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<document id="42DDC5A9820980367C852481626CD9D8" ID-CLB-Dataset="45436" ID-DOI="10.5281/zenodo.211122" ID-GBIF-Dataset="344f8458-2c80-43b3-a96b-3aca02033f88" ID-ISSN="1175-5326" ID-Zenodo-Dep="211122" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460439637460" checkinUser="plazi" docAuthor="Kohlmann, Bert &amp; Solís, Ángel" docDate="2012" docId="7C0D87B7FFB8FFAB8987FF568F56F447" docLanguage="en" docName="zt03193p052.pdf" docOrigin="Zootaxa 3193" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Deltochilum acanthus Kohlmann &amp; Solis, new species" docType="treatment" docVersion="10" lastPageNumber="38" masterDocId="8034FFCFFFB0FFA18910FFC18D46F523" masterDocTitle="New species and revalidations of scarab beetles (Coleoptera: Geotrupidae: Athyreini and Coleoptera: Scarabaeidae: Scarabaeinae) from Costa Rica and Panama" masterLastPageNumber="52" masterPageNumber="28" pageNumber="36" updateTime="1698332701623" updateUser="ExternalLinkService">
<mods:mods id="BC9690C1EBCADD57FF5C82CFEF0AC7DF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="963BFB34C9C903782117CDD72384F879">New species and revalidations of scarab beetles (Coleoptera: Geotrupidae: Athyreini and Coleoptera: Scarabaeidae: Scarabaeinae) from Costa Rica and Panama</mods:title>
</mods:titleInfo>
<mods:name id="B16AD8C32366622A9CA97A7D220CFC27" type="personal">
<mods:role id="F44E85B54723E7095B99CB6216A05AF9">
<mods:roleTerm id="7BE05C6530FF8593AB500E0FF2568CDE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0CC1EF992068D3EBF5F02A6D8621D58F">Kohlmann, Bert</mods:namePart>
</mods:name>
<mods:name id="AD390C7E5E6C20B7CDC74F256837860F" type="personal">
<mods:role id="AB1AA5FEBD274AE286ED1CCF53070ED6">
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<mods:namePart id="7235240E867D2B5923D6ED36EDE2DA7A">Solís, Ángel</mods:namePart>
</mods:name>
<mods:typeOfResource id="B52A534426320F7816F536D973DA9586">text</mods:typeOfResource>
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<mods:title id="BFE9FC932851FC5EC2D2158AABF8CE81">Zootaxa</mods:title>
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<mods:date id="BA0FBBC065504529B76F2E276A09A919">2012</mods:date>
<mods:detail id="B9E252C621EA82396F31C2AE50946A13" type="volume">
<mods:number id="6A52E56D82BAC5A6FE111CC1F0CE05B2">3193</mods:number>
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<mods:classification id="955E797560097D88F3E4EFA86D8A249F">journal article</mods:classification>
<mods:identifier id="4FF87993A163BD494DC7DEC888489A95" type="CLB-Dataset">45436</mods:identifier>
<mods:identifier id="E21FFF2039BE9C62C24745F193EE1A53" type="DOI">10.5281/zenodo.211122</mods:identifier>
<mods:identifier id="ED6F9DB49BAA8229943A306872FE8DC8" type="GBIF-Dataset">344f8458-2c80-43b3-a96b-3aca02033f88</mods:identifier>
<mods:identifier id="7DD2396E91EDF6B11AB488EB829BED23" type="ISSN">1175-5326</mods:identifier>
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<treatment id="7C0D87B7FFB8FFAB8987FF568F56F447" ID-DOI="http://doi.org/10.5281/zenodo.3499664" ID-GBIF-Taxon="119507525" ID-Zenodo-Dep="3499664" LSID="urn:lsid:plazi:treatment:7C0D87B7FFB8FFAB8987FF568F56F447" httpUri="http://treatment.plazi.org/id/7C0D87B7FFB8FFAB8987FF568F56F447" lastPageId="10" lastPageNumber="38" pageId="8" pageNumber="36">
<subSubSection id="BCBE652AFFB8FFA98987FF568C6FF5F1" pageId="8" pageNumber="36" type="nomenclature">
<paragraph id="F41B36A1FFB8FFA98987FF568E76F592" blockId="8.[151,816,151,210]" box="[151,816,151,177]" pageId="8" pageNumber="36">
<heading id="AF5381CDFFB8FFA98987FF568E76F592" bold="true" box="[151,816,151,177]" fontSize="11" level="1" pageId="8" pageNumber="36" reason="1">
<emphasis id="C6D0EAB3FFB8FFA98987FF568E76F592" bold="true" box="[151,816,151,177]" pageId="8" pageNumber="36">
<taxonomicName id="33A44D22FFB8FFA98987FF568FD4F592" ID-CoL="6CJ62" authority="Kohlmann &amp; Solis" authorityName="Kohlmann &amp; Solis" box="[151,658,151,177]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="36" phylum="Arthropoda" rank="species" species="acanthus" status="sp. nov.">
<emphasis id="C6D0EAB3FFB8FFA98987FF568CE5F592" bold="true" box="[151,419,151,177]" italics="true" pageId="8" pageNumber="36">Deltochilum acanthus</emphasis>
Kohlmann &amp; Solís
</taxonomicName>
,
<taxonomicNameLabel id="DDE357C8FFB8FFA98BB0FF568E76F592" box="[672,816,151,177]" pageId="8" pageNumber="36" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="F41B36A1FFB8FFA98987FF7B8C6FF5F1" blockId="8.[151,816,151,210]" box="[151,297,186,210]" pageId="8" pageNumber="36">
<figureCitation id="6C9F2A24FFB8FFA98987FF7B8DA3F5F1" box="[151,229,186,210]" captionStart="FIGURE 6" captionStartId="7.[151,250,1994,2017]" captionTargetBox="[326,1257,693,1954]" captionTargetId="figure@7.[319,1266,687,1962]" captionTargetPageId="7" captionText="FIGURE 6. Dorsal view of a male Deltochilum acanthus Kohlmann &amp; Solís, new species." httpUri="https://zenodo.org/record/211128/files/figure.png" pageId="8" pageNumber="36">Figs. 6</figureCitation>
,
<figureCitation id="6C9F2A24FFB8FFA989E1FF7A8C47F5F1" box="[241,257,187,210]" captionStart="FIGURE 7" captionStartId="8.[151,250,1921,1944]" captionTargetBox="[402,1185,660,1864]" captionTargetId="figure@8.[395,1192,656,1873]" captionTargetPageId="8" captionText="FIGURE 7. Side view of elytra (a, c) and pygidium (e) and dorsal view of left elytra (g) of Deltochilum valgum acropyge, and Deltochilum acanthus Kohlmann &amp; Solís, new species (b, d, f, h). Carinae are delineated." httpUri="https://zenodo.org/record/211129/files/figure.png" pageId="8" pageNumber="36">7</figureCitation>
