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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.949.51269" ID-GBIF-Dataset="e4d2c425-4596-487c-8c4f-b09330b48203" ID-PMC="PMC7381721" ID-Pensoft-Pub="1313-2970-949-1" ID-Pensoft-UUID="303D64593B3B550DBB7CE01AA547409D" ID-PubMed="32765176" ID-ZooBank="93BFA448BB7343CDBC4EA86090DA63D5" ModsDocID="1313-2970-949-1" checkinTime="1594832489974" checkinUser="pensoft" docAuthor="Salata, Sebastian &amp; Fisher, Brian L." docDate="2020" docId="6F63281014765DFA804BD19A2FD0C655" docLanguage="en" docName="ZooKeys 949: 1-185" docOrigin="ZooKeys 949" docSource="http://dx.doi.org/10.3897/zookeys.949.51269" docTitle="Pheidole vony Salata &amp; Fisher 2020, sp. nov." docType="treatment" docUuid="D61A8A41-8E37-4B94-A723-9EABB9A024D6" docUuidSource="ZooBank" docVersion="4" id="303D64593B3B550DBB7CE01AA547409D" lastPageNumber="1" masterDocId="303D64593B3B550DBB7CE01AA547409D" masterDocTitle="Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar" masterLastPageNumber="185" masterPageNumber="1" pageNumber="1" updateTime="1668169097828" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Salata, Sebastian</mods:namePart>
<mods:affiliation>Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-0811-2309</mods:nameIdentifier>
<mods:nameIdentifier type="email">sdsalata@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fisher, Brian L.</mods:namePart>
<mods:affiliation>Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-4653-3270</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>949</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>185</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.949.51269</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.949.51269</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-949-1</mods:identifier>
<mods:identifier type="ZooBank">93BFA448BB7343CDBC4EA86090DA63D5</mods:identifier>
<mods:identifier type="Pensoft-UUID">303D64593B3B550DBB7CE01AA547409D</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="165090174" LSID="urn:lsid:zoobank.org:act:D61A8A41-8E37-4B94-A723-9EABB9A024D6" httpUri="http://treatment.plazi.org/id/6F63281014765DFA804BD19A2FD0C655" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="6F632810-1476-5DFA-804B-D19A2FD0C655" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole vony" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="vony" status="sp. nov.">Pheidole vony</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 62" captionStartId="F62" captionText="Figure 62. Pheidole vony Forel, full-face view (A), profile (C), and dorsal view (E) of minor paratype worker (CASENT 0498053) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498055)." figureDoi="10.3897/zookeys.949.51269.figure62" httpUri="https://binary.pensoft.net/fig/432231" pageId="0" pageNumber="1">Figs 62A-F</figureCitation>
<figureCitation captionStart="Figure 64" captionStartId="F64" captionText="Figure 64. Major worker, hypostomal teeth. Pheidole masoandro sp. nov. (A). Pheidole mavohavoana sp. nov. (B). Pheidole midongy sp. nov. (C). Pheidole mikros sp. nov. (D). Pheidole mivory sp. nov. (E). Pheidole nitidobruna sp. nov. (F). Pheidole parvula sp. nov. (G). Pheidole parvulogibba sp. nov. (H). Pheidole renirano sp. nov. (I). Pheidole sava sp. nov. (J). Pheidole sikorae Forel (K). Pheidole sofia sp. nov. (L). Pheidole sparsa sp. nov. (M). Pheidole tampony sp. nov. (N). Pheidole trichotos sp. nov. (O). Pheidole tsaravoniana sp. nov. (P). Pheidole vadum sp. nov. (Q). Pheidole veteratrix Forel (R). Pheidole volontany sp. nov. (S). Pheidole vony sp. nov. (T)." figureDoi="10.3897/zookeys.949.51269.figure64" httpUri="https://binary.pensoft.net/fig/432233" pageId="0" pageNumber="1">, 64T</figureCitation>
<figureCitation captionStart="Figure 67" captionStartId="F67" captionText="Figure 67. Distribution. Pheidole renirano sp. nov. (A). Pheidole sava sp. nov. (B). Pheidole sikorae Forel (C). Pheidole sofia sp. nov. (D). Pheidole sparsa sp. nov. (E). Pheidole tampony sp. nov. (F). Pheidole trichotos sp. nov. (G). Pheidole tsaravoniana sp. nov. (H). Pheidole vadum sp. nov. (I). Pheidole veteratrix Forel (J). Pheidole volontany sp. nov. (K). Pheidole vony sp. nov. (L)." figureDoi="10.3897/zookeys.949.51269.figure67" httpUri="https://binary.pensoft.net/fig/432236" pageId="0" pageNumber="1">, 67L</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype.</emphasis>
Madagascar. • 1 major worker; Toamasina; Montagne
<normalizedToken originalValue="dAnjanaharibe">d'Anjanaharibe</normalizedToken>
, 19.5 km 27°NNE Ambinanitelo;
<geoCoordinate degrees="15.17833" direction="south" orientation="latitude" precision="1" value="-15.17833">-15.17833</geoCoordinate>
,
<geoCoordinate degrees="49.635" direction="east" orientation="longitude" precision="55" value="49.635">49.635</geoCoordinate>
; alt. 1100 m; 12 Mar 2003; Fisher et al. leg.; montane rainforest, ex rotten tree stump; BLF08197; CASENT0498055, bottom specimen on the pin (CASC).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes.</emphasis>
• 3w., 5s.; same data as for holotype; CASENT0498053, CASENT0872250CASENT0498054 (CASC, MHNG, PBZT).
