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<mods:title id="575E36799B7AD291C9346D7F2325C970">Not so rare snakes: a revision of the Geophis sieboldi group (Colubridae: Dipsadinae) in lower Central America and Colombia</mods:title>
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<paragraph id="8BD236F23548C16FFC5EF95DFA50F977" blockId="35.[888,1379,1739,1827]" box="[898,1369,1739,1762]" pageId="35" pageNumber="596">
<heading id="D09A819E3548C16FFC5EF95DFA50F977" box="[898,1369,1739,1762]" centered="true" fontSize="9" level="2" pageId="35" pageNumber="596" reason="2">
PROVENANCE OF 
<taxonomicName id="4C6D4D713548C16FFB84F95DFA50F977" authority="AND OTHER" box="[1112,1369,1739,1762]" class="Reptilia" family="Dipsadidae" genus="Geophis" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="596" phylum="Chordata" rank="genus">
<emphasis id="B919EAE03548C16FFB84F95DFBCAF974" box="[1112,1219,1739,1762]" italics="true" pageId="35" pageNumber="596">GEOPHIS</emphasis>
AND OTHER
</taxonomicName>
</heading>
</paragraph>
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<paragraph id="8BD236F23548C16FFCA4F978FBE8F8B5" blockId="35.[888,1379,1739,1827]" pageId="35" pageNumber="596">
<heading id="D09A819E3548C16FFCA4F978FBE8F8B5" centered="true" fontSize="9" level="2" pageId="35" pageNumber="596" reason="5">
SNAKES INCORRECTLY CATALOGED FROM ‘ 
<collectingCountry id="F37A76623548C16FFC23F89DFB6CF8B7" box="[1023,1125,1803,1827]" name="Panama" pageId="35" pageNumber="596">PANAMA</collectingCountry>
SABANAS’
</heading>
</paragraph>
<paragraph id="8BD236F23548C168FCE6F8ADFD9CFDBA" blockId="35.[826,1441,1850,1903]" lastBlockId="36.[144,759,198,1903]" lastPageId="36" lastPageNumber="597" pageId="35" pageNumber="596">
Myers (2003) briefly reviewed the listing in the catalogue of the ANSP for 
<taxonomicName id="4C6D4D713548C16FFBE5F8CFFA42F8F8" authorityName=", Dunn" authorityYear="1942" box="[1081,1355,1881,1902]" class="Reptilia" family="Dipsadidae" genus="Geophis" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="596" phylum="Chordata" rank="species" species="brachycephalus">
<emphasis id="B919EAE03548C16FFBE5F8CFFA42F8F8" box="[1081,1355,1881,1902]" italics="true" pageId="35" pageNumber="596">Geophis brachycephalus</emphasis>
</taxonomicName>
(ANSP 24723–24734), 
<taxonomicName id="4C6D4D71354FC168FE92FF50FD2CFF4D" authorityName="Boulenger" authorityYear="1894" box="[334,549,198,219]" class="Reptilia" family="Dipsadidae" genus="Geophis" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="597" phylum="Chordata" rank="species" species="godmani">
<emphasis id="B919EAE0354FC168FE92FF50FD2CFF4D" box="[334,549,198,219]" italics="true" pageId="36" pageNumber="597">Geophis godmani</emphasis>
</taxonomicName>
(ANSP 24722), and 
<emphasis id="B919EAE0354FC168FF16FF73FEACFF6F" box="[202,421,228,250]" italics="true" pageId="36" pageNumber="597">Trimetopon slevini</emphasis>
(ANSP 24717–24719) from ‘ 
<collectingCountry id="F37A7662354FC168FF48FE95FFFAFE8E" box="[148,243,259,280]" name="Panama" pageId="36" pageNumber="597">Panama</collectingCountry>
Sabanas’. As the three species are all upland forms that co-occur in extreme western 
<collectingCountry id="F37A7662354FC168FDB8FEB4FDCBFEA1" box="[612,706,290,311]" name="Panama" pageId="36" pageNumber="597">Panama</collectingCountry>
and all other ‘Sabanas’ material is from lowland central 
<collectingCountry id="F37A7662354FC168FF4CFEC9FFF8FEE2" box="[144,241,351,372]" name="Panama" pageId="36" pageNumber="597">Panama</collectingCountry>
, Myers concluded that their association with that ‘Sabanas’ locality must have been a cataloguing error and that the Academy specimens were part of the 
<collectingCountry id="F37A7662354FC168FF4CFE2DFFE7FE46" box="[144,238,443,464]" name="Panama" pageId="36" pageNumber="597">Panama</collectingCountry>
Snake Census (PSC) collection from 
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: 
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: Finca Lérida (above Alto Quiel). Here we provide further details from Dunn’s notes regarding the provenance of the specimens in question.
