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<document id="DF3A0611844E12DF5BFA2AE64CF5A6BB" ID-DOI="10.5281/zenodo.162803" ID-GBIF-Dataset="f514e964-2196-461c-84ff-64243643bc81" ID-ISSN="1175­5326" ID-Zenodo-Dep="162803" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459445184067" checkinUser="plazi" docAuthor="Erwin, Terry L." docDate="2002" docId="03A287CA7838FFA1FEECFCCE9EDBFE51" docLanguage="en" docName="zt00119.pdf" docOrigin="Zootaxa 119" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Agra schwarzeneggeri Erwin, new species" docType="treatment" docVersion="11" lastPageNumber="47" masterDocId="FF9BFFB27815FF8FFFE4FF829C34FFFF" masterDocTitle="The Beetle Family Carabidae of Costa Rica: Twenty­nine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)" masterLastPageNumber="68" masterPageNumber="1" pageNumber="46" updateTime="1698190273211" updateUser="plazi">
<mods:mods id="17E5AAE81768DB083BB6CE604E05BE12" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4C2EB41649E6F1E82726D5B1BB4D9182">
<mods:title id="4DB48ACF522F7F37374CC454430E8819">The Beetle Family Carabidae of Costa Rica: Twenty­nine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)</mods:title>
</mods:titleInfo>
<mods:name id="57241336B4809A0C234EDB9974C9C048" type="personal">
<mods:role id="7C504E89BF340549D263A67E7E85F07E">
<mods:roleTerm id="EBFBED12B76673622CDAC76A6ADF7B98">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2F63F308D0B067E9BFE409E0BB43938E">Erwin, Terry L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="3CEB23A1F5CE7775C5240EAD98B95337">text</mods:typeOfResource>
<mods:relatedItem id="C4813DE27B7758DD84408544DD2F4E13" type="host">
<mods:titleInfo id="9594371911B2F4CCD1E6C925E6333370">
<mods:title id="5FBC86F2AB6A6327A507E69F7974C854">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="586549929F1E8FDE583D7E4A028EAA08">
<mods:date id="811C7441F6DB13053296BAEF18ED0134">2002</mods:date>
<mods:detail id="63599971C0EBA266404491D48BD18BD2" type="volume">
<mods:number id="17DEC13735E364DA58DED1A129BF2A81">119</mods:number>
</mods:detail>
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<mods:start id="30916A06E2B7FBF8E432A47BD530675C">1</mods:start>
<mods:end id="8AF59E4BCDC437D8BE5B1C52F453959C">68</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="9B01D46167E3E24FE735CD0C2EAB3DCA">journal article</mods:classification>
<mods:identifier id="82725E09B3A1D003AFFD9EB7B1E479CC" type="DOI">10.5281/zenodo.162803</mods:identifier>
<mods:identifier id="24F7CE74C180D201EED9F09870FBCC66" type="GBIF-Dataset">f514e964-2196-461c-84ff-64243643bc81</mods:identifier>
<mods:identifier id="6479B4DECF2DED8ECDB1D88BD0B2C13C" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="46CF7D24166249D9716FCB923EB9153E" type="Zenodo-Dep">162803</mods:identifier>
</mods:mods>
<treatment id="03A287CA7838FFA1FEECFCCE9EDBFE51" ID-DOI="http://doi.org/10.5281/zenodo.6278005" ID-GBIF-Taxon="121065199" ID-Zenodo-Dep="6278005" LSID="urn:lsid:plazi:treatment:03A287CA7838FFA1FEECFCCE9EDBFE51" httpUri="http://treatment.plazi.org/id/03A287CA7838FFA1FEECFCCE9EDBFE51" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
<subSubSection id="C31165577838FFA2FEECFCCE9E02FC71" pageId="45" pageNumber="46" type="nomenclature">
<paragraph id="8BB436DC7838FFA2FEECFCCE9F3EFC9A" blockId="45.[264,778,843,910]" box="[264,778,843,870]" pageId="45" pageNumber="46">