</paragraph>
<caption doi="10.3897/zookeys.949.51269.figure62" httpUri="https://binary.pensoft.net/fig/432231" pageId="0" pageNumber="1" start="Figure 62" startId="F62">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 62.</emphasis>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole vony" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="vony">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole vony</emphasis>
</taxonomicName>
Forel, full-face view (
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
), profile (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
), and dorsal view (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
) of minor paratype worker (CASENT0498053) and full-face view (
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
), profile (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
), and dorsal view (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
) of holotype major worker (CASENT0498055).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Other material.</paragraph>
<paragraph pageId="0" pageNumber="1">
Madagascar. -
<emphasis bold="true" pageId="0" pageNumber="1">Toamasina</emphasis>
: • 3w.; Montagne
<normalizedToken originalValue="dAkirindro">d'Akirindro</normalizedToken>
7.6 km 341°NNW Ambinanitelo;
<geoCoordinate degrees="15.28833" direction="south" orientation="latitude" precision="1" value="-15.28833">-15.28833</geoCoordinate>
,
<geoCoordinate degrees="49.54833" direction="east" orientation="longitude" precision="1" value="49.54833">49.54833</geoCoordinate>
; alt. 600 m; 17 Mar 2003; Fisher et al. leg.; rainforest, ex rotten stick on ground; BLF08317 (CASC). • 1s.; Montagne
<normalizedToken originalValue="dAkirindro">d'Akirindro</normalizedToken>
7.6 km 341°NNW Ambinanitelo;
<geoCoordinate degrees="15.28833" direction="south" orientation="latitude" precision="1" value="-15.28833">-15.28833</geoCoordinate>
,
<geoCoordinate degrees="49.54833" direction="east" orientation="longitude" precision="1" value="49.54833">49.54833</geoCoordinate>
; alt. 600 m; 17 Mar 2003; Fisher et al. leg.; rainforest, ex rotten log; BLF08377 (CASC). • 3w.; Montagne
<normalizedToken originalValue="dAnjanaharibe">d'Anjanaharibe</normalizedToken>
, 19.5 km 27°NNE Ambinanitelo;
<geoCoordinate degrees="15.17833" direction="south" orientation="latitude" precision="1" value="-15.17833">-15.17833</geoCoordinate>
,
<geoCoordinate degrees="49.635" direction="east" orientation="longitude" precision="55" value="49.635">49.635</geoCoordinate>
; alt. 1100 m; 12 Mar 2003; Fisher et al. leg.; montane rainforest, ex rotten log; BLF08172 (CASC). • 3s.; Montagne
<normalizedToken originalValue="dAnjanaharibe">d'Anjanaharibe</normalizedToken>
, 19.5 km 27°NNE Ambinanitelo;
<geoCoordinate degrees="15.17833" direction="south" orientation="latitude" precision="1" value="-15.17833">-15.17833</geoCoordinate>
,
<geoCoordinate degrees="49.635" direction="east" orientation="longitude" precision="55" value="49.635">49.635</geoCoordinate>
; alt. 1100 m; 12 Mar 2003; Fisher et al. leg.; montane rainforest, ex root mat, ground layer; BLF08186 (CASC).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
Moderately large species.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Major workers.</emphasis>
Head in full-face view sub-oval, not widening posteriorly, with anterior and posterior sides slightly convex, in lateral view sub-oval, ventral and dorsal faces convex; sides of the head with moderately dense, moderately long, suberect to erect pilosity; medial frons with moderately dense, thin rugulae, anteriorly longitudinal and interrupted, posteromedial part with irregular rugae, interspaces shiny with sparse and indistinct irregular rugulae; lateral sides with longitudinal anteriorly to irregular posteriorly, thick, and relatively dense rugae with distinctly rugofoveolate interspaces; occipital lobes and area posterolateral from eyes never smooth; scape, when laid back, exceeding the midlength of head by three-fifths of its length; inner hypostomal teeth distinct, large, closely spaced, triangular, with rounded apex directed outward; outer hypostomal teeth lobe-like, lower and more narrow than inner hypostomal teeth, apex directed outward; inner and outer hypostomal teeth closely spaced and not connected by concavity; pronotum and mesonotum predominantly smooth with transverse rugae on dorsum and indistinct rugofoveolae on lateral sides; katepisternum and anepisternum predominantly smooth; propodeum rugofoveolate; body yellow.