<documentid="5536701658FE0831C3545517B0EDA10B"ID-DOI="10.37828/em.2018.16.12"ID-ISSN="2336-9744"ID-Zenodo-Dep="13360980"ID-ZooBank="urn:lsid:zoobank.org:pub:6C3A5085-F624-4D0A-9F10-4EBB9AF19F01"IM.bibliography_approvedBy="felipe"IM.illustrations_approvedBy="felipe"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="guilherme"IM.taxonomicNames_approvedBy="guilherme"IM.treatmentCitations_approvedBy="felipe"IM.treatments_approvedBy="felipe"checkinTime="1720661392767"checkinUser="felipe"docAuthor="Prószyński, Jerzy"docDate="2018"docId="03807147FF91FF9BFDA2FBCDFF211EF6"docLanguage="en"docName="EcolMontenegrina.16.130-179.pdf"docOrigin="Ecologica Montenegrina 16"docSource="http://dx.doi.org/10.37828/em.2018.16.12"docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.3:EcolMontenegrina.2017-.journal_article"docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE"docStyleName="EcolMontenegrina.2017-.journal_article"docStyleVersion="3"docTitle="Emertonius Peckham & Peckham 1892"docType="treatment"docVersion="4"lastPageNumber="165"masterDocId="FFB9093FFFB2FFB8FFA1FFC2FFC41915"masterDocTitle="Review of genera Evarcha and Nigorella, with comments on Emertonius, Padilothorax, Stagetillus, and description of five new genera and two new species (Araneae: Salticidae) *"masterLastPageNumber="179"masterPageNumber="130"pageNumber="165"updateTime="1724330662745"updateUser="ExternalLinkService"zenodo-license-document="CC-BY-4.0">
<mods:titleid="63946B64C62A2B17203F1DB566C858E3">Review of genera Evarcha and Nigorella, with comments on Emertonius, Padilothorax, Stagetillus, and description of five new genera and two new species (Araneae: Salticidae) *</mods:title>
<taxonomicNameid="4C29BBD2FF91FF9BFDE7FBCDFBF71D3C"ID-CoL="4B8V"authority="Peckham & Peckham, 1892"authorityName="Peckham & Peckham"authorityYear="1892"box="[582,1075,1039,1065]"class="Arachnida"family="Salticidae"genus="Emertonius"kingdom="Animalia"order="Araneae"pageId="35"pageNumber="165"phylum="Arthropoda"rank="genus">
<figureCitationid="1312DCD4FF91FF9BFD88FBF2FD241D5E"box="[553,736,1072,1099]"captionStart="Figure 19"captionStartId="31.[151,226,1598,1622]"captionTargetBox="[172,1419,338,1531]"captionTargetId="figure-61@31.[157,1433,855,1192]"captionTargetPageId="31"captionText="Figure 19. Diagnostic criteria of genera of informal group MYRMARACHNINES. A - Emertonius exasperans - type species, A1 - Edward's opinion on identity of Emertonius with Myrmarachne, B - Myrmarachne melanocephala - type species, C - Emertonius exasperans, D - Emertonius shelfordi, E - Myrmage, F - Myrmatheca, G - Toxeus epigealis, H - Toxeus maxillosus, I - Myrmavola, J - Myrmarachne acromegalis, K - Myrmarachne assimilis, L - Myrmarachne biseratensis, M - Myrmarachne cornuta, N - Myrmarachne macrognatha, O - Myrmarachne hanoi. For characters of more species see Figs 21 and 22."figureDoi="http://doi.org/10.5281/zenodo.13361071"httpUri="https://zenodo.org/record/13361071/files/figure.png"pageId="35"pageNumber="165">Figures 19A, C</figureCitation>
<figureCitationid="1312DCD4FF91FF9BFD52FBF2FCCB1D5E"box="[755,783,1072,1099]"captionStart="Figure 21"captionStartId="33.[151,226,2021,2045]"captionTargetBox="[175,1418,531,1982]"captionTargetId="figure-142@33.[161,1428,1226,1562]"captionTargetPageId="33"captionText="Figure 21-I. For figure caption see under Figure 21-II."figureDoi="http://doi.org/10.5281/zenodo.13361077"httpUri="https://zenodo.org/record/13361077/files/figure.png"pageId="35"pageNumber="165">21</figureCitation>