</paragraph>
<paragraph id="8BD236F2354FC168FF74FDA3FED5FCC8" blockId="36.[144,759,198,1903]" pageId="36" pageNumber="597">
Both the ‘ 
<collectingCountry id="F37A7662354FC168FEC3FDA3FE77FDDC" box="[287,382,565,586]" name="Panama" pageId="36" pageNumber="597">Panama</collectingCountry>
Sabanas’ and the Finca Lérida series are part of Herbert C. Clark’s PSC collections. Clark, then Director of the Gorgas Memorial Laboratory, in 
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, instituted a survey of 
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snakes beginning in 1929 and continuing through 1953 (see Dunn, 1947, 1949; Myers, 2003). The intent was to assess the relative medical significance of venomous snakes. Bounties were paid for specimens, usually heads, and specimens accumulated at various centres throughout 
<collectingCountry id="F37A7662354FC168FEAAFCDFFED1FCC8" box="[374,472,841,862]" name="Panama" pageId="36" pageNumber="597">Panama</collectingCountry>
.
</paragraph>
<paragraph id="8BD236F2354FC168FF74FCF1FD8DFA54" blockId="36.[144,759,198,1903]" pageId="36" pageNumber="597">
In 1949, Emmett Reid Dunn published his classic paper on the distribution and abundance of 
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snakes based primarily on the PSC collections (10 
<specimenCount id="9D6BFD7B354FC168FD11FC32FE0FFC4F" pageId="36" pageNumber="597" type="generic">693 specimens</specimenCount>
) and 1073 snakes from natural history museums. The PSC specimens analysed in that report were from west-central and eastern 
<collectingCountry id="F37A7662354FC168FD80FB96FDB3FB83" box="[604,698,1024,1045]" name="Panama" pageId="36" pageNumber="597">Panama</collectingCountry>
and grouped by general locality for this purpose as: Coclé- 
<collectingRegion id="49A9F810354FC168FF4CFBA8FFE7FBC5" box="[144,238,1086,1107]" country="Panama" name="Herrera" pageId="36" pageNumber="597">Herrera</collectingRegion>
, Sabanas, Chagres, and 
<collectingRegion id="49A9F810354FC168FDDAFBABFD51FBC5" box="[518,600,1085,1107]" country="Panama" name="Darien" pageId="36" pageNumber="597">Darién</collectingRegion>
. The Sabanas material (3914 snakes) was from a variety of Pacific lowland localities, both east and west of the 
<collectingCountry id="F37A7662354FC168FD44FBEDFDFFFB06" box="[664,758,1147,1168]" name="Panama" pageId="36" pageNumber="597">Panama</collectingCountry>
Canal, between 
<collectingCountry id="F37A7662354FC168FEB6FB0FFEC5FB38" box="[362,460,1177,1198]" name="Panama" pageId="36" pageNumber="597">Panama</collectingCountry>
: 
<collectingCountry id="F37A7662354FC168FE34FB0FFD43FB38" box="[488,586,1177,1198]" name="Panama" pageId="36" pageNumber="597">Panama</collectingCountry>
: Cañita and Capria, respectively. Many of these have specific locality data but others are catalogued from more inclusive sites (e.g. Chorrera to Arraiijan) or simply 
<collectingCountry id="F37A7662354FC168FF4CFA82FFE7FABF" box="[144,238,1300,1321]" name="Panama" pageId="36" pageNumber="597">Panama</collectingCountry>
Sabanas (usually Sabanas in Dunn’s notes). These localities all lie in an area having a long dry season (mid-December to April) that originally supported scrub, shrub or savanna vegetation but has been anthropogenically modified to an extreme beginning in preconquest times (Bennett, 1968).