<heading id="D0FC81B07838FFA2FEECFCCE9F3EFC9A" bold="true" box="[264,778,843,870]" fontSize="11" level="1" pageId="45" pageNumber="46" reason="1">
<emphasis id="B97FEACE7838FFA2FEECFCCE9F3EFC9A" bold="true" box="[264,778,843,870]" pageId="45" pageNumber="46">
<taxonomicName id="4C0B4D5F7838FFA2FEECFCCE9E59FC9A" ID-CoL="B2X8" authority="Erwin" authorityName="Erwin" box="[264,621,843,870]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="schwarzeneggeri" status="sp. nov.">
<emphasis id="B97FEACE7838FFA2FEECFCCE9E22FC99" bold="true" box="[264,534,844,870]" italics="true" pageId="45" pageNumber="46">Agra schwarzeneggeri</emphasis>
Erwin
</taxonomicName>
,
<taxonomicNameLabel id="A24C57B57838FFA2FD9EFCC99F3EFC9A" box="[634,778,843,869]" pageId="45" pageNumber="46" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB436DC7838FFA2FEECFCF69E02FC71" blockId="45.[264,778,843,910]" box="[264,566,884,910]" pageId="45" pageNumber="46">
(
<figureCitation id="13302A597838FFA2FEF5FCF69D9FFC71" box="[273,427,884,910]" captionStart="FIGURE 20. A" captionStartId="46.[264,367,1318,1342]" captionTargetBox="[434,1149,491,1285]" captionTargetId="figure@46.[433,1153,489,1291]" captionTargetPageId="46" captionText="FIGURE 20. A. schwarzeneggeri, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) middle leg, post­dorsal aspect. Measures (mm): HL = 2.7; WE = 2.3; male femur diameter = 1.1; male SW = 4.0, female SW = 4.6." httpUri="https://zenodo.org/record/162834/files/figure.png" pageId="45" pageNumber="46">Figs. 20a, 20</figureCitation>
b, 20c, 20d)
</paragraph>
</subSubSection>
<subSubSection id="C31165577838FFA2FEECFC439E8EFBFC" pageId="45" pageNumber="46" type="materials_examined">
<paragraph id="8BB436DC7838FFA2FEECFC439E8EFBFC" blockId="45.[264,1326,961,1867]" pageId="45" pageNumber="46">
<emphasis id="B97FEACE7838FFA2FEECFC439DB5FC24" bold="true" box="[264,385,961,987]" pageId="45" pageNumber="46">
<typeStatus id="54B0887E7838FFA2FEECFC439D48FC24" box="[264,380,961,987]" pageId="45" pageNumber="46" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male,
<collectingCountry id="F31C764C7838FFA2FE37FC439EB6FC24" box="[467,642,961,987]" name="Costa Rica" pageId="45" pageNumber="46">COSTA RICA</collectingCountry>
: Cartago,
<taxonomicName id="4C0B4D5F7838FFA2FD13FC439F52FC24" box="[759,870,961,987]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="25" phylum="Arthropoda" rank="species" species="turrialba">Turrialba</taxonomicName>
,
<quantity id="4CF39B397838FFA2FC95FC439F83FC24" box="[881,951,961,987]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="45" pageNumber="46" unit="m" value="650.0">650m</quantity>
,
<geoCoordinate id="EE3F501B7838FFA2FC26FC43982CFC24" box="[962,1048,961,987]" direction="north" orientation="latitude" pageId="45" pageNumber="46" precision="925" value="9.883333">9º 53'N</geoCoordinate>
<geoCoordinate id="EE3F501B7838FFA2FBFBFC4398B7FC24" box="[1055,1155,961,987]" direction="west" orientation="longitude" pageId="45" pageNumber="46" precision="925" value="-83.63333">83º38'W</geoCoordinate>
, February, (H. &amp; A. Howden)(
<collectionCode id="ED1AAE197838FFA2FE26FC6B9E2FFBFC" LSID="urn:lsid:biocol.org:col:34871" box="[450,539,1001,1027]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="45" pageNumber="46">USNM</collectionCode>
: ADP64794).