,
<figureCitation id="6C9F2A24FFB8FFA9881DFF7B8C6FF5F1" box="[269,297,186,210]" captionStart="FIGURE 15" captionStartId="22.[151,250,881,904]" captionTargetBox="[340,1246,195,857]" captionTargetId="figure@22.[340,1246,195,857]" captionTargetPageId="22" captionText="FIGURE 15. Known distribution of Deltochilum acanthus Kohlmann &amp; Solís, new species (triangles) and Deltochilum valgum acropyge (circles). Base map from Google Maps." httpUri="https://zenodo.org/record/211136/files/figure.png" pageId="8" pageNumber="36">15</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="BCBE652AFFB8FFA98987FEC38CA2F441" pageId="8" pageNumber="36" type="diagnosis">
<paragraph id="F41B36A1FFB8FFA98987FEC38CA2F441" blockId="8.[151,1436,258,598]" pageId="8" pageNumber="36">
<emphasis id="C6D0EAB3FFB8FFA98987FEC38C57F438" bold="true" box="[151,273,258,283]" pageId="8" pageNumber="36">Diagnosis.</emphasis>
This species is distinguished from other Costa Rican
<taxonomicName id="33A44D22FFB8FFA98A78FEC28EB4F438" box="[872,1010,259,283]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="C6D0EAB3FFB8FFA98A78FEC28EB4F438" box="[872,1010,259,283]" italics="true" pageId="8" pageNumber="36">Deltochilum</emphasis>
</taxonomicName>
species by the following combination of characters: Pygidium with apex thickened, very acutely angled (90°) and outwardly produced (
<figureCitation id="6C9F2A24FFB8FFA98DFCFEE78876F41D" box="[1260,1328,294,318]" captionStart="FIGURE 7" captionStartId="8.[151,250,1921,1944]" captionTargetBox="[402,1185,660,1864]" captionTargetId="figure@8.[395,1192,656,1873]" captionTargetPageId="8" captionText="FIGURE 7. Side view of elytra (a, c) and pygidium (e) and dorsal view of left elytra (g) of Deltochilum valgum acropyge, and Deltochilum acanthus Kohlmann &amp; Solís, new species (b, d, f, h). Carinae are delineated." httpUri="https://zenodo.org/record/211129/files/figure.png" pageId="8" pageNumber="36">Fig. 7</figureCitation>
f). Apical half of metatibia bent inward.
</paragraph>
</subSubSection>
<subSubSection id="BCBE652AFFB8FFA889D7FEAF8C63F71F" lastPageId="9" lastPageNumber="37" pageId="8" pageNumber="36" type="description">
<paragraph id="F41B36A1FFB8FFA989D7FEAF8CA6F489" blockId="8.[151,1436,258,598]" pageId="8" pageNumber="36">
<emphasis id="C6D0EAB3FFB8FFA989D7FEAF8C92F4A4" bold="true" box="[199,468,366,391]" pageId="8" pageNumber="36">
Description.
<typeStatus id="2B1F8803FFB8FFA98870FEAE8C89F4A4" box="[352,463,367,391]" pageId="8" pageNumber="36" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male (
<figureCitation id="6C9F2A24FFB8FFA98B36FEAF8F2BF4A5" box="[550,621,366,390]" captionStart="FIGURE 6" captionStartId="7.[151,250,1994,2017]" captionTargetBox="[326,1257,693,1954]" captionTargetId="figure@7.[319,1266,687,1962]" captionTargetPageId="7" captionText="FIGURE 6. Dorsal view of a male Deltochilum acanthus Kohlmann &amp; Solís, new species." httpUri="https://zenodo.org/record/211128/files/figure.png" pageId="8" pageNumber="36">Fig. 6</figureCitation>
): Length 11.9 mm. Humeral width 8.0 mm. Body color black, head and pronotum distinctly punctate.
</paragraph>
<paragraph id="F41B36A1FFB8FFA989D7FE778C1CF732" blockId="8.[151,1436,258,598]" pageId="8" pageNumber="36">Head approximately as long as wide; clypeus with two distinct, narrowly separated, upwardly reflexed, pointed teeth, pointed anterior teeth; each clypeal-genal margin with low obtuse teeth; vertex closely punctate, nearly flat; dorsal eyes large.</paragraph>
<paragraph id="F41B36A1FFB8FFA989D7FDDA8EF6F775" blockId="8.[151,1436,258,598]" pageId="8" pageNumber="36">Pronotum with lateral margin angulate at middle, otherwise straight to slightly sinuate before and behind; pronotal surface densely punctate, giving the appearance of a honeycomb.</paragraph>
<caption id="A0DB6629FFB8FFA98987F8408964F295" httpUri="https://zenodo.org/record/211129/files/figure.png" pageId="8" pageNumber="36" targetBox="[402,1185,660,1864]" targetPageId="8">
<paragraph id="F41B36A1FFB8FFA98987F8408964F295" blockId="8.[151,1435,1921,1974]" pageId="8" pageNumber="36">
<emphasis id="C6D0EAB3FFB8FFA98987F8408C55F2B4" bold="true" box="[151,275,1921,1944]" pageId="8" pageNumber="36">FIGURE 7.</emphasis>
Side view of elytra (a, c) and pygidium (e) and dorsal view of left elytra (g) of
<taxonomicName id="33A44D22FFB8FFA98D29F843882CF2BB" box="[1081,1386,1922,1944]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="36" phylum="Arthropoda" rank="subSpecies" species="valgum" subSpecies="acropyge">
<emphasis id="C6D0EAB3FFB8FFA98D29F843882CF2BB" box="[1081,1386,1922,1944]" italics="true" pageId="8" pageNumber="36">Deltochilum valgum acropyge</emphasis>
</taxonomicName>
, and
<taxonomicName id="33A44D22FFB8FFA98987F8618F7AF295" authority="Kohlmann &amp; Solis" authorityName="Kohlmann &amp; Solis" box="[151,572,1952,1974]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="36" phylum="Arthropoda" rank="species" species="acanthus" status="sp. nov.">
<emphasis id="C6D0EAB3FFB8FFA98987F8618C33F295" box="[151,373,1952,1974]" italics="true" pageId="8" pageNumber="36">Deltochilum acanthus</emphasis>
Kohlmann &amp; Solís
</taxonomicName>
,
<taxonomicNameLabel id="DDE357C8FFB8FFA98B56F8608F86F295" box="[582,704,1952,1974]" pageId="8" pageNumber="36" rank="species">new species</taxonomicNameLabel>
(b, d, f, h). Carinae are delineated.
