465 lines
49 KiB
XML
465 lines
49 KiB
XML
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<document id="3F4319E29BEBA71A0651F328C7E924AC" ID-DOI="10.1080/002229300299570" ID-ISSN="1464-5262" ID-Zenodo-Dep="5279608" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630026790407" checkinUser="felipe" docAuthor="Zacharda, Miloslav" docDate="2000" docId="03DE87E2FA4A1245FE1BFBFEFE7DFF7E" docLanguage="en" docName="JNatHist.34.3.463-478.pdf" docOrigin="Journal of Natural History 34 (3)" docSource="http://www.tandfonline.com/doi/abs/10.1080/002229300299570" docStyle="DocumentStyle:B6EF65DE5B4D2965D4D236BA736D172F.2:JNatHist.2000-2002.journal_article.2cover" docStyleId="B6EF65DE5B4D2965D4D236BA736D172F" docStyleName="JNatHist.2000-2002.journal_article.2cover" docStyleVersion="2" docTitle="Troglocheles aggerata Zacharda 2000, n. sp." docType="treatment" docVersion="3" lastPageNumber="476" masterDocId="FFE7FF9AFA411255FF94FA70FF9FFF9B" masterDocTitle="New species of the genus Troglocheles (Acari: Prostigmata: Rhagidiidae) from Oetztal Alps, Tyrol, with a key to adult species of the genus" masterLastPageNumber="478" masterPageNumber="463" pageNumber="472" updateTime="1699138067013" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="006ADDB931EADAF4302384AECCB6C2CF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="29CBD6CF6E8BC93D9A9D6DEA93EACA23">
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<mods:title id="D3972AC5786A314539644E214180E25D">New species of the genus Troglocheles (Acari: Prostigmata: Rhagidiidae) from Oetztal Alps, Tyrol, with a key to adult species of the genus</mods:title>
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</mods:titleInfo>
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<mods:name id="AC4223E1430E84199352CAE60DA88CB7" type="personal">
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<mods:roleTerm id="DADA9F4A1C06F265876C7D1D93AE7561">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6E42270A13D80CA56934C78DF928BE06">Zacharda, Miloslav</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="A7FE383407035F7AE4845B90EA3F4377">text</mods:typeOfResource>
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<mods:relatedItem id="4667315D22399C2F2ABAF938A6FD4DFA" type="host">
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<mods:titleInfo id="CAE48EEB65B272084A3A1E16C1DD45F2">
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<mods:title id="B4017832147A9B1E4E1C81C3D5B436C4">Journal of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part id="AF50F82E0558A889CE0B0BA11D585EC3">
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<mods:date id="BCA91DA292E0A462B67B37BCAA3979E4">2000</mods:date>
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<mods:detail id="239334A2419B207D2CA5FAE5A474B923" type="pubDate">
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<mods:number id="AADEC9C2744690E6B4ED58781407E4C1">2000-03-31</mods:number>
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</mods:detail>
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<mods:detail id="B9B52D6DAF9FE16B2DA8065C302BFBED" type="volume">
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<mods:number id="E8B7BC3100430DD7C359AE301713C05E">34</mods:number>
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</mods:detail>
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<mods:detail id="10DC46EF0E7E249FBEE194B81D7900F9" type="issue">
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<mods:number id="153894454D2D8420F144B2FC9AC42299">3</mods:number>
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</mods:detail>
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<mods:extent id="7BDD1AA37A0752B48BEE96AE33BD5A7A" unit="page">
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<mods:start id="6968C0DB56E42A39679CB1BC2B0149F0">463</mods:start>
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<mods:end id="BA0BA9221CE1AFDD17641B342BB92F64">478</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location id="C211C8CFAB7A5932A86E25DA584FACC2">
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<mods:url id="B732554FDEF4EFF49A609B10241437D1">http://www.tandfonline.com/doi/abs/10.1080/002229300299570</mods:url>
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</mods:location>
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<mods:classification id="E33ADD18CAAB7E006F94DAF7A410A51C">journal article</mods:classification>
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<mods:identifier id="87CC399ECD2D2B4E2F8F9CDE9082E26F" type="DOI">10.1080/002229300299570</mods:identifier>
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<mods:identifier id="CA5365829529401659588D7A25C1E205" type="ISSN">1464-5262</mods:identifier>
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<mods:identifier id="1663A8883799ED4E0EA6926A7073EBEE" type="Zenodo-Dep">5279608</mods:identifier>
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</mods:mods>
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<treatment id="03DE87E2FA4A1245FE1BFBFEFE7DFF7E" ID-DOI="http://doi.org/10.5281/zenodo.5281671" ID-Zenodo-Dep="5281671" LSID="urn:lsid:plazi:treatment:03DE87E2FA4A1245FE1BFBFEFE7DFF7E" httpUri="http://treatment.plazi.org/id/03DE87E2FA4A1245FE1BFBFEFE7DFF7E" lastPageId="16" lastPageNumber="476" pageId="11" pageNumber="472">
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<subSubSection id="C36D657FFA4A125EFE1BFBFEFDEBFE5D" pageId="11" pageNumber="472" type="nomenclature">
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<paragraph id="8BC836F4FA4A125EFE1BFBFEFD55FE3E" blockId="11.[399,714,398,454]" box="[399,714,398,422]" pageId="11" pageNumber="472">
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<heading id="D0808198FA4A125EFE1BFBFEFD55FE3E" box="[399,714,398,422]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="472" reason="3">
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<taxonomicName id="4C774D77FA4A125EFE1BFBFEFD1BFE3D" ID-CoL="a9b9a295-d010-44e2-8546-b7edd625e3e6" authority="Zacharda, 2000" authorityName="Zacharda" authorityYear="2000" box="[399,644,398,422]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="472" phylum="Arthropoda" rank="species" species="aggerata" status="sp. nov.">
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<emphasis id="B903EAE6FA4A125EFE1BFBFEFD1BFE3D" bold="true" box="[399,644,398,422]" italics="true" pageId="11" pageNumber="472">Troglocheles aggerata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A230579DFA4A125EFD18FBFEFD55FE3E" box="[652,714,398,421]" pageId="11" pageNumber="472" rank="species">n. sp.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph id="8BC836F4FA4A125EFE70FBDEFDEBFE5D" blockId="11.[399,714,398,454]" box="[484,628,430,454]" pageId="11" pageNumber="472">(®gures 4±6)</paragraph>
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</subSubSection>
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<caption id="DF08667CFA4A125EFF32FCFBFC2DF93B" ID-DOI="http://doi.org/10.5281/zenodo.5279618" ID-Zenodo-Dep="5279618" box="[166,946,1674,1696]" httpUri="https://zenodo.org/record/5279618/files/figure.png" pageId="11" pageNumber="472" startId="11.[166,182,1675,1696]" targetBox="[94,1017,543,1644]" targetPageId="11">
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<paragraph id="8BC836F4FA4A125EFF32FCFBFC2DF93B" blockId="11.[166,946,1674,1696]" box="[166,946,1674,1696]" pageId="11" pageNumber="472">
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FIG. 4.
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<taxonomicName id="4C774D77FA4A125EFE96FCFAFE42F93B" authority="Zacharda, 2000" authorityName="Zacharda" authorityYear="2000" box="[258,477,1674,1696]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="472" phylum="Arthropoda" rank="species" species="aggerata" status="sp. nov.">
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<emphasis id="B903EAE6FA4A125EFE96FCFAFE42F93B" box="[258,477,1674,1696]" italics="true" pageId="11" pageNumber="472">Troglocheles aggerata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A230579DFA4A125EFE71FCFBFD82F93B" box="[485,541,1675,1696]" pageId="11" pageNumber="472" rank="species">n. sp.</taxonomicNameLabel>
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, adult female: (A) dorsum; (B) venter.
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</paragraph>
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</caption>
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<caption id="DF08667CFA4D1259FFDBFEA9FB96FAB8" ID-DOI="http://doi.org/10.5281/zenodo.5279620" ID-Zenodo-Dep="5279620" httpUri="https://zenodo.org/record/5279620/files/figure.png" pageId="12" pageNumber="473" startId="12.[79,95,1241,1262]" targetBox="[169,952,143,1210]" targetPageId="12">
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<paragraph id="8BC836F4FA4D1259FFDBFEA9FB96FAB8" blockId="12.[79,1035,1241,1316]" pageId="12" pageNumber="473">
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FIG. 5.
