treatments-xml/data/BB/80/54/BB8054E1CA9BF0ADCD640C08E6215786.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.410.7488" ID-GBIF-Dataset="69a473d9-1152-4f3d-8b4e-a9de37e6f4c1" ID-PMC="PMC4042706" ID-Pensoft-Pub="1313-2970-410-23" ID-PubMed="24899848" ID-ZBK="E4C040C1939640F18C12D9518F59F668" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-410-23" ModsDocOrigin="ZooKeys 410" ModsDocTitle="A review of the microgastropod genus Systenostoma Bavay &amp; Dautzenberg, 1908 and a new subterranean species from China (Gastropoda, Pulmonata, Hypselostomatidae)" checkinTime="1451245876428" checkinUser="pensoft" docAuthor="Jochum, Adrienne, Slapnik, Rajko, Kampschulte, Marian, Martels, Gunhild, Heneka, Markus &amp; Pall-Gergely, Barna" docDate="2014" docId="BB8054E1CA9BF0ADCD640C08E6215786" docLanguage="en" docName="ZooKeys 410: 23-40" docOrigin="ZooKeys 410" docSource="http://dx.doi.org/10.3897/zookeys.410.7488" docTitle="Angustopila huoyani Jochum, Slapnik &amp; Pall-Gergely, sp. n." docType="treatment" docUuid="2101F700-9723-422F-B70B-F8D0C2D20345" docUuidSource="ZooBank" docVersion="4" lastPageNumber="28" masterDocId="FFA7FFE8A6732834BC12004CAF57677C" masterDocTitle="A review of the microgastropod genus Systenostoma Bavay &amp; Dautzenberg, 1908 and a new subterranean species from China (Gastropoda, Pulmonata, Hypselostomatidae)" masterLastPageNumber="40" masterPageNumber="23" pageNumber="26" updateTime="1668158519503" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>A review of the microgastropod genus Systenostoma Bavay &amp; Dautzenberg, 1908 and a new subterranean species from China (Gastropoda, Pulmonata, Hypselostomatidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jochum, Adrienne</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Slapnik, Rajko</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kampschulte, Marian</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Martels, Gunhild</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Heneka, Markus</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pall-Gergely, Barna</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>410</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>23</mods:start>
<mods:end>40</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.410.7488</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.410.7488</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-410-23</mods:identifier>
<mods:identifier type="ZBK">E4C040C1939640F18C12D9518F59F668</mods:identifier>
<mods:identifier type="ZooBank">E4C040C1939640F18C12D9518F59F668</mods:identifier>
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<treatment ID-GBIF-Taxon="152053103" LSID="urn:lsid:zoobank.org:act:2101F700-9723-422F-B70B-F8D0C2D20345" httpUri="http://treatment.plazi.org/id/BB8054E1CA9BF0ADCD640C08E6215786" lastPageId="5" lastPageNumber="28" pageId="3" pageNumber="26">
<subSubSection pageId="3" pageNumber="26" type="nomenclature">
<paragraph pageId="3" pageNumber="26">
<taxonomicName LSID="http://zoobank.org/2101F700-9723-422F-B70B-F8D0C2D20345" authority="Jochum, Slapnik &amp; Pall-Gergely" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angustopila huoyani" order="Stylommatophora" pageId="3" pageNumber="26" phylum="Mollusca" rank="species" species="huoyani">
Angustopila huoyani Jochum, Slapnik &amp; 
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
</taxonomicName>
<taxonomicNameLabel pageId="3" pageNumber="26">sp. n.</taxonomicNameLabel>
Figure 4-5, Video 1
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="26" type="type material">
<paragraph pageId="3" pageNumber="26">Type material.</paragraph>
<paragraph pageId="3" pageNumber="26">
China, Hunan, Longshan (龙山县), Huoyan (火焰), Feihu Dong (飞虎洞), (
<geoCoordinate direction="north" orientation="latitude" precision="9" value="29.208834">29°12.53'N</geoCoordinate>
, 
<geoCoordinate direction="east" orientation="longitude" precision="9" value="109.30617">109°18.37'E</geoCoordinate>
, 550 m alt.), soil, leg. Verovnik, 13.04.1997, MCSMNH 50312/1 (holotype), MCSMNH 50312/2-9 (8 paratypes).