</paragraph>
<paragraph id="8BD236F2354FC168FF74FA5DFED6F8F8" blockId="36.[144,759,198,1903]" pageId="36" pageNumber="597">
The Lérida PSC collection was not included in Dunn’s (1949) analysis but all Sabanas snakes collected prior to 1946 were. Significantly, 
<taxonomicName id="4C6D4D71354FC168FD40F99EFE49F9AA" authorityName=", Dunn" authorityYear="1942" class="Reptilia" family="Dipsadidae" genus="Geophis" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="597" phylum="Chordata" rank="species" species="brachycephalus">
<emphasis id="B919EAE0354FC168FD40F99EFE49F9AA" italics="true" pageId="36" pageNumber="597">Geophis brachycephalus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C6D4D71354FC168FE84F9B1FD21F9AA" authorityName="Boulenger" authorityYear="1894" box="[344,552,1575,1596]" class="Reptilia" family="Dipsadidae" genus="Geophis" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="597" phylum="Chordata" rank="species" species="godmani">
<emphasis id="B919EAE0354FC168FE84F9B1FD21F9AA" box="[344,552,1575,1596]" italics="true" pageId="36" pageNumber="597">Geophis godmani</emphasis>
</taxonomicName>
and 
<emphasis id="B919EAE0354FC168FDA9F9B1FDFFF9AA" box="[629,758,1575,1596]" italics="true" pageId="36" pageNumber="597">Trimetopon</emphasis>
are not referred to anywhere in the 1949 paper. However, Dunn (1947) had published an often overlooked report prescient of the 1949 paper on the snakes of Lérida some 2 years earlier based on PSC material (
<specimenCount id="9D6BFD7B354FC168FECCF956FECEF940" box="[272,455,1728,1750]" pageId="36" pageNumber="597" type="generic">533 specimens</specimenCount>
) and seven others in museum collections. This series included 255 
<taxonomicName id="4C6D4D71354FC168FD40F948FE49F884" authorityName=", Dunn" authorityYear="1942" class="Reptilia" family="Dipsadidae" genus="Geophis" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="597" phylum="Chordata" rank="species" species="brachycephalus">
<emphasis id="B919EAE0354FC168FD40F948FE49F884" italics="true" pageId="36" pageNumber="597">Geophis brachycephalus</emphasis>
</taxonomicName>
, three 
<taxonomicName id="4C6D4D71354FC168FE46F96BFD6DF884" authorityName="Boulenger" authorityYear="1894" box="[410,612,1789,1810]" class="Reptilia" family="Dipsadidae" genus="Geophis" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="597" phylum="Chordata" rank="species" species="godmani">
<emphasis id="B919EAE0354FC168FE46F96BFD6DF884" box="[410,612,1789,1810]" italics="true" pageId="36" pageNumber="597">Geophis godmani</emphasis>
</taxonomicName>
and 26 
<emphasis id="B919EAE0354FC168FD10F968FE4DF8A7" italics="true" pageId="36" pageNumber="597">Trimetopon slevini</emphasis>
. The PSC material from Lerida had been accumulated by Tolef B. Mönniche, the owner of the finca, beginning in 1940.
</paragraph>
<paragraph id="8BD236F2354FC168FCE3FF50FC9CFE05" blockId="36.[806,1421,198,403]" pageId="36" pageNumber="597">
A review of Dunn’s notes accumulated over many years, at least beginning in 1928, and organized as a preliminary synopsis of the ophiofauna of lower Central America, clarifies the source of the Sabanas specimens of questionable locality data. Part of Dunn’s entries for 
<taxonomicName id="4C6D4D71354FC168FBE6FEC9FA02FEE2" authorityName=", Dunn" authorityYear="1942" box="[1082,1291,351,372]" class="Reptilia" family="Dipsadidae" genus="Geophis" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="597" phylum="Chordata" rank="species" species="brachycephalus">
<emphasis id="B919EAE0354FC168FBE6FEC9FA02FEE2" box="[1082,1291,351,372]" italics="true" pageId="36" pageNumber="597">G. brachycephalus</emphasis>
</taxonomicName>
list the following:
</paragraph>