</paragraph>
</subSubSection>
<subSubSection id="C31165577838FFA2FEDCFB939D77FB84" pageId="45" pageNumber="46" type="diagnosis">
<paragraph id="8BB436DC7838FFA2FEDCFB939D77FB84" blockId="45.[264,1326,961,1867]" pageId="45" pageNumber="46">
<emphasis id="B97FEACE7838FFA2FEDCFB939D83FBD4" bold="true" box="[312,439,1041,1067]" pageId="45" pageNumber="46">Diagnosis.</emphasis>
The male of this species has an enormously enlarged middle femur (
<figureCitation id="13302A597838FFA2FB1EFB939D13FBAC" captionStart="FIGURE 19. A" captionStartId="44.[264,368,1791,1815]" captionTargetBox="[406,1178,1107,1755]" captionTargetId="figure@44.[405,1181,1089,1765]" captionTargetPageId="44" captionText="FIGURE 19. A. granodeoro, male, dorsal aspect: a) head; b) apex of elytron; c) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.3; WE = 2.2; AL = 5.0." httpUri="https://zenodo.org/record/162832/files/figure.png" pageId="45" pageNumber="46">Fig. 19</figureCitation>
c); females have elytra with brassy­green tint, black forebody and rufous antennomeres 3­11.
</paragraph>
</subSubSection>
<subSubSection id="C31165577838FFA2FEDCFB0B985EFA1C" pageId="45" pageNumber="46" type="description">
<paragraph id="8BB436DC7838FFA2FEDCFB0B985EFA1C" blockId="45.[264,1326,961,1867]" pageId="45" pageNumber="46">
<emphasis id="B97FEACE7838FFA2FEDCFB0B9DE4FB5C" bold="true" box="[312,464,1161,1187]" pageId="45" pageNumber="46">Description.</emphasis>
Color and luster: Black. Elytra with brassy green reflections. Antennomeres 3­4 testaceous, infuscated apically, 5­11 testaceous. Form: Head (
<figureCitation id="13302A597838FFA2FB92FB3398E9FB34" box="[1142,1245,1201,1227]" captionStart="FIGURE 20. A" captionStartId="46.[264,367,1318,1342]" captionTargetBox="[434,1149,491,1285]" captionTargetId="figure@46.[433,1153,489,1291]" captionTargetPageId="46" captionText="FIGURE 20. A. schwarzeneggeri, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) middle leg, post­dorsal aspect. Measures (mm): HL = 2.7; WE = 2.3; male femur diameter = 1.1; male SW = 4.0, female SW = 4.6." httpUri="https://zenodo.org/record/162834/files/figure.png" pageId="45" pageNumber="46">Fig. 20a</figureCitation>
) short behind eyes, that of male tapered and rounded behind, that of female quadrate, vertex dimpled. Prothorax widest at middle, markedly tapered anteriorly in male, somewhat &quot;swollen&quot; at middle in female. Middle leg with markedly enlarged femur (
<figureCitation id="13302A597838FFA2FBBBFAAB988FFABC" box="[1119,1211,1321,1347]" captionStart="FIGURE 20. A" captionStartId="46.[264,367,1318,1342]" captionTargetBox="[434,1149,491,1285]" captionTargetId="figure@46.[433,1153,489,1291]" captionTargetPageId="46" captionText="FIGURE 20. A. schwarzeneggeri, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) middle leg, post­dorsal aspect. Measures (mm): HL = 2.7; WE = 2.3; male femur diameter = 1.1; male SW = 4.0, female SW = 4.6." httpUri="https://zenodo.org/record/162834/files/figure.png" pageId="45" pageNumber="46">Fig. 20</figureCitation>
d). Elytra broad,
<taxonomicName id="4C0B4D5F7838FFA2FEBAFAD39DB7FA94" box="[350,387,1361,1387]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="not">not</taxonomicName>
very convex, apex (
<figureCitation id="13302A597838FFA2FD67FAD39EE9FA94" box="[643,733,1361,1387]" captionStart="FIGURE 20. A" captionStartId="46.[264,367,1318,1342]" captionTargetBox="[434,1149,491,1285]" captionTargetId="figure@46.[433,1153,489,1291]" captionTargetPageId="46" captionText="FIGURE 20. A. schwarzeneggeri, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) middle leg, post­dorsal aspect. Measures (mm): HL = 2.7; WE = 2.3; male femur diameter = 1.1; male SW = 4.0, female SW = 4.6." httpUri="https://zenodo.org/record/162834/files/figure.png" pageId="45" pageNumber="46">Fig. 20</figureCitation>