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="26" type="diagnosis">
<paragraph pageId="3" pageNumber="26">Diagnosis.</paragraph>
<paragraph pageId="3" pageNumber="26">A tiny, thin-shelled conical snail with very deep and narrow umbilicus, 5 shouldered whorls and two apertural denticles (parietal and palatal).</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="27" pageId="3" pageNumber="26" type="description">
<paragraph pageId="3" pageNumber="26">Description.</paragraph>
<paragraph lastPageId="4" lastPageNumber="27" pageId="3" pageNumber="26">
Shell thin, greyish white, semi opaque; conical, widest at its base, with a homogeneous powdery superficial texture and regularly increasing, shouldered whorls separated by deep suture; smooth with no notable spiral or radial sculpture. It is characterized by very fine irregular axial lamellae and reticulating microgranules 
<pageBreakToken pageId="4" pageNumber="27" start="start">producing</pageBreakToken>
the powdery superficial texture; protoconch shows reticulating granules and recognisable radial lines only at the upper part of the first whorl; aperture semi-circular, slightly oblique from ventral view; peristome very slightly thickened and reflexed; parietal callus adnate (attached to the penultimate whorl); aperture with two well-developed but short denticles, one on the parietal and the other on the palatal side; umbilicus very deep and narrow.
</paragraph>
<caption pageId="4" pageNumber="27">
<paragraph pageId="4" pageNumber="27">
Video 1. Micro-CT Video of 
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angustopila huoyani" order="Stylommatophora" pageId="4" pageNumber="27" phylum="Mollusca" rank="species" species="huoyani">Angustopila huoyani</taxonomicName>
sp. n. Video: Markus Heneka. http://www.pensoft.net/J_FILES/1/articles/7488/Jochum_Video_1.avi
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="4" pageNumber="27" type="measurements">
<paragraph pageId="4" pageNumber="27">Measurements.</paragraph>
<paragraph pageId="4" pageNumber="27">See Table 1.</paragraph>
<caption ID-Table-UUID="8F285F054D5BAD995C377CBE8761BB4E" httpUri="http://table.plazi.org/id/8F285F054D5BAD995C377CBE8761BB4E" pageId="4" pageNumber="27">
<paragraph pageId="4" pageNumber="27">
Table 1. Shell measurements (mm) for 
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angustopila huoyani" order="Stylommatophora" pageId="4" pageNumber="27" phylum="Mollusca" rank="species" species="huoyani">Angustopila huoyani</taxonomicName>
sp. n. from the type locality. SH - shell height, SW - shell width, AH - aperture height, AW - aperture width, SW/ 
<normalizedToken originalValue="SH×">SHx</normalizedToken>
100 - shell width shared with shell height and multiplied 100, AW/ 
<normalizedToken originalValue="AH×">AHx</normalizedToken>
100 - aperture width shared with aperture height and multiplied 100).
</paragraph>
</caption>
<paragraph pageId="4" pageNumber="27">
<table pageId="4" pageNumber="27">
<tr pageId="4" pageNumber="27">
<th colspan="1" pageId="4" pageNumber="27" rowspan="1">SH</th>
<th colspan="1" pageId="4" pageNumber="27" rowspan="1">SW</th>
<th colspan="1" pageId="4" pageNumber="27" rowspan="1">AH</th>
<th colspan="1" pageId="4" pageNumber="27" rowspan="1">AW</th>
<th colspan="1" pageId="4" pageNumber="27" rowspan="1">
SW/ 
<normalizedToken originalValue="SH×">SHx</normalizedToken>
100
</th>
<th colspan="1" pageId="4" pageNumber="27" rowspan="1">
AW/ 
<normalizedToken originalValue="AH×">AHx</normalizedToken>
100
</th>
<th colspan="1" pageId="4" pageNumber="27" rowspan="1">specimen</th>
</tr>
</table>
</paragraph>
<caption ID-Table-UUID="AFF530F630C73C5AEE61D76E007B6E64" httpUri="http://table.plazi.org/id/AFF530F630C73C5AEE61D76E007B6E64" pageId="4" pageNumber="27">
<paragraph pageId="4" pageNumber="27">
Table 2. Average. minimum value (min). maximum value (max). variance of values (var) and standard deviation of a set of values (stdev) for 
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angustopila huoyani" order="Stylommatophora" pageId="4" pageNumber="27" phylum="Mollusca" rank="species" species="huoyani">Angustopila huoyani</taxonomicName>
sp. n. N = 30.