<paragraph id="8BD236F2354FC168FCFAFE2BFB31FE46" blockId="36.[806,1421,444,581]" box="[806,1080,444,465]" pageId="36" pageNumber="597">‘Lerida cl 1940 17 + ERD 1</paragraph>
<paragraph id="8BD236F2354FC168FCFAFE4DFC7FFE78" blockId="36.[806,1421,444,581]" box="[806,886,475,494]" pageId="36" pageNumber="597">1941 17</paragraph>
<paragraph id="8BD236F2354FC168FCFAFE6EFC7FFD9D" blockId="36.[806,1421,444,581]" box="[806,886,504,523]" pageId="36" pageNumber="597">1942 35</paragraph>
<paragraph id="8BD236F2354FC168FCFAFD83FC73FDD3" blockId="36.[806,1421,444,581]" pageId="36" pageNumber="597">1943 99 L tot 1947 = 255 1944 45’</paragraph>
<paragraph id="8BD236F2354FC168FCE3FDF9FB85FBD9" blockId="36.[806,1422,622,1501]" pageId="36" pageNumber="597">
In this notation, as throughout Dunn’s notes, cl (= Clark) is used for snakes from the PSC. Year and number of specimens are indicated through 1945. The total of examples from Lérida (L.) through 1947 is also noted and is the same total as listed in Dunn’s 1947 paper. No 
<taxonomicName id="4C6D4D71354FC168FC35FC9EFBB3FC8A" authorityName=", Dunn" authorityYear="1942" box="[1001,1210,775,797]" class="Reptilia" family="Dipsadidae" genus="Geophis" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="597" phylum="Chordata" rank="species" species="brachycephalus">
<emphasis id="B919EAE0354FC168FC35FC9EFBB3FC8A" box="[1001,1210,775,797]" italics="true" pageId="36" pageNumber="597">G. brachycephalus</emphasis>
</taxonomicName>
from Sabanas or 
<collectingCountry id="F37A7662354FC168FCFAFCB0FC8DFCAD" box="[806,900,806,827]" name="Panama" pageId="36" pageNumber="597">Panama</collectingCountry>
Sabanas are listed among the other 69 cited from 
<collectingCountry id="F37A7662354FC168FCB5FCD3FCC2FCCC" box="[873,971,837,858]" name="Panama" pageId="36" pageNumber="597">Panama</collectingCountry>
. The number is actually 65 as three snakes from 
<collectingCountry id="F37A7662354FC168FC66FCF5FB15FCEE" box="[954,1052,867,888]" name="Panama" pageId="36" pageNumber="597">Panama</collectingCountry>
: 
<collectingRegion id="49A9F810354FC168FBF4FCF5FB82FCEF" box="[1064,1163,867,889]" country="Panama" name="Chiriqui" pageId="36" pageNumber="597">Chiriquí</collectingRegion>
: Boquete, in an extensive series (63 snakes) collected by Joseph R. Slevin (1942) (sl in Dunn’s notes) represent other species of 
<taxonomicName id="4C6D4D71354FC168FCFAFC29FB3FFC43" authority="(Downs, 1967)" baseAuthorityName="Downs" baseAuthorityYear="1967" box="[806,1078,959,981]" class="Reptilia" family="Dipsadidae" genus="Geophis" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="597" phylum="Chordata" rank="genus">
<emphasis id="B919EAE0354FC168FCFAFC29FC88FC42" box="[806,897,959,980]" italics="true" pageId="36" pageNumber="597">Geophis</emphasis>
(Downs, 1967)
</taxonomicName>
and include the 
<typeStatus id="54D68850354FC168FAD7FC29FA65FC43" box="[1291,1388,959,981]" pageId="36" pageNumber="597" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C6D4D71354FC168FCFAFC48FCB6FC64" authorityName="Boulenger" authorityYear="1894" box="[806,959,989,1011]" class="Reptilia" family="Dipsadidae" genus="Geophis" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="597" phylum="Chordata" rank="species" species="championi">
<emphasis id="B919EAE0354FC168FCFAFC48FCB6FC64" box="[806,959,989,1011]" italics="true" pageId="36" pageNumber="597">G. championi</emphasis>
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(BMNH 1946.1.1.77). It seems clear from the above evidence and confirmed below that ANSP 2423–2434 catalogued as being from ‘ 
<collectingCountry id="F37A7662354FC168FAF2FB8DFA84FBA6" box="[1326,1421,1051,1072]" name="Panama" pageId="36" pageNumber="597">Panama</collectingCountry>
Sabanas’ are also from Lérida.