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
Head foveolate; area posterolateral from eyes smooth; scape, when laid back, surpassing the posterior head margin by two-fifths of its length; promesonotum moderately low and moderately short; promesonotal groove absent; propodeal spines very small and triangular; mesosoma smooth, sometimes with sparse and indistinct foveolae on lateral sides; body yellow.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
Measurements (
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
= 9): HL: 0.89-1.06 (0.95); HW: 0.83-1.02 (0.92); SL: 0.6-0.69 (0.65); EL: 0.12-0.14 (0.13); WL: 0.94-1.02 (0.98); PSL: 0.15-0.19 (0.16); MTL: 0.57-0.64 (0.6); PNW: 0.38-0.47 (0.42); PTW: 0.12-0.16 (0.13); PPW: 0.19-0.3 (0.24); CI: 100.5-106.4 (104.0); SI: 64.2-76.6 (71.5); PSLI: 15.5-20.0 (17.2); PPI: 47.3-61.5 (55.7); PNI: 44.7-48.5 (46.1); MTI: 59.4-71.7 (66.1).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
In full-face view sub-oval, not widening posteriorly, with anterior and posterior sides slightly convex (Fig.
<figureCitation captionStart="Figure 62" captionStartId="F62" captionText="Figure 62. Pheidole vony Forel, full-face view (A), profile (C), and dorsal view (E) of minor paratype worker (CASENT 0498053) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498055)." figureDoi="10.3897/zookeys.949.51269.figure62" httpUri="https://binary.pensoft.net/fig/432231" pageId="0" pageNumber="1">62B</figureCitation>
). In lateral view sub-oval; ventral and dorsal faces convex; inner hypostomal teeth visible. Sides of the head with moderately dense, moderately long, suberect to erect pilosity; whole head with dense, long, decumbent to erect pilosity. Medial part of frons with moderately dense, thin rugulae, anteriorly longitudinal and interrupted, posteromedial part with irregular rugae, interspaces shiny with sparse, indistinct, and irregular rugulae; lateral sides with longitudinal anteriorly to irregular posteriorly, thick, and relatively dense rugae with distinctly rugofoveolate interspaces. Occipital lobes with sparse, thick and irregular rugae; interspaces with fine rugofoveolae. Gena with relatively dense, thick, and longitudinal rugae and smooth to indistinctly rugofoveolae interspaces. Area posterolateral from eyes with thin and fine rugofoveolae, posteriormost parts with reduced sculpture and smooth notch. Centre of clypeus smooth and shiny, lateral sides with indistinct rugulae; median notch present, moderately wide, and shallow; median longitudinal carina present; lateral longitudinal carinae absent. Scape, when laid back, exceeding the midlength of head by three-fifths of its length; pilosity subdecumbent to erect (Fig.
<figureCitation captionStart="Figure 62" captionStartId="F62" captionText="Figure 62. Pheidole vony Forel, full-face view (A), profile (C), and dorsal view (E) of minor paratype worker (CASENT 0498053) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498055)." figureDoi="10.3897/zookeys.949.51269.figure62" httpUri="https://binary.pensoft.net/fig/432231" pageId="0" pageNumber="1">62B, D</figureCitation>
). Inner hypostomal teeth distinct, large, closely spaced, triangular, with rounded apex directed outward; outer hypostomal teeth lobe-like, lower and more narrow than inner hypostomal teeth, apex directed outward; inner and outer hypostomal teeth closely spaced not connected by concavity (Fig.
