treatments-xml/data/D0/2B/09/D02B095EFFDE0A59FF7CF957FDB4F938.xml

<document id="4083324696D96A9FC8177BEC29896B8A" ID-DOI="10.11646/zootaxa.4995.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="5043649" ID-ZooBank="EE7B9315-2E32-4901-B6FE-D205E9178906" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624998663445" checkinUser="plazi" docAuthor="Joharchi, Omid &amp; Friedrich, Stefan" docDate="2021" docId="D02B095EFFDE0A59FF7CF957FDB4F938" docLanguage="en" docName="zootaxa.4995.1.3.pdf" docOrigin="Zootaxa 4995 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gaeolaelaps acanthopedus Joharchi &amp; Friedrich 2021, sp. nov." docType="treatment" docVersion="3" lastPageNumber="62" masterDocId="2C127126FFDF0A5FFFEBFF91FF93FFD9" masterDocTitle="Two new species of Gaeolaelaps Evans &amp; Till (Acari: Laelapidae) from the Andes Mountains, Peru" masterLastPageNumber="70" masterPageNumber="56" pageNumber="57" updateTime="1698978981213" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="DD9CE5F675CA6E006FD4A786A4108C6A">Two new species of Gaeolaelaps Evans &amp; Till (Acari: Laelapidae) from the Andes Mountains, Peru</mods:title>
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<mods:namePart id="ADD71E1C1A9026F2E6A78F96D770F8ED">Joharchi, Omid</mods:namePart>
<mods:affiliation id="2CF7FE4BE05B9817539FF87972453269">Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, Tyumen, Russia.</mods:affiliation>
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<mods:namePart id="F0997774CC429759883FD7159C142886">Friedrich, Stefan</mods:namePart>
<mods:affiliation id="DF83E47A8E2566353670A6FB9F3F4927">SNSB-Bavarian State Collection of Zoology (ZSM), Munich, Germany.</mods:affiliation>
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<mods:title id="59BE9AEBC63191E2A2F1DEFC2D3D44BB">Zootaxa</mods:title>
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<mods:date id="2767E4F4F0C0373F599834A662DACAFB">2021</mods:date>
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<mods:number id="5CD679F1D89E618620F29A618C049869">2021-06-29</mods:number>
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<mods:number id="87C99C10F460356AD0E23D848D5D1EB3">4995</mods:number>
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<treatment id="D02B095EFFDE0A59FF7CF957FDB4F938" LSID="urn:lsid:plazi:treatment:D02B095EFFDE0A59FF7CF957FDB4F938" httpUri="http://treatment.plazi.org/id/D02B095EFFDE0A59FF7CF957FDB4F938" lastPageId="6" lastPageNumber="62" pageId="1" pageNumber="57">
<subSubSection id="1098EBC3FFDE0A5EFF7CF957FDD3F938" box="[151,576,1734,1761]" pageId="1" pageNumber="57" type="nomenclature">
<paragraph id="583DB848FFDE0A5EFF7CF957FDD3F938" blockId="1.[151,576,1734,1797]" box="[151,576,1734,1761]" pageId="1" pageNumber="57">
<heading id="03750F24FFDE0A5EFF7CF957FDD3F938" bold="true" box="[151,576,1734,1761]" fontSize="11" level="1" pageId="1" pageNumber="57" reason="1">
<emphasis id="6AF6645AFFDE0A5EFF7CF957FDD3F938" bold="true" box="[151,576,1734,1761]" pageId="1" pageNumber="57">
<taxonomicName id="9F82C3CBFFDE0A5EFF7CF957FE48F938" authority="Joharchi &amp; Friedrich, 2021" authorityName="Joharchi &amp; Friedrich" authorityYear="2021" box="[151,475,1734,1761]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="57" phylum="Arthropoda" rank="species" species="acanthopedus" status="sp. nov.">
<emphasis id="6AF6645AFFDE0A5EFF7CF957FE48F938" bold="true" box="[151,475,1734,1761]" italics="true" pageId="1" pageNumber="57">Gaeolaelaps acanthopedus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C5D921FFDE0A5EFE09F956FDD3F938" box="[482,576,1735,1761]" pageId="1" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1098EBC3FFDE0A5EFF7CF97AFEAFF8DC" box="[151,316,1771,1797]" pageId="1" pageNumber="57" type="description">
<paragraph id="583DB848FFDE0A5EFF7CF97AFEAFF8DC" blockId="1.[151,576,1734,1797]" box="[151,316,1771,1797]" pageId="1" pageNumber="57">
(
<figureCitation id="C0B9A4CDFFDE0A5EFF74F97AFEA6F8DC" box="[159,309,1771,1797]" captionStart-0="FIGURES 3–8" captionStart-1="FIGURES 9–15" captionStart-2="FIGURES 16–19" captionStartId-0="3.[151,264,1943,1967]" captionStartId-1="4.[151,264,1943,1967]" captionStartId-2="5.[151,264,1961,1985]" captionTargetBox-0="[191,1395,196,1907]" captionTargetBox-1="[180,1404,191,1913]" captionTargetBox-2="[161,1422,214,1924]" captionTargetId-0="figure-27@3.[179,1408,181,1919]" captionTargetId-1="figure-17@4.[175,1413,181,1920]" captionTargetId-2="figure-27@5.[154,1433,199,1937]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURES 3–8. Gaeolaelaps acanthopedus sp. nov., female. 3, dorsal idiosoma; 4, ventral idiosoma; 5, subcapitulum; 6, supralabral process; 7, epistome; 8, chelicera." captionText-1="FIGURES 9–15. DIC micrographs of Gaeolaelaps acanthopedus sp. nov., female. 9, idiosoma in dorsal view; 10, idiosoma in ventral view; 11, sternal, genital &amp; anal shields; 12, palp, with a focus on apotele; 13, epistome; 14, insemination structures; 15, chelicera." captionText-2="FIGURES 16–19. Gaeolaelaps acanthopedus sp. nov., female. 16, leg I (trochanter-tibia); 17, leg II; 18, leg III; 19, leg IV." figureDoi-0="http://doi.org/10.5281/zenodo.5043654" figureDoi-1="http://doi.org/10.5281/zenodo.5043656" figureDoi-2="http://doi.org/10.5281/zenodo.5043658" httpUri-0="https://zenodo.org/record/5043654/files/figure.png" httpUri-1="https://zenodo.org/record/5043656/files/figure.png" httpUri-2="https://zenodo.org/record/5043658/files/figure.png" pageId="1" pageNumber="57">Figures 3–19</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1098EBC3FFDE0A5EFF7CF8A2FC29F84C" pageId="1" pageNumber="57" type="materials_examined">
<paragraph id="583DB848FFDE0A5EFF7CF8A2FC29F84C" blockId="1.[151,1437,1843,2013]" pageId="1" pageNumber="57">
<emphasis id="6AF6645AFFDE0A5EFF7CF8A2FEDBF894" bold="true" box="[151,328,1843,1869]" pageId="1" pageNumber="57">Type material:</emphasis>
<materialsCitation id="E8EAB215FFDE0A5EFEA4F8A2FA0AF8A8" collectingDate="2016-04-15" collectingDateMax="2016-04-17" collectingDateMin="2016-04-15" collectionCode="MUSM" country="Peru" elevation="2975" latitude="-9.908333" location="Monte Potrero" longLatPrecision="21" longitude="-76.06334" municipality="South America" pageId="1" pageNumber="57" specimenCount="1" stateProvince="Huanuco" typeStatus="holotype">
<typeStatus id="873906EAFFDE0A5EFEA4F8A2FE29F894" box="[335,442,1843,1869]" pageId="1" pageNumber="57" type="holotype">Holotype</typeStatus>
, female, 
<location id="5D5DEE93FFDE0A5EFDCBF8A2FD56F894" LSID="urn:lsid:plazi:treatment:D02B095EFFDE0A59FF7CF957FDB4F938:5D5DEE93FFDE0A5EFDCBF8A2FD56F894" box="[544,709,1843,1869]" country="Peru" latitude="-9.908333" longLatPrecision="21" longitude="-76.06334" municipality="South America" name="Monte Potrero" pageId="1" pageNumber="57" stateProvince="Huanuco">Monte Potrero</location>
, 
<locationDeviation id="498FA16AFFDE0A5EFD3BF8A2FCF6F894" box="[720,869,1843,1869]" location="Molino" pageId="1" pageNumber="57">W of Molino</locationDeviation>
, 
<collectingRegion id="9A4676AAFFDE0A5EFC9AF8A2FB32F894" box="[881,1185,1843,1869]" country="Peru" name="Huanuco" pageId="1" pageNumber="57">Departamento de Huánuco</collectingRegion>
, 
<collectingCountry id="2095F8D8FFDE0A5EFB47F8A2FB71F894" box="[1196,1250,1843,1869]" name="Peru" pageId="1" pageNumber="57">Peru</collectingCountry>
, 
<collectingMunicipality id="B8592232FFDE0A5EFB05F8A2FA0BF894" box="[1262,1432,1843,1869]" pageId="1" pageNumber="57">South America</collectingMunicipality>
, 
<geoCoordinate id="3DB6DE8FFFDE0A5EFF7CF8C6FEB3F8AB" box="[151,288,1879,1906]" degrees="09" direction="south" minutes="54" orientation="latitude" pageId="1" pageNumber="57" precision="15" seconds="30" value="-9.908333">09°54’30’’S</geoCoordinate>
<geoCoordinate id="3DB6DE8FFFDE0A5EFEC3F8C6FE2FF8A8" box="[296,444,1879,1905]" degrees="76" direction="west" minutes="03" orientation="longitude" pageId="1" pageNumber="57" precision="15" seconds="48" value="-76.06334">76°03’48’’W</geoCoordinate>
, 
<date id="2C3C9E88FFDE0A5EFE23F8C6FD00F8A8" box="[456,659,1879,1906]" pageId="1" pageNumber="57" value="2016-04-15" valueMax="2016-04-17" valueMin="2016-04-15">
<collectingDate id="3C786760FFDE0A5EFE23F8C6FD00F8A8" box="[456,659,1879,1906]" pageId="1" pageNumber="57" value="2016-04-15" valueMax="2016-04-17" valueMin="2016-04-15">15–17 April 2016</collectingDate>
</date>
, alt. 