</paragraph>
<paragraph id="8BD236F2354FC168FCE3FBCEFA00FA17" blockId="36.[806,1422,622,1501]" pageId="36" pageNumber="597">
Recall that most snakes from the PSC are heads or heads with a short segment of the body and after identification, unfortunately, Dunn did not keep and/or deposit all of them in museums (see Myers, 2003). He did, however, retain rarer forms and smaller complete specimens. The ANSP contains the following 
<taxonomicName id="4C6D4D71354FC168FC45FA86FB63FAB2" authorityName=", Dunn" authorityYear="1942" box="[921,1130,1295,1317]" class="Reptilia" family="Dipsadidae" genus="Geophis" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="597" phylum="Chordata" rank="species" species="brachycephalus">
<emphasis id="B919EAE0354FC168FC45FA86FB63FAB2" box="[921,1130,1295,1317]" italics="true" pageId="36" pageNumber="597">G. brachycephalus</emphasis>
</taxonomicName>
catalogued from Lérida: ANSP 21699, a specimen collected by Dunn in 1939, and the following PSC specimens: 22922–22938, ANSP 23877–23879, ANSP 24766–24769.
</paragraph>
<paragraph id="8BD236F2354FC168FCE3FA1CFC74FA4B" blockId="36.[806,1422,622,1501]" pageId="36" pageNumber="597">
The situation for 
<taxonomicName id="4C6D4D71354FC168FBC4FA1CFBEDFA09" authorityName="Boulenger" authorityYear="1894" box="[1048,1252,1418,1439]" class="Reptilia" family="Dipsadidae" genus="Geophis" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="597" phylum="Chordata" rank="species" species="godmani">
<emphasis id="B919EAE0354FC168FBC4FA1CFBEDFA09" box="[1048,1252,1418,1439]" italics="true" pageId="36" pageNumber="597">Geophis godmani</emphasis>
</taxonomicName>
is even more clear-cut. The pertinent material in Dunn’s notes is as follows:
</paragraph>
<paragraph id="8BD236F2354FC168FCFAF991FBAFF98C" blockId="36.[806,1190,1543,1562]" box="[806,1190,1543,1562]" pageId="36" pageNumber="597">‘Lerida cl 1940 1h, 1941 1h, 1943 1h.’</paragraph>
<paragraph id="8BD236F2354FC168FCE3F9D5FBA0F8FA" blockId="36.[806,1421,1603,1900]" pageId="36" pageNumber="597">
He makes no mention of other specimens from elsewhere in 
<collectingCountry id="F37A7662354FC168FC0CF9F4FB3BF9E1" box="[976,1074,1634,1655]" name="Panama" pageId="36" pageNumber="597">Panama</collectingCountry>
. The ANSP collections contained only three heads (h in Dunn’s notes) all from the PSC, two from Lérida: ANSP 22910, ANSP 24765, and the putative Sabanas example (ANSP 24722), now lost. Obviously ANSP 24722 is one of the three heads recorded from Lérida by Dunn. Note that typographical errors list ANSP ‘24756’ for one Lérida snake (Downs, 1967) and ANSP ‘24723’ for the ‘Sabanas’ example (Myers, 2003).
</paragraph>
<paragraph id="8BD236F2354EC169FF60FF50FFB7FF6C" blockId="37.[164,778,198,250]" pageId="37" pageNumber="598">
The pertinent Dunn notation for 
<emphasis id="B919EAE0354EC169FDE4FF50FC03FF4D" box="[568,778,198,219]" italics="true" pageId="37" pageNumber="598">Trimetopon slevini</emphasis>
is:
</paragraph>
<paragraph id="8BD236F2354EC169FF78FEB2FD29FEA1" blockId="37.[164,544,290,312]" box="[164,544,290,312]" pageId="37" pageNumber="598">
‘Lerida ANS (2) cl. 