<figureCitation captionStart="Figure 64" captionStartId="F64" captionText="Figure 64. Major worker, hypostomal teeth. Pheidole masoandro sp. nov. (A). Pheidole mavohavoana sp. nov. (B). Pheidole midongy sp. nov. (C). Pheidole mikros sp. nov. (D). Pheidole mivory sp. nov. (E). Pheidole nitidobruna sp. nov. (F). Pheidole parvula sp. nov. (G). Pheidole parvulogibba sp. nov. (H). Pheidole renirano sp. nov. (I). Pheidole sava sp. nov. (J). Pheidole sikorae Forel (K). Pheidole sofia sp. nov. (L). Pheidole sparsa sp. nov. (M). Pheidole tampony sp. nov. (N). Pheidole trichotos sp. nov. (O). Pheidole tsaravoniana sp. nov. (P). Pheidole vadum sp. nov. (Q). Pheidole veteratrix Forel (R). Pheidole volontany sp. nov. (S). Pheidole vony sp. nov. (T)." figureDoi="10.3897/zookeys.949.51269.figure64" httpUri="https://binary.pensoft.net/fig/432233" pageId="0" pageNumber="1">64T</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
In lateral view, promesonotum short, angular, and moderately low, posterior mesonotum moderately steep, mesonotal process indistinct, tubercle-like or absent; promesonotal groove absent; metanotal groove indistinct; propodeal spines moderately long, moderately wide, with acute apex; humeral area laterally weakly produced (Fig.
<figureCitation captionStart="Figure 62" captionStartId="F62" captionText="Figure 62. Pheidole vony Forel, full-face view (A), profile (C), and dorsal view (E) of minor paratype worker (CASENT 0498053) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498055)." figureDoi="10.3897/zookeys.949.51269.figure62" httpUri="https://binary.pensoft.net/fig/432231" pageId="0" pageNumber="1">62D</figureCitation>
). Surface shiny; pronotum and mesonotum predominantly smooth with transverse rugae on dorsum and indistinct rugofoveolae on lateral sides; katepisternum and anepisternum predominantly smooth; propodeum rugofoveolate. Pilosity relatively dense, long, and erect (Fig.
<figureCitation captionStart="Figure 62" captionStartId="F62" captionText="Figure 62. Pheidole vony Forel, full-face view (A), profile (C), and dorsal view (E) of minor paratype worker (CASENT 0498053) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498055)." figureDoi="10.3897/zookeys.949.51269.figure62" httpUri="https://binary.pensoft.net/fig/432231" pageId="0" pageNumber="1">62D, F</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Petiole.</emphasis>
Shiny with fine and dense rugofoveolae; node smooth, low, triangular, with rounded and thin apex, in rear view node dorsoventrally slightly convex; pilosity moderately sparse and erect (Fig.
<figureCitation captionStart="Figure 62" captionStartId="F62" captionText="Figure 62. Pheidole vony Forel, full-face view (A), profile (C), and dorsal view (E) of minor paratype worker (CASENT 0498053) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498055)." figureDoi="10.3897/zookeys.949.51269.figure62" httpUri="https://binary.pensoft.net/fig/432231" pageId="0" pageNumber="1">62D, F</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Postpetiole.</emphasis>
Shiny and smooth; in dorsal view oval, lateral margins medially with two dentate projections; pilosity long, moderately sparse and erect (Fig.
<figureCitation captionStart="Figure 62" captionStartId="F62" captionText="Figure 62. Pheidole vony Forel, full-face view (A), profile (C), and dorsal view (E) of minor paratype worker (CASENT 0498053) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498055)." figureDoi="10.3897/zookeys.949.51269.figure62" httpUri="https://binary.pensoft.net/fig/432231" pageId="0" pageNumber="1">62D, F</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
Shiny and smooth; pilosity moderately dense, long, and erect (Fig.
<figureCitation captionStart="Figure 62" captionStartId="F62" captionText="Figure 62. Pheidole vony Forel, full-face view (A), profile (C), and dorsal view (E) of minor paratype worker (CASENT 0498053) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498055)." figureDoi="10.3897/zookeys.949.51269.figure62" httpUri="https://binary.pensoft.net/fig/432231" pageId="0" pageNumber="1">62D, F</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
Yellow; mandibles and gaster slightly darker (Fig.
<figureCitation captionStart="Figure 62" captionStartId="F62" captionText="Figure 62. Pheidole vony Forel, full-face view (A), profile (C), and dorsal view (E) of minor paratype worker (CASENT 0498053) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498055)." figureDoi="10.3897/zookeys.949.51269.figure62" httpUri="https://binary.pensoft.net/fig/432231" pageId="0" pageNumber="1">62D, F</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
Measurements (
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
= 6): HL: 0.54-0.58 (0.56); HW: 0.45-0.48 (0.47); SL: 0.58-0.61 (0.6); EL: 0.1-0.11 (0.11); WL: 0.69-0.73 (0.71); PSL: 0.07-0.09 (0.07); MTL: 0.48-0.51 (0.49); PNW: 0.31-0.33 (0.32); PTW: 0.07-0.09 (0.08); PPW: 0.12-0.13 (0.13); CI: 117.4-121.1 (118.7); SI: 126.5-131.0 (128.4); PSLI: 12.0-15.0 (13.7); PPI: 58.7-71.1 (63.4); PNI: 65.7-69.0 (67.2); MTI: 101.7-109.2 (104.9).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
Cephalic margin indistinctly convex or straight (Fig.