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<taxonomicName id="4C774D77FA4D1259FF3FFEA9FE19FB75" authority="Zacharda, 2000" authorityName="Zacharda" authorityYear="2000" box="[171,390,1241,1263]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="12" pageNumber="473" phylum="Arthropoda" rank="species" species="aggerata" status="sp. nov.">
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<emphasis id="B903EAE6FA4D1259FF3FFEA9FE19FB75" box="[171,390,1241,1263]" italics="true" pageId="12" pageNumber="473">Troglocheles aggerata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A230579DFA4D1259FE19FEA9FE5BFB75" box="[397,452,1241,1262]" pageId="12" pageNumber="473" rank="species">n. sp.</taxonomicNameLabel>
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, adult female: (A) chelicera, lateral aspect; (B) rhagidial organ I, dorsal aspect; (C) rhagidial organ II, dorsal aspect; (D) palpus, lateral aspect; (E) subcapitulum, ventral aspect; (F) apex of tarsus II with oar-like setae, lateral aspect.
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</paragraph>
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</caption>
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<subSubSection id="C36D657FFA4D125AFFDBFF39FF2EFA3A" lastPageId="15" lastPageNumber="476" pageId="12" pageNumber="473" type="materials_examined">
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<paragraph id="8BC836F4FA4D1259FFDBFF39FEBCFAFA" blockId="12.[79,1035,1353,1569]" box="[79,291,1353,1377]" pageId="12" pageNumber="473">
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<emphasis id="B903EAE6FA4D1259FFDBFF39FEBCFAFA" box="[79,291,1353,1377]" italics="true" pageId="12" pageNumber="473">Material examined</emphasis>
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</paragraph>
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<paragraph id="8BC836F4FA4D1259FFECFF19FEC5F9BB" blockId="12.[79,1035,1353,1569]" pageId="12" pageNumber="473">
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<materialsCitation id="3B1F3CA9FA4D1259FFECFF19FECDF9BA" collectingDate="1991-06-04" collectingDateMax="1992-06-06" collectingDateMin="1991-06-04" collectionCode="V" collectorName="M. Zacharda. & Arachnids & Ottawa & Type No." country="Austria" county="All" elevation="6" location="Obergurgl" municipality="Oetztal Alps" pageId="12" pageNumber="473" specimenCount="1" specimenCount-female="1" stateProvince="Tirol" typeStatus="holotype">
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<collectingCounty id="62A94E78FA4D1259FFECFF19FF04FA1A" box="[120,155,1385,1409]" pageId="12" pageNumber="473">All</collectingCounty>
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from
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<collectingCountry id="F3607664FA4D1259FF75FF19FEAAFA1A" box="[225,309,1385,1409]" name="Austria" pageId="12" pageNumber="473">Austria</collectingCountry>
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,
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<collectingRegion id="49B3F816FA4D1259FED0FF19FE1DFA1A" box="[324,386,1385,1409]" country="Austria" name="Tirol" pageId="12" pageNumber="473">Tyrol</collectingRegion>
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,
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<collectingMunicipality id="6BACAC8EFA4D1259FE04FF19FD81FA1A" box="[400,542,1385,1409]" pageId="12" pageNumber="473">Oetztal Alps</collectingMunicipality>
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,
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<location id="8EA8602FFA4D1259FDB9FF19FD3FFA1A" LSID="urn:lsid:plazi:treatment:03DE87E2FA4A1245FE1BFBFEFE7DFF7E:8EA8602FFA4D1259FDB9FF19FD3FFA1A" box="[557,672,1385,1409]" country="Austria" county="All" municipality="Oetztal Alps" name="Obergurgl" pageId="12" pageNumber="473" stateProvince="Tirol">Obergurgl</location>
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(11ss02¾E, 46ss52¾N).
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<typeStatus id="54CC8856FA4D1259FC1AFF1AFB9EFA1A" box="[910,1025,1385,1409]" pageId="12" pageNumber="473" type="holotype">HOLOTYPE</typeStatus>
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: adult female.
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<collectingMethod id="52364EE3FA4D1259FF78FFF9FD9FFA3A" box="[236,512,1417,1441]" pageId="12" pageNumber="473">Collected in pitfall traps</collectingMethod>
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at
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<quantity id="4C8F9B11FA4D1259FDB2FFF9FDFDFA3A" box="[550,610,1417,1441]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="473" unit="m" value="0.6">
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<elevation id="005AD1C7FA4D1259FDB2FFF9FDFDFA3A" box="[550,610,1417,1441]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="473" unit="m" value="0.6">0.6 m</elevation>
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</quantity>
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in subterranean spaces of talus slope on the southern slope of
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<location id="8EA8602FFA4D1259FEFFFFD9FE6BFA5A" LSID="urn:lsid:plazi:treatment:03DE87E2FA4A1245FE1BFBFEFE7DFF7E:8EA8602FFA4D1259FEFFFFD9FE6BFA5A" box="[363,500,1449,1473]" country="Austria" county="All" municipality="Oetztal Alps" name="Festkogl-mt" pageId="12" pageNumber="473" stateProvince="Tirol">Festkogl-mt</location>
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(
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<date id="FFC91034FA4D1259FD92FFD9FD12FA5A" box="[518,653,1449,1473]" pageId="12" pageNumber="473" value="1991-06-04">
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<collectingDate id="EF8DE9DCFA4D1259FD92FFD9FD12FA5A" box="[518,653,1449,1473]" pageId="12" pageNumber="473" value="1991-06-04">4 June 1991</collectingDate>
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</date>
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to
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<date id="FFC91034FA4D1259FD27FFD9FCA6FA5A" box="[691,825,1449,1473]" pageId="12" pageNumber="473" value="1992-06-06">
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<collectingDate id="EF8DE9DCFA4D1259FD27FFD9FCA6FA5A" box="[691,825,1449,1473]" pageId="12" pageNumber="473" value="1992-06-06">6 June 1992</collectingDate>
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</date>
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); coll.