</paragraph>
</caption>
<paragraph id="F41B36A1FFB9FFA889D7FF568852F5D4" blockId="9.[151,1437,151,2034]" pageId="9" pageNumber="37">
Elytral striae indicated by a double line, line expanded by widely separated, shallow strial punctures; disc intervals opaque, with numerous shallow, very close punctures (
<figureCitation id="6C9F2A24FFB9FFA88A63FF7D8EFFF5F7" box="[883,953,188,212]" captionStart="FIGURE 7" captionStartId="8.[151,250,1921,1944]" captionTargetBox="[402,1185,660,1864]" captionTargetId="figure@8.[395,1192,656,1873]" captionTargetPageId="8" captionText="FIGURE 7. Side view of elytra (a, c) and pygidium (e) and dorsal view of left elytra (g) of Deltochilum valgum acropyge, and Deltochilum acanthus Kohlmann &amp; Solís, new species (b, d, f, h). Carinae are delineated." httpUri="https://zenodo.org/record/211129/files/figure.png" pageId="9" pageNumber="37">Fig. 7</figureCitation>
h); humeral umbone and apices of third to seventh intervals carinate; elytral surface shagreened. Epipleural upper carina interrupted (
<figureCitation id="6C9F2A24FFB9FFA88D9CFF1E899DF5D4" box="[1164,1243,223,247]" captionStart="FIGURE 7" captionStartId="8.[151,250,1921,1944]" captionTargetBox="[402,1185,660,1864]" captionTargetId="figure@8.[395,1192,656,1873]" captionTargetPageId="8" captionText="FIGURE 7. Side view of elytra (a, c) and pygidium (e) and dorsal view of left elytra (g) of Deltochilum valgum acropyge, and Deltochilum acanthus Kohlmann &amp; Solís, new species (b, d, f, h). Carinae are delineated." httpUri="https://zenodo.org/record/211129/files/figure.png" pageId="9" pageNumber="37">Figs. 7</figureCitation>
b, d).
</paragraph>
<paragraph id="F41B36A1FFB9FFA889D7FEC58CDEF447" blockId="9.[151,1437,151,2034]" pageId="9" pageNumber="37">
Pygidium with apex thickened, very acutely angled and outwardly produced (
<figureCitation id="6C9F2A24FFB9FFA88D52FEC589C1F43F" box="[1090,1159,260,284]" captionStart="FIGURE 7" captionStartId="8.[151,250,1921,1944]" captionTargetBox="[402,1185,660,1864]" captionTargetId="figure@8.[395,1192,656,1873]" captionTargetPageId="8" captionText="FIGURE 7. Side view of elytra (a, c) and pygidium (e) and dorsal view of left elytra (g) of Deltochilum valgum acropyge, and Deltochilum acanthus Kohlmann &amp; Solís, new species (b, d, f, h). Carinae are delineated." httpUri="https://zenodo.org/record/211129/files/figure.png" pageId="9" pageNumber="37">Fig. 7</figureCitation>
f); disc nearly flat, shallowly and densely punctate, punctures umbilical. First abdominal sternite forms a rearward projection at the middle of the posterior border.
</paragraph>
<paragraph id="F41B36A1FFB9FFA889D7FEAE8E1BF48F" blockId="9.[151,1437,151,2034]" pageId="9" pageNumber="37">Profemur with margin unmodified; protibia with three apical teeth (the middle one smaller) and numerous serrations on outer margin. Apical half of metatibia bent inwardly.</paragraph>
<paragraph id="F41B36A1FFB9FFA889D7FE768F93F4D7" blockId="9.[151,1437,151,2034]" pageId="9" pageNumber="37">
<emphasis id="C6D0EAB3FFB9FFA889D7FE768C75F4F3" bold="true" box="[199,307,439,464]" pageId="9" pageNumber="37">Allotype.</emphasis>
Female. Length 11.2 mm. Humeral width 7.6 mm. Similar to male, but the first abdominal sternite is evenly arched at the middle of the posterior border.
</paragraph>
<paragraph id="F41B36A1FFB9FFA889D7FE3E8C63F71F" blockId="9.[151,1437,151,2034]" pageId="9" pageNumber="37">
<emphasis id="C6D0EAB3FFB9FFA889D7FE3E8C7DF73B" bold="true" box="[199,315,511,536]" pageId="9" pageNumber="37">Variation.</emphasis>
Fifty-eight specimens examined,
<specimenCount id="E2A2FD28FFB9FFA88BAFFE3E8E62F734" box="[703,804,511,535]" pageId="9" pageNumber="37" type="male">38 males</specimenCount>
and
<specimenCount id="E2A2FD28FFB9FFA88A4DFE3E8E9AF734" box="[861,988,511,535]" pageId="9" pageNumber="37" type="female">20 females</specimenCount>
. Length 10.4–13.0 mm. Humeral width 7.3–8.0 mm.
</paragraph>
</subSubSection>
<subSubSection id="BCBE652AFFB9FFAB89D7FD868802F43F" lastPageId="10" lastPageNumber="38" pageId="9" pageNumber="37" type="materials_examined">
<paragraph id="F41B36A1FFB9FFA889D7FD868DAFF097" blockId="9.[151,1437,151,2034]" pageId="9" pageNumber="37">
<emphasis id="C6D0EAB3FFB9FFA889D7FD868CF6F743" bold="true" box="[199,432,583,608]" pageId="9" pageNumber="37">Examined material</emphasis>
(
<specimenCount id="E2A2FD28FFB9FFA888D3FD868F27F77C" box="[451,609,583,607]" pageId="9" pageNumber="37" type="generic">58 specimens</specimenCount>
).
<emphasis id="C6D0EAB3FFB9FFA88B6BFD898FA8F743" bold="true" box="[635,750,584,608]" pageId="9" pageNumber="37">
<typeStatus id="2B1F8803FFB9FFA88B6BFD898FACF743" box="[635,746,584,608]" pageId="9" pageNumber="37" type="holotype">Holotype</typeStatus>
,
</emphasis>
male:
<collectingCountry id="8CB37631FFB9FFA88A52FD868EABF77C" box="[834,1005,583,607]" name="Costa Rica" pageId="9" pageNumber="37">COSTA RICA</collectingCountry>
. Prov. Puntarenas. Res. Biol. Carara, Est., Quebrada Bonita.
<quantity id="335C9B44FFB9FFA888B8FDAD8CA1F7A7" box="[424,487,620,644]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="37" unit="m" value="50.0">50 m</quantity>
,
<date id="801A1061FFB9FFA888E7FDAD8F27F7A7" box="[503,609,620,644]" pageId="9" pageNumber="37" value="1993-06">jun 1993</date>
. J.C. Saborío. L-N-194500, 469850,
<collectionCode id="92B5AE64FFB9FFA88D03FDAD8904F7A7" LSID="urn:lsid:biocol.org:col:14323" box="[1043,1090,620,644]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="9" pageNumber="37">CRI</collectionCode>
001185073.
<emphasis id="C6D0EAB3FFB9FFA88DC1FDAA8870F7A7" bold="true" box="[1233,1334,619,644]" pageId="9" pageNumber="37">Allotype</emphasis>
, female:
<collectingCountry id="8CB37631FFB9FFA88987FD4E8C00F784" box="[151,326,655,679]" name="Costa Rica" pageId="9" pageNumber="37">COSTA RICA</collectingCountry>
. Prov. Puntarenas. R.B. Carara, Est. Quebrada Bonita.
<quantity id="335C9B44FFB9FFA88AF6FD4E896EF784" box="[998,1064,655,679]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="37" unit="m" value="50.0">50 m</quantity>
,
<date id="801A1061FFB9FFA88D2CFD5189F2F784" box="[1084,1204,655,679]" pageId="9" pageNumber="37" value="1993-11">nov. 1993</date>
. J.C. Saborío. L S 194500_469850, #2470,
<collectionCode id="92B5AE64FFB9FFA888A5FD758CA2F7EF" LSID="urn:lsid:biocol.org:col:14323" box="[437,484,692,716]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="9" pageNumber="37">CRI</collectionCode>
001969700.