<figureCitation captionStart="Figure 62" captionStartId="F62" captionText="Figure 62. Pheidole vony Forel, full-face view (A), profile (C), and dorsal view (E) of minor paratype worker (CASENT 0498053) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498055)." figureDoi="10.3897/zookeys.949.51269.figure62" httpUri="https://binary.pensoft.net/fig/432231" pageId="0" pageNumber="1">62A</figureCitation>
). Pilosity relatively sparse, long, decumbent to suberect. Sculpture foveolate; area posterolateral from eyes smooth. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, surpassing the posterior head margin by two-fifths of its length; pilosity dense, subdecumbent to erect (Fig.
<figureCitation captionStart="Figure 62" captionStartId="F62" captionText="Figure 62. Pheidole vony Forel, full-face view (A), profile (C), and dorsal view (E) of minor paratype worker (CASENT 0498053) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498055)." figureDoi="10.3897/zookeys.949.51269.figure62" httpUri="https://binary.pensoft.net/fig/432231" pageId="0" pageNumber="1">62A, C</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
In lateral view, promesonotum moderately low and moderately short, arched; promesonotal groove absent; metanotal groove indistinct; propodeal spines very small and triangular (Fig.
<figureCitation captionStart="Figure 62" captionStartId="F62" captionText="Figure 62. Pheidole vony Forel, full-face view (A), profile (C), and dorsal view (E) of minor paratype worker (CASENT 0498053) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498055)." figureDoi="10.3897/zookeys.949.51269.figure62" httpUri="https://binary.pensoft.net/fig/432231" pageId="0" pageNumber="1">62C</figureCitation>
). Sculpture smooth, sometimes with sparse and indistinct foveolae on lateral sides. Pilosity very sparse, moderately long, and erect (Fig.
<figureCitation captionStart="Figure 62" captionStartId="F62" captionText="Figure 62. Pheidole vony Forel, full-face view (A), profile (C), and dorsal view (E) of minor paratype worker (CASENT 0498053) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498055)." figureDoi="10.3897/zookeys.949.51269.figure62" httpUri="https://binary.pensoft.net/fig/432231" pageId="0" pageNumber="1">62C, E</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Postpetiole.</emphasis>
Short, low, and relatively flat; with few short, erect setae (Fig.
<figureCitation captionStart="Figure 62" captionStartId="F62" captionText="Figure 62. Pheidole vony Forel, full-face view (A), profile (C), and dorsal view (E) of minor paratype worker (CASENT 0498053) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498055)." figureDoi="10.3897/zookeys.949.51269.figure62" httpUri="https://binary.pensoft.net/fig/432231" pageId="0" pageNumber="1">62C, E</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
With sparse, erect pilosity (Fig.
<figureCitation captionStart="Figure 62" captionStartId="F62" captionText="Figure 62. Pheidole vony Forel, full-face view (A), profile (C), and dorsal view (E) of minor paratype worker (CASENT 0498053) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498055)." figureDoi="10.3897/zookeys.949.51269.figure62" httpUri="https://binary.pensoft.net/fig/432231" pageId="0" pageNumber="1">62C, E</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
Yellow, vertex slightly darker (Fig.
<figureCitation captionStart="Figure 62" captionStartId="F62" captionText="Figure 62. Pheidole vony Forel, full-face view (A), profile (C), and dorsal view (E) of minor paratype worker (CASENT 0498053) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498055)." figureDoi="10.3897/zookeys.949.51269.figure62" httpUri="https://binary.pensoft.net/fig/432231" pageId="0" pageNumber="1">62C, E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">Malagasy for yellow in reference to the body colouration.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
<paragraph pageId="0" pageNumber="1">The species was collected between 600-1100 m in elevation, in rainforest and montane rainforest. Nests were located in rotten logs, root mats, rotten tree stumps, and rotten sticks on the ground.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="comments">
<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole vony" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="vony">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole vony</emphasis>
</taxonomicName>
sp. nov. is a member of a group of species characterised by body colouration bright yellow to orange in majors and yellow in minors, head sub-oval, not widening posteriorly with sides not convex or convex indistinctly, and medial part of frons with longitudinal and interrupted rugae on the anterior part and distinctly irregular rugae on the posterior one. The group includes three taxa:
<taxonomicName lsidName="P. vony" pageId="0" pageNumber="1" rank="species" species="vony">
<emphasis italics="true" pageId="0" pageNumber="1">P. vony</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName lsidName="P. befotaka" pageId="0" pageNumber="1" rank="species" species="befotaka">
<emphasis italics="true" pageId="0" pageNumber="1">P. befotaka</emphasis>
</taxonomicName>
sp. nov., and
<taxonomicName lsidName="P. mamiratra" pageId="0" pageNumber="1" rank="species" species="mamiratra">
<emphasis italics="true" pageId="0" pageNumber="1">P. mamiratra</emphasis>
</taxonomicName>
sp. nov.