b) medially dentate, interneurs 2,4,6,8 with very large setigerous foveae, interneurs of fine close­spaced punctures. Sternum VI (
<figureCitation id="13302A597838FFA2FB5AFAFB9922FA6C" box="[1214,1302,1401,1427]" captionStart="FIGURE 20. A" captionStartId="46.[264,367,1318,1342]" captionTargetBox="[434,1149,491,1285]" captionTargetId="figure@46.[433,1153,489,1291]" captionTargetPageId="46" captionText="FIGURE 20. A. schwarzeneggeri, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) middle leg, post­dorsal aspect. Measures (mm): HL = 2.7; WE = 2.3; male femur diameter = 1.1; male SW = 4.0, female SW = 4.6." httpUri="https://zenodo.org/record/162834/files/figure.png" pageId="45" pageNumber="46">Fig. 20</figureCitation>
c) with deeply circular emargination in female, deeply v­notched in female. Aedeagus (see Notes). Size: medium, 14.0 to 17.0mm in length,
<quantity id="4CF39B397838FFA2FCAAFA4B9FCDFA1C" box="[846,1017,1481,1507]" metricMagnitude="-3" metricUnit="m" metricValue="4.88" metricValueMax="5.16" metricValueMin="4.6" pageId="45" pageNumber="46" unit="mm" value="4.88" valueMax="5.16" valueMin="4.6">4.6 to 5.16mm</quantity>
in width.
</paragraph>
</subSubSection>
<subSubSection id="C31165577838FFA2FEA5FA739D87F8B4" pageId="45" pageNumber="46" type="materials_examined">
<paragraph id="8BB436DC7838FFA2FEA5FA73989AF954" blockId="45.[264,1326,961,1867]" pageId="45" pageNumber="46">
<emphasis id="B97FEACE7838FFA2FEA5FA739EABF9F4" bold="true" box="[321,671,1521,1547]" pageId="45" pageNumber="46">Other specimens examined.</emphasis>
<collectingCountry id="F31C764C7838FFA2FD4DFA739F04F9F4" box="[681,816,1521,1547]" name="Costa Rica" pageId="45" pageNumber="46">Costa Rica</collectingCountry>
: 3
<typeStatus id="54B0887E7838FFA2FCB2FA739FFDF9F4" box="[854,969,1521,1547]" pageId="45" pageNumber="46" type="paratype">paratypes</typeStatus>
, females, from the Cordillera Volcánica Central Conservation Area.
<collectingCountry id="F31C764C7838FFA2FD35F99B9F61F9CC" box="[721,853,1561,1587]" name="Costa Rica" pageId="45" pageNumber="46">Costa Rica</collectingCountry>
, 1f, Cartago, Pavones
<geoCoordinate id="EE3F501B7838FFA2FB86F99B98FDF9CC" box="[1122,1225,1561,1587]" direction="west" orientation="longitude" pageId="45" pageNumber="46" precision="925" value="-83.61667">83°37'W</geoCoordinate>
<geoCoordinate id="EE3F501B7838FFA2FB35F99B9912F9CC" box="[1233,1318,1561,1587]" direction="north" orientation="latitude" pageId="45" pageNumber="46" precision="925" value="9.95">9°57'N</geoCoordinate>
, May, (F.T. Hovore)(
<collectionCode id="ED1AAE197838FFA2FE1DF9C39E73F9A4" box="[505,583,1601,1627]" pageId="45" pageNumber="46">FTHC</collectionCode>
: ADP70530), 1f,
<taxonomicName id="4C0B4D5F7838FFA2FCFAF9C39FB9F9A4" box="[798,909,1601,1627]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="25" phylum="Arthropoda" rank="species" species="turrialba">Turrialba</taxonomicName>
, 3.0 km SEIICA/
<taxonomicName id="4C0B4D5F7838FFA2FB85F9C39883F9A4" box="[1121,1207,1601,1627]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="51" phylum="Arthropoda" rank="species" species="catie">
<collectionCode id="ED1AAE197838FFA2FB85F9C39883F9A4" LSID="urn:lsid:biocol.org:col:15110" box="[1121,1207,1601,1627]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15110" name="Tropical Agricultural Research and Training Center (CATIE)" pageId="45" pageNumber="46">CATIE</collectionCode>
