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<document id="EF6EB4C26F100B59D5E0AF30955C19CE" ID-DOI="10.18590/euscorpius.2011.vol2011.iss129.1" ID-ISSN="1536-9307" ID-Zenodo-Dep="12782418" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721381368639" checkinUser="felipe" docAuthor="Kovařík, František" docDate="2011" docId="03FB8791A07D4778FE98D17A0993854F" docLanguage="en" docName="Euscorpius.129.1-18.pdf" docOrigin="Euscorpius 129 (129)" docSource="https://mds.marshall.edu/euscorpius/vol2011/iss129/1/" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Pandinus Thorell 1876" docType="treatment" docVersion="2" lastPageNumber="1" masterDocId="FFC2FFE9A07F477AFFFBD55B0B0E8339" masterDocTitle="A review of the subgenus Pandinus Thorell, 1876 with descriptions of two new species from Uganda and Ethiopia (Scorpiones: Scorpionidae)" masterLastPageNumber="18" masterPageNumber="1" pageNumber="1" updateTime="1721382788196" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="A6A63EDBFA2D4496E7C56BB2661C6C6B">A review of the subgenus Pandinus Thorell, 1876 with descriptions of two new species from Uganda and Ethiopia (Scorpiones: Scorpionidae)</mods:title>
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<mods:namePart id="83A540BB7364A581D8E9169B3A43A7B5">Kovařík, František</mods:namePart>
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<mods:title id="7279FEA9EC059E9841B6F0F22ABACA08">Euscorpius</mods:title>
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<taxonomicName id="4C524D04A07D4778FE98D17A09718705" ID-CoL="63JDT" authority="Thorell, 1876" authorityName="Thorell" authorityYear="1876" box="[355,639,1057,1084]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis id="B926EA95A07D4778FE98D17A0AD98705" bold="true" box="[355,471,1057,1084]" italics="true" pageId="2" pageNumber="1">Pandinus</emphasis>
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<bibRefCitation id="EFC34B76A07D4778FE24D17909718705" author="THORELL" box="[479,639,1058,1084]" pageId="2" pageNumber="1" pagination="1 - 15" refId="ref7761" refString="THORELL, T. 1876. On the classification of Scorpions. Annals and Magazine of Natural History, 4 (17): 1 - 15." type="journal article" year="1876">Thorell, 1876</bibRefCitation>
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(
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<figureCitation id="13692A02A07D4778FE4CD11809248763" box="[439,554,1091,1114]" captionStart-0="Figures 1–5" captionStart-1="Figures 6–9" captionStart-10="Figure 42" captionStart-2="Figures 10–13" captionStart-3="Figures 14–16" captionStart-4="Figures 17–19" captionStart-5="Figures 20–22" captionStart-6="Figures 23–28" captionStart-7="Figure 29" captionStart-8="Figures 30–32" captionStart-9="Figures 33–41" captionStartId-0="3.[192,278,1871,1894]" captionStartId-1="5.[192,278,1860,1883]" captionStartId-10="16.[192,268,1821,1844]" captionStartId-2="6.[192,278,1846,1869]" captionStartId-3="7.[192,278,1786,1809]" captionStartId-4="9.[192,278,1832,1855]" captionStartId-5="10.[192,278,1862,1885]" captionStartId-6="12.[192,278,1846,1869]" captionStartId-7="13.[192,268,1230,1253]" captionStartId-8="14.[192,278,1856,1879]" captionStartId-9="15.[192,278,1768,1791]" captionTargetBox-0="[204,1394,193,1835]" captionTargetBox-1="[236,1405,167,1829]" captionTargetBox-10="[196,1437,169,1789]" captionTargetBox-2="[224,1401,197,1813]" captionTargetBox-3="[213,1417,167,1753]" captionTargetBox-4="[211,1417,176,1801]" captionTargetBox-5="[211,1417,167,1831]" captionTargetBox-6="[205,1424,193,1813]" captionTargetBox-7="[210,1424,195,1198]" captionTargetBox-8="[191,1438,159,1825]" captionTargetBox-9="[251,1403,194,1737]" captionTargetId-0="figure-49@3.[204,1408,192,1841]" captionTargetId-1="figure-6@5.[237,1404,168,1828]" captionTargetId-10="figure-14@16.[192,1440,165,1800]" captionTargetId-2="figure-10@6.[192,1437,192,1816]" captionTargetId-3="figure-107@7.[214,1416,168,1752]" captionTargetId-4="figure-6@9.[212,1416,177,1800]" captionTargetId-5="figure-14@10.[212,1416,168,1830]" captionTargetId-6="figure-14@12.[192,1440,192,1816]" captionTargetId-7="figure-6@13.[192,1440,192,1200]" captionTargetId-8="figure-14@14.[192,1437,160,1824]" captionTargetId-9="figure-61@15.[216,1418,191,1752]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-10="16" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="9" captionTargetPageId-5="10" captionTargetPageId-6="12" captionTargetPageId-7="13" captionTargetPageId-8="14" captionTargetPageId-9="15" captionText-0="Figures 1–5: Pandinus mazuchi sp. n., ♀ (92.5 mm) holotype. 1–2. dorsal and ventral views. 