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<collectionCode id="ED66AE31FA4D1259FC18FFDAFC3DFA5A" box="[908,930,1450,1473]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="473" type="Museum">V</collectionCode>
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. RuÊzÏicÏka and
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<collectorName id="26825322FA4D1259FF17FFBAFE80FA7A" box="[131,287,1481,1505]" pageId="12" pageNumber="473">M. Zacharda.</collectorName>
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<typeStatus id="54CC8856FA4D1259FEBCFFBAFE00FA7A" box="[296,415,1481,1505]" pageId="12" pageNumber="473" type="paratype">PARATYPES</typeStatus>
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:
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<specimenCount id="9D71FD7DFA4D1259FE24FFBAFDA8FA7A" box="[432,567,1481,1505]" pageId="12" pageNumber="473" type="female">two females</specimenCount>
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, same data as holotype; deposited in the
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<location id="8EA8602FFA4D1259FFDBFF99FDBEF99B" LSID="urn:lsid:plazi:treatment:03DE87E2FA4A1245FE1BFBFEFE7DFF7E:8EA8602FFA4D1259FFDBFF99FDBEF99B" box="[79,545,1513,1537]" country="Austria" county="All" municipality="Oetztal Alps" name="Canadian National Collection of Insects" pageId="12" pageNumber="473" stateProvince="Tirol">Canadian National Collection of Insects</location>
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and
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<collectorName id="26825322FA4D1259FDCBFF99FD4CF99A" box="[607,723,1513,1537]" pageId="12" pageNumber="473">Arachnids</collectorName>
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,
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<collectorName id="26825322FA4D1259FD71FF99FCA7F99A" box="[741,824,1513,1537]" pageId="12" pageNumber="473">Ottawa</collectorName>
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, in micropreparations,
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<collectorName id="26825322FA4D1259FF03FC79FE9CF9BB" box="[151,259,1545,1569]" pageId="12" pageNumber="473">Type No.</collectorName>
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22,407
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BC836F4FA4D1259FFDBFC3AFF24F9FA" blockId="12.[79,1034,1610,1700]" box="[79,187,1610,1633]" pageId="12" pageNumber="473">
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<emphasis id="B903EAE6FA4D1259FFDBFC3AFF24F9FA" box="[79,187,1610,1633]" italics="true" pageId="12" pageNumber="473">D iagnosis</emphasis>
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</paragraph>
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<paragraph id="8BC836F4FA4D1258FFECFC19FF56F99A" blockId="12.[79,1034,1610,1700]" lastBlockId="13.[79,1035,1321,1537]" lastPageId="13" lastPageNumber="474" pageId="12" pageNumber="473">
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Prosoma, in comparison with opisthosoma, small, narrow; lateral shoulders of opisthosoma with setae c
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<subScript id="17F334B1FA4D1259FEE8FCE3FE1AF93F" attach="left" box="[380,389,1683,1700]" fontSize="7" pageId="12" pageNumber="473">2</subScript>
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striking and protruding laterally. Bothridial setae sc
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<subScript id="17F334B1FA4D1259FB95FCE3FB95F93F" attach="left" box="[1025,1034,1683,1700]" fontSize="7" pageId="12" pageNumber="473">1</subScript>
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short, slightly overlap disjugal suture. Opisthosoma enlarged, ventral epimeres I±
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<collectionCode id="ED66AE31FA4C1258FC79FF5AFB95FADA" box="[1005,1034,1322,1345]" pageId="13" pageNumber="474">IV</collectionCode>
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strikingly small. Dorsal opisthosomal setae c
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<subScript id="17F334B1FA4C1258FDDAFF24FDC8FAFE" attach="left" box="[590,599,1364,1381]" fontSize="7" pageId="13" pageNumber="474">1</subScript>
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, d
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<subScript id="17F334B1FA4C1258FDECFF24FD1EFAFE" attach="left" box="[632,641,1364,1381]" fontSize="7" pageId="13" pageNumber="474">1</subScript>
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, e
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<subScript id="17F334B1FA4C1258FD0BFF24FD37FAFE" attach="left" box="[671,680,1364,1381]" fontSize="7" pageId="13" pageNumber="474">1</subScript>
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, f
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|
|
<subScript id="17F334B1FA4C1258FD51FF24FD51FAFE" attach="left" box="[709,718,1364,1381]" fontSize="7" pageId="13" pageNumber="474">1</subScript>
|
||
|
|
short; insertions of setae f
|
||
|
|
<subScript id="17F334B1FA4C1258FB95FF24FB95FAFE" attach="right" box="[1025,1034,1364,1381]" fontSize="7" pageId="13" pageNumber="474">1</subScript>
|
||
|
|
distal to f
|
||
|
|
<subScript id="17F334B1FA4C1258FF51FF03FF51FA1F" attach="right" box="[197,206,1395,1412]" fontSize="7" pageId="13" pageNumber="474">2</subScript>
|
||
|
|
, setae f
|
||
|
|
<subScript id="17F334B1FA4C1258FEA5FF03FEA5FA1F" attach="left" box="[305,314,1395,1412]" fontSize="7" pageId="13" pageNumber="474">1</subScript>
|
||
|
|
do not reach insertions of f
|
||
|
|
<subScript id="17F334B1FA4C1258FD19FF03FD09FA1F" attach="left" box="[653,662,1395,1412]" fontSize="7" pageId="13" pageNumber="474">2</subScript>
|
||
|
|
. Chelicera with relatively short and robust digits; ®xed digit ¯attened laterally and broad; masticatory surface of ®xed digit distinctly convex. Rhagidial organs I and
|
||
|
|
<collectionCode id="ED66AE31FA4C1258FD3CFFDAFD23FA5A" box="[680,700,1450,1473]" pageId="13" pageNumber="474">II</collectionCode>
|
||
|
|
each consist of four separate oblique rhagidial solenidia and famulus positioned laterad of proximal rhagidial solenidion.
|
||
|
|
</paragraph>
|
||
|
|
<caption id="DF08667CFA4C1258FFDBFEC6FD35FA9B" ID-DOI="http://doi.org/10.5281/zenodo.5279622" ID-Zenodo-Dep="5279622" httpUri="https://zenodo.org/record/5279622/files/figure.png" pageId="13" pageNumber="474" startId="13.[79,95,1206,1227]" targetBox="[87,1031,144,1176]" targetPageId="13">
|
||
|
|
<paragraph id="8BC836F4FA4C1258FFDBFEC6FD35FA9B" blockId="13.[79,1035,1205,1280]" pageId="13" pageNumber="474">
|
||
|
|
FIG. 6.
|
||
|
|
<taxonomicName id="4C774D77FA4C1258FF3FFEC5FE18FB50" authority="Zacharda, 2000" authorityName="Zacharda" authorityYear="2000" box="[171,391,1205,1227]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="13" pageNumber="474" phylum="Arthropoda" rank="species" species="aggerata" status="sp. nov.">
|
||
|
|
<emphasis id="B903EAE6FA4C1258FF3FFEC5FE18FB50" box="[171,391,1205,1227]" italics="true" pageId="13" pageNumber="474">Troglocheles aggerata</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
<taxonomicNameLabel id="A230579DFA4C1258FE1BFEC6FE57FB50" box="[399,456,1206,1227]" pageId="13" pageNumber="474" rank="species">n. sp.</taxonomicNameLabel>
|
||
|
|
, adult female, setal arrangement on legs, lateral aspect: (A) leg I; (B) leg II; (C) leg III; (D) leg IV (
|
||
|
|
<emphasis id="B903EAE6FA4C1258FDF9FEA0FDE1FB7D" bold="true" box="[621,638,1232,1254]" italics="true" pageId="13" pageNumber="474">v</emphasis>
|
||
|
|
,
|
||
|
|
<emphasis id="B903EAE6FA4C1258FD19FEA0FD05FB7D" bold="true" box="[653,666,1232,1254]" italics="true" pageId="13" pageNumber="474">Q</emphasis>
|
||
|
|
,
|
||
|
|
<emphasis id="B903EAE6FA4C1258FD3EFEA0FD28FB7D" bold="true" box="[682,695,1232,1254]" italics="true" pageId="13" pageNumber="474">s</emphasis>
|
||
|
|
and
|
||
|
|
<emphasis id="B903EAE6FA4C1258FD7BFEA0FD67FB7D" bold="true" box="[751,760,1232,1254]" italics="true" pageId="13" pageNumber="474">e</emphasis>
|
||
|
|
designations for solenidia on tarsus, tibia, genu and for famulus, respectively).
|
||
|
|
</paragraph>
|
||
|
|
</caption>
|
||
|
|
<paragraph id="8BC836F4FA4C1258FFDBFC5AFF50F9DA" blockId="13.[79,1034,1578,1697]" box="[79,207,1578,1601]" pageId="13" pageNumber="474">
|
||
|
|
<emphasis id="B903EAE6FA4C1258FFDBFC5AFF50F9DA" box="[79,207,1578,1601]" italics="true" pageId="13" pageNumber="474">D escription</emphasis>
|
||
|
|
</paragraph>
|
||
|
|
<paragraph id="8BC836F4FA4C1258FFECFC39FD93F91B" blockId="13.[79,1034,1578,1697]" pageId="13" pageNumber="474">
|
||
|
|
Adult female (three examined). Length of idiosoma
|
||
|
|
<quantity id="4C8F9B11FA4C1258FD45FC39FCC3F9FA" box="[721,860,1609,1633]" metricMagnitude="3" metricUnit="m" metricValue="1.3599999999999999" metricValueMax="1.44" metricValueMin="1.28" pageId="13" pageNumber="474" unit="m" value="1360.0" valueMax="1440.0" valueMin="1280.0">
|
||
|
|
1280±1440
|
||
|
|
<emphasis id="B903EAE6FA4C1258FCDAFC39FCC3F9FA" bold="true" box="[846,860,1609,1633]" italics="true" pageId="13" pageNumber="474">m</emphasis>
|
||
|
|
</quantity>
|
||
|
|
m. Ratio of leg I length to idiosomal length 0.73±0.93.