<taxonomicNameid="4C29BBD2FF91FF9BFE9DFBB7FC641D9B"ID-CoL="39GP4"authority="Peckham & Peckham, 1892"authorityName="Peckham & Peckham"authorityYear="1892"box="[316,928,1139,1166]"class="Arachnida"family="Salticidae"genus="Emertonius"kingdom="Animalia"order="Araneae"pageId="35"pageNumber="165"phylum="Arthropoda"rank="species"species="exasperans">
<figureCitationid="1312DCD4FF91FF9BFD5DFB57FCB31DA5"box="[764,887,1173,1200]"captionStart="Figure 19"captionStartId="31.[151,226,1598,1622]"captionTargetBox="[172,1419,338,1531]"captionTargetId="figure-61@31.[157,1433,855,1192]"captionTargetPageId="31"captionText="Figure 19. Diagnostic criteria of genera of informal group MYRMARACHNINES. A - Emertonius exasperans - type species, A1 - Edward's opinion on identity of Emertonius with Myrmarachne, B - Myrmarachne melanocephala - type species, C - Emertonius exasperans, D - Emertonius shelfordi, E - Myrmage, F - Myrmatheca, G - Toxeus epigealis, H - Toxeus maxillosus, I - Myrmavola, J - Myrmarachne acromegalis, K - Myrmarachne assimilis, L - Myrmarachne biseratensis, M - Myrmarachne cornuta, N - Myrmarachne macrognatha, O - Myrmarachne hanoi. For characters of more species see Figs 21 and 22."figureDoi="http://doi.org/10.5281/zenodo.13361071"httpUri="https://zenodo.org/record/13361071/files/figure.png"pageId="35"pageNumber="165">Figs. 19A</figureCitation>
<figureCitationid="1312DCD4FF91FF9BFC28FB57FC6D1DA5"box="[905,937,1173,1200]"captionStart="Figure 21"captionStartId="33.[151,226,2021,2045]"captionTargetBox="[175,1418,531,1982]"captionTargetId="figure-142@33.[161,1428,1226,1562]"captionTargetPageId="33"captionText="Figure 21-I. For figure caption see under Figure 21-II."figureDoi="http://doi.org/10.5281/zenodo.13361077"httpUri="https://zenodo.org/record/13361077/files/figure.png"pageId="35"pageNumber="165">21</figureCitation>
<figureCitationid="1312DCD4FF91FF9BFC1BFB57FC1C1DA5"box="[954,984,1173,1200]"captionStart="Figure 25"captionStartId="38.[151,226,1916,1940]"captionTargetBox="[161,1393,300,1863]"captionTargetId="figure-30@38.[151,1416,760,1095]"captionTargetPageId="38"captionText="Figure 25-I. For figure caption see under Figure 25-II."figureDoi="http://doi.org/10.5281/zenodo.13361097"httpUri="https://zenodo.org/record/13361097/files/figure.png"pageId="35"pageNumber="165">25</figureCitation>
<figureCitationid="1312DCD4FF91FF9BFDBCFB75FD431DC7"box="[541,647,1207,1234]"captionStart="Figure 19"captionStartId="31.[151,226,1598,1622]"captionTargetBox="[172,1419,338,1531]"captionTargetId="figure-61@31.[157,1433,855,1192]"captionTargetPageId="31"captionText="Figure 19. Diagnostic criteria of genera of informal group MYRMARACHNINES. A - Emertonius exasperans - type species, A1 - Edward's opinion on identity of Emertonius with Myrmarachne, B - Myrmarachne melanocephala - type species, C - Emertonius exasperans, D - Emertonius shelfordi, E - Myrmage, F - Myrmatheca, G - Toxeus epigealis, H - Toxeus maxillosus, I - Myrmavola, J - Myrmarachne acromegalis, K - Myrmarachne assimilis, L - Myrmarachne biseratensis, M - Myrmarachne cornuta, N - Myrmarachne macrognatha, O - Myrmarachne hanoi. For characters of more species see Figs 21 and 22."figureDoi="http://doi.org/10.5281/zenodo.13361071"httpUri="https://zenodo.org/record/13361071/files/figure.png"pageId="35"pageNumber="165">Fig. 19B</figureCitation>
and elsewhere in literature). Palps confront to general appearance of 180 species of informal group MYRMARACHNINES, with minor differences, insufficient to separate genera and/or species. Internal structure of epigyne (
