treatments-xml/data/03/B7/5F/03B75F43F06FFFBAFF4877C13317208A.xml

<document id="135693C37C03634C8393FC43CE1351DA" ID-DOI="10.11646/zootaxa.5567.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="14703735" ID-ZooBank="AF6254F6-CE26-41FE-BB85-A3FFD16FAA39" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737371447294" checkinUser="plazi" docAuthor="VIRAKTAMATH, C. A., WEBB, M. D. &amp; YESHWANTH, H. M." docDate="2025" docId="03B75F43F06FFFBAFF4877C13317208A" docLanguage="en" docName="zootaxa.5567.1.1.pdf" docOrigin="Zootaxa 5567 (1)" docSource="https://doi.org/10.11646/zootaxa.5567.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Panghoella nera VIRAKTAMATH, WEBB &amp; YESHWANTH, 2025, sp. nov." docType="treatment" docVersion="2" lastPageNumber="39" masterDocId="FF8E273BF049FF9CFFDF7756364D2343" masterDocTitle="Review of the Monobazus-group leafhopper genera (Hemiptera: Cicadellidae: Deltocephalinae) from the Indian subcontinent with description of five new genera and twenty-eight new species" masterLastPageNumber="105" masterPageNumber="1" pageNumber="39" updateTime="1737389185710" updateUser="ExternalLinkService">
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<mods:title id="CBE751674A88A0B55E0FA04659718816">Review of the Monobazus-group leafhopper genera (Hemiptera: Cicadellidae: Deltocephalinae) from the Indian subcontinent with description of five new genera and twenty-eight new species</mods:title>
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<mods:namePart id="664049E5D669C0AAAAA1D5DE233E4282">VIRAKTAMATH, C. A.</mods:namePart>
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<mods:namePart id="F11A17A48C323076D3E22977D2AB218D">WEBB, M. D.</mods:namePart>
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<subSubSection id="C304BDDEF06FFFBAFF4877C1378823F2" box="[151,453,151,178]" pageId="38" pageNumber="39" type="nomenclature">
<paragraph id="8BA1EE55F06FFFBAFF4877C1378823F2" blockId="38.[151,453,151,213]" box="[151,453,151,178]" pageId="38" pageNumber="39">
<heading id="D0E95939F06FFFBAFF4877C1378823F2" bold="true" box="[151,453,151,178]" fontSize="11" level="2" pageId="38" pageNumber="39" reason="4">
<emphasis id="B96A3247F06FFFBAFF4877C1378823F2" bold="true" box="[151,453,151,178]" pageId="38" pageNumber="39">
<taxonomicName id="4C1E95D6F06FFFBAFF4877C1372D23F2" authorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" authorityYear="2025" box="[151,352,151,178]" class="Insecta" family="Cicadellidae" genus="Panghoella" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="nera" status="sp. nov.">
<emphasis id="B96A3247F06FFFBAFF4877C1372D23F2" bold="true" box="[151,352,151,178]" italics="true" pageId="38" pageNumber="39">Panghoella nera</emphasis>
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<taxonomicNameLabel id="A2598F3CF06FFFBAFEB777C1378823F2" box="[360,453,151,177]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C304BDDEF06FFFBAFF4877ED3269201E" pageId="38" pageNumber="39" type="description">
<paragraph id="8BA1EE55F06FFFBAFF4877ED375B2396" blockId="38.[151,453,151,213]" box="[151,278,187,213]" pageId="38" pageNumber="39">
<figureCitation id="1325F2D0F06FFFBAFF4877ED375B2396" box="[151,278,187,213]" captionStart="FIGURES 48" captionStartId="89.[151,264,1834,1858]" captionTargetBox="[156,1430,181,1806]" captionTargetId="figure-17@89.[151,1435,181,1811]" captionTargetPageId="89" captionText="FIGURES 48A–I. Panghoella nera sp. nov.: A–B, holotype male habitus, dorsal and lateral views; C, face; D, pygofer, lateral view; E, valve and subgenital plates ventral view; F, style, dorsal view; G–H, connective and aedeagus ventral and dorsal views; I, aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14703875" httpUri="https://zenodo.org/record/14703875/files/figure.png" pageId="38" pageNumber="39">Figs 48A–I</figureCitation>