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole vony" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="vony">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole vony</emphasis>
</taxonomicName>
sp. nov. is known from two localities in Toamasina: Montagne
<normalizedToken originalValue="dAnjanaharibe">d'Anjanaharibe</normalizedToken>
and Montagne
<normalizedToken originalValue="dAkirindro">d'Akirindro</normalizedToken>
, and its distribution
<normalizedToken originalValue="doesnt">doesn't</normalizedToken>
overlap with two remaining members of the group. Morphologically
<taxonomicName lsidName="P. vony" pageId="0" pageNumber="1" rank="species" species="vony">
<emphasis italics="true" pageId="0" pageNumber="1">P. vony</emphasis>
</taxonomicName>
sp. nov. is most similar to
<taxonomicName lsidName="P. mamiratra" pageId="0" pageNumber="1" rank="species" species="mamiratra">
<emphasis italics="true" pageId="0" pageNumber="1">P. mamiratra</emphasis>
</taxonomicName>
sp. nov., known only from Station
<normalizedToken originalValue="Forestière">Forestiere</normalizedToken>
Angavokely in Antananarivo. Major workers of
<taxonomicName lsidName="P. vony" pageId="0" pageNumber="1" rank="species" species="vony">
<emphasis italics="true" pageId="0" pageNumber="1">P. vony</emphasis>
</taxonomicName>
sp. nov. differ from
<taxonomicName lsidName="P. mamiratra" pageId="0" pageNumber="1" rank="species" species="mamiratra">
<emphasis italics="true" pageId="0" pageNumber="1">P. mamiratra</emphasis>
</taxonomicName>
sp. nov. in medial part of frons with moderately dense and thin rugae and inner hypostomal teeth large and triangular, with rounded apex directed outward and outer hypostomal teeth lobe-like, lower, and more narrow than inner hypostomal teeth; minors differ in more distinctly foveolate head sculpture. However, minor and major workers of
<taxonomicName lsidName="P. vony" pageId="0" pageNumber="1" rank="species" species="vony">
<emphasis italics="true" pageId="0" pageNumber="1">P. vony</emphasis>
</taxonomicName>
sp. nov. can be also confused with
<taxonomicName lsidName="P. sparsa" pageId="0" pageNumber="1" rank="species" species="sparsa">
<emphasis italics="true" pageId="0" pageNumber="1">P. sparsa</emphasis>
</taxonomicName>
sp. nov., known from Bemanevika in Mahajanga, and
<taxonomicName lsidName="P. hazo" pageId="0" pageNumber="1" rank="species" species="hazo">
<emphasis italics="true" pageId="0" pageNumber="1">P. hazo</emphasis>
</taxonomicName>
sp. nov., described from the vicinity of Antananarivo. Majors of
<taxonomicName lsidName="P. vony" pageId="0" pageNumber="1" rank="species" species="vony">
<emphasis italics="true" pageId="0" pageNumber="1">P. vony</emphasis>
</taxonomicName>
sp. nov. differ from
<taxonomicName lsidName="P. sparsa" pageId="0" pageNumber="1" rank="species" species="sparsa">
<emphasis italics="true" pageId="0" pageNumber="1">P. sparsa</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName lsidName="P. hazo" pageId="0" pageNumber="1" rank="species" species="hazo">
<emphasis italics="true" pageId="0" pageNumber="1">P. hazo</emphasis>
</taxonomicName>
sp. nov. in median frons with thinner and irregular rugae and more irregular sculpture on lateral sides of frons; minor workers differ in smaller propodeal spines, lack of promesonotal groove, strongly reduced sculpture of mesosoma, and head lacking additional, indistinct rugulae.
</paragraph>
<caption doi="10.3897/zookeys.949.51269.figure63" httpUri="https://binary.pensoft.net/fig/432232" pageId="0" pageNumber="1" start="Figure 63" startId="F63">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 63.</emphasis>
Major worker, hypostomal teeth.