<figureCitationid="1312DCD4FF91FF9BFCADFB38FCA11C00"box="[780,869,1274,1301]"captionStart="Figure 19"captionStartId="31.[151,226,1598,1622]"captionTargetBox="[172,1419,338,1531]"captionTargetId="figure-61@31.[157,1433,855,1192]"captionTargetPageId="31"captionText="Figure 19. Diagnostic criteria of genera of informal group MYRMARACHNINES. A - Emertonius exasperans - type species, A1 - Edward's opinion on identity of Emertonius with Myrmarachne, B - Myrmarachne melanocephala - type species, C - Emertonius exasperans, D - Emertonius shelfordi, E - Myrmage, F - Myrmatheca, G - Toxeus epigealis, H - Toxeus maxillosus, I - Myrmavola, J - Myrmarachne acromegalis, K - Myrmarachne assimilis, L - Myrmarachne biseratensis, M - Myrmarachne cornuta, N - Myrmarachne macrognatha, O - Myrmarachne hanoi. For characters of more species see Figs 21 and 22."figureDoi="http://doi.org/10.5281/zenodo.13361071"httpUri="https://zenodo.org/record/13361071/files/figure.png"pageId="35"pageNumber="165">Figs 19</figureCitation>
<figureCitationid="1312DCD4FF91FF9BFEECFADEFE601C22"box="[333,420,1308,1335]"captionStart="Figure 19"captionStartId="31.[151,226,1598,1622]"captionTargetBox="[172,1419,338,1531]"captionTargetId="figure-61@31.[157,1433,855,1192]"captionTargetPageId="31"captionText="Figure 19. Diagnostic criteria of genera of informal group MYRMARACHNINES. A - Emertonius exasperans - type species, A1 - Edward's opinion on identity of Emertonius with Myrmarachne, B - Myrmarachne melanocephala - type species, C - Emertonius exasperans, D - Emertonius shelfordi, E - Myrmage, F - Myrmatheca, G - Toxeus epigealis, H - Toxeus maxillosus, I - Myrmavola, J - Myrmarachne acromegalis, K - Myrmarachne assimilis, L - Myrmarachne biseratensis, M - Myrmarachne cornuta, N - Myrmarachne macrognatha, O - Myrmarachne hanoi. For characters of more species see Figs 21 and 22."figureDoi="http://doi.org/10.5281/zenodo.13361071"httpUri="https://zenodo.org/record/13361071/files/figure.png"pageId="35"pageNumber="165">Figs 19</figureCitation>
<taxonomicNameid="4C29BBD2FF91FF9BFE86FAFDFE6A1C4D"authorityName="Peckham & Peckham"authorityYear="1892"box="[295,430,1343,1368]"class="Arachnida"family="Salticidae"genus="Emertonius"kingdom="Animalia"order="Araneae"pageId="35"pageNumber="165"phylum="Arthropoda"rank="genus">
<figureCitationid="1312DCD4FF91FF9BFD21FA9DFD341C6F"box="[640,752,1375,1402]"captionStart="Figure 19"captionStartId="31.[151,226,1598,1622]"captionTargetBox="[172,1419,338,1531]"captionTargetId="figure-61@31.[157,1433,855,1192]"captionTargetPageId="31"captionText="Figure 19. Diagnostic criteria of genera of informal group MYRMARACHNINES. A - Emertonius exasperans - type species, A1 - Edward's opinion on identity of Emertonius with Myrmarachne, B - Myrmarachne melanocephala - type species, C - Emertonius exasperans, D - Emertonius shelfordi, E - Myrmage, F - Myrmatheca, G - Toxeus epigealis, H - Toxeus maxillosus, I - Myrmavola, J - Myrmarachne acromegalis, K - Myrmarachne assimilis, L - Myrmarachne biseratensis, M - Myrmarachne cornuta, N - Myrmarachne macrognatha, O - Myrmarachne hanoi. For characters of more species see Figs 21 and 22."figureDoi="http://doi.org/10.5281/zenodo.13361071"httpUri="https://zenodo.org/record/13361071/files/figure.png"pageId="35"pageNumber="165">Figs 19A</figureCitation>
