackknife values over 51 are reported for tree one of eight." captionText-1="Fig. 2. Tree resulting from implied weighting analysis using K = 12 with characters unambiguously optimized. Symmetric resampling values over 51 are represented for trees obtained with K = 3, K = 6, K = 9, and K = 12 analyses. Characters and character states are described in pp. 11–22. Specimens not to scale." figureDoi-0="http://doi.org/10.5281/zenodo.5846638" figureDoi-1="http://doi.org/10.5281/zenodo.5846640" httpUri-0="https://zenodo.org/record/5846638/files/figure.png" httpUri-1="https://zenodo.org/record/5846640/files/figure.png" pageId="78" pageNumber="79">Figs 1–2</figureCitation>
,
<figureCitation id="11CF23E3FFC5FFCAFD12011AFCA9B822" box="[757,773,517,543]" captionStart="Fig" captionStartId="33.[189,231,1954,1980]" captionTargetBox="[189,1398,597,1914]" captionTargetId="figure-95@33.[189,1398,597,1914]" captionTargetPageId="33" captionText="Fig. 8. Dorsal habitus of holotypes of Voconia Stål, 1866 species with distribution: purple = Afrotropical region, green = Australasian region, blue = Neotropical region, pink = Oriental region. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5846652" httpUri="https://zenodo.org/record/5846652/files/figure.png" pageId="78" pageNumber="79">8</figureCitation>
,
<figureCitation id="11CF23E3FFC5FFCAFCF5011AFC9EB822" box="[786,818,517,543]" captionStart="Fig" captionStartId="39.[188,231,1955,1981]" captionTargetBox="[189,1398,882,1916]" captionTargetId="figure-204@39.[189,1399,882,1916]" captionTargetPageId="39" captionText="Fig. 10. Lateral habitus of species of Voconia Stål, 1866. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5846656" httpUri="https://zenodo.org/record/5846656/files/figure.png" pageId="78" pageNumber="79">10</figureCitation>
,
<figureCitation id="11CF23E3FFC5FFCAFCD9011AFC26B822" box="[830,906,517,543]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="74.[191,234,1981,2007]" captionStartId-1="75.[188,231,1989,2015]" captionTargetBox-0="[191,1399,689,1939]" captionTargetBox-1="[189,1397,760,1946]" captionTargetId-0="figure-149@74.[191,1399,689,1939]" captionTargetId-1="figure-180@75.[189,1397,760,1946]" captionTargetPageId-0="74" captionTargetPageId-1="75" captionText-0="Fig. 13. Pygophore of species of Voconia Stål, 1866. Scale bars = 0.5 mm." captionText-1="Fig. 14. Aedeagus of species of Gerbelius Distant, 1903 and Voconia Stål, 1866. Scale bars = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5846666" figureDoi-1="http://doi.org/10.5281/zenodo.5846668" httpUri-0="https://zenodo.org/record/5846666/files/figure.png" httpUri-1="https://zenodo.org/record/5846668/files/figure.png" pageId="78" pageNumber="79">13–14</figureCitation>
,
<figureCitation id="11CF23E3FFC5FFCAFC71011AFC1FB822" box="[918,947,517,543]" captionStart="Fig" captionStartId="85.[189,232,1988,2014]" captionTargetBox="[189,1398,872,1948]" captionTargetId="figure-209@85.[189,1398,872,1948]" captionTargetPageId="85" captionText="Fig. 17. The distribution of species of Voconia Stål, 1866 in Africa." figureDoi="http://doi.org/10.5281/zenodo.5846674" httpUri="https://zenodo.org/record/5846674/files/figure.png" pageId="78" pageNumber="79">17</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C1EE6CEDFFC5FFCAFF5A0159FC30B902" pageId="78" pageNumber="79" type="diagnosis">
<paragraph id="894B3F66FFC5FFCAFF5A0159FE9AB85D" blockId="78.[189,310,582,608]" box="[189,310,582,608]" pageId="78" pageNumber="79">