<emphasis id="C6D0EAB3FFB9FFA88B64FD758FB4F7EF" bold="true" box="[628,754,692,716]" pageId="9" pageNumber="37">
<typeStatus id="2B1F8803FFB9FFA88B64FD758FA8F7EF" box="[628,750,692,716]" pageId="9" pageNumber="37" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCountry id="8CB37631FFB9FFA88BECFD758EE1F7EF" box="[764,935,692,716]" name="Costa Rica" pageId="9" pageNumber="37">COSTA RICA</collectingCountry>
. Prov. Puntarenas. Centro Juvenil Tropical. Alrededor de la Estación.
<specimenCount id="E2A2FD28FFB9FFA888D7FD168F48F7CC" box="[455,526,727,751]" pageId="9" pageNumber="37" type="generic">
<quantity id="335C9B44FFB9FFA888D7FD168F4CF7CC" box="[455,522,727,751]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="37" unit="m" value="100.0">100m</quantity>
.
</specimenCount>
<date id="801A1061FFB9FFA88B08FD168F89F7CC" box="[536,719,727,751]" pageId="9" pageNumber="37" value="1997-07-05" valueMax="1997-07-12" valueMin="1997-07-05">5–12 JUL 1997</date>
. M. Lobo. Colecta Nocturna. L_S_294700_517100 #47733,
<specimenCount id="E2A2FD28FFB9FFA88C9EFD1688DAF7CC" box="[1422,1436,727,751]" pageId="9" pageNumber="37" type="female">1</specimenCount>
female; Estación Esquinas, 0 m, Península de Osa,
<date id="801A1061FFB9FFA88BC8FD3D8E00F637" box="[728,838,764,788]" pageId="9" pageNumber="37" value="1993-04">Abr 1993</date>
. M. Segura, L-S 301400_542200,
<specimenCount id="E2A2FD28FFB9FFA88DDBFD3D899FF637" box="[1227,1241,764,788]" pageId="9" pageNumber="37" type="female">1</specimenCount>
female; Estación Quebrada Bonita,
<quantity id="335C9B44FFB9FFA88873FCDE8CD8F61B" box="[355,414,799,824]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="37" unit="m" value="50.0">50 m</quantity>
, Res. Biol. Carara,
<date id="801A1061FFB9FFA88B68FCDE8E65F614" box="[632,803,799,824]" pageId="9" pageNumber="37" value="1992-07-01" valueMax="1992-07-29" valueMin="1992-07-01">1 a 29 jul 1992</date>
, R. Guzmán, L- N 194500_469850,
<specimenCount id="E2A2FD28FFB9FFA88DABFCDE898FF614" box="[1211,1225,799,823]" pageId="9" pageNumber="37" type="male">1</specimenCount>
male;
<date id="801A1061FFB9FFA88C05FCDE8839F614" box="[1301,1407,799,823]" pageId="9" pageNumber="37" value="1993-10">Oct 1993</date>
. J. C. Saborío, L N 194500_469850 # 2396,
<specimenCount id="E2A2FD28FFB9FFA88B74FC858F34F67F" box="[612,626,836,860]" pageId="9" pageNumber="37" type="female">1</specimenCount>
female;
<date id="801A1061FFB9FFA88BC3FC858E7EF67F" box="[723,824,836,860]" pageId="9" pageNumber="37" value="1993-09">Set 1993</date>
. J. Saborío, L N 194500_469850 # 2354,
<specimenCount id="E2A2FD28FFB9FFA88C18FC858850F67F" box="[1288,1302,836,860]" pageId="9" pageNumber="37" type="male">2</specimenCount>
males;
<date id="801A1061FFB9FFA88C7CFC848D94F65E" pageId="9" pageNumber="37" value="1993-11">Nov 1993</date>
, R. Guzmán, L N 194500_469850 # 2447,
<specimenCount id="E2A2FD28FFB9FFA88BD6FCA48F92F65E" box="[710,724,869,893]" pageId="9" pageNumber="37" type="male">2</specimenCount>
males;
<date id="801A1061FFB9FFA88A3FFCA78EDCF65E" box="[815,922,869,893]" pageId="9" pageNumber="37" value="1993-06">Jun 1993</date>
, R. Guzmán, L N 194500_469850 # 2202,
<specimenCount id="E2A2FD28FFB9FFA88C9EFCA488DAF65E" box="[1422,1436,869,893]" pageId="9" pageNumber="37" type="male">1</specimenCount>
male;
<date id="801A1061FFB9FFA889CCFC478CCCF6BD" box="[220,394,902,926]" pageId="9" pageNumber="37" value="1992-09-02" valueMax="1992-09-23" valueMin="1992-09-02">2 a 23 set 1992</date>
, R. Guzmán, L-N 194500_469850,
<specimenCount id="E2A2FD28FFB9FFA88A0DFC478E6DF6BD" box="[797,811,902,926]" pageId="9" pageNumber="37" type="male">3</specimenCount>
males; agos 1993, R.M. Guzmán, L N 194500_469850 # 2297,
<specimenCount id="E2A2FD28FFB9FFA889E1FC668DB9F69C" box="[241,255,935,959]" pageId="9" pageNumber="37" type="female">1</specimenCount>
female;
<date id="801A1061FFB9FFA88871FC698C88F69C" box="[353,462,935,959]" pageId="9" pageNumber="37" value="1994-01">Ene 1994</date>
, R. M. Guzmán, L N 194500_469850 # 2572,
<specimenCount id="E2A2FD28FFB9FFA88AC6FC668EA2F69C" box="[982,996,935,959]" pageId="9" pageNumber="37" type="female">1</specimenCount>
female;
<date id="801A1061FFB9FFA88D56FC6689F7F69C" box="[1094,1201,935,959]" pageId="9" pageNumber="37" value="1994-02">Feb 1994</date>
, R. M. Guzmán, L N 194500_469850 # 2613,
<specimenCount id="E2A2FD28FFB9FFA888A1FC088CF9F6C2" box="[433,447,969,993]" pageId="9" pageNumber="37" type="male">1</specimenCount>
male;
<date id="801A1061FFB9FFA88B1DFC0B8FC4F6C2" box="[525,642,969,994]" pageId="9" pageNumber="37" value="1994-05">May 1994</date>
, R. M. Guzmán, L N 194500_469850 # 2914,
<specimenCount id="E2A2FD28FFB9FFA88D84FC0889E4F6C2" box="[1172,1186,969,993]" pageId="9" pageNumber="37" type="male">3</specimenCount>
males,
<specimenCount id="E2A2FD28FFB9FFA88DEAFC088826F6C2" box="[1274,1376,969,993]" pageId="9" pageNumber="37" type="female">1 female</specimenCount>
;
<date id="801A1061FFB9FFA88C7CFC0B8D94F121" pageId="9" pageNumber="37" value="1994-08">Ago 1994</date>
, R. M. Guzmán, L N 194500_469850 # 3163,
<specimenCount id="E2A2FD28FFB9FFA88A56FC2B8E12F121" box="[838,852,1002,1026]" pageId="9" pageNumber="37" type="female">1</specimenCount>
female;