|
||
|
|
</paragraph>
|
||
|
|
<paragraph id="8BC836F4FA4C125BFFECFCF8FE19FA5A" blockId="13.[79,1034,1578,1697]" lastBlockId="14.[79,1036,843,1700]" lastPageId="14" lastPageNumber="475" pageId="13" pageNumber="474">
|
||
|
|
<emphasis id="B903EAE6FA4C1258FFECFCF8FE9EF93B" box="[120,257,1672,1696]" italics="true" pageId="13" pageNumber="474">Gnathosom a</emphasis>
|
||
|
|
. Subcapitulum slender, oval, subtriangular (®gure 5E); ratio of length to breadth 1.50±1.66; distal hypostomal lips with spiniform internal malar and serrate external malar processes; adoral setae nude; proximal subcapitular setae ciliate, external pair slightly longer than internal pair. Chelicerae slender, robust, dorsal surface with distinct saddle-shaped depression distal to level of bases of digits (®gure 5A); cheliceral digits long and robust; ®xed digit ¯attened laterally and broad, its masticatory surface smooth and with distinct broad convex rim arising opposite insertion of distal cheliceral seta; masticatory surface of movable digit smooth. Chelicerae with two setae inserted distad of articulation of movable digit; tip of proximal seta not reaching insertion of distal seta; tip of distal seta slightly overlaps apex of ®xed digit. Length of chelicera
|
||
|
|
<quantity id="4C8F9B11FA4F125BFD9EFE1AFDE5FB19" box="[522,634,1130,1154]" metricMagnitude="2" metricUnit="m" metricValue="2.85" metricValueMax="2.9" metricValueMin="2.8" pageId="14" pageNumber="475" unit="m" value="285.0" valueMax="290.0" valueMin="280.0">
|
||
|
|
280±290
|
||
|
|
<emphasis id="B903EAE6FA4F125BFDF8FE1AFDE5FB19" bold="true" box="[620,634,1130,1154]" italics="true" pageId="14" pageNumber="475">m</emphasis>
|
||
|
|
</quantity>
|
||
|
|
m, dorsoventra l width
|
||
|
|
<quantity id="4C8F9B11FA4F125BFCE9FE1AFC72FB19" box="[893,1005,1130,1154]" metricMagnitude="2" metricUnit="m" metricValue="1.1199999999999999" metricValueMax="1.15" metricValueMin="1.09" pageId="14" pageNumber="475" unit="m" value="112.0" valueMax="115.0" valueMin="109.0">
|
||
|
|
109±115
|
||
|
|
<emphasis id="B903EAE6FA4F125BFC4BFE1AFC72FB19" bold="true" box="[991,1005,1130,1154]" italics="true" pageId="14" pageNumber="475">m</emphasis>
|
||
|
|
</quantity>
|
||
|
|
m, length of movable digit
|
||
|
|
<quantity id="4C8F9B11FA4F125BFEFDFEFAFE46FB39" box="[361,473,1162,1186]" metricMagnitude="2" metricUnit="m" metricValue="1.105" metricValueMax="1.1199999999999999" metricValueMin="1.09" pageId="14" pageNumber="475" unit="m" value="110.5" valueMax="112.0" valueMin="109.0">
|
||
|
|
109±112
|
||
|
|
<emphasis id="B903EAE6FA4F125BFE5FFEFAFE46FB39" bold="true" box="[459,473,1162,1186]" italics="true" pageId="14" pageNumber="475">m</emphasis>
|
||
|
|
</quantity>
|
||
|
|
m, length of proximal and distal cheliceral setae 23 and
|
||
|
|
<quantity id="4C8F9B11FA4F125BFF31FEDAFF64FB59" box="[165,251,1194,1218]" metricMagnitude="1" metricUnit="m" metricValue="3.8" metricValueMax="4.3" metricValueMin="3.3" pageId="14" pageNumber="475" unit="m" value="38.0" valueMax="43.0" valueMin="33.0">
|
||
|
|
33±43
|
||
|
|
<emphasis id="B903EAE6FA4F125BFF79FEDAFF64FB59" bold="true" box="[237,251,1194,1218]" italics="true" pageId="14" pageNumber="475">m</emphasis>
|
||
|
|
</quantity>
|
||
|
|
m, respectively, distance between their insertions
|
||
|
|
<quantity id="4C8F9B11FA4F125BFCBAFEDAFC1AFB59" box="[814,901,1194,1218]" metricMagnitude="1" metricUnit="m" metricValue="2.8" metricValueMax="3.0" metricValueMin="2.6" pageId="14" pageNumber="475" unit="m" value="28.0" valueMax="30.0" valueMin="26.0">
|
||
|
|
26±30
|
||
|
|
<emphasis id="B903EAE6FA4F125BFCE3FEDAFC1AFB59" bold="true" box="[887,901,1194,1218]" italics="true" pageId="14" pageNumber="475">m</emphasis>
|
||
|
|
</quantity>
|
||
|
|
m. Ratio of cheliceral length to dorsoventral width 2.43±2.57; ratio of length of movable digit to length of chelicera 0.38±0.40; ratio of length of movable digit to dorsoventral width of chelicera 0.97±1.00. Palpus robust, with relatively slender tarsus (®gure 5D); ratio of length to width of tarsus 2.58±3.00. Length of palpal trochanter, femorogenu, tibia and tarsus 33±36, 105±115, 30 ±46 and
|
||
|
|
<quantity id="4C8F9B11FA4F125BFDF0FF39FD57FAFA" box="[612,712,1353,1377]" metricMagnitude="2" metricUnit="m" metricValue="1.02" metricValueMax="1.05" metricValueMin="0.99" pageId="14" pageNumber="475" unit="m" value="102.0" valueMax="105.0" valueMin="99.0">
|
||
|
|
99±105
|
||
|
|
<emphasis id="B903EAE6FA4F125BFD2EFF39FD57FAFA" bold="true" box="[698,712,1353,1377]" italics="true" pageId="14" pageNumber="475">m</emphasis>
|
||
|
|
</quantity>
|
||
|
|
m, respectively. Number of setae and solenidia (in brackets) on palpal trochanter, femorogenu, tibia and tarsus 0-2-3-10(1), respectively. Two ciliated setae on palpal tibia strikingly short, tarsal solenidion spiniform, erect.
|
||
|
|
</paragraph>
|
||
|
|
<caption id="DF08667CFA4F125BFFDBF8C9FE65FC85" ID-DOI="http://doi.org/10.5281/zenodo.5279624" ID-Zenodo-Dep="5279624" httpUri="https://zenodo.org/record/5279624/files/figure.png" pageId="14" pageNumber="475" startId="14.[79,95,697,718]" targetBox="[84,1031,141,667]" targetPageId="14">
|
||
|
|
<paragraph id="8BC836F4FA4F125BFFDBF8C9FE65FC85" blockId="14.[79,1034,696,798]" pageId="14" pageNumber="475">
|
||
|
|
FIG. 7. Geographic distribution of
|
||
|
|
<taxonomicName id="4C774D77FA4F125BFE7FF8C8FD5EFD55" box="[491,705,696,718]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="14" pageNumber="475" phylum="Arthropoda" rank="species" species="undetermined">
|
||
|
|
<emphasis id="B903EAE6FA4F125BFE7FF8C8FDF7FD55" box="[491,616,696,718]" italics="true" pageId="14" pageNumber="475">Troglocheles</emphasis>
|
||
|
|
species
|
||
|
|
</taxonomicName>
|
||
|
|
in Europe: 1,
|
||
|
|
<taxonomicName id="4C774D77FA4F125BFC1CF8C9FB9BFD55" baseAuthorityName="Willmann" baseAuthorityYear="1932" box="[904,1028,697,718]" class="Arachnida" family="Rhagidiidae" genus="Rhagidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="14" pageNumber="475" phylum="Arthropoda" rank="species" species="strasseri">
|
||
|
|
<emphasis id="B903EAE6FA4F125BFC1CF8C9FB9BFD55" box="[904,1028,697,718]" italics="true" pageId="14" pageNumber="475">T. strasseri</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
; 2,
|
||
|
|
<taxonomicName id="4C774D77FA4F125BFF3EF8A4FEB7FD72" box="[170,296,723,745]" pageId="14" pageNumber="475" rank="species" species="tiroliensis">
|
||
|
|
<emphasis id="B903EAE6FA4F125BFF3EF8A4FEB7FD72" box="[170,296,723,745]" italics="true" pageId="14" pageNumber="475">T. tiroliensis</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
; 3,
|
||
|
|
<taxonomicName id="4C774D77FA4F125BFEC5F8A4FDFFFD72" authority="sensu Rack, 1974" authorityName="Rack" authorityYear="1974" box="[337,608,723,745]" class="Arachnida" family="Rhagidiidae" genus="Rhagidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="14" pageNumber="475" phylum="Arthropoda" rank="species" species="gineti">