<heading id="D203880AFFC5FFCAFF5A0159FE9AB85D" bold="true" box="[189,310,582,608]" fontSize="11" level="3" pageId="78" pageNumber="79" reason="3">
<emphasis id="BB80E374FFC5FFCAFF5A0159FE9AB85D" bold="true" box="[189,310,582,608]" pageId="78" pageNumber="79">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="894B3F66FFC5FFCAFF5A016AFC30B902" blockId="78.[188,1399,629,832]" pageId="78" pageNumber="79">
Distinguished from other species of
<taxonomicName id="4EF444E5FFC5FFCAFD660169FD4CB8B2" authorityName="Stal" authorityYear="1866" box="[641,736,630,655]" class="Insecta" family="Reduviidae" genus="Voconia" kingdom="Animalia" order="Hemiptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="BB80E374FFC5FFCAFD660169FD4CB8B2" box="[641,736,630,655]" italics="true" pageId="78" pageNumber="79">Voconia</emphasis>
</taxonomicName>
by the following combination of characters: coarsely granulose head and pronotum; maxillary plates not adjacent to and far surpass clypeal apex; and antennifers with a long, lateral spine. This species is recognized from the macropterous morph of
<taxonomicName id="4EF444E5FFC5FFCAFF5A01FFFEC9B8C4" authorityName="Villiers" authorityYear="1964" box="[189,357,735,761]" class="Insecta" family="Reduviidae" genus="Pseudocethera" kingdom="Animalia" order="Hemiptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="schoutedeni">
<emphasis id="BB80E374FFC5FFCAFF5A01FFFEC9B8C4" box="[189,357,735,761]" italics="true" pageId="78" pageNumber="79">V. schoutedeni</emphasis>
</taxonomicName>
by the yellow head and pronotum; the maxillary plates in lateral view are not wide, about the width of the scape; scutellar spine is subhorizontal; and the hemelytra is not blackish, it is a light brown with pale coloration at the base and apex of the corium.
</paragraph>
</subSubSection>
<subSubSection id="C1EE6CEDFFC5FFCAFF5A0078FC8CB9CB" pageId="78" pageNumber="79" type="etymology">
<paragraph id="894B3F66FFC5FFCAFF5A0078FEE8B9BC" blockId="78.[189,324,871,897]" box="[189,324,871,897]" pageId="78" pageNumber="79">
<heading id="D203880AFFC5FFCAFF5A0078FEE8B9BC" bold="true" box="[189,324,871,897]" fontSize="11" level="3" pageId="78" pageNumber="79" reason="3">
<emphasis id="BB80E374FFC5FFCAFF5A0078FEE8B9BC" bold="true" box="[189,324,871,897]" pageId="78" pageNumber="79">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="894B3F66FFC5FFCAFF5A008AFC8CB9CB" blockId="78.[189,1399,917,1014]" pageId="78" pageNumber="79">
The specific epithet is the Latin adjective ‘
<taxonomicName id="4EF444E5FFC5FFCAFD4C008AFD50B992" authorityName="Castillo &amp; Rédei &amp; Weirauch" authorityYear="2022" box="[683,764,917,943]" class="Insecta" family="Reduviidae" genus="Voconia" kingdom="Animalia" order="Hemiptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="tridens">
<emphasis id="BB80E374FFC5FFCAFD4C008AFD50B992" box="[683,764,917,943]" italics="true" pageId="78" pageNumber="79">tridens</emphasis>
</taxonomicName>
, -, -’, meaning ‘three-toothed, three-pronged’, derived from the prefix ‘
<emphasis id="BB80E374FFC5FFCAFE6F00A6FE08B9EF" box="[392,420,953,978]" italics="true" pageId="78" pageNumber="79">tri</emphasis>
-’ combined with the Latin noun ‘
<emphasis id="BB80E374FFC5FFCAFCA400A7FCD5B9EF" box="[835,889,952,978]" italics="true" pageId="78" pageNumber="79">dens</emphasis>
’, referring to the elongated and separated maxillary plates and clypeus that form three prongs.