<subScript id="17E934B7354EC169FEB4FEB2FE96FEA1" attach="right" box="[360,415,292,311]" fontSize="5" pageId="37" pageNumber="598">19411</subScript>
- 
<subScript id="17E934B7354EC169FE77FEB2FEE8FEAE" attach="left" box="[427,481,292,312]" fontSize="5" pageId="37" pageNumber="598">19427</subScript>
<subScript id="17E934B7354EC169FE36FEB2FD29FEA1" attach="none" box="[490,544,292,311]" fontSize="5" pageId="37" pageNumber="598">19439</subScript>
</paragraph>
<paragraph id="8BD236F2354EC169FF60FEC9FE01FCB7" blockId="37.[164,779,351,955]" pageId="37" pageNumber="598">
<materialsCitation id="3B053CAF354EC169FF60FEC9FE0CFCB7" accessionNumber="CAS 78938" collectionCode="PSC, ANSP, T, CAS" country="Panama" county="Sabanas'" location="Clearly" municipality="Academy" pageId="37" pageNumber="598" specimenCode="ANSP 21700-21701, ANSP 22939-22940, ANSP 23875, ANSP 24770-24772, ANSP 24717-24719" specimenCount="2" stateProvince="Chiriqui" typeStatus="holotype">
The notation 
<subScript id="17E934B7354EC169FE8AFEF6FE9CFEE0" attach="left" box="[342,405,352,374]" fontSize="5" pageId="37" pageNumber="598">19439</subScript>
apparently refers to the number of specimens collected by the 
<collectionCode id="ED7CAE37354EC169FE24FEE8FD23FE05" box="[504,554,382,403]" pageId="37" pageNumber="598">PSC</collectionCode>
from 1940 to 1943. ANS (2) apparently refers to the 
<specimenCount id="9D6BFD7B354EC169FD82FE0BFC02FE24" box="[606,779,413,434]" pageId="37" pageNumber="598" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
(
<specimenCode id="DBCB9E89354EC169FF77FE2DFE98FE47" box="[171,401,443,465]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="37" pageNumber="598">ANSP 21700–21701</specimenCode>
) collected in 1939 prior to when the 
<collectionCode id="ED7CAE37354EC169FF0CFE4CFE0BFE79" box="[208,258,474,495]" pageId="37" pageNumber="598">PSC</collectionCode>
began obtaining material from the finca or to 
<specimenCode id="DBCB9E89354EC169FF78FE6FFE86FD98" box="[164,399,505,526]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="37" pageNumber="598">ANSP 22939–22940</specimenCode>
. The 
<collectionCode id="ED7CAE37354EC169FE0AFE6FFD15FD98" box="[470,540,505,526]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="37" pageNumber="598">ANSP</collectionCode>
catalogue lists four other 
<collectionCode id="ED7CAE37354EC169FF2CFD81FE2BFDBA" box="[240,290,535,556]" pageId="37" pageNumber="598">PSC</collectionCode>
Lérida specimens: 
<specimenCode id="DBCB9E89354EC169FDCAFD81FDBBFDBB" box="[534,690,535,557]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="37" pageNumber="598">ANSP 23875</specimenCode>
, 
<specimenCode id="DBCB9E89354EC169FD19FD81FE33FDDD" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="37" pageNumber="598">ANSP 24770–24772</specimenCode>
and 
<specimenCount id="9D6BFD7B354EC169FEA8FDA0FD25FDDA" box="[372,556,566,588]" pageId="37" pageNumber="598" type="generic">three specimens</specimenCount>
of 
<emphasis id="B919EAE0354EC169FD93FDA0FDBCFDDD" box="[591,693,566,587]" italics="true" pageId="37" pageNumber="598">
<collectionCode id="ED7CAE37354EC169FD93FDA0FD68FDDD" box="[591,609,566,587]" name="Tavera, Department of Geology and Geophysics" pageId="37" pageNumber="598">T</collectionCode>
. slevini
</emphasis>
(
<specimenCode id="DBCB9E89354EC169FD18FDA0FE35FDFC" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="37" pageNumber="598">ANSP 24717–24719</specimenCode>
) are catalogued from ‘ 
<collectingCountry id="F37A7662354EC169FDE1FDC2FD95FDFF" box="[573,668,596,617]" name="Panama" pageId="37" pageNumber="598">Panama</collectingCountry>
<collectingCounty id="62B34E7E354EC169FD79FDC2FC02FDFC" box="[677,779,596,618]" pageId="37" pageNumber="598">Sabanas’</collectingCounty>
for a total of 11 
<collectingMunicipality id="6BB6AC88354EC169FEB2FDE5FEDFFD1F" box="[366,470,627,649]" pageId="37" pageNumber="598">Academy</collectingMunicipality>
specimens of the species. 