<quantity id="9F7A15ADFFDE0A5EFD21F8C6FCFAF8A8" box="[714,873,1879,1906]" metricMagnitude="3" metricUnit="m" metricValue="2.975" metricValueMax="3.1" metricValueMin="2.85" pageId="1" pageNumber="57" unit="m" value="2975.0" valueMax="3100.0" valueMin="2850.0">
<elevation id="D3AF5F7BFFDE0A5EFD21F8C6FCFAF8A8" box="[714,873,1879,1906]" metricMagnitude="3" metricUnit="m" metricValue="2.975" metricValueMax="3.1" metricValueMin="2.85" pageId="1" pageNumber="57" unit="m" value="2975.0" valueMax="3100.0" valueMin="2850.0">2850–3100 m</elevation>
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, leg. local collectors, from soil-litter (in 
<collectionCode id="3E93208DFFDE0A5EFADDF8C6FA02F8A8" box="[1334,1425,1879,1905]" pageId="1" pageNumber="57">MUSM</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="E8EAB215FFDE0A5EFF7CF8EAFC25F84C" box="[151,950,1915,1941]" collectingDate="2016-04-15" collectingDateMax="2016-04-17" collectingDateMin="2016-04-15" collectionCode="ZSM, TSUMZ" country="Peru" elevation="2975" latitude="-9.908333" location="Monte Potrero" longLatPrecision="21" longitude="-76.06334" municipality="South America" pageId="1" pageNumber="57" specimenCount="1" specimenCount-female="1" stateProvince="Huanuco" typeStatus="paratype">
<typeStatus id="873906EAFFDE0A5EFF7CF8EAFE94F84C" box="[151,263,1915,1941]" pageId="1" pageNumber="57" type="paratype">Paratypes</typeStatus>
, 
<specimenCount id="4E8473C1FFDE0A5EFEF9F8EAFE23F84C" box="[274,432,1915,1941]" pageId="1" pageNumber="57" type="female">seven females</specimenCount>
same data as holotype (in 
<collectionCode id="3E93208DFFDE0A5EFD30F8EAFC86F84C" box="[731,789,1915,1941]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="1" pageNumber="57" type="Museum">ZSM</collectionCode>
and 
<collectionCode id="3E93208DFFDE0A5EFCA1F8EAFC3DF84C" box="[842,942,1915,1941]" pageId="1" pageNumber="57">TSUMZ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="1098EBC3FFDE0A5DFF2CF80EFAFCFD73" lastPageId="2" lastPageNumber="58" pageId="1" pageNumber="57" type="diagnosis">
<paragraph id="583DB848FFDE0A5DFF2CF80EFAFCFD73" blockId="1.[151,1437,1843,2013]" lastBlockId="2.[151,1437,151,682]" lastPageId="2" lastPageNumber="58" pageId="1" pageNumber="57">
<emphasis id="6AF6645AFFDE0A5EFF2CF80EFEA9F860" bold="true" box="[199,314,1951,1977]" pageId="1" pageNumber="57">Diagnosis</emphasis>
(adult female). Dorsal shield with reticulation, more distinct in opisthonotal and lateral regions, with 39 pairs of smooth setae, 22 pairs of podonotal setae and 17 pairs of opisthonotal setae, including two pairs of 
<emphasis id="6AF6645AFFDE0A5EFA6BF855FA0FF804" box="[1408,1436,1988,2013]" italics="true" pageId="1" pageNumber="57">Zx</emphasis>
setae and two unpaired setae 
<emphasis id="6AF6645AFFDD0A5DFE34FF06FE6BFF69" box="[479,504,151,176]" italics="true" pageId="2" pageNumber="58">Jx</emphasis>
between 
<emphasis id="6AF6645AFFDD0A5DFD88FF06FDEEFF68" box="[611,637,151,177]" italics="true" pageId="2" pageNumber="58">J2</emphasis>
and 
<emphasis id="6AF6645AFFDD0A5DFD5FFF06FD5CFF68" box="[692,719,151,177]" italics="true" pageId="2" pageNumber="58">J3</emphasis>
, length of setae decreasing from anterior to posterior, 
<emphasis id="6AF6645AFFDD0A5DFAC4FF06FAD7FF68" box="[1327,1348,151,177]" italics="true" pageId="2" pageNumber="58">j1</emphasis>
longest and 
<emphasis id="6AF6645AFFDD0A5DFF2DFF2AFF70FF0C" box="[198,227,187,213]" italics="true" pageId="2" pageNumber="58">Z5</emphasis>
shorter than 
<emphasis id="6AF6645AFFDD0A5DFE98FF2AFE1BFF0C" box="[371,392,187,213]" italics="true" pageId="2" pageNumber="58">j1</emphasis>
, shield with six pairs of discernible gland pores (
<emphasis id="6AF6645AFFDD0A5DFC4CFF2AFC7EFF0C" box="[935,1005,187,213]" italics="true" pageId="2" pageNumber="58">gd1–2</emphasis>
, 
<emphasis id="6AF6645AFFDD0A5DFC11FF2AFBD3FF0D" box="[1018,1088,186,213]" italics="true" pageId="2" pageNumber="58">gd5–6</emphasis>
, 
<emphasis id="6AF6645AFFDD0A5DFBA6FF2AFB00FF0C" box="[1101,1171,187,213]" italics="true" pageId="2" pageNumber="58">gd8–9</emphasis>
). Presternal area lightly sclerotised, punctate, with a few transverse curved lines, sternal shield with reticulate ornamentation throughout, except central and posterior parts smooth (or faintly reticulated), anterior margin almost straight but posterior margin slightly concave, remnants of gland pores 
<emphasis id="6AF6645AFFDD0A5DFD7EFEB6FD2EFE98" box="[661,701,295,321]" italics="true" pageId="2" pageNumber="58">gv1</emphasis>
apparent near posterior shield margin, ratio of shield length/width ≃ 0.79. Genital shield flask-shaped, elongated and widened posteriorly behind setae 
<emphasis id="6AF6645AFFDD0A5DFB8AFEDAFB11FEBC" box="[1121,1154,331,357]" italics="true" pageId="2" pageNumber="58">st5</emphasis>
, surface reticulated with irregular, mostly longitudinal lines in anterior region, posteriorly comprising 7–8 elongated oblique irregular cells enclosed behind an inverted V-shaped ridge, anterior margin of shield convex, overlapping posterior area of sternal shield, extending at level of poroids 
<emphasis id="6AF6645AFFDD0A5DFDC5FE26FDC3FE08" box="[558,592,439,465]" italics="true" pageId="2" pageNumber="58">iv2</emphasis>
, with length/width ratio ≃ 2.2. Anal shield pear-shaped, slightly wider than long, with length/width ratio ≃ 0.96, anal opening positioned somewhat centrally on the shield; cribrum extending anterolaterally slightly beyond postanal seta. Peritremes long, extending anteriorly to mid-level of coxae I. Soft opisthogastric cuticle with 10 pairs of setae. Internal malae fringed, with two pairs of projections. Supralabral process with a very long apicoventral projection. Fixed digit of chelicera with eight teeth, including an offset subapical tooth (gabelzahn). Most leg setae on tarsi II–IV spine-like (strongly thickened and apically tapered), ventral setae on tibiae II–IV, genua II–IV and setae 
<emphasis id="6AF6645AFFDD0A5DFDAEFD1EFDF2FD70" box="[581,609,655,681]" italics="true" pageId="2" pageNumber="58">ad</emphasis>
, 
<emphasis id="6AF6645AFFDD0A5DFD84FD1EFD16FD70" box="[623,645,655,681]" italics="true" pageId="2" pageNumber="58">pl</emphasis>
on trochanter IV spur-like (strongly thickened and apically blunt).
</paragraph>
</subSubSection>
<subSubSection id="1098EBC3FFDD0A59FF7CFD46FD3EFBC8" lastPageId="6" lastPageNumber="62" pageId="2" pageNumber="58" type="description">
<paragraph id="583DB848FFDD0A5DFF7CFD46FE5BFD28" blockId="2.[151,1437,727,2013]" box="[151,456,727,753]" pageId="2" pageNumber="58">
<emphasis id="6AF6645AFFDD0A5DFF7CFD46FE11FD28" bold="true" box="[151,386,727,753]" pageId="2" pageNumber="58">
<emphasis id="6AF6645AFFDD0A5DFF7CFD46FE8EFD28" bold="true" box="[151,285,727,753]" italics="true" pageId="2" pageNumber="58">Description</emphasis>
. Female
</emphasis>
(n=8)
</paragraph>
<paragraph id="583DB848FFDD0A5DFF2CFD6AFA03FC34" blockId="2.[151,1437,727,2013]" pageId="2" pageNumber="58">
<emphasis id="6AF6645AFFDD0A5DFF2CFD6AFE11FCCC" box="[199,386,763,789]" italics="true" pageId="2" pageNumber="58">Dorsal idiosoma</emphasis>
(
<figureCitation id="C0B9A4CDFFDD0A5DFE64FD6AFE46FCCC" box="[399,469,763,789]" captionStart="FIGURES 3–8" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[191,1395,196,1907]" captionTargetId="figure-27@3.[179,1408,181,1919]" captionTargetPageId="3" captionText="FIGURES 3–8. Gaeolaelaps acanthopedus sp. nov., female. 3, dorsal idiosoma; 4, ventral idiosoma; 5, subcapitulum; 6, supralabral process; 7, epistome; 8, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5043654" httpUri="https://zenodo.org/record/5043654/files/figure.png" pageId="2" pageNumber="58">Figs 3</figureCitation>
&amp; 
<figureCitation id="C0B9A4CDFFDD0A5DFE1DFD6AFD95FCCC" box="[502,518,763,789]" captionStart="FIGURES 9–15" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[180,1404,191,1913]" captionTargetId="figure-17@4.[175,1413,181,1920]" captionTargetPageId="4" captionText="FIGURES 9–15. DIC micrographs of Gaeolaelaps acanthopedus sp. nov., female. 9, idiosoma in dorsal view; 10, idiosoma in ventral view; 11, sternal, genital &amp; anal shields; 12, palp, with a focus on apotele; 13, epistome; 14, insemination structures; 15, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5043656" httpUri="https://zenodo.org/record/5043656/files/figure.png" pageId="2" pageNumber="58">9</figureCitation>
). Dorsal shield suboval, large, 674–702 long, 390–396 wide, covering most of dorsal idiosoma; reticulate throughout, more distinct in opisthonotal and lateral regions (
<figureCitation id="C0B9A4CDFFDD0A5DFBA1FC8EFB03FCE0" box="[1098,1168,799,825]" captionStart="FIGURES 3–8" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[191,1395,196,1907]" captionTargetId="figure-27@3.[179,1408,181,1919]" captionTargetPageId="3" captionText="FIGURES 3–8. Gaeolaelaps acanthopedus sp. nov., female. 3, dorsal idiosoma; 4, ventral idiosoma; 5, subcapitulum; 6, supralabral process; 7, epistome; 8, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5043654" httpUri="https://zenodo.org/record/5043654/files/figure.png" pageId="2" pageNumber="58">Figs 3</figureCitation>
&amp; 
<figureCitation id="C0B9A4CDFFDD0A5DFB58FC8EFB50FCE0" box="[1203,1219,799,825]" captionStart="FIGURES 9–15" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[180,1404,191,1913]" captionTargetId="figure-17@4.[175,1413,181,1920]" captionTargetPageId="4" captionText="FIGURES 9–15. DIC micrographs of Gaeolaelaps acanthopedus sp. nov., female. 9, idiosoma in dorsal view; 10, idiosoma in ventral view; 11, sternal, genital &amp; anal shields; 12, palp, with a focus on apotele; 13, epistome; 14, insemination structures; 15, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5043656" httpUri="https://zenodo.org/record/5043656/files/figure.png" pageId="2" pageNumber="58">9</figureCitation>