</paragraph>
<paragraph id="8BA1EE55F06FFFBAFF487652355D22C9" blockId="38.[151,1437,259,970]" pageId="38" pageNumber="39">
<emphasis id="B96A3247F06FFFBAFF487652375D225E" box="[151,272,260,285]" italics="true" pageId="38" pageNumber="39">Diagnosis.</emphasis>
Structure and coloration very similar to 
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<emphasis id="B96A3247F06FFFBAFD3176523535225E" box="[750,888,259,285]" italics="true" pageId="38" pageNumber="39">P. maculosa</emphasis>
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but differing in the straight aedeagal shaft and thinner basal processes. Apex of shaft angularly produced dorsally as in the case of 
<taxonomicName id="4C1E95D6F06FFFBAFB84767E33D12202" authority="(Xing &amp; Li)" authorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" authorityYear="2025" baseAuthorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" baseAuthorityYear="2025" box="[1115,1436,295,321]" class="Insecta" family="Cicadellidae" genus="Panghoella" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="suiyangensis">
<emphasis id="B96A3247F06FFFBAFB84767E33452202" box="[1115,1288,296,321]" italics="true" pageId="38" pageNumber="39">P. suiyangensis</emphasis>
(Xing &amp; Li)
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another species in the 
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<emphasis id="B96A3247F06FFFBAFE53761A345C2226" box="[396,529,331,357]" italics="true" pageId="38" pageNumber="39">P. maculosa</emphasis>
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species group, but the latter species differs from the new species in having curved aedeagal shaft with more or less sinuate basal processes.
</paragraph>
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<emphasis id="B96A3247F06FFFBAFF1876C2370622EE" box="[199,331,404,429]" italics="true" pageId="38" pageNumber="39">Description</emphasis>
. Coloration and structure similar to that in 
<taxonomicName id="4C1E95D6F06FFFBAFCEF76C235FB22EE" authorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" authorityYear="2025" baseAuthorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" baseAuthorityYear="2025" box="[816,950,403,429]" class="Insecta" family="Cicadellidae" genus="Panghoella" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="B96A3247F06FFFBAFCEF76C235FB22EE" box="[816,950,403,429]" italics="true" pageId="38" pageNumber="39">P. maculosa</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA1EE55F06FFFBAFF1876E1377021C6" blockId="38.[151,1437,259,970]" pageId="38" pageNumber="39">
<emphasis id="B96A3247F06FFFBAFF1876E137262292" box="[199,363,439,465]" italics="true" pageId="38" pageNumber="39">Male genitalia</emphasis>
. Pygofer about 1.4× as long as height in lateral view, posterior margin rounded, (without a dorsal marginal process). Subgenital plates about 3× as long as wide at base, basal sclerotized area 0.45× as long as total length. Style and connective as in 
<taxonomicName id="4C1E95D6F06FFFBAFDC0755634EA215A" authorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" authorityYear="2025" baseAuthorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" baseAuthorityYear="2025" box="[543,679,511,537]" class="Insecta" family="Cicadellidae" genus="Panghoella" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="B96A3247F06FFFBAFDC0755634EA215A" box="[543,679,511,537]" italics="true" pageId="38" pageNumber="39">P. maculosa</emphasis>
</taxonomicName>
. Aedeagus with dorsal apodeme and shaft subequal, shaft more or less straight, slightly nearer apically in lateral view and with short triangular expansion on lateral surface near apex, basal pair of processes thin needle-like, slightly sinuate at base and then more or less straight and shorter than shaft in lateral view.