<figureCitationid="1312DCD4FF91FF9BFC27FA43FC301C89"box="[902,1012,1409,1436]"captionStart="Figure 19"captionStartId="31.[151,226,1598,1622]"captionTargetBox="[172,1419,338,1531]"captionTargetId="figure-61@31.[157,1433,855,1192]"captionTargetPageId="31"captionText="Figure 19. Diagnostic criteria of genera of informal group MYRMARACHNINES. A - Emertonius exasperans - type species, A1 - Edward's opinion on identity of Emertonius with Myrmarachne, B - Myrmarachne melanocephala - type species, C - Emertonius exasperans, D - Emertonius shelfordi, E - Myrmage, F - Myrmatheca, G - Toxeus epigealis, H - Toxeus maxillosus, I - Myrmavola, J - Myrmarachne acromegalis, K - Myrmarachne assimilis, L - Myrmarachne biseratensis, M - Myrmarachne cornuta, N - Myrmarachne macrognatha, O - Myrmarachne hanoi. For characters of more species see Figs 21 and 22."figureDoi="http://doi.org/10.5281/zenodo.13361071"httpUri="https://zenodo.org/record/13361071/files/figure.png"pageId="35"pageNumber="165">Fig. 19B</figureCitation>
). The most important difference is internal structure of epigyne, in
<taxonomicNameid="4C29BBD2FF91FF9BFD82FA66FD6E1CA8"authorityName="Peckham & Peckham"authorityYear="1892"box="[547,682,1444,1469]"class="Arachnida"family="Salticidae"genus="Emertonius"kingdom="Animalia"order="Araneae"pageId="35"pageNumber="165"phylum="Arthropoda"rank="genus">
<figureCitationid="1312DCD4FF91FF9BFCEAFA07FC6C1CF5"box="[843,936,1477,1504]"captionStart="Figure 19"captionStartId="31.[151,226,1598,1622]"captionTargetBox="[172,1419,338,1531]"captionTargetId="figure-61@31.[157,1433,855,1192]"captionTargetPageId="31"captionText="Figure 19. Diagnostic criteria of genera of informal group MYRMARACHNINES. A - Emertonius exasperans - type species, A1 - Edward's opinion on identity of Emertonius with Myrmarachne, B - Myrmarachne melanocephala - type species, C - Emertonius exasperans, D - Emertonius shelfordi, E - Myrmage, F - Myrmatheca, G - Toxeus epigealis, H - Toxeus maxillosus, I - Myrmavola, J - Myrmarachne acromegalis, K - Myrmarachne assimilis, L - Myrmarachne biseratensis, M - Myrmarachne cornuta, N - Myrmarachne macrognatha, O - Myrmarachne hanoi. For characters of more species see Figs 21 and 22."figureDoi="http://doi.org/10.5281/zenodo.13361071"httpUri="https://zenodo.org/record/13361071/files/figure.png"pageId="35"pageNumber="165">Fig. 19</figureCitation>
the spermathecae have characteristic form of two parallel, sclerotized “pipe- like” structures, running medially along the whole length of epigyne, with striking detour in anterior third of the "pipe " in a form of a loop, entangled knot, or double spiral (sample of rich variety of these is shown on
<figureCitationid="1312DCD4FF91FF9BFAB3F9E8FAAF1F50"box="[1298,1387,1578,1605]"captionStart="Figure 19"captionStartId="31.[151,226,1598,1622]"captionTargetBox="[172,1419,338,1531]"captionTargetId="figure-61@31.[157,1433,855,1192]"captionTargetPageId="31"captionText="Figure 19. Diagnostic criteria of genera of informal group MYRMARACHNINES. A - Emertonius exasperans - type species, A1 - Edward's opinion on identity of Emertonius with Myrmarachne, B - Myrmarachne melanocephala - type species, C - Emertonius exasperans, D - Emertonius shelfordi, E - Myrmage, F - Myrmatheca, G - Toxeus epigealis, H - Toxeus maxillosus, I - Myrmavola, J - Myrmarachne acromegalis, K - Myrmarachne assimilis, L - Myrmarachne biseratensis, M - Myrmarachne cornuta, N - Myrmarachne macrognatha, O - Myrmarachne hanoi. For characters of more species see Figs 21 and 22."figureDoi="http://doi.org/10.5281/zenodo.13361071"httpUri="https://zenodo.org/record/13361071/files/figure.png"pageId="35"pageNumber="165">Figs 19</figureCitation>
J-O, 22A-E1). Transparent, membranous copulatory ducts, very entangled and long, running from copulatory openings to junction with spermathecae, are usually overlooked and not mentioned in descriptions because they are visible only on cleared and stained (by Chlorazol Black B) microscope slides (even perfect photographs by Yamasaki do not display them -