<date id="801A1061FFB9FFA88ADEFC2B8902F121" box="[974,1092,1002,1026]" pageId="9" pageNumber="37" value="1994-10">Oct 1994</date>
. J.C. Saborío, Desconocido L_N_469850_194500 #3288,
<specimenCount id="E2A2FD28FFB9FFA888FAFBCD8CBEF107" box="[490,504,1036,1060]" pageId="9" pageNumber="37" type="female">1</specimenCount>
female;
<date id="801A1061FFB9FFA88B4CFBCC8F81F107" box="[604,711,1036,1060]" pageId="9" pageNumber="37" value="1996-06">Jun 1996</date>
. R. Guzmán. L_N_195250_469850 #7648,
<specimenCount id="E2A2FD28FFB9FFA88DA4FBCD8984F107" box="[1204,1218,1036,1060]" pageId="9" pageNumber="37" type="male">1</specimenCount>
male;
<date id="801A1061FFB9FFA88C00FBCD883EF107" box="[1296,1400,1036,1060]" pageId="9" pageNumber="37" value="1994-09">Set 1994</date>
. R. M. Guzmán, L_N_194500,469850 #3214,
<specimenCount id="E2A2FD28FFB9FFA88B68FBEC8FC0F166" box="[632,646,1069,1093]" pageId="9" pageNumber="37" type="male">2</specimenCount>
males,
<specimenCount id="E2A2FD28FFB9FFA88BCEFBEC8E02F166" box="[734,836,1069,1093]" pageId="9" pageNumber="37" type="female">1 female</specimenCount>
;
<date id="801A1061FFB9FFA88A5FFBEF8E83F166" box="[847,965,1069,1094]" pageId="9" pageNumber="37" value="1994-05">May 1994</date>
. J. Saborío. L_N_470000_195200 #2849,
<specimenCount id="E2A2FD28FFB9FFA88987FB8F8DE3F145" box="[151,165,1102,1126]" pageId="9" pageNumber="37" type="male">4</specimenCount>
males,
<specimenCount id="E2A2FD28FFB9FFA88810FB8F8C21F145" box="[256,359,1102,1126]" pageId="9" pageNumber="37" type="female">1 female</specimenCount>
; Estación Sirena,
<quantity id="335C9B44FFB9FFA88B26FB8F8FD2F145" box="[566,660,1102,1126]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="9" pageNumber="37" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">0–100m</quantity>
, P. N. Corcovado,
<date id="801A1061FFB9FFA88A7EFB8E8EA7F145" box="[878,993,1102,1127]" pageId="9" pageNumber="37" value="1991-08">Ago 1991</date>
, J. C. Saborío, L S
<date id="801A1061FFB9FFA88DD8FB8F885BF145" box="[1224,1309,1102,1126]" pageId="9" pageNumber="37" value="00-05-27">270500</date>
_508300,
<specimenCount id="E2A2FD28FFB9FFA88C9EFB8F88DAF145" box="[1422,1436,1102,1126]" pageId="9" pageNumber="37" type="female">1</specimenCount>
female; G. Fonseca,
<date id="801A1061FFB9FFA88869FBB18CA6F1A4" box="[377,480,1135,1159]" pageId="9" pageNumber="37" value="1991-06">Jun 1991</date>
, L- S
<date id="801A1061FFB9FFA88B33FBAE8F3EF1A4" box="[547,632,1135,1159]" pageId="9" pageNumber="37" value="00-05-27">270500</date>
_508300,
<specimenCount id="E2A2FD28FFB9FFA88BF6FBAE8FB2F1A4" box="[742,756,1135,1159]" pageId="9" pageNumber="37" type="male">1</specimenCount>
male;
<date id="801A1061FFB9FFA88A2FFBB18EE0F1A4" box="[831,934,1135,1159]" pageId="9" pageNumber="37" value="1991-06">Jun 1991</date>
, J. C. Saborío, L- S
<date id="801A1061FFB9FFA88D96FBAE899DF1A4" box="[1158,1243,1135,1159]" pageId="9" pageNumber="37" value="00-05-27">270500</date>
_508300,
<specimenCount id="E2A2FD28FFB9FFA88C59FBAE8811F1A4" box="[1353,1367,1135,1159]" pageId="9" pageNumber="37" type="male">1</specimenCount>
male;
<date id="801A1061FFB9FFA88987FB508C47F18A" box="[151,257,1169,1193]" pageId="9" pageNumber="37" value="1993-10">Oct 1993</date>
. G. Fonseca, L S
<date id="801A1061FFB9FFA888D4FB508F5EF18A" box="[452,536,1169,1193]" pageId="9" pageNumber="37" value="00-05-27">270500</date>
_508300 # 2380,
<specimenCount id="E2A2FD28FFB9FFA88BCCFB508FACF18A" box="[732,746,1169,1193]" pageId="9" pageNumber="37" type="male">1</specimenCount>
male;
<date id="801A1061FFB9FFA88A27FB538ED8F18A" box="[823,926,1169,1193]" pageId="9" pageNumber="37" value="1990-01">Jan 1990</date>
, G. Fonseca, L_S_
<date id="801A1061FFB9FFA88D7DFB508985F18A" box="[1133,1219,1169,1193]" pageId="9" pageNumber="37" value="00-05-27">270500</date>
_508300,
<specimenCount id="E2A2FD28FFB9FFA88C23FB508807F18A" box="[1331,1345,1169,1193]" pageId="9" pageNumber="37" type="female">1</specimenCount>
female; G. Fonseca,
<date id="801A1061FFB9FFA88836FB738CD3F1E9" box="[294,405,1202,1226]" pageId="9" pageNumber="37" value="1991-04">Abr 1991</date>
, L- S
<date id="801A1061FFB9FFA888F2FB738F71F1E9" box="[482,567,1202,1226]" pageId="9" pageNumber="37" value="00-05-27">270500</date>
_508300,
<specimenCount id="E2A2FD28FFB9FFA88BB9FB738FF1F1E9" box="[681,695,1202,1226]" pageId="9" pageNumber="37" type="male">1</specimenCount>
male,
<specimenCount id="E2A2FD28FFB9FFA88A19FB738E37F1E9" box="[777,881,1202,1226]" pageId="9" pageNumber="37" type="female">1 female</specimenCount>
; G. Fonseca,
<date id="801A1061FFB9FFA88D1EFB73893DF1E9" box="[1038,1147,1202,1226]" pageId="9" pageNumber="37" value="1989-10">Oct 1989</date>
, L- S
<date id="801A1061FFB9FFA88DD7FB73885AF1E9" box="[1223,1308,1202,1226]" pageId="9" pageNumber="37" value="00-05-27">270500</date>
_508300,
<specimenCount id="E2A2FD28FFB9FFA88C9EFB7388DAF1E9" box="[1422,1436,1202,1226]" pageId="9" pageNumber="37" type="female">1</specimenCount>
female; F. Quesada,
<date id="801A1061FFB9FFA88891FB148CADF1CF" box="[385,491,1236,1260]" pageId="9" pageNumber="37" value="1990-06">Jun 1990</date>
, L- S
<date id="801A1061FFB9FFA88B23FB158FCEF1CF" box="[563,648,1236,1260]" pageId="9" pageNumber="37" value="00-05-27">270500</date>