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole alina" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="alina">Pheidole alina</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole ambohimanga" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ambohimanga">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole ambohimanga</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole analavelona" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="analavelona">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole analavelona</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole andohahela" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="andohahela">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole andohahela</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole anomala" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="anomala">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole anomala</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole anosyenne" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="anosyenne">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole anosyenne</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole antranohofa" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="antranohofa">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole antranohofa</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole beanka" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="beanka">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole beanka</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole befotaka" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="befotaka">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole befotaka</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">I</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole dasos" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dasos">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole dasos</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">J</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole flavominuta" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="flavominuta">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole flavominuta</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">K</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole gracilis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole gracilis</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">L</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole haboka" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="haboka">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole haboka</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">M</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole havoana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="havoana">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole havoana</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">N</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole hazo" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hazo">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole hazo</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">O</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole itremo" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="itremo">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole itremo</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">P</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole joffreville" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="joffreville">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole joffreville</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">Q</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole kely" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kely">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole kely</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">R</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole lavasoa" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lavasoa">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole lavasoa</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">S</emphasis>
).
<taxonomicName authorityName="Forel" authorityYear="1892" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole litigiosa" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="litigiosa">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole litigiosa</emphasis>
</taxonomicName>
Forel (
<emphasis bold="true" pageId="0" pageNumber="1">T</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole mahamavo" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mahamavo">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole mahamavo</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">U</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole mainty" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mainty">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole mainty</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">V</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole mamiratra" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mamiratra">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole mamiratra</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">W</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole manantenina" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="manantenina">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole manantenina</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">X</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.949.51269.figure64" httpUri="https://binary.pensoft.net/fig/432233" pageId="0" pageNumber="1" start="Figure 64" startId="F64">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 64.</emphasis>
Major worker, hypostomal teeth.
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole masoandro" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="masoandro">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole masoandro</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole mavohavoana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mavohavoana">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole mavohavoana</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole midongy" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="midongy">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole midongy</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole mikros" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mikros">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole mikros</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole mivory" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mivory">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole mivory</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole nitidobruna" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nitidobruna">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole nitidobruna</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole parvula" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="parvula">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole parvula</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole parvulogibba" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="parvulogibba">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole parvulogibba</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole renirano" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="renirano">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole renirano</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">I</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole sava" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sava">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole sava</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">J</emphasis>
).
<taxonomicName authorityName="Forel" authorityYear="1891" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole sikorae" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sikorae">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole sikorae</emphasis>
</taxonomicName>
Forel (
<emphasis bold="true" pageId="0" pageNumber="1">K</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole sofia" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sofia">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole sofia</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">L</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole sparsa" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sparsa">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole sparsa</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">M</emphasis>
).
<taxonomicName class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole</emphasis>
</taxonomicName>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="tampony" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tampony">
<emphasis italics="true" pageId="0" pageNumber="1">tampony</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">N</emphasis>
).
<taxonomicName class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole</emphasis>
</taxonomicName>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="trichotos" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="trichotos">
<emphasis italics="true" pageId="0" pageNumber="1">trichotos</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">O</emphasis>
).
<taxonomicName class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole</emphasis>
</taxonomicName>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="tsaravoniana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tsaravoniana">
<emphasis italics="true" pageId="0" pageNumber="1">tsaravoniana</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">P</emphasis>
).
<taxonomicName class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole</emphasis>
</taxonomicName>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="vadum" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="vadum">
<emphasis italics="true" pageId="0" pageNumber="1">vadum</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">Q</emphasis>
).
<taxonomicName class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole</emphasis>
</taxonomicName>
<taxonomicName authorityName="Forel" authorityYear="1891" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="veteratrix" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="veteratrix">
<emphasis italics="true" pageId="0" pageNumber="1">veteratrix</emphasis>
</taxonomicName>
Forel (
<emphasis bold="true" pageId="0" pageNumber="1">R</emphasis>
).
<taxonomicName class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole</emphasis>
</taxonomicName>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="volontany" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="volontany">
<emphasis italics="true" pageId="0" pageNumber="1">volontany</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">S</emphasis>
).
<taxonomicName class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole</emphasis>
</taxonomicName>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="vony" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="vony">
<emphasis italics="true" pageId="0" pageNumber="1">vony</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">T</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.949.51269.figure65" httpUri="https://binary.pensoft.net/fig/432234" pageId="0" pageNumber="1" start="Figure 65" startId="F65">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 65.</emphasis>
Distribution.