</paragraph>
</subSubSection>
<subSubSection id="C1EE6CEDFFC5FFCAFF5A0702FB11BE91" pageId="78" pageNumber="79" type="materials_examined">
<paragraph id="894B3F66FFC5FFCAFF5A0702FEC0BE0A" blockId="78.[189,364,1053,1079]" box="[189,364,1053,1079]" pageId="78" pageNumber="79">
<heading id="D203880AFFC5FFCAFF5A0702FEC0BE0A" bold="true" box="[189,364,1053,1079]" fontSize="11" level="3" pageId="78" pageNumber="79" reason="3">
<emphasis id="BB80E374FFC5FFCAFF5A0702FEC0BE0A" bold="true" box="[189,364,1053,1079]" pageId="78" pageNumber="79">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="894B3F66FFC5FFCAFF0D0753FB11BE91" blockId="78.[189,1399,1100,1197]" pageId="78" pageNumber="79">
<materialsCitation id="399C353BFFC5FFCAFF0D0753FB11BE91" ID-GBIF-Occurrence="3436573356" collectingDate="1962-11-23" collectingDateMax="1962-11-24" collectingDateMin="1962-11-23" collectionCode="AMNH" collectorName="R. Linnavuori" country="Sudan" latitude="12.25" location="Abu Hashim-Galegu" longLatPrecision="776" longitude="34.69" municipality="Abu Hashim-Galegu" pageId="78" pageNumber="79" specimenCode="AMNH_PBI 00170712" specimenCount="1" specimenCount-male="1" stateProvince="Blue Nile" typeStatus="holotype">
<emphasis id="BB80E374FFC5FFCAFF0D0753FEF0BE5B" bold="true" box="[234,348,1100,1126]" pageId="78" pageNumber="79">
<typeStatus id="564F81C4FFC5FFCAFF0D0753FEF0BE5B" box="[234,348,1100,1126]" pageId="78" pageNumber="79" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F1E37FF6FFC5FFCAFF5A0770FE8EBEB4" box="[189,290,1135,1161]" name="Sudan" pageId="78" pageNumber="79">SUDAN</collectingCountry>
•
<specimenCount id="9FF2F4EFFFC5FFCAFED90770FEF8BEB4" box="[318,340,1135,1161]" pageId="78" pageNumber="79" type="male">♂</specimenCount>
(dissected pygophore and aedeagus in vial);
<collectingRegion id="4B30F184FFC5FFCAFC960770FC4BBEB4" box="[881,999,1135,1161]" country="Sudan" name="An Nil al Azraq" pageId="78" pageNumber="79">Blue Nile</collectingRegion>
,
<collectingMunicipality id="692FA51CFFC5FFCAFC140770FB40BEB4" box="[1011,1260,1135,1161]" country="Sudan" latitude="12.25" longLatPrecision="776" longitude="34.69" name="Abu Hashim-Galegu" pageId="78" pageNumber="79" stateProvince="An Nil al Azraq">Abu Hashim-Galegu</collectingMunicipality>
; [
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12.25
<emphasis id="BB80E374FFC5FFCAFAA20770FAFDBEB4" bold="true" box="[1349,1361,1135,1161]" pageId="78" pageNumber="79">°</emphasis>
N
</geoCoordinate>
,
<geoCoordinate id="ECC059A1FFC5FFCAFF5A078DFE84BE91" box="[189,296,1169,1196]" degrees="34.69" direction="east" orientation="longitude" pageId="78" pageNumber="79" precision="555" value="34.69">
34.69
<emphasis id="BB80E374FFC5FFCAFF18078DFEA7BE91" bold="true" box="[255,267,1170,1196]" pageId="78" pageNumber="79">°</emphasis>
E
</geoCoordinate>
];
<date id="FD4A19A6FFC5FFCAFEDA078DFDA7BE91" box="[317,523,1170,1197]" pageId="78" pageNumber="79" value="1962-11-23" valueMax="1962-11-24" valueMin="1962-11-23">
<collectingDate id="ED0EE04EFFC5FFCAFEDA078DFDA7BE91" box="[317,523,1170,1197]" pageId="78" pageNumber="79" value="1962-11-23" valueMax="1962-11-24" valueMin="1962-11-23">23–24 Nov. 1962</collectingDate>
</date>
;
<collectorName id="24015AB0FFC5FFCAFDF0078CFD10BE91" box="[535,700,1170,1197]" pageId="78" pageNumber="79">R. Linnavuori</collectorName>
leg.; USI:
<specimenCode id="D952971DFFC5FFCAFCDC078DFBFDBE91" box="[827,1105,1170,1196]" pageId="78" pageNumber="79">AMNH_PBI 00170712</specimenCode>
;