). Shield bearing 39 pairs of smooth setae, 22 pairs of podonotal setae (34–51) and 17 pairs of opisthonotal setae (25–36), including two pairs of 
<emphasis id="6AF6645AFFDD0A5DFF19FCF6FE9DFC59" box="[242,270,871,896]" italics="true" pageId="2" pageNumber="58">Zx</emphasis>
setae and two unpaired setae 
<emphasis id="6AF6645AFFDD0A5DFDB3FCF6FDE2FC59" box="[600,625,871,896]" italics="true" pageId="2" pageNumber="58">Jx</emphasis>
between 
<emphasis id="6AF6645AFFDD0A5DFD31FCF6FD67FC58" box="[730,756,871,897]" italics="true" pageId="2" pageNumber="58">J2</emphasis>
and 
<emphasis id="6AF6645AFFDD0A5DFCC1FCF6FCD6FC58" box="[810,837,871,897]" italics="true" pageId="2" pageNumber="58">J3</emphasis>
, lengths of setae decreasing from anterior to posterior and from central to lateral, 
<emphasis id="6AF6645AFFDD0A5DFE2EFC1AFE48FC7C" box="[453,475,907,933]" italics="true" pageId="2" pageNumber="58">j1</emphasis>
(37–39) longest and 
<emphasis id="6AF6645AFFDD0A5DFD2CFC1AFD73FC7C" box="[711,736,907,933]" italics="true" pageId="2" pageNumber="58">z1</emphasis>
(17–21) shortest, 
<emphasis id="6AF6645AFFDD0A5DFC41FC1AFC56FC7C" box="[938,965,907,933]" italics="true" pageId="2" pageNumber="58">J5</emphasis>
also short (24–26) and 
<emphasis id="6AF6645AFFDD0A5DFB20FC1AFB7AFC7C" box="[1227,1257,907,933]" italics="true" pageId="2" pageNumber="58">Z5</emphasis>
(30–32) shorter than 
<emphasis id="6AF6645AFFDD0A5DFF24FC3EFF77FC10" box="[207,228,943,969]" italics="true" pageId="2" pageNumber="58">j1</emphasis>
. Shield with about 22 pairs of discernible pore-like structures, including 16 poroids (
<emphasis id="6AF6645AFFDD0A5DFB4AFC3EFB56FC10" box="[1185,1221,943,969]" italics="true" pageId="2" pageNumber="58">id1</emphasis>
, 
<emphasis id="6AF6645AFFDD0A5DFB38FC3EFB64FC10" box="[1235,1271,943,969]" italics="true" pageId="2" pageNumber="58">id2</emphasis>
, 
<emphasis id="6AF6645AFFDD0A5DFAEEFC3EFAC9FC11" box="[1285,1370,942,969]" italics="true" pageId="2" pageNumber="58">id4–id6</emphasis>
, 
<emphasis id="6AF6645AFFDD0A5DFA82FC3EFF5CFC35" italics="true" pageId="2" pageNumber="58">idm - idm6</emphasis>
, 
<emphasis id="6AF6645AFFDD0A5DFF36FC42FF6DFC34" box="[221,254,979,1005]" italics="true" pageId="2" pageNumber="58">is1</emphasis>
, 
<emphasis id="6AF6645AFFDD0A5DFEE7FC42FEBDFC34" box="[268,302,979,1005]" italics="true" pageId="2" pageNumber="58">idx</emphasis>
, 
<emphasis id="6AF6645AFFDD0A5DFED7FC42FEFBFC34" box="[316,360,979,1005]" italics="true" pageId="2" pageNumber="58">idl1</emphasis>
, 
<emphasis id="6AF6645AFFDD0A5DFE9DFC42FE32FC34" box="[374,417,979,1005]" italics="true" pageId="2" pageNumber="58">idl3</emphasis>
, 
<emphasis id="6AF6645AFFDD0A5DFE44FC42FE49FC34" box="[431,474,979,1005]" italics="true" pageId="2" pageNumber="58">idl4</emphasis>
) and six gland openings (
<emphasis id="6AF6645AFFDD0A5DFD16FC42FCD0FC34" box="[765,835,979,1005]" italics="true" pageId="2" pageNumber="58">gd1–2</emphasis>
, 
<emphasis id="6AF6645AFFDD0A5DFCBAFC42FC04FC35" box="[849,919,978,1005]" italics="true" pageId="2" pageNumber="58">gd5–6</emphasis>
, 
<emphasis id="6AF6645AFFDD0A5DFC4EFC42FC78FC34" box="[933,1003,979,1005]" italics="true" pageId="2" pageNumber="58">gd8–9</emphasis>
), others indistinct, see 
<figureCitation id="C0B9A4CDFFDD0A5DFB04FC42FACBFC34" box="[1263,1368,979,1005]" captionStart="FIGURES 3–8" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[191,1395,196,1907]" captionTargetId="figure-27@3.[179,1408,181,1919]" captionTargetPageId="3" captionText="FIGURES 3–8. Gaeolaelaps acanthopedus sp. nov., female. 3, dorsal idiosoma; 4, ventral idiosoma; 5, subcapitulum; 6, supralabral process; 7, epistome; 8, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5043654" httpUri="https://zenodo.org/record/5043654/files/figure.png" pageId="2" pageNumber="58">Figures 3</figureCitation>
&amp; 
<figureCitation id="C0B9A4CDFFDD0A5DFA90FC42FA1FFC34" box="[1403,1420,979,1005]" captionStart="FIGURES 9–15" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[180,1404,191,1913]" captionTargetId="figure-17@4.[175,1413,181,1920]" captionTargetPageId="4" captionText="FIGURES 9–15. DIC micrographs of Gaeolaelaps acanthopedus sp. nov., female. 9, idiosoma in dorsal view; 10, idiosoma in ventral view; 11, sternal, genital &amp; anal shields; 12, palp, with a focus on apotele; 13, epistome; 14, insemination structures; 15, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5043656" httpUri="https://zenodo.org/record/5043656/files/figure.png" pageId="2" pageNumber="58">9</figureCitation>
.
</paragraph>
<paragraph id="583DB848FFDD0A5DFF2CFC66FD6BF84C" blockId="2.[151,1437,727,2013]" pageId="2" pageNumber="58">
<emphasis id="6AF6645AFFDD0A5DFF2CFC66FE14FBC8" box="[199,391,1015,1041]" italics="true" pageId="2" pageNumber="58">Ventral idiosoma</emphasis>
(
<figureCitation id="C0B9A4CDFFDD0A5DFE7DFC66FE73FBC8" box="[406,480,1015,1041]" captionStart="FIGURES 3–8" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[191,1395,196,1907]" captionTargetId="figure-27@3.[179,1408,181,1919]" captionTargetPageId="3" captionText="FIGURES 3–8. Gaeolaelaps acanthopedus sp. nov., female. 3, dorsal idiosoma; 4, ventral idiosoma; 5, subcapitulum; 6, supralabral process; 7, epistome; 8, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5043654" httpUri="https://zenodo.org/record/5043654/files/figure.png" pageId="2" pageNumber="58">Figs 4</figureCitation>
, 
<figureCitation id="C0B9A4CDFFDD0A5DFE06FC66FDDFFBC8" box="[493,588,1015,1041]" captionStart="FIGURES 9–15" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[180,1404,191,1913]" captionTargetId="figure-17@4.[175,1413,181,1920]" captionTargetPageId="4" captionText="FIGURES 9–15. DIC micrographs of Gaeolaelaps acanthopedus sp. nov., female. 9, idiosoma in dorsal view; 10, idiosoma in ventral view; 11, sternal, genital &amp; anal shields; 12, palp, with a focus on apotele; 13, epistome; 14, insemination structures; 15, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5043656" httpUri="https://zenodo.org/record/5043656/files/figure.png" pageId="2" pageNumber="58">10 &amp; 11</figureCitation>
). Tritosternum with paired pilose laciniae (98–110), fused basally (10–15), columnar base 41–43 × 18–20 wide; presternal area lightly punctate, with transverse curved lines; sternal shield length 163–167, maximum width 204–211, narrowest between coxae II (125–128), with reticulate ornamentation throughout, except central and posterior parts smooth (or faintly reticulated), anterior margin almost straight, posterior margin slightly concave; bearing three pairs of smooth setae (
<emphasis id="6AF6645AFFDD0A5DFC7FFB16FC27FB78" box="[916,948,1159,1185]" italics="true" pageId="2" pageNumber="58">st1</emphasis>
54–57, 
<emphasis id="6AF6645AFFDD0A5DFBFBFB16FBA2FB78" box="[1040,1073,1159,1185]" italics="true" pageId="2" pageNumber="58">st2</emphasis>
54–57, 
<emphasis id="6AF6645AFFDD0A5DFB66FB16FB3DFB78" box="[1165,1198,1159,1185]" italics="true" pageId="2" pageNumber="58">st3</emphasis>
49–51), 
<emphasis id="6AF6645AFFDD0A5DFAFFFB16FAA7FB78" box="[1300,1332,1159,1185]" italics="true" pageId="2" pageNumber="58">st2</emphasis>
reaching to base of 
<emphasis id="6AF6645AFFDD0A5DFEE5FB3AFEA3FB1C" box="[270,304,1195,1221]" italics="true" pageId="2" pageNumber="58">st3</emphasis>
, and two pairs of slit-like poroids (lyrifissures), a pair of lyrifissures (
<emphasis id="6AF6645AFFDD0A5DFBA6FB3AFBFCFB1C" box="[1101,1135,1195,1221]" italics="true" pageId="2" pageNumber="58">iv1</emphasis>
) adjacent to setae 
<emphasis id="6AF6645AFFDD0A5DFAAFFB3AFAF6FB1C" box="[1348,1381,1195,1221]" italics="true" pageId="2" pageNumber="58">st1</emphasis>
, and another pair of lyrifissures (
<emphasis id="6AF6645AFFDD0A5DFE3CFB5EFE69FB30" box="[471,506,1231,1257]" italics="true" pageId="2" pageNumber="58">iv2</emphasis>
) between 
<emphasis id="6AF6645AFFDD0A5DFD9AFB5EFD01FB30" box="[625,658,1231,1257]" italics="true" pageId="2" pageNumber="58">st2</emphasis>
and 
<emphasis id="6AF6645AFFDD0A5DFD27FB5EFD7EFB30" box="[716,749,1231,1257]" italics="true" pageId="2" pageNumber="58">st3</emphasis>
; remnants of gland pores 
<emphasis id="6AF6645AFFDD0A5DFBF1FB5EFBD1FB30" box="[1050,1090,1231,1257]" italics="true" pageId="2" pageNumber="58">gv1</emphasis>
apparent near posterior shield margin. Metasternal setae 
<emphasis id="6AF6645AFFDD0A5DFE2CFB62FE7BFAD4" box="[455,488,1267,1293]" italics="true" pageId="2" pageNumber="58">st4</emphasis>
(37–40) and metasternal poroids located on soft integument; metasternal platelets absent. Endopodal plates between coxae I–II (bearing gland pores 
<emphasis id="6AF6645AFFDD0A5DFC93FA86FC33FAE8" box="[888,928,1303,1329]" italics="true" pageId="2" pageNumber="58">gvb</emphasis>
) and II–III completely fused to sternal shield, endopodal plates III/IV elongate, narrow and curved. Genital shield flask-shaped, elongate and widened posteriorly behind setae 
<emphasis id="6AF6645AFFDD0A5DFECCFACEFED4FAA0" box="[295,327,1375,1401]" italics="true" pageId="2" pageNumber="58">st5</emphasis>
, length 322–330, maximum width 143–149, anterior margin of shield convex, overlapping posterior area of sternal shield, extending at level of poroids 
<emphasis id="6AF6645AFFDD0A5DFD3AFA12FD67FA44" box="[721,756,1411,1437]" italics="true" pageId="2" pageNumber="58">iv2</emphasis>
, posterior margin rounded, surface reticulated with irregular, mostly longitudinal lines in anterior region, posteriorly comprising 7–8 elongated oblique irregular cells enclosed behind an inverted V-shaped ridge (