|
||
|
|
<emphasis id="B903EAE6FA4F125BFEC5F8A4FE39FD72" box="[337,422,724,745]" italics="true" pageId="14" pageNumber="475">T. gineti</emphasis>
|
||
|
|
sensu
|
||
|
|
<bibRefCitation id="EFE64B05FA4F125BFE79F8A3FDFFFD72" author="RACK, G." box="[493,608,723,745]" pageId="14" pageNumber="475" pagination="128" refId="ref7266" refString="RACK, G., 1974, Milben (Acarina) aus HoEhlen der Schwabischen Alb und einiger anderen HoEhlen, Jahreshefte der Gesellschaft fuEr Naturkunde in W uErttemberg, 129, 128 ± 137." type="journal article" year="1974">Rack, 1974</bibRefCitation>
|
||
|
|
</taxonomicName>
|
||
|
|
; 4,
|
||
|
|
<taxonomicName id="4C774D77FA4F125BFD1EF8A4FCBEFD72" baseAuthorityName="Willmann" baseAuthorityYear="1953" box="[650,801,723,745]" pageId="14" pageNumber="475" rank="species" species="vornatscheri">
|
||
|
|
<emphasis id="B903EAE6FA4F125BFD1EF8A4FCBEFD72" box="[650,801,723,745]" italics="true" pageId="14" pageNumber="475">T. vornatscheri</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
; 5,
|
||
|
|
<emphasis id="B903EAE6FA4F125BFCDEF8A4FCC0FD72" box="[842,863,724,745]" italics="true" pageId="14" pageNumber="475">T.</emphasis>
|
||
|
|
spp; 6,
|
||
|
|
<taxonomicName id="4C774D77FA4F125BFC21F8A4FEE2FC98" authority="sensu Cooreman, 1959" authorityName="sensu Cooreman" authorityYear="1959" class="Arachnida" family="Rhagidiidae" genus="Rhagidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="14" pageNumber="475" phylum="Arthropoda" rank="species" species="gineti">
|
||
|
|
<emphasis id="B903EAE6FA4F125BFC21F8A4FB95FD72" box="[949,1034,724,745]" italics="true" pageId="14" pageNumber="475">T. gineti</emphasis>
|
||
|
|
sensu Cooreman, 1959
|
||
|
|
</taxonomicName>
|
||
|
|
; 7,
|
||
|
|
<taxonomicName id="4C774D77FA4F125BFE38F89EFD8EFC98" authorityName="Zacharda" authorityYear="1987" box="[428,529,749,771]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="14" pageNumber="475" phylum="Arthropoda" rank="species" species="vandeli">
|
||
|
|
<emphasis id="B903EAE6FA4F125BFE38F89EFD8EFC98" box="[428,529,749,771]" italics="true" pageId="14" pageNumber="475">T. vandeli</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
; 8,
|
||
|
|
<taxonomicName id="4C774D77FA4F125BFDD4F89EFD4BFC98" authority="Zacharda, 2000" authorityName="Zacharda" authorityYear="2000" box="[576,724,749,771]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="14" pageNumber="475" phylum="Arthropoda" rank="species" species="archetypica" status="sp. nov.">
|
||
|
|
<emphasis id="B903EAE6FA4F125BFDD4F89EFD4BFC98" box="[576,724,749,771]" italics="true" pageId="14" pageNumber="475">T. archetypica</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
<taxonomicNameLabel id="A230579DFA4F125BFD4AF89EFC86FC98" box="[734,793,750,771]" pageId="14" pageNumber="475" rank="species">n. sp.</taxonomicNameLabel>
|
||
|
|
; 9,
|
||
|
|
<taxonomicName id="4C774D77FA4F125BFCD3F89EFC21FC98" authority="Zacharda, 2000" authorityName="Zacharda" authorityYear="2000" box="[839,958,750,771]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="14" pageNumber="475" phylum="Arthropoda" rank="species" species="aggerata" status="sp. nov.">
|
||
|
|
<emphasis id="B903EAE6FA4F125BFCD3F89EFC21FC98" box="[839,958,750,771]" italics="true" pageId="14" pageNumber="475">T. aggerata</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
<taxonomicNameLabel id="A230579DFA4F125BFC5DF89EFB9BFC98" box="[969,1028,750,771]" pageId="14" pageNumber="475" rank="species">n. sp.</taxonomicNameLabel>
|
||
|
|
; 10,
|
||
|
|
<taxonomicName id="4C774D77FA4F125BFF22F979FEB5FC85" authorityName="Lombardini" authorityYear="1951" box="[182,298,777,798]" class="Arachnida" family="Rhagidiidae" genus="Rhagidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="14" pageNumber="475" phylum="Arthropoda" rank="species" species="conciana">
|
||
|
|
<emphasis id="B903EAE6FA4F125BFF22F979FEB5FC85" box="[182,298,777,798]" italics="true" pageId="14" pageNumber="475">T. conciana</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
; 11,
|
||
|
|
<taxonomicName id="4C774D77FA4F125BFEF4F979FE6AFC85" authorityName="Turk" authorityYear="1972" box="[352,501,776,798]" class="Arachnida" family="Rhagidiidae" genus="Rhagidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="14" pageNumber="475" phylum="Arthropoda" rank="species" species="odontochela">
|
||
|
|
<emphasis id="B903EAE6FA4F125BFEF4F979FE6AFC85" box="[352,501,776,798]" italics="true" pageId="14" pageNumber="475">T. odontochela</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
.
|
||
|
|
</paragraph>
|
||
|
|
</caption>
|
||
|
|
<paragraph id="8BC836F4FA4F125BFFECFFB9FD94F9DB" blockId="14.[79,1036,843,1700]" pageId="14" pageNumber="475">
|
||
|
|
<emphasis id="B903EAE6FA4F125BFFECFFB9FF70FA7A" box="[120,239,1481,1505]" italics="true" pageId="14" pageNumber="475">Prodorsum</emphasis>
|
||
|
|
. Naso well-developed, with pair of internal vertical setae v
|
||
|
|
<subScript id="17F334B1FA4F125BFCEAFFA3FC18FA7F" attach="left" box="[894,903,1491,1508]" fontSize="7" pageId="14" pageNumber="475">1</subScript>
|
||
|
|
. Bothridial setae sc
|
||
|
|
<subScript id="17F334B1FA4F125BFF3EFF83FF2CF99F" attach="left" box="[170,179,1523,1540]" fontSize="7" pageId="14" pageNumber="475">1</subScript>
|
||
|
|
®liform, ®nely pubescent, relatively short, overlap disjugal suture only slightly (®gure 4A). Setae v
|
||
|
|
<subScript id="17F334B1FA4F125BFE1FFC63FE0BF9BF" attach="left" box="[395,404,1555,1572]" fontSize="7" pageId="14" pageNumber="475">2</subScript>
|
||
|
|
strikingly short. Length of setae: v
|
||
|
|
<subScript id="17F334B1FA4F125BFCA3FC63FCDFF9BF" attach="left" box="[823,832,1555,1572]" fontSize="7" pageId="14" pageNumber="475">1</subScript>
|
||
|
|
46±49, v
|
||
|
|
<subScript id="17F334B1FA4F125BFC39FC63FC29F9BF" attach="left" box="[941,950,1555,1572]" fontSize="7" pageId="14" pageNumber="475">2</subScript>
|
||
|
|
30, sc
|
||
|
|
<subScript id="17F334B1FA4F125BFB95FC63FB95F9BF" attach="left" box="[1025,1034,1555,1572]" fontSize="7" pageId="14" pageNumber="475">1</subScript>
|
||
|
|
102± 115, sc
|
||
|
|
<subScript id="17F334B1FA4F125BFF40FC43FF42F9DF" attach="left" box="[212,221,1587,1604]" fontSize="7" pageId="14" pageNumber="475">2</subScript>
|
||
|
|
<quantity id="4C8F9B11FA4F125BFF73FC59FEC7F9DA" box="[231,344,1577,1601]" metricMagnitude="2" metricUnit="m" metricValue="1.135" metricValueMax="1.15" metricValueMin="1.1199999999999999" pageId="14" pageNumber="475" unit="m" value="113.5" valueMax="115.0" valueMin="112.0">
|
||
|
|
112±115
|
||
|
|
<emphasis id="B903EAE6FA4F125BFEDEFC59FEC7F9DA" bold="true" box="[330,344,1577,1601]" italics="true" pageId="14" pageNumber="475">m</emphasis>
|
||
|
|
</quantity>
|
||
|
|
m. Eyes absent.