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole alina" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="alina">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole alina</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole ambohimanga" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ambohimanga">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole ambohimanga</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole analavelona" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="analavelona">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole analavelona</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole andohahela" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="andohahela">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole andohahela</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole anomala" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="anomala">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole anomala</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole anosyenne" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="anosyenne">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole anosyenne</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole antranohofa" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="antranohofa">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole antranohofa</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole beanka" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="beanka">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole beanka</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole befotaka" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="befotaka">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole befotaka</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">I</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole dasos" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dasos">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole dasos</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">J</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole flavominuta" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="flavominuta">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole flavominuta</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">K</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole gracilis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole gracilis</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">L</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole haboka" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="haboka">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole haboka</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">M</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole havoana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="havoana">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole havoana</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">N</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole hazo" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hazo">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole hazo</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">O</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole itremo" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="itremo">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole itremo</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">P</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole joffreville" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="joffreville">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole joffreville</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">Q</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole kely" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kely">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole kely</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">R</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole lavasoa" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lavasoa">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole lavasoa</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">S</emphasis>
).
<taxonomicName authorityName="Forel" authorityYear="1892" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole litigiosa" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="litigiosa">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole litigiosa</emphasis>
</taxonomicName>
Forel (
<emphasis bold="true" pageId="0" pageNumber="1">T</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.949.51269.figure66" httpUri="https://binary.pensoft.net/fig/432235" pageId="0" pageNumber="1" start="Figure 66" startId="F66">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 66.</emphasis>
Distribution.
<taxonomicName class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole</emphasis>
</taxonomicName>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="mahamavo" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mahamavo">
<emphasis italics="true" pageId="0" pageNumber="1">mahamavo</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
).
<taxonomicName class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole</emphasis>
</taxonomicName>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="mainty" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mainty">
<emphasis italics="true" pageId="0" pageNumber="1">mainty</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
).
<taxonomicName class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole</emphasis>
</taxonomicName>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="mamiratra" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mamiratra">
<emphasis italics="true" pageId="0" pageNumber="1">mamiratra</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole manantenina" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="manantenina">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole manantenina</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
).
<taxonomicName class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole</emphasis>
</taxonomicName>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="masoandro" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="masoandro">
<emphasis italics="true" pageId="0" pageNumber="1">masoandro</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole mavohavoana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mavohavoana">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole mavohavoana</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
).
<taxonomicName class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole</emphasis>
</taxonomicName>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="midongy" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="midongy">
<emphasis italics="true" pageId="0" pageNumber="1">midongy</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole mikros" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mikros">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole mikros</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
).
<taxonomicName class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole</emphasis>
</taxonomicName>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="mivory" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mivory">
<emphasis italics="true" pageId="0" pageNumber="1">mivory</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">I</emphasis>
).
<taxonomicName class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole</emphasis>
</taxonomicName>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="nitidobruna" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nitidobruna">
<emphasis italics="true" pageId="0" pageNumber="1">nitidobruna</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">J</emphasis>
).
<taxonomicName class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole</emphasis>
</taxonomicName>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="parvula" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="parvula">
<emphasis italics="true" pageId="0" pageNumber="1">parvula</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">K</emphasis>
).
<taxonomicName class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole</emphasis>
</taxonomicName>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="parvulogibba" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="parvulogibba">
<emphasis italics="true" pageId="0" pageNumber="1">parvulogibba</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">L</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.949.51269.figure67" httpUri="https://binary.pensoft.net/fig/432236" pageId="0" pageNumber="1" start="Figure 67" startId="F67">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 67.</emphasis>
Distribution.
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole renirano" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="renirano">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole renirano</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole sava" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sava">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole sava</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
).
<taxonomicName authorityName="Forel" authorityYear="1891" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole sikorae" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sikorae">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole sikorae</emphasis>
</taxonomicName>
Forel (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole sofia" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sofia">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole sofia</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole sparsa" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sparsa">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole sparsa</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole tampony" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tampony">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole tampony</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole trichotos" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="trichotos">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole trichotos</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole tsaravoniana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tsaravoniana">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole tsaravoniana</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole vadum" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="vadum">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole vadum</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">I</emphasis>
).
<taxonomicName authorityName="Forel" authorityYear="1891" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole veteratrix" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="veteratrix">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole veteratrix</emphasis>
</taxonomicName>
Forel (
<emphasis bold="true" pageId="0" pageNumber="1">J</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole volontany" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="volontany">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole volontany</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">K</emphasis>
).
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole vony" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="vony">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole vony</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="1">L</emphasis>
).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>