<figureCitation id="C0B9A4CDFFDD0A5DFDDAFA5AFDE9FA3C" box="[561,634,1483,1509]" captionStart="FIGURES 3–8" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[191,1395,196,1907]" captionTargetId="figure-27@3.[179,1408,181,1919]" captionTargetPageId="3" captionText="FIGURES 3–8. Gaeolaelaps acanthopedus sp. nov., female. 3, dorsal idiosoma; 4, ventral idiosoma; 5, subcapitulum; 6, supralabral process; 7, epistome; 8, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5043654" httpUri="https://zenodo.org/record/5043654/files/figure.png" pageId="2" pageNumber="58">Figs 4</figureCitation>
&amp; 
<figureCitation id="C0B9A4CDFFDD0A5DFD49FA5AFD2CFA3C" box="[674,703,1483,1509]" captionStart="FIGURES 9–15" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[180,1404,191,1913]" captionTargetId="figure-17@4.[175,1413,181,1920]" captionTargetPageId="4" captionText="FIGURES 9–15. DIC micrographs of Gaeolaelaps acanthopedus sp. nov., female. 9, idiosoma in dorsal view; 10, idiosoma in ventral view; 11, sternal, genital &amp; anal shields; 12, palp, with a focus on apotele; 13, epistome; 14, insemination structures; 15, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5043656" httpUri="https://zenodo.org/record/5043656/files/figure.png" pageId="2" pageNumber="58">11</figureCitation>
). Paragenital poroids 
<emphasis id="6AF6645AFFDD0A5DFC51FA5AFC4FFA3C" box="[954,988,1483,1509]" italics="true" pageId="2" pageNumber="58">iv5</emphasis>
located on soft cuticle lateral to shield near seta 
<emphasis id="6AF6645AFFDD0A5DFEE8FA7EFEB0F9D0" box="[259,291,1519,1545]" italics="true" pageId="2" pageNumber="58">st5</emphasis>
. Anal shield pear-shaped, rounded anteriorly, length 84–93, width 93–96, anterior half lineate-reticulate, para-anal setae (27–30) longer than post-anal seta (22–25), cribrum with 3–4 irregular rows of spicules and a pair of anterior arms extending slightly beyond postanal seta (
<figureCitation id="C0B9A4CDFFDD0A5DFCA5F9A6FC06F988" box="[846,917,1591,1617]" captionStart="FIGURES 3–8" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[191,1395,196,1907]" captionTargetId="figure-27@3.[179,1408,181,1919]" captionTargetPageId="3" captionText="FIGURES 3–8. Gaeolaelaps acanthopedus sp. nov., female. 3, dorsal idiosoma; 4, ventral idiosoma; 5, subcapitulum; 6, supralabral process; 7, epistome; 8, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5043654" httpUri="https://zenodo.org/record/5043654/files/figure.png" pageId="2" pageNumber="58">Figs 4</figureCitation>
&amp; 
<figureCitation id="C0B9A4CDFFDD0A5DFC52F9A6FC45F988" box="[953,982,1591,1617]" captionStart="FIGURES 9–15" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[180,1404,191,1913]" captionTargetId="figure-17@4.[175,1413,181,1920]" captionTargetPageId="4" captionText="FIGURES 9–15. DIC micrographs of Gaeolaelaps acanthopedus sp. nov., female. 9, idiosoma in dorsal view; 10, idiosoma in ventral view; 11, sternal, genital &amp; anal shields; 12, palp, with a focus on apotele; 13, epistome; 14, insemination structures; 15, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5043656" httpUri="https://zenodo.org/record/5043656/files/figure.png" pageId="2" pageNumber="58">11</figureCitation>
); anal gland pores (
<emphasis id="6AF6645AFFDD0A5DFB5FF9A6FB4FF988" box="[1204,1244,1591,1617]" italics="true" pageId="2" pageNumber="58">gv3</emphasis>
) on anterolateral margin of anal shield. Primary (most lateral) metapodal platelets suboval (27–30 long × 13–10 wide), secondary and tertiary (18–22 long × 5–8 wide) more median, smaller, oriented obliquely. Soft opisthogastric cuticle with ten pairs of smooth setae (
<emphasis id="6AF6645AFFDD0A5DFE77F932FE6BF964" box="[412,504,1699,1725]" italics="true" pageId="2" pageNumber="58">Jv1–Jv5</emphasis>
, 
<emphasis id="6AF6645AFFDD0A5DFDECF932FDFAF964" box="[519,617,1699,1725]" italics="true" pageId="2" pageNumber="58">Zv1–Zv4</emphasis>
&amp; R) [
<emphasis id="6AF6645AFFDD0A5DFD55F932FC89F964" box="[702,794,1699,1725]" italics="true" pageId="2" pageNumber="58">Jv1–Jv3</emphasis>
&amp; 
<emphasis id="6AF6645AFFDD0A5DFCAAF932FC37F964" box="[833,932,1699,1725]" italics="true" pageId="2" pageNumber="58">Zv1–Zv2</emphasis>
(29–35), 
<emphasis id="6AF6645AFFDD0A5DFBFFF932FBE2F964" box="[1044,1137,1699,1725]" italics="true" pageId="2" pageNumber="58">Jv4–Jv5</emphasis>
&amp; 
<emphasis id="6AF6645AFFDD0A5DFB7CF932FB6AF964" box="[1175,1273,1699,1725]" italics="true" pageId="2" pageNumber="58">Zv4–Zv5</emphasis>
(23–26)], and four pairs of poroids (three 
<emphasis id="6AF6645AFFDD0A5DFE20F956FE7EF939" box="[459,493,1735,1760]" italics="true" pageId="2" pageNumber="58">ivo</emphasis>
&amp; 
<emphasis id="6AF6645AFFDD0A5DFDFAF956FDA0F939" box="[529,563,1735,1760]" italics="true" pageId="2" pageNumber="58">ivp</emphasis>
). Exopodal platelets between coxae II–III subtriangular, elongate, narrow and curved, those behind coxae III–IV and parapodal platelets fused, surrounding coxa IV, gland pores 
<emphasis id="6AF6645AFFDD0A5DFB36F97AFA95F8DC" box="[1245,1286,1771,1797]" italics="true" pageId="2" pageNumber="58">gv2</emphasis>
on parapodal platelets (
<figureCitation id="C0B9A4CDFFDD0A5DFEEEF89EFEDFF8F0" box="[261,332,1807,1833]" captionStart="FIGURES 3–8" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[191,1395,196,1907]" captionTargetId="figure-27@3.[179,1408,181,1919]" captionTargetPageId="3" captionText="FIGURES 3–8. Gaeolaelaps acanthopedus sp. nov., female. 3, dorsal idiosoma; 4, ventral idiosoma; 5, subcapitulum; 6, supralabral process; 7, epistome; 8, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5043654" httpUri="https://zenodo.org/record/5043654/files/figure.png" pageId="2" pageNumber="58">Fig. 4</figureCitation>
). Peritrematal shield well developed, bearing one gland pores (
<emphasis id="6AF6645AFFDD0A5DFBF0F89EFBD6F8F0" box="[1051,1093,1807,1833]" italics="true" pageId="2" pageNumber="58">gd3</emphasis>
) and one poroid (
<emphasis id="6AF6645AFFDD0A5DFAFFF89EFAABF8F0" box="[1300,1336,1807,1833]" italics="true" pageId="2" pageNumber="58">id3</emphasis>
) at level near coxae II–III, poststigmatic extension of shield more or less wide, with two pairs of poroids (
<emphasis id="6AF6645AFFDD0A5DFB3CF8A2FB68F894" box="[1239,1275,1843,1869]" italics="true" pageId="2" pageNumber="58">id7</emphasis>
&amp; 
<emphasis id="6AF6645AFFDD0A5DFAF5F8A2FAA7F895" box="[1310,1332,1843,1868]" italics="true" pageId="2" pageNumber="58">ip</emphasis>
) and one pair of gland pores (
<emphasis id="6AF6645AFFDD0A5DFE93F8C6FE07F8A9" box="[376,404,1879,1904]" italics="true" pageId="2" pageNumber="58">gp</emphasis>
); anterior region of peritrematal shield fused to anterior margin of dorsal shield behind setae 
<emphasis id="6AF6645AFFDD0A5DFF7CF8EAFF23F84C" box="[151,176,1915,1941]" italics="true" pageId="2" pageNumber="58">z1</emphasis>
. Peritremes long, extending to mid-level of coxae I.
</paragraph>
<paragraph id="583DB848FFDD0A59FF2CF80EFEE0FE2C" blockId="2.[151,1437,727,2013]" lastBlockId="6.[151,1437,151,1761]" lastPageId="6" lastPageNumber="62" pageId="2" pageNumber="58">
<taxonomicName id="9F82C3CBFFDD0A5DFF2CF80EFEC5F860" authorityName="Krefft" authorityYear="1871" box="[199,342,1951,1977]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="2" pageNumber="58" phylum="Nematoda" rank="genus">
<emphasis id="6AF6645AFFDD0A5DFF2CF80EFEC5F860" box="[199,342,1951,1977]" italics="true" pageId="2" pageNumber="58">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="C0B9A4CDFFDD0A5DFE8CF80EFE5DF860" box="[359,462,1950,1977]" captionStart="FIGURES 3–8" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[191,1395,196,1907]" captionTargetId="figure-27@3.[179,1408,181,1919]" captionTargetPageId="3" captionText="FIGURES 3–8. Gaeolaelaps acanthopedus sp. nov., female. 3, dorsal idiosoma; 4, ventral idiosoma; 5, subcapitulum; 6, supralabral process; 7, epistome; 8, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5043654" httpUri="https://zenodo.org/record/5043654/files/figure.png" pageId="2" pageNumber="58">Figs 5–8</figureCitation>
, 
<figureCitation id="C0B9A4CDFFDD0A5DFE30F80EFDF9F860" box="[475,618,1950,1977]" captionStart="FIGURES 9–15" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[180,1404,191,1913]" captionTargetId="figure-17@4.[175,1413,181,1920]" captionTargetPageId="4" captionText="FIGURES 9–15. DIC micrographs of Gaeolaelaps acanthopedus sp. nov., female. 9, idiosoma in dorsal view; 10, idiosoma in ventral view; 11, sternal, genital &amp; anal shields; 12, palp, with a focus on apotele; 13, epistome; 14, insemination structures; 15, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5043656" httpUri="https://zenodo.org/record/5043656/files/figure.png" pageId="2" pageNumber="58">12, 13 &amp; 15</figureCitation>
). Epistome subtriangular and irregularly denticulate (
<figureCitation id="C0B9A4CDFFDD0A5DFB20F80EFA87F860" box="[1227,1300,1951,1977]" captionStart="FIGURES 3–8" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[191,1395,196,1907]" captionTargetId="figure-27@3.[179,1408,181,1919]" captionTargetPageId="3" captionText="FIGURES 3–8. Gaeolaelaps acanthopedus sp. nov., female. 3, dorsal idiosoma; 4, ventral idiosoma; 5, subcapitulum; 6, supralabral process; 7, epistome; 8, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5043654" httpUri="https://zenodo.org/record/5043654/files/figure.png" pageId="2" pageNumber="58">Figs 7</figureCitation>
&amp; 
<figureCitation id="C0B9A4CDFFDD0A5DFAD0F80EFAC9F860" box="[1339,1370,1951,1977]" captionStart="FIGURES 9–15" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[180,1404,191,1913]" captionTargetId="figure-17@4.[175,1413,181,1920]" captionTargetPageId="4" captionText="FIGURES 9–15. DIC micrographs of Gaeolaelaps acanthopedus sp. nov., female. 9, idiosoma in dorsal view; 10, idiosoma in ventral view; 11, sternal, genital &amp; anal shields; 12, palp, with a focus on apotele; 13, epistome; 14, insemination structures; 15, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5043656" httpUri="https://zenodo.org/record/5043656/files/figure.png" pageId="2" pageNumber="58">13</figureCitation>