_508300,
<specimenCount id="E2A2FD28FFB9FFA88BE9FB158E41F1CF" box="[761,775,1236,1260]" pageId="9" pageNumber="37" type="male">1</specimenCount>
male; Refugio de Vida Silvestre Golfito, Estación Naranjales,
<quantity id="335C9B44FFB9FFA889CAFB348C0DF02E" box="[218,331,1269,1293]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="9" pageNumber="37" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">0 – 100m</quantity>
,
<date id="801A1061FFB9FFA88848FB348CBCF02E" box="[344,506,1269,1294]" pageId="9" pageNumber="37" value="2004-04-25">25 ABR 2004</date>
, W. Porras, Libre, L_S_289900_553450 #76842,
<specimenCount id="E2A2FD28FFB9FFA88D23FB348907F02E" box="[1075,1089,1269,1293]" pageId="9" pageNumber="37" type="male">1</specimenCount>
male;
<date id="801A1061FFB9FFA88D81FB348836F02E" box="[1169,1392,1269,1294]" pageId="9" pageNumber="37" value="2004-04-22" valueMax="2004-04-27" valueMin="2004-04-22">22 – 27 ABR 2004</date>
, W. Porras, B. Gamboa, D. Briceño, M. Moraga, Amarilla, L_S_289900_553450 #76946,
<specimenCount id="E2A2FD28FFB9FFA88D74FAD78934F00D" box="[1124,1138,1302,1326]" pageId="9" pageNumber="37" type="male">2</specimenCount>
males,
<specimenCount id="E2A2FD28FFB9FFA88DDAFAD7887DF00D" box="[1226,1339,1302,1326]" pageId="9" pageNumber="37" type="female">3 females</specimenCount>
; Rancho Quemado,
<quantity id="335C9B44FFB9FFA88803FAF68C18F06C" box="[275,350,1335,1359]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="37" unit="m" value="200.0">200 m</quantity>
, Península de Osa.
<date id="801A1061FFB9FFA88B2FFAF68E57F06C" box="[575,785,1335,1360]" pageId="9" pageNumber="37" value="1993-05-12" valueMax="1993-05-24" valueMin="1993-05-12">12 a 24 may 1993</date>
. A. Gutiérrez, L S 292500_511000,
<specimenCount id="E2A2FD28FFB9FFA88DA1FAF689F9F06C" box="[1201,1215,1335,1359]" pageId="9" pageNumber="37" type="male">1</specimenCount>
male;
<date id="801A1061FFB9FFA88C1FFAF6883AF06C" box="[1295,1404,1335,1359]" pageId="9" pageNumber="37" value="1990-10">Oct 1990</date>
. F. Quesada. L-S 292500,511000,
<specimenCount id="E2A2FD28FFB9FFA888E7FA988F43F052" box="[503,517,1369,1393]" pageId="9" pageNumber="37" type="male">1</specimenCount>
male;
<date id="801A1061FFB9FFA88B44FA988E4BF052" box="[596,781,1369,1393]" pageId="9" pageNumber="37" value="1993-10-11" valueMax="1993-10-28" valueMin="1993-10-11">11–28 Oct 1993</date>
, A. H. Gutiérrez, L S 292500_511000 # 2409,
<specimenCount id="E2A2FD28FFB9FFA88C37FA988873F052" box="[1319,1333,1369,1393]" pageId="9" pageNumber="37" type="male">2</specimenCount>
males,
<specimenCount id="E2A2FD28FFB9FFA88C9EFA988DA1F0B1" pageId="9" pageNumber="37" type="female">1 female</specimenCount>
; Río Agujas. Estación Agujas. Send. Ajo.
<specimenCount id="E2A2FD28FFB9FFA88BD4FABB8E4DF0B1" box="[708,779,1402,1426]" pageId="9" pageNumber="37" type="generic">
<quantity id="335C9B44FFB9FFA88BD4FABB8E42F0B1" box="[708,772,1402,1426]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="37" unit="m" value="300.0">300m</quantity>
.
</specimenCount>
<date id="801A1061FFB9FFA88A02FABB8E8AF0B1" box="[786,972,1402,1426]" pageId="9" pageNumber="37" value="1998-01-06" valueMax="1998-01-12" valueMin="1998-01-06">6–12 ENE 1998</date>
. M. Lobo. L_S_276750_526550 #49736,
<specimenCount id="E2A2FD28FFB9FFA88987FA5D8DE3F097" box="[151,165,1436,1460]" pageId="9" pageNumber="37" type="male">1</specimenCount>
male.
</paragraph>
<paragraph id="F41B36A1FFB9FFA889D7FA7F88CCF0DA" blockId="9.[151,1437,151,2034]" pageId="9" pageNumber="37">
<collectingCountry id="8CB37631FFB9FFA889D7FA7F8C04F0F5" box="[199,322,1470,1494]" name="Panama" pageId="9" pageNumber="37">PANAMA</collectingCountry>
. Canal Zone, Barro Colorado, Poscher’s Peninsula,
<date id="801A1061FFB9FFA88A9DFA7C894CF0F6" box="[909,1034,1469,1493]" pageId="9" pageNumber="37" value="1986-06-06">6 jun 1986</date>
. H. Wolda,
<specimenCount id="E2A2FD28FFB9FFA88D80FA7C89A6F0F6" box="[1168,1248,1469,1493]" pageId="9" pageNumber="37" type="male">1 male</specimenCount>
;
<date id="801A1061FFB9FFA88DFCFA7C8833F0F6" box="[1260,1397,1469,1493]" pageId="9" pageNumber="37" value="1986-06-11">11 jun 1986</date>
. H. Wolda,
<specimenCount id="E2A2FD28FFB9FFA889FDFA208C14F0DA" box="[237,338,1505,1529]" pageId="9" pageNumber="37" type="female">1 female</specimenCount>
;
<date id="801A1061FFB9FFA8884DFA208CA0F0DA" box="[349,486,1505,1529]" pageId="9" pageNumber="37" value="1986-06-18">18 jun 1986</date>
. H. Wolda,
<specimenCount id="E2A2FD28FFB9FFA88B7AFA208FFEF0DA" box="[618,696,1505,1529]" pageId="9" pageNumber="37" type="male">1 male</specimenCount>
;
<date id="801A1061FFB9FFA88BD4FA208E0DF0DA" box="[708,843,1505,1529]" pageId="9" pageNumber="37" value="1986-06-25">25 jun 1986</date>
, H. Wolda,
<specimenCount id="E2A2FD28FFB9FFA88AC0FA208959F0DA" box="[976,1055,1505,1529]" pageId="9" pageNumber="37" type="male">1 male</specimenCount>
;
<date id="801A1061FFB9FFA88D3AFA2089F2F0DA" box="[1066,1204,1505,1529]" pageId="9" pageNumber="37" value="1987-09-11">11 sep 1987</date>
. H. Wolda,
<specimenCount id="E2A2FD28FFB9FFA88C27FA2088C0F0DA" box="[1335,1414,1505,1529]" pageId="9" pageNumber="37" type="male">1 male</specimenCount>
.