|
||
|
|
</paragraph>
|
||
|
|
<paragraph id="8BC836F4FA4F125AFFECFC39FD31FE9E" blockId="14.[79,1036,843,1700]" lastBlockId="15.[79,1036,142,1091]" lastPageId="15" lastPageNumber="476" pageId="14" pageNumber="475">
|
||
|
|
<emphasis id="B903EAE6FA4F125BFFECFC39FEFAF9FA" box="[120,357,1609,1633]" italics="true" pageId="14" pageNumber="475">Opisthosomal dorsum</emphasis>
|
||
|
|
. Complement and arrangement of dorsal setae and cupules typical for
|
||
|
|
<taxonomicName id="4C774D77FA4F125BFF5BFC19FEC6F91A" authorityName="Oudemans" authorityYear="1922" box="[207,345,1641,1665]" class="Arachnida" family="Rhagidiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="14" pageNumber="475" phylum="Arthropoda" rank="family">Rhagidiidae</taxonomicName>
|
||
|
|
(®gure 4A); three pairs of cupules; ia positioned laterally at level about midway between setae c
|
||
|
|
<subScript id="17F334B1FA4F125BFE71FCE3FE71F93F" attach="left" box="[485,494,1683,1700]" fontSize="7" pageId="14" pageNumber="475">1</subScript>
|
||
|
|
and d
|
||
|
|
<subScript id="17F334B1FA4F125BFDACFCE3FDDEF93F" attach="left" box="[568,577,1683,1700]" fontSize="7" pageId="14" pageNumber="475">1</subScript>
|
||
|
|
, im lateral and just anterior to setae e
|
||
|
|
<subScript id="17F334B1FA4F125BFC63FCE3FB9FF93F" attach="left" box="[1015,1024,1683,1700]" fontSize="7" pageId="14" pageNumber="475">1</subScript>
|
||
|
|
, ip laterally at level about two-thirds of interval from setae e
|
||
|
|
<subScript id="17F334B1FA4E125AFD6BFAE8FC97FF32" attach="left" box="[767,776,152,169]" fontSize="7" pageId="15" pageNumber="476">1</subScript>
|
||
|
|
to f
|
||
|
|
<subScript id="17F334B1FA4E125AFCA9FAE8FCD9FF32" attach="right" box="[829,838,152,169]" fontSize="7" pageId="15" pageNumber="476">1</subScript>
|
||
|
|
. Setae c
|
||
|
|
<subScript id="17F334B1FA4E125AFC3DFAE8FC2DFF32" attach="left" box="[937,946,152,169]" fontSize="7" pageId="15" pageNumber="476">1</subScript>
|
||
|
|
, d
|
||
|
|
<subScript id="17F334B1FA4E125AFC40FAE8FC42FF32" attach="left" box="[980,989,152,169]" fontSize="7" pageId="15" pageNumber="476">1</subScript>
|
||
|
|
, e
|
||
|
|
<subScript id="17F334B1FA4E125AFC6FFAE8FB96FF32" attach="left" box="[1019,1033,152,169]" fontSize="7" pageId="15" pageNumber="476">1,</subScript>
|
||
|
|
f
|
||
|
|
<subScript id="17F334B1FA4E125AFFCEFAC8FFFCFF52" attach="left" box="[90,99,184,201]" fontSize="7" pageId="15" pageNumber="476">1</subScript>
|
||
|
|
reach 0.30, 0.26, 0.23±0.26, 0.30 of distance to insertion of successive setae. Setae f
|
||
|
|
<subScript id="17F334B1FA4E125AFFCEFAA8FFFCFF72" attach="left" box="[90,99,216,233]" fontSize="7" pageId="15" pageNumber="476">1</subScript>
|
||
|
|
strikingly distant from f
|
||
|
|
<subScript id="17F334B1FA4E125AFE18FAA8FE0AFF72" attach="right" box="[396,405,216,233]" fontSize="7" pageId="15" pageNumber="476">2</subScript>
|
||
|
|
. Length of setae: c
|
||
|
|
<subScript id="17F334B1FA4E125AFDE9FAA8FD19FF72" attach="left" box="[637,646,216,233]" fontSize="7" pageId="15" pageNumber="476">1</subScript>
|
||
|
|
46±63, c
|
||
|
|
<subScript id="17F334B1FA4E125AFD60FAA8FD62FF72" attach="left" box="[756,765,216,233]" fontSize="7" pageId="15" pageNumber="476">2</subScript>
|
||
|
|
105±112, d
|
||
|
|
<subScript id="17F334B1FA4E125AFC1EFAA8FC0CFF72" attach="left" box="[906,915,216,233]" fontSize="7" pageId="15" pageNumber="476">1</subScript>
|
||
|
|
53±59,
|
||
|
|
<geoCoordinate id="EE435033FA4E125AFC61FABEFFF6FE9E" degrees="1" direction="east" minutes="59" orientation="longitude" pageId="15" pageNumber="476" precision="925" value="1.9833333">
|
||
|
|
e
|
||
|
|
<subScript id="17F334B1FA4E125AFB95FAA8FB95FF72" attach="left" box="[1025,1034,216,233]" fontSize="7" pageId="15" pageNumber="476">1</subScript>
|
||
|
|
59
|
||
|
|
</geoCoordinate>
|
||
|
|
± 63, f
|
||
|
|
<subScript id="17F334B1FA4E125AFF3BFA88FF27FE92" attach="left" box="[175,184,248,265]" fontSize="7" pageId="15" pageNumber="476">1</subScript>
|
||
|
|
72±79, f
|
||
|
|
<subScript id="17F334B1FA4E125AFEB5FA88FEB5FE92" attach="left" box="[289,298,248,265]" fontSize="7" pageId="15" pageNumber="476">2</subScript>
|
||
|
|
56±59, h
|
||
|
|
<subScript id="17F334B1FA4E125AFE03FA88FE3FFE92" attach="left" box="[407,416,248,265]" fontSize="7" pageId="15" pageNumber="476">1</subScript>
|
||
|
|
112±115, h
|
||
|
|
<subScript id="17F334B1FA4E125AFDBCFA88FDAEFE92" attach="left" box="[552,561,248,265]" fontSize="7" pageId="15" pageNumber="476">2</subScript>
|
||
|
|
<quantity id="4C8F9B11FA4E125AFDAEFA9DFD0EFE9E" box="[570,657,237,261]" metricMagnitude="1" metricUnit="m" metricValue="6.9" metricValueMax="7.2" metricValueMin="6.6" pageId="15" pageNumber="476" unit="m" value="69.0" valueMax="72.0" valueMin="66.0">
|
||
|
|
66±72
|
||
|
|
<emphasis id="B903EAE6FA4E125AFD17FA9DFD0EFE9E" bold="true" box="[643,657,237,261]" italics="true" pageId="15" pageNumber="476">m</emphasis>
|
||
|
|
</quantity>
|
||
|
|
m.
|
||
|
|
</paragraph>
|
||
|
|
<paragraph id="8BC836F4FA4E125AFFECFB7DFD24FEDE" blockId="15.[79,1036,142,1091]" pageId="15" pageNumber="476">
|
||
|
|
<emphasis id="B903EAE6FA4E125AFFECFB7DFF77FEBE" box="[120,232,269,293]" italics="true" pageId="15" pageNumber="476">Podosoma</emphasis>
|
||
|
|
. Coxisternal plates (epimeres) I,
|
||
|
|
<collectionCode id="ED66AE31FA4E125AFD17FB7EFD08FEBE" box="[643,663,270,293]" pageId="15" pageNumber="476">II</collectionCode>
|
||
|
|
,
|
||
|
|
<collectionCode id="ED66AE31FA4E125AFD39FB7EFD54FEBE" box="[685,715,270,293]" pageId="15" pageNumber="476">III</collectionCode>
|
||
|
|
,
|
||
|
|
<collectionCode id="ED66AE31FA4E125AFD76FB7EFD60FEBE" box="[738,767,270,293]" pageId="15" pageNumber="476">IV</collectionCode>
|
||
|
|
strikingly small, with 3-1-4-3 ®nely pubescent setae, respectively (®gure 4B).