</paragraph>
<paragraph id="8BA1EE55F06FFFBAFF1875C63741218D" blockId="38.[151,1437,259,970]" pageId="38" pageNumber="39">
<emphasis id="B96A3247F06FFFBAFF1875C6372421EA" box="[199,361,656,681]" italics="true" pageId="38" pageNumber="39">Measurements</emphasis>
. Male 
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long, 
<quantity id="4CE643B0F06FFFBAFD8775D934E221E9" box="[600,687,655,682]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="38" pageNumber="39" unit="mm" value="1.3">1.3 mm</quantity>
wide across eyes and 
<quantity id="4CE643B0F06FFFBAFC7475D9324F21E9" box="[939,1026,655,682]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="38" pageNumber="39" unit="mm" value="1.5">1.5 mm</quantity>
wide across posterolateral angles of pronotum.
</paragraph>
<paragraph id="8BA1EE55F06FFFBAFF18758137482056" blockId="38.[151,1437,259,970]" pageId="38" pageNumber="39">
<emphasis id="B96A3247F06FFFBAFF18758137D721B2" box="[199,410,727,753]" italics="true" pageId="38" pageNumber="39">Material examined</emphasis>
. 
<materialsCitation id="3B76E408F06FFFBAFE77758137482056" collectingDate="1935-03-06" collectionCode="BMNH" collectorName="M. Steele" country="India" elevation="1631" location="INDIA" pageId="38" pageNumber="39" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectingCountry id="F309AEC5F06FFFBAFE77758137B121B2" box="[424,508,727,753]" name="India" pageId="38" pageNumber="39">INDIA</collectingCountry>
: 
<typeStatus id="54A550F7F06FFFBAFDDA758134D421B2" box="[517,665,727,753]" pageId="38" pageNumber="39" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="9D1825DCF06FFFBAFD40758E34FB21B1" box="[671,694,728,754]" count="1" pageId="38" pageNumber="39" type="male">♂</specimenCount>
, [BURMA]: Mishmi Hills: Chhaglon, 
<quantity id="4CE643B0F06FFFBAFBBC758132E221B1" box="[1123,1199,727,754]" metricMagnitude="3" metricUnit="m" metricValue="1.63068" pageId="38" pageNumber="39" unit="ft" value="5350.0">
<elevation id="00330966F06FFFBAFBBC758132E221B1" box="[1123,1199,727,754]" metricMagnitude="3" metricUnit="m" metricValue="1.63068" pageId="38" pageNumber="39" unit="ft" value="5350.0">5350ft</elevation>
</quantity>
, 
<date id="FFA0C895F06FFFBAFB667581336A21B1" box="[1209,1319,727,754]" pageId="38" pageNumber="39" value="1935-03-06">
<collectingDate id="EFE4317DF06FFFBAFB667581336A21B1" box="[1209,1319,727,754]" pageId="38" pageNumber="39" value="1935-03-06">6.iii.1935</collectingDate>
</date>
, 
<collectorName id="26EB8B83F06FFFBAFAED758E33D121B2" box="[1330,1436,727,754]" pageId="38" pageNumber="39">M. Steele</collectorName>
(
<collectionCode id="ED0F7690F06FFFBAFF4075AD36B42056" box="[159,249,763,789]" pageId="38" pageNumber="39">BMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BA1EE55F06FFFBAFF1874493269201E" blockId="38.[151,1437,259,970]" pageId="38" pageNumber="39">
<emphasis id="B96A3247F06FFFBAFF1874493772207A" box="[199,319,799,825]" italics="true" pageId="38" pageNumber="39">Etymology</emphasis>
. The species name (nera) is a word in the Kannada language meaning straight, referring to the straight aedeagal shaft and ventral processes of the aedeagus; noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C304BDDEF06FFFBAFF1874313317208A" pageId="38" pageNumber="39" type="discussion">
<paragraph id="8BA1EE55F06FFFBAFF1874313317208A" blockId="38.[151,1437,259,970]" pageId="38" pageNumber="39">
<emphasis id="B96A3247F06FFFBAFF187431377920C2" bold="true" box="[199,308,871,897]" pageId="38" pageNumber="39">Remarks</emphasis>
. 