</paragraph>
<paragraph id="F41B36A1FFB9FFA889D7F9C48E55F362" blockId="9.[151,1437,151,2034]" pageId="9" pageNumber="37">
<emphasis id="C6D0EAB3FFB9FFA889D7F9C48C6CF33D" bold="true" box="[199,298,1541,1566]" pageId="9" pageNumber="37">Habitat.</emphasis>
The species has been collected with flight interception traps in tropical rain forest, ranging from
<quantity id="335C9B44FFB9FFA88C90F9C48D98F362" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="9" pageNumber="37" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">0– 100 m</quantity>
altitude, during the months of April to November.
</paragraph>
<paragraph id="F41B36A1FFB9FFA889D7F98C8CB1F3AA" blockId="9.[151,1437,151,2034]" pageId="9" pageNumber="37">
<emphasis id="C6D0EAB3FFB9FFA889D7F98C8F35F345" bold="true" box="[199,627,1613,1638]" pageId="9" pageNumber="37">
Geographical distribution (
<figureCitation id="6C9F2A24FFB9FFA88B1FF98C8F22F345" box="[527,612,1613,1638]" captionStart="FIGURE 15" captionStartId="22.[151,250,881,904]" captionTargetBox="[340,1246,195,857]" captionTargetId="figure@22.[340,1246,195,857]" captionTargetPageId="22" captionText="FIGURE 15. Known distribution of Deltochilum acanthus Kohlmann &amp; Solís, new species (triangles) and Deltochilum valgum acropyge (circles). Base map from Google Maps." httpUri="https://zenodo.org/record/211136/files/figure.png" pageId="9" pageNumber="37">Fig. 15</figureCitation>
).
</emphasis>
This species is known so far from the Pacific rain forest of
<collectingCountry id="8CB37631FFB9FFA88C30F98C88DAF346" box="[1312,1436,1613,1637]" name="Costa Rica" pageId="9" pageNumber="37">Costa Rica</collectingCountry>
and the Canal Zone of
<collectingCountry id="8CB37631FFB9FFA88887F9B38CB2F3AA" box="[407,500,1650,1673]" name="Panama" pageId="9" pageNumber="37">Panama</collectingCountry>
.
</paragraph>
<paragraph id="F41B36A1FFB9FFA889D7F9548923F3F2" blockId="9.[151,1437,151,2034]" pageId="9" pageNumber="37">
<emphasis id="C6D0EAB3FFB9FFA889D7F9548C91F38D" bold="true" box="[199,471,1685,1710]" pageId="9" pageNumber="37">Chorological affinities.</emphasis>
<taxonomicName id="33A44D22FFB9FFA888CFF9548F90F38E" box="[479,726,1685,1709]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="37" phylum="Arthropoda" rank="species" species="acanthus">
<emphasis id="C6D0EAB3FFB9FFA888CFF9548F90F38E" box="[479,726,1685,1709]" italics="true" pageId="9" pageNumber="37">Deltochilum acanthus</emphasis>
</taxonomicName>
seems to show a geographic vicariant pattern in relation to the similar species
<taxonomicName id="33A44D22FFB9FFA88853F97B8F60F3F2" box="[323,550,1721,1745]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="37" phylum="Arthropoda" rank="subSpecies" species="valgum" subSpecies="acropyge">
<emphasis id="C6D0EAB3FFB9FFA88853F97B8F60F3F2" box="[323,550,1721,1745]" italics="true" pageId="9" pageNumber="37">D. valgum acropyge</emphasis>
</taxonomicName>
, which inhabits the Caribbean slope of
<collectingCountry id="8CB37631FFB9FFA88AF3F9788927F3F2" box="[995,1121,1721,1745]" name="Costa Rica" pageId="9" pageNumber="37">Costa Rica</collectingCountry>
.
</paragraph>
<paragraph id="F41B36A1FFB9FFA889D7F91C8930F28A" blockId="9.[151,1437,151,2034]" pageId="9" pageNumber="37">
<emphasis id="C6D0EAB3FFB9FFA889D7F91C8CB2F3D5" bold="true" box="[199,500,1757,1782]" pageId="9" pageNumber="37">Taxonomic relationships.</emphasis>
It would appear that the new species originated from a vicariant event, when the Talamanca range rose up approximately 3 million years ago, isolating the rain forest on the Pacific coast from the rainforests on the Caribbean coast. This mechanism seems to account for the origin of a great number of other animal vicariant species, examined in
<bibRefCitation id="90354B50FFB9FFA88B31F8888EC5F242" author="Kohlmann" box="[545,899,1865,1889]" pageId="9" pageNumber="37" refString="Kohlmann, B. &amp; Wilkinson, J. (2007) The Tarcoles Line: biogeographic effects of the Talamanca Range in lower Central America. Giornale Italiano di Entomologia, 54, 1 - 30." type="journal article" year="2007">Kohlmann &amp; Wilkinson (2007)</bibRefCitation>
. We believe that
<taxonomicName id="33A44D22FFB9FFA88D53F88B89FEF242" box="[1091,1208,1865,1889]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="37" phylum="Arthropoda" rank="species" species="valgum">
<emphasis id="C6D0EAB3FFB9FFA88D53F88B89FEF242" box="[1091,1208,1865,1889]" italics="true" pageId="9" pageNumber="37">D. valgum</emphasis>
</taxonomicName>
needs to be studied, and that its different subspecies represent a species complex in need of hierarchical revaluation. We therefore describe
<taxonomicName id="33A44D22FFB9FFA889EBF8538CC4F28A" box="[251,386,1937,1961]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="37" phylum="Arthropoda" rank="species" species="acanthus">
<emphasis id="C6D0EAB3FFB9FFA889EBF8538CC4F28A" box="[251,386,1937,1961]" italics="true" pageId="9" pageNumber="37">D. acanthus</emphasis>
</taxonomicName>
as a species and not as a subspecies, in anticipation of this process.