|
||
|
|
</paragraph>
|
||
|
|
<paragraph id="8BC836F4FA4E125AFFECFB3DFEA6FE7F" blockId="15.[79,1036,142,1091]" pageId="15" pageNumber="476">
|
||
|
|
<emphasis id="B903EAE6FA4E125AFFECFB3DFE86FEFE" box="[120,281,333,357]" italics="true" pageId="15" pageNumber="476">Genital region</emphasis>
|
||
|
|
. Genital valves each with ®ve ®nely pubescent setae of similar length,
|
||
|
|
<quantity id="4C8F9B11FA4E125AFF30FB1DFF64FE1E" box="[164,251,365,389]" metricMagnitude="1" metricUnit="m" metricValue="2.65" metricValueMax="3.0" metricValueMin="2.3" pageId="15" pageNumber="476" unit="m" value="26.5" valueMax="30.0" valueMin="23.0">
|
||
|
|
23±30
|
||
|
|
<emphasis id="B903EAE6FA4E125AFF79FB1DFF64FE1E" bold="true" box="[237,251,365,389]" italics="true" pageId="15" pageNumber="476">m</emphasis>
|
||
|
|
</quantity>
|
||
|
|
m, arranged evenly along medial edge of valve (®gure 4B). Five pairs of aggenital (paragenital) setae of similar length,
|
||
|
|
<quantity id="4C8F9B11FA4E125AFD14FBFDFD48FE3E" box="[640,727,397,421]" metricMagnitude="1" metricUnit="m" metricValue="3.45" metricValueMax="3.6" metricValueMin="3.3" pageId="15" pageNumber="476" unit="m" value="34.5" valueMax="36.0" valueMin="33.0">
|
||
|
|
33±36
|
||
|
|
<emphasis id="B903EAE6FA4E125AFD5DFBFDFD48FE3E" bold="true" box="[713,727,397,421]" italics="true" pageId="15" pageNumber="476">m</emphasis>
|
||
|
|
</quantity>
|
||
|
|
m. Length of genital valves
|
||
|
|
<quantity id="4C8F9B11FA4E125AFFDBFBDDFF5FFE5E" box="[79,192,429,453]" metricMagnitude="2" metricUnit="m" metricValue="1.4" metricValueMax="1.48" metricValueMin="1.3199999999999998" pageId="15" pageNumber="476" unit="m" value="140.0" valueMax="148.0" valueMin="132.0">
|
||
|
|
132± 148
|
||
|
|
<emphasis id="B903EAE6FA4E125AFF26FBDDFF5FFE5E" bold="true" box="[178,192,429,453]" italics="true" pageId="15" pageNumber="476">m</emphasis>
|
||
|
|
</quantity>
|
||
|
|
m. Cupules ih positioned ventrolaterally, almost laterad of posteriormost pair of adanal setae.
|
||
|
|
</paragraph>
|
||
|
|
<paragraph id="8BC836F4FA4E125AFFECFB9DFD89FBD9" blockId="15.[79,1036,142,1091]" pageId="15" pageNumber="476">
|
||
|
|
<emphasis id="B903EAE6FA4E125AFFECFB9DFF2CFD9F" box="[120,179,493,516]" italics="true" pageId="15" pageNumber="476">L egs.</emphasis>
|
||
|
|
Leg I
|
||
|
|
<quantity id="4C8F9B11FA4E125AFE9CFB9DFE1AFD9E" box="[264,389,493,517]" metricMagnitude="3" metricUnit="m" metricValue="1.0879999999999999" metricValueMax="1.2" metricValueMin="0.976" pageId="15" pageNumber="476" unit="m" value="1088.0" valueMax="1200.0" valueMin="976.0">
|
||
|
|
976±1200
|
||
|
|
<emphasis id="B903EAE6FA4E125AFEE3FB9DFE1AFD9E" bold="true" box="[375,389,493,517]" italics="true" pageId="15" pageNumber="476">m</emphasis>
|
||
|
|
</quantity>
|
||
|
|
m long, 0.73±0.93 as long as idiosoma. Empodia of all legs setulose, broadly oval in dorsoventral view, slightly longer than claws; claws each with small ventrobasal clawlet. Number of setae (solenidia and famulus
|
||
|
|
<emphasis id="B903EAE6FA4E125AFB95F85DFB94FDDE" bold="true" box="[1025,1035,557,581]" italics="true" pageId="15" pageNumber="476">e</emphasis>
|
||
|
|
bracketed), respectively, on legs I-II-III-IV are: trochanters 1-1-2-2; basifemora + telofemora 5 +5-5 +5-4 +4-3 +4; genua 10 (1)-7 (1)-6(1)-6; tibiae 12 (2)-7(2)-5 (2)- 5 (1); tarsi 17 (4 +
|
||
|
|
<emphasis id="B903EAE6FA4E125AFEB7F8FCFEB2FD3F" bold="true" box="[291,301,652,676]" italics="true" pageId="15" pageNumber="476">e</emphasis>
|
||
|
|
)-16(4 +
|
||
|
|
<emphasis id="B903EAE6FA4E125AFE07F8FCFE02FD3F" bold="true" box="[403,413,652,676]" italics="true" pageId="15" pageNumber="476">e</emphasis>
|
||
|
|
)-14-13 (®gure 6). Genua I and
|
||
|
|
<collectionCode id="ED66AE31FA4E125AFC99F8FDFCBEFD3F" box="[781,801,653,676]" pageId="15" pageNumber="476">II</collectionCode>
|
||
|
|
each with one erect spiniform distoventral solenidion, genu
|
||
|
|
<collectionCode id="ED66AE31FA4E125AFD83F8DDFDA9FD5F" box="[535,566,685,708]" pageId="15" pageNumber="476">III</collectionCode>
|
||
|
|
with one small spiniform dorsoproximal solenidion. Tibia I with one dorsoproximal erect spiniform solenidion, and one dorsodistal rhagidial solenidion; tibia
|
||
|
|
<collectionCode id="ED66AE31FA4E125AFD98F89DFDBFFC9F" box="[524,544,749,772]" pageId="15" pageNumber="476">II</collectionCode>
|
||
|
|
with one laterodorsal proximal spiniform erect solenidion, and one lanceolate dorsodistal solenidion recessed in a deep pit with a small surface pore; tibia
|
||
|
|
<collectionCode id="ED66AE31FA4E125AFE57F95CFE7DFCD8" box="[451,482,812,835]" pageId="15" pageNumber="476">III</collectionCode>
|
||
|
|
with two erect spiniform laterodorsal proximal solenidia arranged in tandem; tibia
|
||
|
|
<collectionCode id="ED66AE31FA4E125AFD87F93CFDAFFCF8" box="[531,560,844,867]" pageId="15" pageNumber="476">IV</collectionCode>
|
||
|
|
with one erect spiniform laterodorsal solenidion. Tarsi
|
||
|
|
<collectionCode id="ED66AE31FA4E125AFE89F91CFEAEFC18" box="[285,305,876,899]" pageId="15" pageNumber="476">II</collectionCode>
|
||
|
|
,
|
||
|
|
<collectionCode id="ED66AE31FA4E125AFED0F91CFEFDFC18" box="[324,354,876,899]" pageId="15" pageNumber="476">III</collectionCode>
|
||
|
|
,
|
||
|
|
<collectionCode id="ED66AE31FA4E125AFEE2F91CFE0CFC18" box="[374,403,876,899]" pageId="15" pageNumber="476">IV</collectionCode>
|
||
|
|
each with two distolateral oar-like ciliated subapical setae (®gure 5F). Tarsus I slender, its tip abruptly truncated in lateral view, length to width ratio 4.87±5.64, with four rhagidial solenidia lying obliquely in separate depressions, stellate famulus
|
||
|
|
<emphasis id="B903EAE6FA4E125AFE09F9BBFE38FC78" bold="true" box="[413,423,971,995]" italics="true" pageId="15" pageNumber="476">e</emphasis>
|
||
|
|
inserted antaxially laterad of ®rst proximal rhagidial solenidion (®gure 5B). Tarsus
|
||
|
|
<collectionCode id="ED66AE31FA4E125AFE26F99BFE59FB99" box="[434,454,1003,1026]" pageId="15" pageNumber="476">II</collectionCode>
|
||
|
|
with four rhagidial solenidia in separate oblique depressions and small spiniform famulus
|
||
|
|
<emphasis id="B903EAE6FA4E125AFDACFE7BFDDDFBB8" bold="true" box="[568,578,1035,1059]" italics="true" pageId="15" pageNumber="476">e</emphasis>
|
||
|
|
inserted antaxially laterad of ®rst and second proximal solenidion (®gure 5C).