). Hypostomal groove with six transverse rows of denticles, each row with 7–15 small denticles, with smooth anterior and posterior transverse lines, groove very slightly wider anteriorly. Hypostome with four pairs of smooth setae, 
<emphasis id="6AF6645AFFD90A59FA6BFF06FA0FFF68" box="[1408,1436,151,177]" italics="true" pageId="6" pageNumber="62">h3</emphasis>
(55–53)&gt; 
<emphasis id="6AF6645AFFD90A59FEE7FF2AFEBBFF0C" box="[268,296,187,213]" italics="true" pageId="6" pageNumber="62">h1</emphasis>
(50–53)&gt; 
<emphasis id="6AF6645AFFD90A59FE4FFF2AFE2DFF0D" box="[420,446,187,212]" italics="true" pageId="6" pageNumber="62">pc</emphasis>
(37–39)&gt; 
<emphasis id="6AF6645AFFD90A59FDD1FF2AFDC5FF0C" box="[570,598,187,213]" italics="true" pageId="6" pageNumber="62">h2</emphasis>
(32–34) (
<figureCitation id="C0B9A4CDFFD90A59FD28FF2AFC94FF0C" box="[707,775,186,213]" captionStart="FIGURES 3–8" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[191,1395,196,1907]" captionTargetId="figure-27@3.[179,1408,181,1919]" captionTargetPageId="3" captionText="FIGURES 3–8. Gaeolaelaps acanthopedus sp. nov., female. 3, dorsal idiosoma; 4, ventral idiosoma; 5, subcapitulum; 6, supralabral process; 7, epistome; 8, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5043654" httpUri="https://zenodo.org/record/5043654/files/figure.png" pageId="6" pageNumber="62">Fig. 5</figureCitation>
). Corniculi robust and horn-like, extending slightly beyond palp trochanter. Supralabral process with a very long apicoventral projection (
<figureCitation id="C0B9A4CDFFD90A59FBEFFF4EFBDAFF23" box="[1028,1097,223,250]" captionStart="FIGURES 3–8" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[191,1395,196,1907]" captionTargetId="figure-27@3.[179,1408,181,1919]" captionTargetPageId="3" captionText="FIGURES 3–8. Gaeolaelaps acanthopedus sp. nov., female. 3, dorsal idiosoma; 4, ventral idiosoma; 5, subcapitulum; 6, supralabral process; 7, epistome; 8, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5043654" httpUri="https://zenodo.org/record/5043654/files/figure.png" pageId="6" pageNumber="62">Fig. 6</figureCitation>
). Internal malae with one pair of fimbriate median projections, flanked by lobes with fimbriate anterior margin; labrum with pilose surface (
<figureCitation id="C0B9A4CDFFD90A59FA84FE92FF34FE98" captionStart="FIGURES 3–8" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[191,1395,196,1907]" captionTargetId="figure-27@3.[179,1408,181,1919]" captionTargetPageId="3" captionText="FIGURES 3–8. Gaeolaelaps acanthopedus sp. nov., female. 3, dorsal idiosoma; 4, ventral idiosoma; 5, subcapitulum; 6, supralabral process; 7, epistome; 8, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5043654" httpUri="https://zenodo.org/record/5043654/files/figure.png" pageId="6" pageNumber="62">Fig. 5</figureCitation>
). Chaetotaxy of palps (
<figureCitation id="C0B9A4CDFFD90A59FE4CFEB6FE68FE98" box="[423,507,295,321]" captionStart="FIGURES 9–15" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[180,1404,191,1913]" captionTargetId="figure-17@4.[175,1413,181,1920]" captionTargetPageId="4" captionText="FIGURES 9–15. DIC micrographs of Gaeolaelaps acanthopedus sp. nov., female. 9, idiosoma in dorsal view; 10, idiosoma in ventral view; 11, sternal, genital &amp; anal shields; 12, palp, with a focus on apotele; 13, epistome; 14, insemination structures; 15, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5043656" httpUri="https://zenodo.org/record/5043656/files/figure.png" pageId="6" pageNumber="62">Fig. 14</figureCitation>
): trochanter 2, femur 5, genu 6, tibia 14, tarsus 15, all setae smooth; palp femur with seta 
<emphasis id="6AF6645AFFD90A59FF27FEDAFF74FEBC" box="[204,231,331,357]" italics="true" pageId="6" pageNumber="62">d3</emphasis>
thickened and 
<emphasis id="6AF6645AFFD90A59FE7CFEDAFE3EFEBC" box="[407,429,331,357]" italics="true" pageId="6" pageNumber="62">al</emphasis>
thickened; palp genu with 
<emphasis id="6AF6645AFFD90A59FD0CFEDAFC98FEBC" box="[743,779,331,357]" italics="true" pageId="6" pageNumber="62">al1</emphasis>
stout, 
<emphasis id="6AF6645AFFD90A59FCB2FEDAFCEEFEBC" box="[857,893,331,357]" italics="true" pageId="6" pageNumber="62">al2</emphasis>
thickened and apically club-shaped; palp tarsal claw two-tined (
<figureCitation id="C0B9A4CDFFD90A59FEBAFEFEFE35FE50" box="[337,422,367,393]" captionStart="FIGURES 9–15" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[180,1404,191,1913]" captionTargetId="figure-17@4.[175,1413,181,1920]" captionTargetPageId="4" captionText="FIGURES 9–15. DIC micrographs of Gaeolaelaps acanthopedus sp. nov., female. 9, idiosoma in dorsal view; 10, idiosoma in ventral view; 11, sternal, genital &amp; anal shields; 12, palp, with a focus on apotele; 13, epistome; 14, insemination structures; 15, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5043656" httpUri="https://zenodo.org/record/5043656/files/figure.png" pageId="6" pageNumber="62">Fig. 12</figureCitation>
). Fixed digit of chelicera with an offset distal tooth (gabelzahn), followed by seven variously sized teeth, a setaceous pilus dentilis, dorsal cheliceral setae prostrate, arthrodial membrane with a rounded flap and normal filaments; cheliceral dorsal and lateral lyrifissures distinct; movable digit with two relatively large teeth (
<figureCitation id="C0B9A4CDFFD90A59FF37FE4AFEB0FE2C" box="[220,291,475,501]" captionStart="FIGURES 3–8" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[191,1395,196,1907]" captionTargetId="figure-27@3.[179,1408,181,1919]" captionTargetPageId="3" captionText="FIGURES 3–8. Gaeolaelaps acanthopedus sp. nov., female. 3, dorsal idiosoma; 4, ventral idiosoma; 5, subcapitulum; 6, supralabral process; 7, epistome; 8, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5043654" httpUri="https://zenodo.org/record/5043654/files/figure.png" pageId="6" pageNumber="62">Figs 8</figureCitation>
&amp; 
<figureCitation id="C0B9A4CDFFD90A59FEACFE4BFEF5FE2C" box="[327,358,474,501]" captionStart="FIGURES 9–15" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[180,1404,191,1913]" captionTargetId="figure-17@4.[175,1413,181,1920]" captionTargetPageId="4" captionText="FIGURES 9–15. DIC micrographs of Gaeolaelaps acanthopedus sp. nov., female. 9, idiosoma in dorsal view; 10, idiosoma in ventral view; 11, sternal, genital &amp; anal shields; 12, palp, with a focus on apotele; 13, epistome; 14, insemination structures; 15, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5043656" httpUri="https://zenodo.org/record/5043656/files/figure.png" pageId="6" pageNumber="62">15</figureCitation>
).
</paragraph>
<caption id="0CFDE8C0FFDC0A5CFF7CF806FDBCF80D" ID-DOI="http://doi.org/10.5281/zenodo.5043654" ID-Zenodo-Dep="5043654" httpUri="https://zenodo.org/record/5043654/files/figure.png" pageId="3" pageNumber="59" startId="3.[151,264,1943,1967]" targetBox="[191,1395,196,1907]" targetPageId="3">
<paragraph id="583DB848FFDC0A5CFF7CF806FDBCF80D" blockId="3.[151,1436,1943,2004]" pageId="3" pageNumber="59">
<emphasis id="6AF6645AFFDC0A5CFF7CF806FEA9F876" bold="true" box="[151,314,1943,1967]" pageId="3" pageNumber="59">FIGURES 3–8.</emphasis>
<taxonomicName id="9F82C3CBFFDC0A5CFED4F809FDC2F869" authority="Joharchi &amp; Friedrich, 2021" authorityName="Joharchi &amp; Friedrich" authorityYear="2021" box="[319,593,1944,1968]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="59" phylum="Arthropoda" rank="species" species="acanthopedus" status="sp. nov.">
<emphasis id="6AF6645AFFDC0A5CFED4F809FDC2F869" box="[319,593,1944,1968]" italics="true" pageId="3" pageNumber="59">Gaeolaelaps acanthopedus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C5D921FFDC0A5CFDBCF809FD3BF876" box="[599,680,1944,1968]" pageId="3" pageNumber="59" rank="species">
<emphasis id="6AF6645AFFDC0A5CFDBCF809FD32F869" bold="true" box="[599,673,1944,1968]" pageId="3" pageNumber="59">sp. nov</emphasis>
.
</taxonomicNameLabel>
, female. 3, dorsal idiosoma; 4, ventral idiosoma; 5, subcapitulum; 6, supralabral process; 7, epistome; 8, chelicera.
</paragraph>
</caption>
<caption id="0CFDE8C0FFDB0A5BFF7CF806FEB3F821" ID-DOI="http://doi.org/10.5281/zenodo.5043656" ID-Zenodo-Dep="5043656" httpUri="https://zenodo.org/record/5043656/files/figure.png" pageId="4" pageNumber="60" startId="4.[151,264,1943,1967]" targetBox="[180,1404,191,1913]" targetPageId="4">
<paragraph id="583DB848FFDB0A5BFF7CF806FEB3F821" blockId="4.[151,1436,1943,2040]" pageId="4" pageNumber="60">
<emphasis id="6AF6645AFFDB0A5BFF7CF806FEDAF876" bold="true" box="[151,329,1943,1967]" pageId="4" pageNumber="60">FIGURES 9–15.</emphasis>
DIC micrographs of 
<taxonomicName id="9F82C3CBFFDB0A5BFDCCF809FCA8F869" authority="Joharchi &amp; Friedrich, 2021" authorityName="Joharchi &amp; Friedrich" authorityYear="2021" box="[551,827,1944,1968]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="60" phylum="Arthropoda" rank="species" species="acanthopedus" status="sp. nov.">
<emphasis id="6AF6645AFFDB0A5BFDCCF809FCA8F869" box="[551,827,1944,1968]" italics="true" pageId="4" pageNumber="60">Gaeolaelaps acanthopedus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C5D921FFDB0A5BFCA8F809FC05F876" box="[835,918,1944,1968]" pageId="4" pageNumber="60" rank="species">
<emphasis id="6AF6645AFFDB0A5BFCA8F809FC03F869" bold="true" box="[835,912,1944,1968]" pageId="4" pageNumber="60">sp. nov</emphasis>
.
</taxonomicNameLabel>
, female. 9, idiosoma in dorsal view; 10, idiosoma in ventral view; 11, sternal, genital &amp; anal shields; 12, palp, with a focus on apotele; 13, epistome; 14, insemination structures; 15, chelicera.