</taxonomicName>
,
<geoCoordinate id="EE3F501B7838FFA2FB22F9C39919F9A4" box="[1222,1325,1601,1627]" direction="west" orientation="longitude" pageId="45" pageNumber="46" precision="925" value="-83.63333">83°38'W</geoCoordinate>
<geoCoordinate id="EE3F501B7838FFA2FEECF9EB9D69F97C" box="[264,349,1641,1667]" direction="north" orientation="latitude" pageId="45" pageNumber="46" precision="925" value="9.883333">9°53'N</geoCoordinate>
, April, (E. Giesbert)(
<collectionCode id="ED1AAE197838FFA2FD8DF9EB9E82F97C" box="[617,694,1641,1667]" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="45" pageNumber="46">FSCA</collectionCode>
: ADP4420), 1f, August, (D.R. Whitehead)(
<collectionCode id="ED1AAE197838FFA2FB36F9EB991DF97C" LSID="urn:lsid:biocol.org:col:34871" box="[1234,1321,1641,1667]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="45" pageNumber="46">USNM</collectionCode>
: ADP61717),
<quantity id="4CF39B397838FFA2FE40F9139DF8F954" box="[420,460,1681,1707]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="45" pageNumber="46" unit="m" value="1.0">1m</quantity>
, February, (H.F. Howden &amp; A. Howden)(
<collectionCode id="ED1AAE197838FFA2FC52F913983BF954" LSID="urn:lsid:biocol.org:col:34404" box="[950,1039,1681,1707]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34404" name="University of Alberta, E.H. Strickland Entomological Museum" pageId="45" pageNumber="46">UASM</collectionCode>
: ADP54794).
</paragraph>
<paragraph id="8BB436DC7838FFA2FEDCF93B9D87F8B4" blockId="45.[264,1326,961,1867]" pageId="45" pageNumber="46">
<emphasis id="B97FEACE7838FFA2FEDCF93B9DCBF92C" bold="true" box="[312,511,1721,1747]" pageId="45" pageNumber="46">Specific epithet.</emphasis>
The specific epithet,
<taxonomicName id="4C0B4D5F7838FFA2FD1BF9389FF2F92C" box="[767,966,1722,1747]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="schwarzeneggeri">
<emphasis id="B97FEACE7838FFA2FD1BF9389FF2F92C" box="[767,966,1722,1747]" italics="true" pageId="45" pageNumber="46">schwarzeneggeri</emphasis>
</taxonomicName>
is the Latinized genitive form of the family name of the actor, Arnold Schwarzenegger, in reference to the markedly developed (biceps­like) middle femora of the males of this species reminiscent of the actor’s physic.
</paragraph>
</subSubSection>
<subSubSection id="C3116557783BFFA1FEDCFE999EDBFE51" pageId="46" pageNumber="47" type="description">
<paragraph id="8BB436DC783BFFA1FEDCFE999EDBFE51" blockId="46.[264,1323,283,430]" pageId="46" pageNumber="47">
<emphasis id="B97FEACE783BFFA1FEDCFE999DB1FECA" bold="true" box="[312,389,283,309]" pageId="46" pageNumber="47">Notes.</emphasis>
The genitalia of the dissected male is missing from the vial pinned beneath the specimen, thus has
<taxonomicName id="4C0B4D5F783BFFA1FE0FFEC69E24FEA1" box="[491,528,324,350]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="not">not</taxonomicName>
been illustrated. The aedeagus of
<taxonomicName id="4C0B4D5F783BFFA1FC44FEC698C9FEA1" authority="Straneo 1982" authorityName="Straneo" authorityYear="1982" box="[928,1277,324,350]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="perrinae">
<emphasis id="B97FEACE783BFFA1FC44FEC69873FEA2" box="[928,1095,324,349]" italics="true" pageId="46" pageNumber="47">Agra perrinae</emphasis>
(Straneo 1982)
</taxonomicName>
has a moderately long ostium, apex of phallus a simple diamond­shaped lobe which is markedly arched dorsally, parameres glabrous.
</paragraph>
</subSubSection>
</treatment>
</document>
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