</paragraph>
</caption>
<paragraph pageId="4" pageNumber="27">
<table pageId="4" pageNumber="27">
<tr pageId="4" pageNumber="27">
<th colspan="1" pageId="4" pageNumber="27" rowspan="1">SH</th>
<th colspan="1" pageId="4" pageNumber="27" rowspan="1">SW</th>
<th colspan="1" pageId="4" pageNumber="27" rowspan="1">AH</th>
<th colspan="1" pageId="4" pageNumber="27" rowspan="1">AW</th>
<th colspan="1" pageId="4" pageNumber="27" rowspan="1">
SW/ 
<normalizedToken originalValue="SH×">SHx</normalizedToken>
100
</th>
<th colspan="1" pageId="4" pageNumber="27" rowspan="1">
AW/ 
<normalizedToken originalValue="AH×">AHx</normalizedToken>
100
</th>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="28" pageId="4" pageNumber="27" type="differential diagnosis">
<paragraph pageId="4" pageNumber="27">Differential diagnosis.</paragraph>
<paragraph pageId="5" pageNumber="28">
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tonkinospira defixa" order="Stylommatophora" pageId="5" pageNumber="28" phylum="Mollusca" rank="species" species="defixa">
<pageBreakToken pageId="5" pageNumber="28" start="start">Tonkinospira</pageBreakToken>
defixa
</taxonomicName>
, 
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tonkinospira pulverea" order="Stylommatophora" pageId="5" pageNumber="28" phylum="Mollusca" rank="species" species="pulverea">Tonkinospira pulverea</taxonomicName>
, 
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tonkinospira pauperrima" order="Stylommatophora" pageId="5" pageNumber="28" phylum="Mollusca" rank="species" species="pauperrima">Tonkinospira pauperrima</taxonomicName>
and 
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tonkinospira depressa" order="Stylommatophora" pageId="5" pageNumber="28" phylum="Mollusca" rank="species" species="depressa">Tonkinospira depressa</taxonomicName>
are much larger than the new species, have reticulated sculpture and lack denticles in the aperture. Moreover, 
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tonkinospira defixa" order="Stylommatophora" pageId="5" pageNumber="28" phylum="Mollusca" rank="species" species="defixa">Tonkinospira defixa</taxonomicName>
has a more depressed spire, fewer whorls, wider umbilicus and slightly keeled body whorl; 
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tonkinospira pulverea" order="Stylommatophora" pageId="5" pageNumber="28" phylum="Mollusca" rank="species" species="pulverea">Tonkinospira pulverea</taxonomicName>
has fewer whorls, a comparatively larger aperture, somewhat keeled, wider body whorl and its umbilicus is partly closed by the apertural margin; 
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tonkinospira pauperrima" order="Stylommatophora" pageId="5" pageNumber="28" phylum="Mollusca" rank="species" species="pauperrima">Tonkinospira pauperrima</taxonomicName>
shows increased bulging in whorl configuration; 
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tonkinospira depressa" order="Stylommatophora" pageId="5" pageNumber="28" phylum="Mollusca" rank="species" species="depressa">Tonkinospira depressa</taxonomicName>
has a lower spire, slightly keeled body whorl and a large aperture without denticles. 
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Hypselostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypselostoma" order="Stylommatophora" pageId="5" pageNumber="28" phylum="Mollusca" rank="genus">Hypselostoma</taxonomicName>
(?) edentata also lacks denticles in the aperture and possesses a very wide, laterally compressed body whorl. 
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angustopila tamlod" order="Stylommatophora" pageId="5" pageNumber="28" phylum="Mollusca" rank="species" species="tamlod">Angustopila tamlod</taxonomicName>
, the most similar species, is slightly smaller, has fewer whorls, wider umbilicus and obvious spiral striation on the teleoconch. 
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angustopila concava" order="Stylommatophora" pageId="5" pageNumber="28" phylum="Mollusca" rank="species" species="concava">Angustopila concava</taxonomicName>
has a much wider body whorl than that of 
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angustopila huoyani" order="Stylommatophora" pageId="5" pageNumber="28" phylum="Mollusca" rank="species" species="huoyani">Angustopila huoyani</taxonomicName>
, has weaker apertural denticles and prominent spiral sculpture. 
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angustopila elevata" order="Stylommatophora" pageId="5" pageNumber="28" phylum="Mollusca" rank="species" species="elevata">Angustopila elevata</taxonomicName>
has no denticles in the aperture and possesses a wider umbilicus and spirally striated shell. 
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angustopila neglecta" order="Stylommatophora" pageId="5" pageNumber="28" phylum="Mollusca" rank="species" species="neglecta">Angustopila neglecta</taxonomicName>
(see also notes under that species) has a wider umbilicus and more rapidly growing whorls, resulting in a comparatively wider body whorl than in the new species. 
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angustopila neglecta" order="Stylommatophora" pageId="5" pageNumber="28" phylum="Mollusca" rank="species" species="neglecta">Angustopila neglecta</taxonomicName>
also has spiral lines on the shell and its sinulus is wider. Shell characters and ecological information of all 
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angustopila" order="Stylommatophora" pageId="5" pageNumber="28" phylum="Mollusca" rank="genus">Angustopila</taxonomicName>
species are presented in Table 3.
</paragraph>
<caption pageId="5" pageNumber="28">
<paragraph pageId="5" pageNumber="28">
Table 3. The most important morphological traits and ecological information for 
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angustopila huoyani" order="Stylommatophora" pageId="5" pageNumber="28" phylum="Mollusca" rank="species" species="huoyani">Angustopila huoyani</taxonomicName>
sp. n. and its congeners extracted from the literature.