<figureCitationid="1312DCD4FF91FF9BFD5EF973FC9F1FD9"box="[767,859,1713,1740]"captionStart="Figure 19"captionStartId="31.[151,226,1598,1622]"captionTargetBox="[172,1419,338,1531]"captionTargetId="figure-61@31.[157,1433,855,1192]"captionTargetPageId="31"captionText="Figure 19. Diagnostic criteria of genera of informal group MYRMARACHNINES. A - Emertonius exasperans - type species, A1 - Edward's opinion on identity of Emertonius with Myrmarachne, B - Myrmarachne melanocephala - type species, C - Emertonius exasperans, D - Emertonius shelfordi, E - Myrmage, F - Myrmatheca, G - Toxeus epigealis, H - Toxeus maxillosus, I - Myrmavola, J - Myrmarachne acromegalis, K - Myrmarachne assimilis, L - Myrmarachne biseratensis, M - Myrmarachne cornuta, N - Myrmarachne macrognatha, O - Myrmarachne hanoi. For characters of more species see Figs 21 and 22."figureDoi="http://doi.org/10.5281/zenodo.13361071"httpUri="https://zenodo.org/record/13361071/files/figure.png"pageId="35"pageNumber="165">Figs 19</figureCitation>
<taxonomicNameid="4C29BBD2FF91FF9BFD48F8D5FCB41E25"authorityName="Peckham & Peckham"authorityYear="1892"box="[745,880,1815,1840]"class="Arachnida"family="Salticidae"genus="Emertonius"kingdom="Animalia"order="Araneae"pageId="35"pageNumber="165"phylum="Arthropoda"rank="genus">
and other "lesser genera" require further comparative research in order to distinguish it from intra-populational variation, whose sample is shown on
<figureCitationid="1312DCD4FF91FF9BFF36F89BFF221E61"box="[151,230,1881,1908]"captionStart="Figure 26"captionStartId="39.[151,226,1800,1824]"captionTargetBox="[151,1430,1207,1748]"captionTargetId="figure-278@39.[151,1436,1198,1763]"captionTargetPageId="39"captionText="Figure 26-I. For figure caption see under Figure 26-II."figureDoi="http://doi.org/10.5281/zenodo.13361103"httpUri="https://zenodo.org/record/13361103/files/figure.png"pageId="35"pageNumber="165">Fig 26</figureCitation>
<taxonomicNameid="4C29BBD2FF91FF9BFE10F8B9FC771E85"authority="Peckham & Peckham, 1892"authorityName="Peckham & Peckham"authorityYear="1892"box="[433,947,1914,1936]"class="Arachnida"family="Salticidae"genus="Emertonius"kingdom="Animalia"order="Araneae"pageId="35"pageNumber="165"phylum="Arthropoda"rank="species"species="exasperans">
<taxonomicNameid="4C29BBD2FF91FF9BFE0BF855FC191EB9"authority="(Edmunds & Proszynski, 2003)"authorityName="Prószyński"authorityYear="2018"baseAuthorityName="Edmunds & Proszynski"baseAuthorityYear="2003"box="[426,989,1942,1964]"class="Arachnida"family="Salticidae"genus="Emertonius"kingdom="Animalia"order="Araneae"pageId="35"pageNumber="165"phylum="Arthropoda"rank="species"species="malayanus"status="comb. nov.">
<bibRefCitationid="EFB8BDA0FF91FF9BFD00F854FC121EB9"author="Edmunds M. & Proszynski J."box="[673,982,1942,1964]"pageId="35"pageNumber="165"pagination="297 - 323"refId="ref26908"refString="Edmunds M. & Proszynski J. (2003). On a collection of Myrmarachne spiders (Araneae: Salticidae) from peninsular Malaya. Bulletin of the British Arachnological Society 12: 297 - 323."type="journal article"year="2003">Edmunds & Prószyński, 2003</bibRefCitation>
<taxonomicNameid="4C29BBD2FF91FF9BFCEAF870FAF31ED2"authority="(Peckham & Peckham, 1907)"authorityName="Prószyński"authorityYear="2018"baseAuthorityName="Peckham & Peckham"baseAuthorityYear="1907"box="[843,1335,1969,1991]"class="Arachnida"family="Salticidae"genus="Emertonius"kingdom="Animalia"order="Araneae"pageId="35"pageNumber="165"phylum="Arthropoda"rank="species"species="shelfordi"status="comb. nov.">