<location id="8EB26029354EC169FF78FD07FFFEFD31" LSID="urn:lsid:plazi:treatment:03C487E43548C169FC5EF95DFEA8F8F9:8EB26029354EC169FF78FD07FFFEFD31" box="[164,247,657,679]" country="Panama" county="Sabanas'" municipality="Academy" name="Clearly" pageId="37" pageNumber="598" stateProvince="Chiriqui">Clearly</location>
, the ‘ 
<location id="8EB26029354EC169FEEBFD07FE94FD31" LSID="urn:lsid:plazi:treatment:03C487E43548C169FC5EF95DFEA8F8F9:8EB26029354EC169FEEBFD07FE94FD31" box="[311,413,657,679]" country="Panama" county="Sabanas'" municipality="Academy" name="Sabanas'" pageId="37" pageNumber="598" stateProvince="Chiriqui">Sabanas’</location>
specimens are part of the nine collected for the 
<collectionCode id="ED7CAE37354EC169FEA8FD26FEAFFD53" box="[372,422,688,709]" pageId="37" pageNumber="598">PSC</collectionCode>
from Lérida. The only other specimen of this species listed in 
<location id="8EB26029354EC169FDF6FD59FD72FD72" LSID="urn:lsid:plazi:treatment:03C487E43548C169FC5EF95DFEA8F8F9:8EB26029354EC169FDF6FD59FD72FD72" box="[554,635,719,740]" country="Panama" county="Sabanas'" municipality="Academy" name="Dunn's" pageId="37" pageNumber="598" stateProvince="Chiriqui">Dunn’s</location>
notes is the 
<typeStatus id="54D68850354EC169FF78FD7BFE0CFC95" box="[164,261,749,771]" pageId="37" pageNumber="598" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED7CAE37354EC169FEC4FD7BFE45FC94" box="[280,332,749,770]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="37" pageNumber="598" type="Museum">CAS</collectionCode>
78938) from 
<collectingCountry id="F37A7662354EC169FE2EFD78FD5DFC95" box="[498,596,750,771]" name="Panama" pageId="37" pageNumber="598">Panama</collectingCountry>
: 
<collectingRegion id="49A9F810354EC169FDB8FD7BFDCEFC95" box="[612,711,749,771]" country="Panama" name="Chiriqui" pageId="37" pageNumber="598">Chiriquí</collectingRegion>
: near 
<location id="8EB26029354EC169FF78FC9AFE0CFCB7" LSID="urn:lsid:plazi:treatment:03C487E43548C169FC5EF95DFEA8F8F9:8EB26029354EC169FF78FC9AFE0CFCB7" box="[164,261,780,801]" country="Panama" county="Sabanas'" municipality="Academy" name="Boquete" pageId="37" pageNumber="598" stateProvince="Chiriqui">Boquete</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8BD236F2354EC169FF60FCBCFED6FC2C" blockId="37.[164,779,351,955]" pageId="37" pageNumber="598">
Although not noted by previous authors, another upland snake, 
<taxonomicName id="4C6D4D71354EC169FE8FFCDCFD0AFCC8" baseAuthorityName="Cope" baseAuthorityYear="1875" box="[339,515,841,863]" class="Insecta" family="Thyrididae" genus="Ninia" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="598" phylum="Arthropoda" rank="species" species="psephota">
<emphasis id="B919EAE0354EC169FE8FFCDCFD0AFCC8" box="[339,515,841,863]" italics="true" pageId="37" pageNumber="598">Ninia psephota</emphasis>
</taxonomicName>
, is represented in the Academy collections by a PSC specimen (ANSP 24720) purportedly from ‘ 
<collectingCountry id="F37A7662354EC169FE09FC10FD3DFC0D" box="[469,564,902,923]" name="Panama" pageId="37" pageNumber="598">Panama</collectingCountry>
Sabanas’. Dunn’s notes on 
<taxonomicName id="4C6D4D71354EC169FED3FC30FE9CFC2C" baseAuthorityName="Cope" baseAuthorityYear="1875" box="[271,405,933,955]" class="Insecta" family="Thyrididae" genus="Ninia" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="598" phylum="Arthropoda" rank="species" species="psephota">
<emphasis id="B919EAE0354EC169FED3FC30FE9CFC2C" box="[271,405,933,955]" italics="true" pageId="37" pageNumber="598">N. psephota</emphasis>
</taxonomicName>
state:
</paragraph>
<paragraph id="8BD236F2354EC169FF78FC72FEACFB83" blockId="37.[164,778,996,1045]" pageId="37" pageNumber="598">‘Lerida cl. 1940 31941 31942 13 1944 4 [a line has been drawn through these] Lerida 38’</paragraph>
<paragraph id="8BD236F2354EC169FF60FBA8FD67FB5B" blockId="37.[164,779,1085,1903]" pageId="37" pageNumber="598">
No mention is made of any members of this species from elsewhere in 
<collectingCountry id="F37A7662354EC169FE5BFBCAFEE1FBE7" box="[391,488,1116,1137]" name="Panama" pageId="37" pageNumber="598">Panama</collectingCountry>
, except the series of 61 snakes from nearby Boquete reported on by Slevin (1942). The number of snakes of this species listed from Lérida by Dunn (1947) is also 38.