</paragraph>
</caption>
<paragraph pageId="5" pageNumber="28">
<table pageId="5" pageNumber="28">
<tr pageId="5" pageNumber="28">
<th colspan="1" pageId="5" pageNumber="28" rowspan="1">
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angustopila concava" order="Stylommatophora" pageId="5" pageNumber="28" phylum="Mollusca" rank="species" species="concava">Angustopila concava</taxonomicName>
</th>
<th colspan="1" pageId="5" pageNumber="28" rowspan="1">
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angustopila elevata" order="Stylommatophora" pageId="5" pageNumber="28" phylum="Mollusca" rank="species" species="elevata">Angustopila elevata</taxonomicName>
</th>
<th colspan="1" pageId="5" pageNumber="28" rowspan="1">
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angustopila huoyani" order="Stylommatophora" pageId="5" pageNumber="28" phylum="Mollusca" rank="species" species="huoyani">Angustopila huoyani</taxonomicName>
sp. n.
</th>
<th colspan="1" pageId="5" pageNumber="28" rowspan="1">
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angustopila neglecta" order="Stylommatophora" pageId="5" pageNumber="28" phylum="Mollusca" rank="species" species="neglecta">Angustopila neglecta</taxonomicName>
</th>
<th colspan="1" pageId="5" pageNumber="28" rowspan="1">
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angustopila tamlod" order="Stylommatophora" pageId="5" pageNumber="28" phylum="Mollusca" rank="species" species="tamlod">Angustopila tamlod</taxonomicName>
</th>
</tr>
<tr pageId="5" pageNumber="28">
<td colspan="1" pageId="5" pageNumber="28" rowspan="1">Shell colour</td>
</tr>
<tr pageId="5" pageNumber="28">
<td colspan="1" pageId="5" pageNumber="28" rowspan="1">Teleoconch sculpture</td>
</tr>
<tr pageId="5" pageNumber="28">
<td colspan="1" pageId="5" pageNumber="28" rowspan="1">Protoconch sculpture</td>
</tr>
<tr pageId="5" pageNumber="28">
<td colspan="1" pageId="5" pageNumber="28" rowspan="1">Aperture shape/ peristome</td>
</tr>
<tr pageId="5" pageNumber="28">
<td colspan="1" pageId="5" pageNumber="28" rowspan="1">Aperture dentition</td>
</tr>
<tr pageId="5" pageNumber="28">
<td colspan="1" pageId="5" pageNumber="28" rowspan="1">Umbilicus</td>
</tr>
<tr pageId="5" pageNumber="28">
<td colspan="1" pageId="5" pageNumber="28" rowspan="1">Whorl number</td>
</tr>
<tr pageId="5" pageNumber="28">
<td colspan="1" pageId="5" pageNumber="28" rowspan="1">Shell height (mm)</td>
</tr>
<tr pageId="5" pageNumber="28">
<td colspan="1" pageId="5" pageNumber="28" rowspan="1">Ecology</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="28" type="etymology">
<paragraph pageId="5" pageNumber="28">Etymology.</paragraph>
<paragraph pageId="5" pageNumber="28">The new species is named after the Gorges of Huoyan, where the type locality is located.</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="28" type="distribution">
<paragraph pageId="5" pageNumber="28">Distribution.</paragraph>
<paragraph pageId="5" pageNumber="28">The new species is known from the type locality only.</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="28" type="ecology">
<paragraph pageId="5" pageNumber="28">Ecology.</paragraph>
<paragraph pageId="5" pageNumber="28">
The new species is known only from the Feihu Dong (&quot;Cave of the Wind Tiger&quot;). 
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angustopila huoyani" order="Stylommatophora" pageId="5" pageNumber="28" phylum="Mollusca" rank="species" species="huoyani">Angustopila huoyani</taxonomicName>
were culled from samples of rocky-loamy substrate collected in the entrance corridor of the cave. It is highly likely that the distribution of 
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angustopila huoyani" order="Stylommatophora" pageId="5" pageNumber="28" phylum="Mollusca" rank="species" species="huoyani">Angustopila huoyani</taxonomicName>
sp. n. is restricted to this cave only.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="28" type="conservational status">
<paragraph pageId="5" pageNumber="28">Conservational status.</paragraph>
<paragraph pageId="5" pageNumber="28">
Our knowledge of the biogeography of the genus is very limited. However, we assume that most 
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angustopila" order="Stylommatophora" pageId="5" pageNumber="28" phylum="Mollusca" rank="genus">Angustopila</taxonomicName>
, especially the cave-dwelling species, are narrow-range endemics. Since extreme endemism always makes species vulnerable to human encroachment, this species warrants conservation priority. Currently, no direct threats are known.
</paragraph>
</subSubSection>
</treatment>
</document>