</paragraph>
</caption>
<caption id="0CFDE8C0FFDA0A5AFF7CF838FAEFF818" ID-DOI="http://doi.org/10.5281/zenodo.5043658" ID-Zenodo-Dep="5043658" box="[151,1404,1961,1985]" httpUri="https://zenodo.org/record/5043658/files/figure.png" pageId="5" pageNumber="61" startId="5.[151,264,1961,1985]" targetBox="[161,1422,214,1924]" targetPageId="5">
<paragraph id="583DB848FFDA0A5AFF7CF838FAEFF818" blockId="5.[151,1404,1961,1985]" box="[151,1404,1961,1985]" pageId="5" pageNumber="61">
<emphasis id="6AF6645AFFDA0A5AFF7CF838FEC7F818" bold="true" box="[151,340,1961,1985]" pageId="5" pageNumber="61">FIGURES 16–19.</emphasis>
<taxonomicName id="9F82C3CBFFDA0A5AFEB1F838FDFDF818" authority="Joharchi &amp; Friedrich, 2021" authorityName="Joharchi &amp; Friedrich" authorityYear="2021" box="[346,622,1961,1985]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="61" phylum="Arthropoda" rank="species" species="acanthopedus" status="sp. nov.">
<emphasis id="6AF6645AFFDA0A5AFEB1F838FDFDF818" box="[346,622,1961,1985]" italics="true" pageId="5" pageNumber="61">Gaeolaelaps acanthopedus</emphasis>
</taxonomicName>
<emphasis id="6AF6645AFFDA0A5AFD9FF838FD57F818" bold="true" box="[628,708,1961,1985]" pageId="5" pageNumber="61">
<taxonomicNameLabel id="71C5D921FFDA0A5AFD9FF838FD57F818" box="[628,708,1961,1985]" pageId="5" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female. 16, leg I (trochanter-tibia); 17, leg II; 18, leg III; 19, leg IV.
</paragraph>
</caption>
<paragraph id="583DB848FFD90A59FF2CFE6EFDA6FC7C" blockId="6.[151,1437,151,1761]" pageId="6" pageNumber="62">
<emphasis id="6AF6645AFFD90A59FF2CFE6EFF6FFDC1" box="[199,252,511,536]" italics="true" pageId="6" pageNumber="62">Legs</emphasis>
(
<figureCitation id="C0B9A4CDFFD90A59FEE6FE6EFE01FDC3" box="[269,402,511,538]" captionStart="FIGURES 16–19" captionStartId="5.[151,264,1961,1985]" captionTargetBox="[161,1422,214,1924]" captionTargetId="figure-27@5.[154,1433,199,1937]" captionTargetPageId="5" captionText="FIGURES 16–19. Gaeolaelaps acanthopedus sp. nov., female. 16, leg I (trochanter-tibia); 17, leg II; 18, leg III; 19, leg IV." figureDoi="http://doi.org/10.5281/zenodo.5043658" httpUri="https://zenodo.org/record/5043658/files/figure.png" pageId="6" pageNumber="62">Figs 16–19</figureCitation>
). Legs II (517–528) and III (494–503) short, I (640–664) and IV (741–768) longer. Chaetotaxy normal for free-living 
<taxonomicName id="9F82C3CBFFD90A59FE03FDB2FDF4FDE4" box="[488,615,547,573]" class="Arachnida" family="Laelapidae" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="62" phylum="Arthropoda" rank="family">Laelapidae</taxonomicName>
: Leg I (
<figureCitation id="C0B9A4CDFFD90A59FD28FDB2FC8AFDE7" box="[707,793,547,574]" captionStart="FIGURES 16–19" captionStartId="5.[151,264,1961,1985]" captionTargetBox="[161,1422,214,1924]" captionTargetId="figure-27@5.[154,1433,199,1937]" captionTargetPageId="5" captionText="FIGURES 16–19. Gaeolaelaps acanthopedus sp. nov., female. 16, leg I (trochanter-tibia); 17, leg II; 18, leg III; 19, leg IV." figureDoi="http://doi.org/10.5281/zenodo.5043658" httpUri="https://zenodo.org/record/5043658/files/figure.png" pageId="6" pageNumber="62">Fig. 16</figureCitation>
): coxa 0 0/1, 0/1 0, trochanter 1 0/1, 1/2 1 (
<emphasis id="6AF6645AFFD90A59FAFCFDB2FAACFDE4" box="[1303,1343,547,573]" italics="true" pageId="6" pageNumber="62">pv1</emphasis>
slightly thickened), femur 2 2/1, 3/3 2 (
<emphasis id="6AF6645AFFD90A59FE1EFDD6FD8CFDB8" box="[501,543,583,609]" italics="true" pageId="6" pageNumber="62">ad1</emphasis>
&amp; 
<emphasis id="6AF6645AFFD90A59FDA8FDD6FD0EFDB8" box="[579,669,583,609]" italics="true" pageId="6" pageNumber="62">pd1 pd3</emphasis>
thickened and inserted on small tubercles), genu 2 3/2, 3/1 2, tibia 2 3/2, 3/1 2. Leg II (
<figureCitation id="C0B9A4CDFFD90A59FE8CFDFAFE28FD5C" box="[359,443,619,645]" captionStart="FIGURES 16–19" captionStartId="5.[151,264,1961,1985]" captionTargetBox="[161,1422,214,1924]" captionTargetId="figure-27@5.[154,1433,199,1937]" captionTargetPageId="5" captionText="FIGURES 16–19. Gaeolaelaps acanthopedus sp. nov., female. 16, leg I (trochanter-tibia); 17, leg II; 18, leg III; 19, leg IV." figureDoi="http://doi.org/10.5281/zenodo.5043658" httpUri="https://zenodo.org/record/5043658/files/figure.png" pageId="6" pageNumber="62">Fig. 17</figureCitation>
): coxa 0 0/1, 0/1 0, trochanter 1 0/2, 0/1 1 (
<emphasis id="6AF6645AFFD90A59FC43FDFAFC2DFD5C" box="[936,958,619,645]" italics="true" pageId="6" pageNumber="62">al</emphasis>
slightly thickened), femur 2 3/1, 2/2 1 (
<emphasis id="6AF6645AFFD90A59FA69FDFAFF4DFD70" italics="true" pageId="6" pageNumber="62">av &amp; pd2</emphasis>
thickened and inserted on small tubercles), genu 2 3/1, 2/1 2 (
<emphasis id="6AF6645AFFD90A59FC70FD1EFC26FD71" box="[923,949,655,680]" italics="true" pageId="6" pageNumber="62">av</emphasis>
thickened and inserted on small tubercles), tibia 2 2/1, 2/1 2 (all ventral setae spur-like). Leg III (
<figureCitation id="C0B9A4CDFFD90A59FD06FD22FCD3FD14" box="[749,832,691,717]" captionStart="FIGURES 16–19" captionStartId="5.[151,264,1961,1985]" captionTargetBox="[161,1422,214,1924]" captionTargetId="figure-27@5.[154,1433,199,1937]" captionTargetPageId="5" captionText="FIGURES 16–19. Gaeolaelaps acanthopedus sp. nov., female. 16, leg I (trochanter-tibia); 17, leg II; 18, leg III; 19, leg IV." figureDoi="http://doi.org/10.5281/zenodo.5043658" httpUri="https://zenodo.org/record/5043658/files/figure.png" pageId="6" pageNumber="62">Fig. 18</figureCitation>
): coxa 0 0/1, 0/1 0, trochanter 1 1/1, 0/1 1 (
<emphasis id="6AF6645AFFD90A59FACEFD22FAD2FD14" box="[1317,1345,691,717]" italics="true" pageId="6" pageNumber="62">ad</emphasis>
slightly thickened), femur 1 2/1, 1/0 1, genu 2 2/1, 2/1 1 (all ventral setae spur-like, 
<emphasis id="6AF6645AFFD90A59FC08FD46FB9EFD28" box="[995,1037,727,753]" italics="true" pageId="6" pageNumber="62">ad1</emphasis>
&amp; 
<emphasis id="6AF6645AFFD90A59FBDAFD46FBC8FD28" box="[1073,1115,727,753]" italics="true" pageId="6" pageNumber="62">pd1</emphasis>
inserted on small tubercles), tibia: 2 1/1, 2/1 1 (all ventral setae spur-like, 
<emphasis id="6AF6645AFFD90A59FD7DFD6AFD2CFCCC" box="[662,703,763,789]" italics="true" pageId="6" pageNumber="62">pd2</emphasis>
inserted on small tubercles). Leg IV (
<figureCitation id="C0B9A4CDFFD90A59FB86FD6AFB52FCCC" box="[1133,1217,763,789]" captionStart="FIGURES 16–19" captionStartId="5.[151,264,1961,1985]" captionTargetBox="[161,1422,214,1924]" captionTargetId="figure-27@5.[154,1433,199,1937]" captionTargetPageId="5" captionText="FIGURES 16–19. Gaeolaelaps acanthopedus sp. nov., female. 16, leg I (trochanter-tibia); 17, leg II; 18, leg III; 19, leg IV." figureDoi="http://doi.org/10.5281/zenodo.5043658" httpUri="https://zenodo.org/record/5043658/files/figure.png" pageId="6" pageNumber="62">Fig. 19</figureCitation>
): coxa 0 0/1, 0/0 0, trochanter 1 1/1, 0/1 1 (
<emphasis id="6AF6645AFFD90A59FE48FC8EFE2CFCE0" box="[419,447,799,825]" italics="true" pageId="6" pageNumber="62">ad</emphasis>
&amp; 
<emphasis id="6AF6645AFFD90A59FE0FFC8EFE69FCE0" box="[484,506,799,825]" italics="true" pageId="6" pageNumber="62">pl</emphasis>
spur-like), femur 1 2/1, 1/0 1 (
<emphasis id="6AF6645AFFD90A59FCB1FC8EFC17FCE0" box="[858,900,799,825]" italics="true" pageId="6" pageNumber="62">ad1</emphasis>
, 
<emphasis id="6AF6645AFFD90A59FC78FC8EFC2EFCE0" box="[915,957,799,825]" italics="true" pageId="6" pageNumber="62">ad2</emphasis>
and 
<emphasis id="6AF6645AFFD90A59FC1EFC8EFB82FCE0" box="[1013,1041,799,825]" italics="true" pageId="6" pageNumber="62">pd</emphasis>
thickened &amp; 
<emphasis id="6AF6645AFFD90A59FB40FC8EFB46FCE0" box="[1195,1237,799,825]" italics="true" pageId="6" pageNumber="62">ad1</emphasis>
, 
<emphasis id="6AF6645AFFD90A59FB08FC8EFA9DFCE0" box="[1251,1294,799,825]" italics="true" pageId="6" pageNumber="62">ad2</emphasis>
on small tubercles), genu 2 2/1, 3/0 1 (
<emphasis id="6AF6645AFFD90A59FE21FCD2FE77FC85" box="[458,484,835,860]" italics="true" pageId="6" pageNumber="62">av</emphasis>
spur-like), tibia 2 1/1, 3/1 2 (av, 
<emphasis id="6AF6645AFFD90A59FCBFFCD2FCFDFC85" box="[852,878,835,860]" italics="true" pageId="6" pageNumber="62">pv</emphasis>
&amp; 
<emphasis id="6AF6645AFFD90A59FC79FCD2FC25FC84" box="[914,950,835,861]" italics="true" pageId="6" pageNumber="62">pl1</emphasis>
spur-like). Tarsi II–IV with 18 setae (3 3/2, 3/2 3 + 
<emphasis id="6AF6645AFFD90A59FF1AFCF6FE81FC59" box="[241,274,871,896]" italics="true" pageId="6" pageNumber="62">mv</emphasis>
, 
<emphasis id="6AF6645AFFD90A59FECAFCF6FED0FC58" box="[289,323,871,897]" italics="true" pageId="6" pageNumber="62">md</emphasis>
); most setae spine-like, see 
<figureCitation id="C0B9A4CDFFD90A59FD6FFCF6FCB0FC58" box="[644,803,871,897]" captionStart="FIGURES 16–19" captionStartId="5.[151,264,1961,1985]" captionTargetBox="[161,1422,214,1924]" captionTargetId="figure-27@5.[154,1433,199,1937]" captionTargetPageId="5" captionText="FIGURES 16–19. Gaeolaelaps acanthopedus sp. nov., female. 16, leg I (trochanter-tibia); 17, leg II; 18, leg III; 19, leg IV." figureDoi="http://doi.org/10.5281/zenodo.5043658" httpUri="https://zenodo.org/record/5043658/files/figure.png" pageId="6" pageNumber="62">Figures 17 19</figureCitation>
. All pretarsi with well-developed paired claws, rounded pulvilli and normal ambulacral stalk.