<collectionCode id="EFE5A7A3FFC5FFCAFBBC078DFB15BE91" box="[1115,1209,1170,1196]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="78" pageNumber="79">AMNH</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C1EE6CEDFFC5FFD4FF5A07CCFE95BE3C" lastPageId="80" lastPageNumber="81" pageId="78" pageNumber="79" type="description">
<paragraph id="894B3F66FFC5FFCAFF5A07CCFEE2BED0" blockId="78.[189,334,1235,1261]" box="[189,334,1235,1261]" pageId="78" pageNumber="79">
<heading id="D203880AFFC5FFCAFF5A07CCFEE2BED0" bold="true" box="[189,334,1235,1261]" fontSize="11" level="3" pageId="78" pageNumber="79" reason="3">
<emphasis id="BB80E374FFC5FFCAFF5A07CCFEE2BED0" bold="true" box="[189,334,1235,1261]" pageId="78" pageNumber="79">Description</emphasis>
</heading>
</paragraph>
<paragraph id="894B3F66FFC5FFCAFF0D061DFE15BF21" blockId="78.[189,711,1282,1344]" box="[234,441,1282,1308]" pageId="78" pageNumber="79">
<emphasis id="BB80E374FFC5FFCAFF0D061DFE86BF21" bold="true" box="[234,298,1282,1308]" pageId="78" pageNumber="79">Male</emphasis>
(
<figureCitation id="11CF23E3FFC5FFCAFEDE061DFE29BF21" box="[313,389,1282,1308]" captionStart="Fig" captionStartId="33.[189,231,1954,1980]" captionTargetBox="[189,1398,597,1914]" captionTargetId="figure-95@33.[189,1398,597,1914]" captionTargetPageId="33" captionText="Fig. 8. Dorsal habitus of holotypes of Voconia Stål, 1866 species with distribution: purple = Afrotropical region, green = Australasian region, blue = Neotropical region, pink = Oriental region. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5846652" httpUri="https://zenodo.org/record/5846652/files/figure.png" pageId="78" pageNumber="79">Figs 8</figureCitation>
,
<figureCitation id="11CF23E3FFC5FFCAFE75061DFE1CBF21" box="[402,432,1282,1308]" captionStart="Fig" captionStartId="39.[188,231,1955,1981]" captionTargetBox="[189,1398,882,1916]" captionTargetId="figure-204@39.[189,1399,882,1916]" captionTargetPageId="39" captionText="Fig. 10. Lateral habitus of species of Voconia Stål, 1866. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5846656" httpUri="https://zenodo.org/record/5846656/files/figure.png" pageId="78" pageNumber="79">10</figureCitation>
)
</paragraph>
<paragraph id="894B3F66FFC5FFCAFF5A0639FD6BBF7D" blockId="78.[189,711,1282,1344]" box="[189,711,1317,1344]" pageId="78" pageNumber="79">BODY LENGTH. About 8.0 mm; macropterous.</paragraph>
<paragraph id="894B3F66FFC5FFCAFF5A0678FD62BC30" blockId="78.[188,1400,1383,1550]" pageId="78" pageNumber="79">COLORATION. Head: yellow; postocular region with red patches adjacent to medial ocellar margin; labium as head. Thorax: uniformly yellow; scutellum yellow. Hemelytron (abducted): clavus uniformly yellow; corium proximal half mostly yellow, distal half brown with yellow spot at distal apex; membrane dark with pale V-shaped marking along R and M veins. Legs: yellow. Abdomen: yellow, dorsal laterotergites with a pale brown stripe on posterior margin.</paragraph>
<paragraph id="894B3F66FFC5FFCAFF5A052AFE97BD1F" blockId="78.[189,1400,1589,1826]" pageId="78" pageNumber="79">INTEGUMENT AND VESTITURE. Head and pronotum: coarsely granulose with dense, short setigerous tubercles; interocular region with two pairs of stout setigerous tubercles paramedially; antennifer with long lateral spine; morphologically ventral surface of labium with sparse, short macrosetae. Thorax: anterolateral angles of pronotal collar without macrosetae or setigerous tubercles; scutellar lateral carinae with pubescent tubercles. Hemelytron: corium with short macrosetae. Legs: posterior row of protuberances on mid and hind femora with three large spines on distal half. Abdomen: ventral surface pubescent.</paragraph>