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<emphasis id="B96A3247F06FFFBAFE9A743E37DB20C2" box="[325,406,872,897]" italics="true" pageId="38" pageNumber="39">P. nera</emphasis>
</taxonomicName>
<emphasis id="B96A3247F06FFFBAFE7F743137B020C2" bold="true" box="[416,509,871,897]" pageId="38" pageNumber="39">
<taxonomicNameLabel id="A2598F3CF06FFFBAFE7F743137B020C2" box="[416,509,871,897]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is a part of the species complex of 
<taxonomicName id="4C1E95D6F06FFFBAFC7D743E326120C2" authorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" authorityYear="2025" baseAuthorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" baseAuthorityYear="2025" box="[930,1068,871,897]" class="Insecta" family="Cicadellidae" genus="Panghoella" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="B96A3247F06FFFBAFC7D743E326120C2" box="[930,1068,871,897]" italics="true" pageId="38" pageNumber="39">P. maculosa</emphasis>
</taxonomicName>
and differs from the two forms illustrated here (
<figureCitation id="1325F2D0F06FFFBAFE9474DD37D320E6" box="[331,414,907,933]" captionStart="FIGURES 47" captionStartId="88.[151,264,1859,1883]" captionTargetBox="[163,1426,186,1830]" captionTargetId="figure-26@88.[158,1430,181,1835]" captionTargetPageId="88" captionText="FIGURES 47A–M. Panghoella maculosa (Distant), male genitalia: A, pygofer and anal segments, lateral view; B, valve and subgenital plate, ventral view; C, connective, dorsal view; D–E, style, dorsal view and magnified view of apophysis; F, aedeagus ventral view; G, connective and aedeagus, lateral view; H–J, syntype male from Kurseong: H, aedeagus lateral view; I, apex of aedeagal shaft lateral view magnified; J, aedeagus, ventral view; male from Nepal: K–L aedeagus, lateral and ventral view, M, style dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14703873" httpUri="https://zenodo.org/record/14703873/files/figure.png" pageId="38" pageNumber="39">Figs 47</figureCitation>
GHK) in having a straighter aedeagal shaft and the basal processes of the aedeagus slenderer and shorter than aedeagal shaft (
<figureCitation id="1325F2D0F06FFFBAFDDE74F93411208A" box="[513,604,943,969]" captionStart="FIGURES 48" captionStartId="89.[151,264,1834,1858]" captionTargetBox="[156,1430,181,1806]" captionTargetId="figure-17@89.[151,1435,181,1811]" captionTargetPageId="89" captionText="FIGURES 48A–I. Panghoella nera sp. nov.: A–B, holotype male habitus, dorsal and lateral views; C, face; D, pygofer, lateral view; E, valve and subgenital plates ventral view; F, style, dorsal view; G–H, connective and aedeagus ventral and dorsal views; I, aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14703875" httpUri="https://zenodo.org/record/14703875/files/figure.png" pageId="38" pageNumber="39">Fig. 48I</figureCitation>
) and the male pygofer lacking a dorsal marginal process (
<figureCitation id="1325F2D0F06FFFBAFB3A74F93303208A" box="[1253,1358,943,969]" captionStart="FIGURES 48" captionStartId="89.[151,264,1834,1858]" captionTargetBox="[156,1430,181,1806]" captionTargetId="figure-17@89.[151,1435,181,1811]" captionTargetPageId="89" captionText="FIGURES 48A–I. Panghoella nera sp. nov.: A–B, holotype male habitus, dorsal and lateral views; C, face; D, pygofer, lateral view; E, valve and subgenital plates ventral view; F, style, dorsal view; G–H, connective and aedeagus ventral and dorsal views; I, aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14703875" httpUri="https://zenodo.org/record/14703875/files/figure.png" pageId="38" pageNumber="39">Fig. 48D</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
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