</paragraph>
<paragraph id="F41B36A1FFB9FFAB89D7F8748802F43F" blockId="9.[151,1437,151,2034]" lastBlockId="10.[151,1436,151,356]" lastPageId="10" lastPageNumber="38" pageId="9" pageNumber="37">
<taxonomicName id="33A44D22FFB9FFA889D7F8748CF8F2EE" box="[199,446,1973,1997]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="37" phylum="Arthropoda" rank="species" species="acanthus">
<emphasis id="C6D0EAB3FFB9FFA889D7F8748CF8F2EE" box="[199,446,1973,1997]" italics="true" pageId="9" pageNumber="37">Deltochilum acanthus</emphasis>
</taxonomicName>
can be easily separated from
<taxonomicName id="33A44D22FFB9FFA88A1DF8778EC4F2EE" box="[781,898,1973,1997]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="37" phylum="Arthropoda" rank="species" species="valgum">
<emphasis id="C6D0EAB3FFB9FFA88A1DF8778EC4F2EE" box="[781,898,1973,1997]" italics="true" pageId="9" pageNumber="37">D. valgum</emphasis>
</taxonomicName>
by its pygidium, which has the thickened apex, very acutely angled and outwardly produced (
<figureCitation id="6C9F2A24FFB9FFA88B88F8188F9DF2D2" box="[664,731,2009,2033]" captionStart="FIGURE 7" captionStartId="8.[151,250,1921,1944]" captionTargetBox="[402,1185,660,1864]" captionTargetId="figure@8.[395,1192,656,1873]" captionTargetPageId="8" captionText="FIGURE 7. Side view of elytra (a, c) and pygidium (e) and dorsal view of left elytra (g) of Deltochilum valgum acropyge, and Deltochilum acanthus Kohlmann &amp; Solís, new species (b, d, f, h). Carinae are delineated." httpUri="https://zenodo.org/record/211129/files/figure.png" pageId="9" pageNumber="37">Fig. 7</figureCitation>
f), whereas
<taxonomicName id="33A44D22FFB9FFA88A4EF81B8E92F2D2" box="[862,980,2009,2033]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="37" phylum="Arthropoda" rank="species" species="valgum">
<emphasis id="C6D0EAB3FFB9FFA88A4EF81B8E92F2D2" box="[862,980,2009,2033]" italics="true" pageId="9" pageNumber="37">D. valgum</emphasis>
</taxonomicName>
has a much less thickened and projected pygidium (
<figureCitation id="6C9F2A24FFBAFFAB8803FF568C1EF58C" box="[275,344,151,175]" captionStart="FIGURE 7" captionStartId="8.[151,250,1921,1944]" captionTargetBox="[402,1185,660,1864]" captionTargetId="figure@8.[395,1192,656,1873]" captionTargetPageId="8" captionText="FIGURE 7. Side view of elytra (a, c) and pygidium (e) and dorsal view of left elytra (g) of Deltochilum valgum acropyge, and Deltochilum acanthus Kohlmann &amp; Solís, new species (b, d, f, h). Carinae are delineated." httpUri="https://zenodo.org/record/211129/files/figure.png" pageId="10" pageNumber="38">Fig. 7</figureCitation>
e). There are also differences in dorsal punctation: the pronotum in
<taxonomicName id="33A44D22FFBAFFAB8D4EFF5889AEF593" box="[1118,1256,152,176]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="38" phylum="Arthropoda" rank="species" species="acanthus">
<emphasis id="C6D0EAB3FFBAFFAB8D4EFF5889AEF593" box="[1118,1256,152,176]" italics="true" pageId="10" pageNumber="38">D. acanthus</emphasis>
</taxonomicName>
is very densely punctured, producing the effect of a honeycomb, whereas in
<taxonomicName id="33A44D22FFBAFFAB8A56FF7C8EFDF5F7" box="[838,955,188,212]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="38" phylum="Arthropoda" rank="species" species="valgum">
<emphasis id="C6D0EAB3FFBAFFAB8A56FF7C8EFDF5F7" box="[838,955,188,212]" italics="true" pageId="10" pageNumber="38">D. valgum</emphasis>
</taxonomicName>
the punctures are more spaced, by at least the length of one puncture. The elytral punctures are also different, in
<taxonomicName id="33A44D22FFBAFFAB8ABAFF208974F5DB" box="[938,1074,224,248]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="38" phylum="Arthropoda" rank="species" species="acanthus">
<emphasis id="C6D0EAB3FFBAFFAB8ABAFF208974F5DB" box="[938,1074,224,248]" italics="true" pageId="10" pageNumber="38">D. acanthus</emphasis>
</taxonomicName>
punctation is dense and the elytral striae are broad (
<figureCitation id="6C9F2A24FFBAFFAB886FFEC58C84F43F" box="[383,450,260,284]" captionStart="FIGURE 7" captionStartId="8.[151,250,1921,1944]" captionTargetBox="[402,1185,660,1864]" captionTargetId="figure@8.[395,1192,656,1873]" captionTargetPageId="8" captionText="FIGURE 7. Side view of elytra (a, c) and pygidium (e) and dorsal view of left elytra (g) of Deltochilum valgum acropyge, and Deltochilum acanthus Kohlmann &amp; Solís, new species (b, d, f, h). Carinae are delineated." httpUri="https://zenodo.org/record/211129/files/figure.png" pageId="10" pageNumber="38">Fig. 7</figureCitation>
h), whereas
<taxonomicName id="33A44D22FFBAFFAB8B59FEC48FF9F43F" box="[585,703,260,284]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="38" phylum="Arthropoda" rank="species" species="valgum">
<emphasis id="C6D0EAB3FFBAFFAB8B59FEC48FF9F43F" box="[585,703,260,284]" italics="true" pageId="10" pageNumber="38">D. valgum</emphasis>
</taxonomicName>
is less densely punctured and the striae are thin (
<figureCitation id="6C9F2A24FFBAFFAB8DF3FEC58861F43F" box="[1251,1319,260,284]" captionStart="FIGURE 7" captionStartId="8.[151,250,1921,1944]" captionTargetBox="[402,1185,660,1864]" captionTargetId="figure@8.[395,1192,656,1873]" captionTargetPageId="8" captionText="FIGURE 7. Side view of elytra (a, c) and pygidium (e) and dorsal view of left elytra (g) of Deltochilum valgum acropyge, and Deltochilum acanthus Kohlmann &amp; Solís, new species (b, d, f, h). Carinae are delineated." httpUri="https://zenodo.org/record/211129/files/figure.png" pageId="10" pageNumber="38">
Fig.
<quantity id="335C9B44FFBAFFAB8C08FEC58861F43F" box="[1304,1319,260,284]" metricMagnitude="-3" metricUnit="kg" metricValue="7.0" pageId="10" pageNumber="38" unit="g" value="7.0">7</quantity>
</figureCitation>
g).
</paragraph>
</subSubSection>
<subSubSection id="BCBE652AFFBAFFAB89D7FEE98F56F447" pageId="10" pageNumber="38" type="etymology">
<paragraph id="F41B36A1FFBAFFAB89D7FEE98F56F447" blockId="10.[151,1436,151,356]" pageId="10" pageNumber="38">
<emphasis id="C6D0EAB3FFBAFFAB89D7FEE98C08F463" bold="true" box="[199,334,296,320]" pageId="10" pageNumber="38">Etymology.</emphasis>
<taxonomicName id="33A44D22FFBAFFAB8844FEE98CFAF463" box="[340,444,296,320]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="50" phylum="Arthropoda" rank="species" species="acanthus">
<emphasis id="C6D0EAB3FFBAFFAB8844FEE98CFAF463" box="[340,444,296,320]" italics="true" pageId="10" pageNumber="38">Acanthus</emphasis>
</taxonomicName>
(ĸανθοζ = acanthos), a Latinized Greek noun in apposition, meaning thorn, making reference to the spiny pygidial apex.
</paragraph>
</subSubSection>
</treatment>
</document>
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