<paragraph id="894B3F66FFC5FFD4FF5A0455FB92B9A1" blockId="78.[189,1399,1865,1998]" lastBlockId="80.[189,1401,265,925]" lastPageId="80" lastPageNumber="81" pageId="78" pageNumber="79">
STRUCTURE. Head: elongate, about 1.3 times as long as wide; anteocular region two-fifths of head length, longer than postocular region (measured to anterior margin of neck); postocular region in dorsal view longer than eye, lateral margins subrectangular; scape not reaching head apex; pedicel about 0.9 times length of head width; maxillary plates ellipsoidal, not adjacent to or longer than clypeus; apices of maxillary plates in dorsal view diverging; clypeus in dorsal view narrower than maxillary plates; clypeal apex round, narrowed; interocular glabrous markings V-shaped, joined medially at interocular sulcus; interocular sulcus in dorsal view curved and smoothly rounded anteriorly; eye width in dorsal view narrower than synthlipsis; eye reaching ventral head margin in lateral view; ventrolateral swelling of buccula with acute protrusion surpassing buccular margin, small setigerous tubercles along margin of protrusion; labial segment I in lateral view straight, surpassing posteroventral eye margin; morphologically dorsal surface of labial segment II nearly straight, about 0.4 times length of segment I. Thorax: pronotal collar in dorsal view wide medially with anterolateral angles long, flared laterally; anterior pronotal lobe about 0.7 times length of posterior pronotal lobe, lateral margins almost straight; glabrous markings on pronotum deeply depressed; median apodeme depression of pronotum shallow, elongated longitudinally; scutellar spine long, subhorizontal; anteriad-directed process of prosternum with paramedial lobes; anterior margin of stridulitrum not elongated into protuberance; proepimeron with smoothly rounded posteroventral margin. Legs: fossula spongiosa present on fore and mid legs. Pygophore (
<figureCitation id="11CF23E3FFDBFFD4FEB401CDFE03B8D0" box="[339,431,722,749]" captionStart="Fig" captionStartId="74.[191,234,1981,2007]" captionTargetBox="[191,1399,689,1939]" captionTargetId="figure-149@74.[191,1399,689,1939]" captionTargetPageId="74" captionText="Fig. 13. Pygophore of species of Voconia Stål, 1866. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5846666" httpUri="https://zenodo.org/record/5846666/files/figure.png" pageId="80" pageNumber="81">Fig. 13</figureCitation>
): transverse bridge with triangular posterior margin; posterior region of ventral surface of pygophore in lateral view straight, not swollen; short median apical process sharply bent posteriorly in lateral view; lateral pygophore margin with protuberance; posterior pygophore margin with clustered macrosetae on protuberance; parameres round, apex tapered into squarely rounded tip. Aedeagus: endosoma almost entirely covered with spicules; apex of dorsal phallothecal sclerite tongueshaped in dorsal view; basal plate extension about 4.6 times as long as wide.
</paragraph>
<caption id="DD8B6FEEFFC4FFCBFF5A047DFC39BD40" ID-DOI="http://doi.org/10.5281/zenodo.5846670" ID-Zenodo-Dep="5846670" box="[189,917,1890,1917]" httpUri="https://zenodo.org/record/5846670/files/figure.png" pageId="79" pageNumber="80" startId="79.[189,232,1890,1916]" targetBox="[189,1398,280,1838]" targetPageId="79">
<paragraph id="894B3F66FFC4FFCBFF5A047DFC39BD40" blockId="79.[189,917,1890,1917]" box="[189,917,1890,1917]" pageId="79" pageNumber="80">
<emphasis id="BB80E374FFC4FFCBFF5A047DFEB6BD41" bold="true" box="[189,282,1890,1916]" pageId="79" pageNumber="80">Fig. 15.</emphasis>
Pygophore of outgroup species. Scale bars = 0.5 mm.