</paragraph>
<paragraph id="8BD236F2354EC169FF60FB40FDFCF9AA" blockId="37.[164,779,1085,1903]" pageId="37" pageNumber="598">
A single specimen of 
<emphasis id="B919EAE0354EC169FE1AFB40FDA7FB7D" box="[454,686,1238,1259]" italics="true" pageId="37" pageNumber="598">Liophis epinephelus</emphasis>
(ANSP 24721) listed in the Academy catalogue as being from ‘ 
<collectingCountry id="F37A7662354EC169FF74FA82FE0EFABF" box="[168,263,1300,1321]" name="Panama" pageId="37" pageNumber="598">Panama</collectingCountry>
Sabanas’ has not been mentioned by previous authors. A typed label in the jar containing this specimen reads ‘Finca Lerida, 
<collectingRegion id="49A9F810354EC169FDD9FAC7FD6EFAF1" box="[517,615,1361,1383]" country="Panama" name="Chiriqui" pageId="37" pageNumber="598">Chiriqui</collectingRegion>
, PSC’. Dunn’s notes record 52 (
<quantity id="4C959B17354EC169FEBAFAF9FEABFA13" box="[358,418,1391,1413]" metricMagnitude="0" metricUnit="m" metricValue="1.4478" pageId="37" pageNumber="598" unit="in" value="57.0">57 in</quantity>
Dunn, 1947) specimens of this species from Lérida and 19 heads from Sabanas. As the Academy specimen is complete and has associated locality data, it must have been collected at Lérida. In an obvious 
<emphasis id="B919EAE0354EC169FEF3FA7CFE71FA69" box="[303,376,1514,1535]" italics="true" pageId="37" pageNumber="598">lapsus</emphasis>
Dunn (1947) lists the number of specimens of this form from Lérida as 31, duplicating the number recorded for 
<emphasis id="B919EAE0354EC169FE16F9B1FDFCF9AA" box="[458,757,1575,1596]" italics="true" pageId="37" pageNumber="598">
<taxonomicName id="4C6D4D71354EC169FE16F9B1FD68F9AA" box="[458,609,1575,1596]" class="Reptilia" family="Colubridae" genus="Lampropeltis" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="598" phylum="Chordata" rank="genus">Lampropeltis</taxonomicName>
triangulum.
</emphasis>
</paragraph>
<paragraph id="8BD236F2354EC169FF60F9D3FEA8F8F9" blockId="37.[164,779,1085,1903]" pageId="37" pageNumber="598">
Significantly, the catalogue numbers for all the reputed ‘ 
<collectingCountry id="F37A7662354EC169FED6F9F2FE60F9EF" box="[266,361,1636,1657]" name="Panama" pageId="37" pageNumber="598">Panama</collectingCountry>
Sabanas’ PSC specimens that obviously came from Lérida are in a single sequence (ANSP 24717–24734). The previous series, ANSP 24703–24716, are PSC snakes recorded in Dunn’s notes and the Academy catalogue as from ‘Sabanas’. We can only conclude that there was a cataloguing error where 
<collectingCountry id="F37A7662354EC169FE9DF88AFE96F8A7" box="[321,415,1820,1841]" name="Panama" pageId="37" pageNumber="598">Panama</collectingCountry>
Sabanas was entered as the locality for the next following 
<specimenCount id="9D6BFD7B354EC169FDD5F8ACFDADF8C6" box="[521,676,1850,1872]" pageId="37" pageNumber="598" type="generic">18 specimens</specimenCount>
actually collected from Lérida.
</paragraph>
</subSubSection>
</treatment>
</document>