3. Pectinal area. 4. Movable fin-" captionText-1="Figures 6–9: Pandinus mazuchi sp. n., ♀ holotype photographed at the type locality (6–7) and the type locality, Ethiopia, Jijiga env., 09°20'15.8"N 42°42'17.5"E, 2100 m a.s.l. (8–9)." captionText-10="Figure 42: Distribution map of the subgenus Pandinus. Localities of P. imperator and P. gambiensis are after Prendini (2004: 255, fig. 13), localities of other species are new records except Hargeysa (type locality of P. smithi). Type locality of P. phillipsii corresponds with new records (Goolis Mts.). Other localities of P. phillipsii and P. smithi cited by older authors (Borelli, 1919; Pocock, 1896, 1897, and 1900) are not included in the map. Exact distributions of these species need to be further investigated." captionText-2="Figures 10–13: Pandinus phillipsii (Pocock, 1896). 10–11. ♂ (125 mm), dorsal and ventral views, Somaliland, 15 km N of Sheikh, Goolis Mts., 09°58.927'N 45°10.377'E, 1247 m a.s.l. 12–13. ♀ (109 mm), dorsal and ventral views, Somaliland, Sheikh, 09°57'25.9"N 45°09'52.2"E, 1492 m a.s.l." captionText-3="Figures 14–16: Pandinus phillipsii (Pocock, 1896). Juvenile (80 mm) at the locality Somaliland, Sheikh, Goolis Mts., 09°56'23"N 45°11'14.2"E, 1439 m a.s.l. The large photo (14) is an overall view after overturning a smaller boulder. The smallest photo (15) shows a shallow depression under the boulder in which the scorpion was hiding and underwent an ecdysis (the same spot is shown in the large photo by an arrow). Small arrow points to a partial exuvia of this specimen. The lower photo (16) shows the entire juvenile after removal from the hiding place." captionText-4="Figures 17–19: Pandinus phillipsii (Pocock, 1896). 17
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)
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</paragraph>
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<paragraph id="8BED3687A07D4778FF3BD1DC092887C3" blockId="2.[192,801,1159,1274]" pageId="2" pageNumber="1">
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<treatmentCitation id="0AF31096A07D4778FF3BD1DC090487A7" author="THORELL" box="[192,522,1159,1182]" page="12" pageId="2" pageNumber="1" year="1876">
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<taxonomicName id="4C524D04A07D4778FF3BD1DC090487A7" ID-CoL="63JDT" authority="Thorell, 1876: 12" authorityName="Thorell" authorityPageNumber="12" authorityYear="1876" box="[192,522,1159,1182]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis id="B926EA95A07D4778FF3BD1DC0A2B87A7" box="[192,293,1159,1182]" italics="true" pageId="2" pageNumber="1">Pandinus</emphasis>
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<bibRefCitation id="EFC34B76A07D4778FECCD1DC090487A7" author="THORELL" box="[311,522,1159,1182]" pageId="2" pageNumber="1" pagination="1 - 15" refId="ref7761" refString="THORELL, T. 1876. On the classification of Scorpions. Annals and Magazine of Natural History, 4 (17): 1 - 15." type="journal article" year="1876">Thorell, 1876: 12</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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;
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<treatmentCitation id="0AF31096A07D4778FDDAD1DC081387A7" author="KRAEPELIN" box="[545,797,1159,1182]" page="116" pageId="2" pageNumber="1" year="1899">
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<bibRefCitation id="EFC34B76A07D4778FDDAD1DC081387A7" author="KRAEPELIN" box="[545,797,1159,1182]" pageId="2" pageNumber="1" refId="ref7357" refString="KRAEPELIN, K. 1899. Scorpiones und Pedipalpi. In F. Dahl (ed.), Das Tierreich. Herausgegeben von der Deutschen Zoologischen Gesellschaft. Berlin: R. Friedlander und Sohn Verlag, 8. Lieferung. 265 pp." type="book" year="1899">Kraepelin, 1899: 116</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0AF31096A07D4778FF3BD1FE0A968785" author="VACHON" box="[192,408,1189,1212]" page="953" pageId="2" pageNumber="1" year="1974">
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<bibRefCitation id="EFC34B76A07D4778FF3BD1FE0A968785" author="VACHON" box="[192,408,1189,1212]" pageId="2" pageNumber="1" pagination="857 - 958" refId="ref7847" refString="VACHON, M. 1974. Etude des caracteres utilises pour classer les familles et les genres de Scorpions. Bulletin du Museum national d´Histoire naturelle, Paris, 140: 857 - 958." type="journal article" year="1974">Vachon, 1974: 953</bibRefCitation>
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</treatmentCitation>
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, figs. 113-118;
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<treatmentCitation id="0AF31096A07D4778FDB7D1FE08138785" author="SISSOM" box="[588,797,1189,1212]" page="136" pageId="2" pageNumber="1" year="1990">
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<bibRefCitation id="EFC34B76A07D4778FDB7D1FE08138785" author="SISSOM" box="[588,797,1189,1212]" pageId="2" pageNumber="1" pagination="64 - 160" refId="ref7714" refString="SISSOM, W. D. 1990. Systematics, biogeography and paleontology. Pp. 64 - 160. In: Polis, G. A. (ed.): The Biology of Scorpions. Stanford: Stanford University Psress, 587 pp." type="book chapter" year="1990">Sissom, 1990: 136</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0AF31096A07D4778FF3BD19F0A6087E2" author="FET" box="[192,366,1220,1243]" page="248" pageId="2" pageNumber="1" year="1997">