</paragraph>
</caption>
<paragraph id="894B3F66FFDBFFD4FF0D00DBFEE9B9E3" blockId="80.[189,325,964,1025]" box="[234,325,964,990]" pageId="80" pageNumber="81">
<emphasis id="BB80E374FFDBFFD4FF0D00DBFEE9B9E3" bold="true" box="[234,325,964,990]" pageId="80" pageNumber="81">Female</emphasis>
</paragraph>
<paragraph id="894B3F66FFDBFFD4FF5A00F8FE95BE3C" blockId="80.[189,325,964,1025]" box="[189,313,999,1025]" pageId="80" pageNumber="81">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C1EE6CEDFFDBFFD4FF5A0737FC6DBE4C" pageId="80" pageNumber="81" type="distribution">
<paragraph id="894B3F66FFDBFFD4FF5A0737FE68BE7E" blockId="80.[189,452,1064,1091]" box="[189,452,1064,1091]" pageId="80" pageNumber="81">
<emphasis id="BB80E374FFDBFFD4FF5A0737FEFABE7F" bold="true" box="[189,342,1064,1090]" pageId="80" pageNumber="81">Distribution</emphasis>
(
<figureCitation id="11CF23E3FFDBFFD4FE810736FE17BE7E" box="[358,443,1065,1091]" captionStart="Fig" captionStartId="85.[189,232,1988,2014]" captionTargetBox="[189,1398,872,1948]" captionTargetId="figure-209@85.[189,1398,872,1948]" captionTargetPageId="85" captionText="Fig. 17. The distribution of species of Voconia Stål, 1866 in Africa." figureDoi="http://doi.org/10.5281/zenodo.5846674" httpUri="https://zenodo.org/record/5846674/files/figure.png" pageId="80" pageNumber="81">Fig. 17</figureCitation>
)
</paragraph>
<paragraph id="894B3F66FFDBFFD4FF5A0748FC6DBE4C" blockId="80.[189,961,1111,1138]" box="[189,961,1111,1138]" pageId="80" pageNumber="81">
Only known from the
<typeStatus id="564F81C4FFDBFFD4FE250747FE59BE4F" box="[450,501,1112,1138]" pageId="80" pageNumber="81">type</typeStatus>
locality in Northeastern Africa,
<collectingCountry id="F1E37FF6FFDBFFD4FC970748FC11BE4C" box="[880,957,1111,1137]" name="Sudan" pageId="80" pageNumber="81">Sudan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C1EE6CEDFFDBFFD4FF5A0786FCD1BF76" pageId="80" pageNumber="81" type="discussion">
<paragraph id="894B3F66FFDBFFD4FF5A0786FE83BE8E" blockId="80.[189,303,1177,1203]" box="[189,303,1177,1203]" pageId="80" pageNumber="81">
<heading id="D203880AFFDBFFD4FF5A0786FE83BE8E" bold="true" box="[189,303,1177,1203]" fontSize="11" level="3" pageId="80" pageNumber="81" reason="3">
<emphasis id="BB80E374FFDBFFD4FF5A0786FE83BE8E" bold="true" box="[189,303,1177,1203]" pageId="80" pageNumber="81">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="894B3F66FFDBFFD4FF5A07D8FCD1BF76" blockId="80.[189,1399,1223,1355]" pageId="80" pageNumber="81">
The antennifer spine on the right side of the head of the
<typeStatus id="564F81C4FFDBFFD4FCB607D8FC14BEDC" box="[849,952,1223,1249]" pageId="80" pageNumber="81" type="holotype">holotype</typeStatus>
is absent. We assume it was damaged since three species,
<taxonomicName id="4EF444E5FFDBFFD4FE4907F4FDF5BF39" authorityName="Villiers" authorityYear="1964" box="[430,601,1258,1284]" class="Insecta" family="Reduviidae" genus="Pseudocethera" kingdom="Animalia" order="Hemiptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="schoutedeni">
<emphasis id="BB80E374FFDBFFD4FE4907F4FDF5BF39" box="[430,601,1258,1284]" italics="true" pageId="80" pageNumber="81">V. schoutedeni</emphasis>
</taxonomicName>
,
<taxonomicName id="4EF444E5FFDBFFD4FD8D07F4FD4EBF39" authorityName="Villiers" authorityYear="1963" box="[618,738,1258,1284]" class="Insecta" family="Reduviidae" genus="Pseudocethera" kingdom="Animalia" order="Hemiptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="monodi">
<emphasis id="BB80E374FFDBFFD4FD8D07F4FD4EBF39" box="[618,738,1258,1284]" italics="true" pageId="80" pageNumber="81">V. monodi</emphasis>
</taxonomicName>
, and
<taxonomicName id="4EF444E5FFDBFFD4FCC007F4FC2FBF39" authorityName="Villiers" authorityYear="1963" box="[807,899,1258,1284]" class="Insecta" family="Reduviidae" genus="Pseudocethera" kingdom="Animalia" order="Hemiptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="ifana">
<emphasis id="BB80E374FFDBFFD4FCC007F4FC2FBF39" box="[807,899,1258,1284]" italics="true" pageId="80" pageNumber="81">V. ifana</emphasis>
</taxonomicName>
, have similar long spines that are paired. The pygophore of specimen AMNH_PBI 00170712 was used for DNA extraction, but we did not get sufficient DNA in our NGS library for Illumina sequencing.
</paragraph>
</subSubSection>
</treatment>
</document>
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