</paragraph>
<paragraph id="583DB848FFD90A59FF2CFC3EFD3EFBC8" blockId="6.[151,1437,151,1761]" pageId="6" pageNumber="62">
<emphasis id="6AF6645AFFD90A59FF2CFC3EFE48FC10" bold="true" box="[199,475,943,969]" italics="true" pageId="6" pageNumber="62">Insemination structures</emphasis>
(
<figureCitation id="C0B9A4CDFFD90A59FE00FC3EFDADFC10" box="[491,574,943,969]" captionStart="FIGURES 9–15" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[180,1404,191,1913]" captionTargetId="figure-17@4.[175,1413,181,1920]" captionTargetPageId="4" captionText="FIGURES 9–15. DIC micrographs of Gaeolaelaps acanthopedus sp. nov., female. 9, idiosoma in dorsal view; 10, idiosoma in ventral view; 11, sternal, genital &amp; anal shields; 12, palp, with a focus on apotele; 13, epistome; 14, insemination structures; 15, chelicera." figureDoi="http://doi.org/10.5281/zenodo.5043656" httpUri="https://zenodo.org/record/5043656/files/figure.png" pageId="6" pageNumber="62">Fig. 14</figureCitation>
). Laelapid-type sperm access system, tubulus long, wider at the solenostome level of coxa III and entering sacculus via a pair of circular openings. Sacculus irregular, the proximal ends of the tubulus slightly swollen at junction with ramus.
</paragraph>
</subSubSection>
<subSubSection id="1098EBC3FFD90A59FF2CFB8AFEF3FB80" pageId="6" pageNumber="62" type="etymology">
<paragraph id="583DB848FFD90A59FF2CFB8AFEF3FB80" blockId="6.[151,1437,151,1761]" pageId="6" pageNumber="62">
<emphasis id="6AF6645AFFD90A59FF2CFB8AFEDDFBEC" bold="true" box="[199,334,1051,1077]" pageId="6" pageNumber="62">Etymology.</emphasis>
The name 
<taxonomicName id="9F82C3CBFFD90A59FE20FB8AFDF4FBEC" authorityName="Joharchi &amp; Friedrich" authorityYear="2021" box="[459,615,1051,1077]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="62" phylum="Arthropoda" rank="species" species="acanthopedus">
<emphasis id="6AF6645AFFD90A59FE20FB8AFDF4FBEC" box="[459,615,1051,1077]" italics="true" pageId="6" pageNumber="62">acanthopedus</emphasis>
</taxonomicName>
(Latin 
<emphasis id="6AF6645AFFD90A59FD53FB8AFC86FBEC" box="[696,789,1051,1077]" italics="true" pageId="6" pageNumber="62">Acantho</emphasis>
, spur + 
<emphasis id="6AF6645AFFD90A59FC9AFB8AFC0AFBEC" box="[881,921,1051,1077]" italics="true" pageId="6" pageNumber="62">ped</emphasis>
, foot) refers to the spur/spine-like appearance of some leg setae.
</paragraph>
</subSubSection>
<subSubSection id="1098EBC3FFD90A59FF2CFBF2FDB4F938" pageId="6" pageNumber="62" type="discussion">
<paragraph id="583DB848FFD90A59FF2CFBF2FDB4F938" blockId="6.[151,1437,151,1761]" pageId="6" pageNumber="62">
<emphasis id="6AF6645AFFD90A59FF2CFBF2FEA8FBA4" bold="true" box="[199,315,1123,1149]" pageId="6" pageNumber="62">Remarks.</emphasis>
The new species resembles 
<taxonomicName id="9F82C3CBFFD90A59FD90FBF2FCEFFBA4" authority="(Karg, 2000)" baseAuthorityName="Karg" baseAuthorityYear="2000" box="[635,892,1123,1149]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="62" phylum="Arthropoda" rank="species" species="loksai">
<emphasis id="6AF6645AFFD90A59FD90FBF2FD1CFBA4" box="[635,655,1123,1149]" italics="true" pageId="6" pageNumber="62">G</emphasis>
. 
<emphasis id="6AF6645AFFD90A59FD75FBF2FD72FBA4" box="[670,737,1123,1149]" italics="true" pageId="6" pageNumber="62">loksai</emphasis>
(
<bibRefCitation id="3C13C5B9FFD90A59FD1BFBF2FCE7FBA4" author="Karg, W." box="[752,884,1123,1149]" pageId="6" pageNumber="62" pagination="243 - 262" refId="ref8243" refString="Karg, W. (2000) Zur Systematik der Raubmilbenfamilien Hypoaspididae v. Vitzthum, 1941 und Rhodacaridae Oudemans, 1902 (Acarina, Parasitiformes) mit neuen Arten aus Sud- und Mittelamerika. Mitteilungen aus dem Museum fur Naturkunde in Berlin, Zoologische Reihe, 76, 243 - 262. https: // doi. org / 10.1002 / mmnz. 4850760207" type="journal article" year="2000">Karg, 2000</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="9F82C3CBFFD90A59FC6CFBF2FB19FBA4" authority="(Karg, 1982)" baseAuthorityName="Karg" baseAuthorityYear="1982" box="[903,1162,1123,1149]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="62" phylum="Arthropoda" rank="species" species="cerata">
<emphasis id="6AF6645AFFD90A59FC6CFBF2FC08FBA4" box="[903,923,1123,1149]" italics="true" pageId="6" pageNumber="62">G</emphasis>
. 
<emphasis id="6AF6645AFFD90A59FC41FBF2FC61FBA5" box="[938,1010,1123,1148]" italics="true" pageId="6" pageNumber="62">cerata</emphasis>
(
<bibRefCitation id="3C13C5B9FFD90A59FBEAFBF2FB12FBA4" author="Karg, W." box="[1025,1153,1123,1149]" pageId="6" pageNumber="62" pagination="233 - 256" refId="ref8196" refString="Karg, W. (1982) Zur Kenntnis der Raubmilbeng Hypoaspis Canestrini, 1884 (Acarina, Parasitiformes). Mitteilungen aus dem Zoologischen Museum in Berlin, 58, 233 - 256. https: // doi. org / 10.1002 / mmnz. 4830580204" type="journal article" year="1982">Karg, 1982</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="9F82C3CBFFD90A59FB2AFBF2FF5CFB78" authority="Hyatt, 1964" authorityName="Hyatt" authorityYear="1964" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="62" phylum="Arthropoda" rank="species" species="circularis">
<emphasis id="6AF6645AFFD90A59FB2AFBF2FB46FBA4" box="[1217,1237,1123,1149]" italics="true" pageId="6" pageNumber="62">G</emphasis>
. 
<emphasis id="6AF6645AFFD90A59FB0FFBF2FAC3FBA4" box="[1252,1360,1123,1149]" italics="true" pageId="6" pageNumber="62">circularis</emphasis>
<bibRefCitation id="3C13C5B9FFD90A59FABCFBF2FF5CFB78" author="Hyatt, K. H." pageId="6" pageNumber="62" pagination="465 - 509" refId="ref7926" refString="Hyatt, K. H. (1964) A collection of Mesostigmata (Acari) associated with Coleoptera and Hemiptera in Venezuela. Bulletin of the British Museum (Natural History), Zoology, 11 (7), 465 - 509. https: // doi. org / 10.5962 / bhl. part. 4723" type="journal article" year="1964">Hyatt, 1964</bibRefCitation>
</taxonomicName>
described from 
<collectingCountry id="2095F8D8FFD90A59FE7BFB16FE7EFB78" box="[400,493,1159,1185]" name="Ecuador" pageId="6" pageNumber="62">Ecuador</collectingCountry>
and 
<collectingCountry id="2095F8D8FFD90A59FDC1FB16FD0EFB78" box="[554,669,1159,1185]" name="Venezuela" pageId="6" pageNumber="62">Venezuela</collectingCountry>
(South America), respectively, in having similarly shaped and reticulated genital shield, the similar length of dorsal shield setae and body size. In both 
<taxonomicName id="9F82C3CBFFD90A59FB81FB3AFB43FB1C" baseAuthorityName="Karg" baseAuthorityYear="2000" box="[1130,1232,1195,1221]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="62" phylum="Arthropoda" rank="species" species="loksai">
<emphasis id="6AF6645AFFD90A59FB81FB3AFBEDFB1C" box="[1130,1150,1195,1221]" italics="true" pageId="6" pageNumber="62">G</emphasis>
. 
<emphasis id="6AF6645AFFD90A59FB66FB3AFB43FB1C" box="[1165,1232,1195,1221]" italics="true" pageId="6" pageNumber="62">loksai</emphasis>
</taxonomicName>
and 
<taxonomicName id="9F82C3CBFFD90A59FAE3FB3AFAE0FB1D" baseAuthorityName="Karg" baseAuthorityYear="1982" box="[1288,1395,1195,1221]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="62" phylum="Arthropoda" rank="species" species="cerata">
<emphasis id="6AF6645AFFD90A59FAE3FB3AFA8FFB1C" box="[1288,1308,1195,1221]" italics="true" pageId="6" pageNumber="62">G</emphasis>
. 
<emphasis id="6AF6645AFFD90A59FAC0FB3AFAE0FB1D" box="[1323,1395,1195,1220]" italics="true" pageId="6" pageNumber="62">cerata</emphasis>
</taxonomicName>
the peritremes short, reaching to the mid-level of coxae II, while in 
<taxonomicName id="9F82C3CBFFD90A59FCB6FB5EFB88FB30" authorityName="Joharchi &amp; Friedrich" authorityYear="2021" box="[861,1051,1231,1257]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="62" phylum="Arthropoda" rank="species" species="acanthopedus">
<emphasis id="6AF6645AFFD90A59FCB6FB5EFCE2FB30" box="[861,881,1231,1257]" italics="true" pageId="6" pageNumber="62">G</emphasis>
. 
<emphasis id="6AF6645AFFD90A59FC94FB5EFB88FB30" box="[895,1051,1231,1257]" italics="true" pageId="6" pageNumber="62">acanthopedus</emphasis>
</taxonomicName>
the peritremes are long, extending to the mid-level of coxae I. 
<taxonomicName id="9F82C3CBFFD90A59FE22FB62FD6AFAD4" authorityName="Joharchi &amp; Friedrich" authorityYear="2021" box="[457,761,1267,1293]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="62" phylum="Arthropoda" rank="species" species="acanthopedus">
<emphasis id="6AF6645AFFD90A59FE22FB62FD6AFAD4" box="[457,761,1267,1293]" italics="true" pageId="6" pageNumber="62">Gaeolaelaps acanthopedus</emphasis>
</taxonomicName>
most resembles 
<taxonomicName id="9F82C3CBFFD90A59FC59FB62FBD3FAD4" authorityName="Hyatt" authorityYear="1964" box="[946,1088,1267,1293]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="62" phylum="Arthropoda" rank="species" species="circularis">
<emphasis id="6AF6645AFFD90A59FC59FB62FC55FAD4" box="[946,966,1267,1293]" italics="true" pageId="6" pageNumber="62">G</emphasis>
. 