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<bibRefCitation id="EFC34B76A07D4778FF3BD19F0A6087E2" author="FET" box="[192,366,1220,1243]" pageId="2" pageNumber="1" pagination="245 - 250" refId="ref6958" refString="FET, V. 1997. Notes on the taxonomy of some Old World scorpions (Scorpiones: Buthidae, Chactidae, Ischnuridae, Scorpionidae). Journal of Arachnology, 25: 245 - 250." type="book chapter" year="1997">Fet, 1997: 248</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0AF31096A07D4778FE79D19F093E87E2" author="FET" box="[386,560,1220,1243]" page="465" pageId="2" pageNumber="1" year="2000">
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<bibRefCitation id="EFC34B76A07D4778FE79D19F093E87E2" author="FET" box="[386,560,1220,1243]" pageId="2" pageNumber="1" refId="ref6995" refString="FET, V., W. D. SISSOM, G. LOWE & M. E. BRAUNWALDER. 2000. Catalog of the Scorpions of the World (1758 - 1998). New York: The New York Entomological Society, 689 pp." type="book" year="2000">Fet, 2000: 465</bibRefCitation>
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</treatmentCitation>
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; Prendini, 2000: 44;
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<treatmentCitation id="0AF31096A07D4778FF3BD1B80A8B87C3" author="KOVARIK" box="[192,389,1251,1274]" page="50" pageId="2" pageNumber="1" year="2009">
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<bibRefCitation id="EFC34B76A07D4778FF3BD1B80A8B87C3" author="KOVARIK" box="[192,389,1251,1274]" pageId="2" pageNumber="1" refId="ref7185" refString="KOVARIK, F. 2009. Illustrated catalog of scorpions. Part I. Introductory remarks; keys to families and genera; subfamily Scorpioninae with keys to Heterometrus and Pandinus species. Prague: Clairon Production, 170 pp." type="book" year="2009">Kovařík, 2009: 50</bibRefCitation>
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</treatmentCitation>
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, figs. 284-420.
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</paragraph>
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</subSubSection>
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.
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<taxonomicName id="4C524D04A07D4778FEA5D07D09D38604" authority="C. L. Koch, 1841" authorityName="C. L. Koch" authorityYear="1841" box="[350,733,1318,1341]" class="Arachnida" family="Buthidae" genus="Buthus" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="imperator">
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<emphasis id="B926EA95A07D4778FEA5D07D09148604" box="[350,538,1318,1341]" italics="true" pageId="2" pageNumber="1">Buthus imperator</emphasis>
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C. L.
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<bibRefCitation id="EFC34B76A07D4778FDA6D07D09D38604" author="KOCH" box="[605,733,1318,1341]" pageId="2" pageNumber="1" pagination="695 - 698" refId="ref7043" refString="KOCH, C. L. 1841. Die Arachniden, 9 (1842): 108 pp. (Figs. 695 - 698), Nurnberg." type="book chapter" year="1841">Koch, 1841</bibRefCitation>
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</taxonomicName>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C348650CA07D4778FF3BD0310993854F" pageId="2" pageNumber="1" type="diagnosis">
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<paragraph id="8BED3687A07D4778FF3BD0310993854F" blockId="2.[192,802,1386,1654]" pageId="2" pageNumber="1">
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<smallCapsWord id="8D0BA05BA07D4778FF3BD0310A3886B9" baselines="1403,1403" box="[192,310,1386,1409]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="2" pageNumber="1">DIAGNOSIS</smallCapsWord>
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. Total length
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<quantity id="4CAA9B62A07D4778FE25D031096686B8" box="[478,616,1386,1409]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" metricValueMax="2.2" metricValueMin="0.6" pageId="2" pageNumber="1" unit="mm" value="140.0" valueMax="220.0" valueMin="60.0">60–220 mm</quantity>
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. Pedipalp femur with 3 trichobothria, only one of them on internal surface. Pedipalp patella with 13–16 external and numerous (usually about 30) ventral trichobothria. Retrolateral pedal spurs absent. Lateroapical margins of tarsi produced into rounded lobes. Metasomal segments I–IV with paired ventral submedian carinae. Stridulatory organ located on opposing surfaces of pedipalp coxa and first leg. Telson without subaculear tubercle.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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