<emphasis id="6AF6645AFFD90A59FC3FFB62FBD3FAD4" box="[980,1088,1267,1293]" italics="true" pageId="6" pageNumber="62">circularis</emphasis>
</taxonomicName>
(redescribed by 
<bibRefCitation id="3C13C5B9FFD90A59FB12FB62FF6EFAE8" author="Marticorena, J. L. M. &amp; Moreira, G. F. &amp; de Moraes, G. J." pageId="6" pageNumber="62" pagination="333 - 348" refId="ref8618" refString="Marticorena, J. L. M., Moreira, G. F. &amp; de Moraes, G. J. (2020) Mites of the genus Gaeolaelaps (Acari: Laelapidae) from southern Brazil, with description of two new species. Zootaxa, 4772 (2), 333 - 348. https: // doi. org / 10.11646 / zootaxa. 4772.2.5" type="journal article" year="2020">
Marticorena 
<emphasis id="6AF6645AFFD90A59FA63FB62FF3EFAE8" italics="true" pageId="6" pageNumber="62">et al</emphasis>
., 2020
</bibRefCitation>
), due to its similarly shaped and reticulated genital shield and spine like setae on different segments of legs (especially those on tarsi II–IV). 
<taxonomicName id="9F82C3CBFFD90A59FDE2FAAAFCA9FA8C" authorityName="Joharchi &amp; Friedrich" authorityYear="2021" box="[521,826,1339,1365]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="62" phylum="Arthropoda" rank="species" species="acanthopedus">
<emphasis id="6AF6645AFFD90A59FDE2FAAAFCA9FA8C" box="[521,826,1339,1365]" italics="true" pageId="6" pageNumber="62">Gaeolaelaps acanthopedus</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="9F82C3CBFFD90A59FC25FAAAFBCFFA8C" authorityName="Hyatt" authorityYear="1964" box="[974,1116,1339,1365]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="62" phylum="Arthropoda" rank="species" species="circularis">
<emphasis id="6AF6645AFFD90A59FC25FAAAFC71FA8C" box="[974,994,1339,1365]" italics="true" pageId="6" pageNumber="62">G</emphasis>
. 
<emphasis id="6AF6645AFFD90A59FC1BFAAAFBCFFA8C" box="[1008,1116,1339,1365]" italics="true" pageId="6" pageNumber="62">circularis</emphasis>
</taxonomicName>
by having 39 pairs of dorsal setae and much larger body size (674–702 long, 390–396 wide), while in 
<taxonomicName id="9F82C3CBFFD90A59FC2CFACEFBC6FAA0" authorityName="Hyatt" authorityYear="1964" box="[967,1109,1375,1401]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="62" phylum="Arthropoda" rank="species" species="circularis">
<emphasis id="6AF6645AFFD90A59FC2CFACEFC48FAA0" box="[967,987,1375,1401]" italics="true" pageId="6" pageNumber="62">G</emphasis>
. 
<emphasis id="6AF6645AFFD90A59FC02FACEFBC6FAA0" box="[1001,1109,1375,1401]" italics="true" pageId="6" pageNumber="62">circularis</emphasis>
</taxonomicName>
the dorsal shield has 33 pairs of setae and smaller body size (see 
<bibRefCitation id="3C13C5B9FFD90A59FDF7FA12FCBFFA44" author="Marticorena, J. L. M. &amp; Moreira, G. F. &amp; de Moraes, G. J." box="[540,812,1411,1437]" pageId="6" pageNumber="62" pagination="333 - 348" refId="ref8618" refString="Marticorena, J. L. M., Moreira, G. F. &amp; de Moraes, G. J. (2020) Mites of the genus Gaeolaelaps (Acari: Laelapidae) from southern Brazil, with description of two new species. Zootaxa, 4772 (2), 333 - 348. https: // doi. org / 10.11646 / zootaxa. 4772.2.5" type="journal article" year="2020">
Marticorena 
<emphasis id="6AF6645AFFD90A59FD40FA12FD48FA44" box="[683,731,1411,1437]" italics="true" pageId="6" pageNumber="62">et al</emphasis>
., 2020
</bibRefCitation>
). 
<taxonomicName id="9F82C3CBFFD90A59FCD6FA12FBFEFA44" authorityName="Joharchi &amp; Friedrich" authorityYear="2021" box="[829,1133,1411,1437]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="62" phylum="Arthropoda" rank="species" species="acanthopedus">
<emphasis id="6AF6645AFFD90A59FCD6FA12FBFEFA44" box="[829,1133,1411,1437]" italics="true" pageId="6" pageNumber="62">Gaeolaelaps acanthopedus</emphasis>
</taxonomicName>
can easily be distinguished from all other South American species of 
<taxonomicName id="9F82C3CBFFD90A59FD82FA36FD64FA18" box="[617,759,1447,1473]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="6AF6645AFFD90A59FD82FA36FD64FA18" box="[617,759,1447,1473]" italics="true" pageId="6" pageNumber="62">Gaeolaelaps</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="3C13C5B9FFD90A59FCD9FA36FC45FA18" author="Moreira, G. F." box="[818,982,1447,1473]" pageId="6" pageNumber="62" refId="ref8688" refString="Moreira, G. F. (2014) Taxonomic studies of Laelapidae mites (Acari: Mesostigmata: Laelapidae) and their use in combination with entomopathogenic Nematodes (Rhabditida: Steinernematidae, Heterorhabditida) to control Frankliniella occidentalis (Thysanoptera: Thripidae). Doctorate Thesis in Agronomia (Entomologia Agricola) at University Estadual Paulista, Campus de Jaboticaba, 521 pp." type="book" year="2014">Moreira, 2014</bibRefCitation>
; 
<bibRefCitation id="3C13C5B9FFD90A59FC0BFA36FB63FA18" author="Marticorena, J. L. M. &amp; Moreira, G. F. &amp; de Moraes, G. J." box="[992,1264,1447,1473]" pageId="6" pageNumber="62" pagination="333 - 348" refId="ref8618" refString="Marticorena, J. L. M., Moreira, G. F. &amp; de Moraes, G. J. (2020) Mites of the genus Gaeolaelaps (Acari: Laelapidae) from southern Brazil, with description of two new species. Zootaxa, 4772 (2), 333 - 348. https: // doi. org / 10.11646 / zootaxa. 4772.2.5" type="journal article" year="2020">
Marticorena 
<emphasis id="6AF6645AFFD90A59FB9BFA36FB32FA18" box="[1136,1185,1447,1473]" italics="true" pageId="6" pageNumber="62">et al</emphasis>
., 2020
</bibRefCitation>
) by the following combination of characters: (1) most leg setae on tarsi II–IV spine-like (strongly thickened and apically tapered), ventral setae on tibiae II–IV, genua II–IV and setae 
<emphasis id="6AF6645AFFD90A59FD25FA7EFD79F9D0" box="[718,746,1519,1545]" italics="true" pageId="6" pageNumber="62">ad</emphasis>
, 
<emphasis id="6AF6645AFFD90A59FD1CFA7EFC9EF9D0" box="[759,781,1519,1545]" italics="true" pageId="6" pageNumber="62">pl</emphasis>
on trochanter IV spur-like (strongly thickened and apically blunt); (2) genital shield flask-shaped, elongated (anterior margin of shield extending at level of poroids 
<emphasis id="6AF6645AFFD90A59FAA2F982FAF8F9F4" box="[1353,1387,1555,1581]" italics="true" pageId="6" pageNumber="62">iv2</emphasis>
; (3) 
<emphasis id="6AF6645AFFD90A59FF7CF9A6FF3EF988" box="[151,173,1591,1617]" italics="true" pageId="6" pageNumber="62">j1</emphasis>
longest dorsal setae and 
<emphasis id="6AF6645AFFD90A59FE22F9A6FE74F988" box="[457,487,1591,1617]" italics="true" pageId="6" pageNumber="62">Z5</emphasis>
shorter than 
<emphasis id="6AF6645AFFD90A59FD97F9A6FD02F988" box="[636,657,1591,1617]" italics="true" pageId="6" pageNumber="62">j1</emphasis>
(vs. 
<emphasis id="6AF6645AFFD90A59FD21F9A6FD7BF988" box="[714,744,1591,1617]" italics="true" pageId="6" pageNumber="62">Z5</emphasis>
longest dorsal setae in both 
<taxonomicName id="9F82C3CBFFD90A59FBC0F9A6FB05F989" baseAuthorityName="Karg" baseAuthorityYear="1982" box="[1067,1174,1591,1617]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="62" phylum="Arthropoda" rank="species" species="cerata">
<emphasis id="6AF6645AFFD90A59FBC0F9A6FBACF988" box="[1067,1087,1591,1617]" italics="true" pageId="6" pageNumber="62">G</emphasis>
. 
<emphasis id="6AF6645AFFD90A59FBA5F9A6FB05F989" box="[1102,1174,1591,1616]" italics="true" pageId="6" pageNumber="62">cerata</emphasis>
</taxonomicName>
and 
<taxonomicName id="9F82C3CBFFD90A59FB26F9A6FAA0F988" baseAuthorityName="Karg" baseAuthorityYear="2000" box="[1229,1331,1591,1617]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="62" phylum="Arthropoda" rank="species" species="loksai">
<emphasis id="6AF6645AFFD90A59FB26F9A6FB72F988" box="[1229,1249,1591,1617]" italics="true" pageId="6" pageNumber="62">G</emphasis>
. 
<emphasis id="6AF6645AFFD90A59FB1BF9A6FAA0F988" box="[1264,1331,1591,1617]" italics="true" pageId="6" pageNumber="62">loksai</emphasis>
</taxonomicName>
); and (4) peritremes long, extending anteriorly to mid-level of coxae I (vs. in both 
<taxonomicName id="9F82C3CBFFD90A59FC3BF9CAFBA8F9AD" baseAuthorityName="Karg" baseAuthorityYear="1982" box="[976,1083,1627,1653]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="62" phylum="Arthropoda" rank="species" species="cerata">
<emphasis id="6AF6645AFFD90A59FC3BF9CAFC77F9AC" box="[976,996,1627,1653]" italics="true" pageId="6" pageNumber="62">G</emphasis>
. 
<emphasis id="6AF6645AFFD90A59FC18F9CAFBA8F9AD" box="[1011,1083,1627,1652]" italics="true" pageId="6" pageNumber="62">cerata</emphasis>
</taxonomicName>
and 
<taxonomicName id="9F82C3CBFFD90A59FB99F9CAFB4BF9AC" baseAuthorityName="Karg" baseAuthorityYear="2000" box="[1138,1240,1627,1653]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="62" phylum="Arthropoda" rank="species" species="loksai">
<emphasis id="6AF6645AFFD90A59FB99F9CAFB15F9AC" box="[1138,1158,1627,1653]" italics="true" pageId="6" pageNumber="62">G</emphasis>
. 
<emphasis id="6AF6645AFFD90A59FB7EF9CAFB4BF9AC" box="[1173,1240,1627,1653]" italics="true" pageId="6" pageNumber="62">loksai</emphasis>
</taxonomicName>
peritremes short, reaches to middle of coxa II); (5) tarsus IV without elongated setae; (6) fixed digit of chelicera with eight teeth; (7) palp tarsal apotele two-tined; (8) sternal shield poroid (
<emphasis id="6AF6645AFFD90A59FCEAF932FCB0F964" box="[769,803,1699,1725]" italics="true" pageId="6" pageNumber="62">iv2</emphasis>
), slit-like (lyrifissures); and (9) supralabral process with a very long apicoventral projection.
</paragraph>
</subSubSection>
</treatment>
</document>