|
||
|
|
</paragraph>
|
||
|
|
<paragraph id="8BC836F4FA4E125AFFDBFE1AFF2DFB19" blockId="15.[79,1035,1130,1250]" box="[79,178,1130,1154]" pageId="15" pageNumber="476">
|
||
|
|
<emphasis id="B903EAE6FA4E125AFFDBFE1AFF2DFB19" box="[79,178,1130,1154]" italics="true" pageId="15" pageNumber="476">A nities</emphasis>
|
||
|
|
</paragraph>
|
||
|
|
<paragraph id="8BC836F4FA4E125AFFECFEFBFCFCFB79" blockId="15.[79,1035,1130,1250]" pageId="15" pageNumber="476">
|
||
|
|
<taxonomicName id="4C774D77FA4E125AFFECFEFBFF62FB39" authority="Zacharda, 2000" authorityName="Zacharda" authorityYear="2000" box="[120,253,1163,1186]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="15" pageNumber="476" phylum="Arthropoda" rank="species" species="aggerata" status="sp. nov.">
|
||
|
|
<emphasis id="B903EAE6FA4E125AFFECFEFBFF62FB39" box="[120,253,1163,1186]" italics="true" pageId="15" pageNumber="476">T. aggerata</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
<taxonomicNameLabel id="A230579DFA4E125AFE99FEFBFED0FB39" box="[269,335,1163,1186]" pageId="15" pageNumber="476" rank="species">n. sp.</taxonomicNameLabel>
|
||
|
|
di ers from other known
|
||
|
|
<taxonomicName id="4C774D77FA4E125AFD0AFEFAFC1BFB39" box="[670,900,1162,1186]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="15" pageNumber="476" phylum="Arthropoda" rank="species" species="undetermined">
|
||
|
|
<emphasis id="B903EAE6FA4E125AFD0AFEFAFCB9FB39" box="[670,806,1162,1186]" italics="true" pageId="15" pageNumber="476">Troglocheles</emphasis>
|
||
|
|
species
|
||
|
|
</taxonomicName>
|
||
|
|
in (1) the proportions of prosoma and opisthosoma, (2) relatively smaller epimeres I±
|
||
|
|
<collectionCode id="ED66AE31FA4E125AFC2EFEDBFC48FB59" box="[954,983,1195,1218]" pageId="15" pageNumber="476">IV</collectionCode>
|
||
|
|
, (3) the morphology of the chelicera with the robust ¯attened ®xed digit.
|
||
|
|
</paragraph>
|
||
|
|
<paragraph id="8BC836F4FA4E125AFFDBFF7AFF58FAB9" blockId="15.[79,1034,1290,1377]" box="[79,199,1290,1314]" pageId="15" pageNumber="476">
|
||
|
|
<emphasis id="B903EAE6FA4E125AFFDBFF7AFF58FAB9" box="[79,199,1290,1314]" italics="true" pageId="15" pageNumber="476">Etymology</emphasis>
|
||
|
|
</paragraph>
|
||
|
|
<paragraph id="8BC836F4FA4E125AFFECFF59FD3CFAFA" blockId="15.[79,1034,1290,1377]" pageId="15" pageNumber="476">
|
||
|
|
The name
|
||
|
|
<taxonomicName id="4C774D77FA4E125AFF62FF5AFECAFADA" authorityName="Zacharda" authorityYear="2000" box="[246,341,1322,1345]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="15" pageNumber="476" phylum="Arthropoda" rank="species" species="aggerata">
|
||
|
|
<emphasis id="B903EAE6FA4E125AFF62FF5AFECAFADA" box="[246,341,1322,1345]" italics="true" pageId="15" pageNumber="476">aggerata</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
means inhabiting talus slopes, and this reēcts the habitat where this species was discovered. Feminine gender.
|
||
|
|
</paragraph>
|
||
|
|
<paragraph id="8BC836F4FA4E125AFFDBFFF9FF2EFA3A" blockId="15.[79,1035,1417,1696]" box="[79,177,1417,1441]" pageId="15" pageNumber="476">
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<emphasis id="B903EAE6FA4E125AFFDBFFF9FF2EFA3A" box="[79,177,1417,1441]" italics="true" pageId="15" pageNumber="476">Remarks</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D657FFA4E1245FFECFFD9FE7DFF7E" lastPageId="16" lastPageNumber="477" pageId="15" pageNumber="476" type="reference_group">
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<paragraph id="8BC836F4FA4E1245FFECFFD9FE7DFF7E" blockId="15.[79,1035,1417,1696]" lastBlockId="16.[79,1035,142,230]" lastPageId="16" lastPageNumber="477" pageId="15" pageNumber="476">
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<taxonomicName id="4C774D77FA4E125AFFECFFD9FEF5FA5A" ID-CoL="a9b9a295-d010-44e2-8546-b7edd625e3e6" authority="Zacharda, 2000" authorityName="Zacharda" authorityYear="2000" box="[120,362,1449,1473]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="15" pageNumber="476" phylum="Arthropoda" rank="species" species="aggerata" status="sp. nov.">
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<emphasis id="B903EAE6FA4E125AFFECFFD9FEF5FA5A" box="[120,362,1449,1473]" italics="true" pageId="15" pageNumber="476">Troglocheles aggerata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A230579DFA4E125AFEE5FFDAFE31FA5A" box="[369,430,1450,1473]" pageId="15" pageNumber="476" rank="species">n. sp.</taxonomicNameLabel>
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was only collected in spaces between talus fragments in a cool and moist talus slope in the alpine subnival zone of the Oetztal Alps. The occurrence of this species seems to be con®ned to this biotope. Speci®c morphological adaptations such as (1) the loss of eyes; (2) the extremely ®ne whitish integument; (3) relative reduction of the prosoma and legs; (4) relative enlargement of the opisthosoma can be viewed as characters that suggest movement in con®ned spaces, i.e. edaphic or hemiedaphic habitats. They resemble edaphomorphism s as introduced by
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<bibRefCitation id="EFE64B05FA4E125AFFE6FCF8FEB0F93B" author="ZACHARDA, M." box="[114,303,1672,1696]" pageId="15" pageNumber="476" pagination="514" refId="ref7655" refString="ZACHARDA, M., 1979, The evaluation of the morphological characters in Rhagidiidae, in J. G. Rodriguez (ed.) Recent Advances in Acarology, Vol. II (New York: Academic Press), pp. 509 ± 514." type="journal article" year="1979">Zacharda (1979)</bibRefCitation>
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and broadly discussed by
|
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<bibRefCitation id="EFE64B05FA4E125AFDCCFCF8FB9EF93B" author="LINDQUIST, E. E. & PALACIOS-VARGAS, J. G." box="[600,1025,1672,1696]" pageId="15" pageNumber="476" pagination="341" refId="ref7160" refString="LINDQUIST, E. E. and PALACIOS-VARGAS, J. G., 1991, Proterorhagiidae (Acari: Endeostigmata), a new family of rhagidiid-like mites from Mexico, Acarologia, 32 (4), 341 ± 363." type="journal article" year="1991">Lindquist and Palacios-Vargas (1991)</bibRefCitation>
|
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. In the light of these derivative modi®cations
|
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<taxonomicName id="4C774D77FA511245FDE5FAFFFD6AFF3D" ID-CoL="a9b9a295-d010-44e2-8546-b7edd625e3e6" authorityName="Zacharda" authorityYear="2000" box="[625,757,143,166]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="16" pageNumber="477" phylum="Arthropoda" rank="species" species="aggerata">
|
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<emphasis id="B903EAE6FA511245FDE5FAFFFD6AFF3D" box="[625,757,143,166]" italics="true" pageId="16" pageNumber="477">T. aggerata</emphasis>
|
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|
</taxonomicName>
|
||
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represents a derivative lineage adapted to special environmental conditions in con®ned spaces between fragments of the alpine talus slope.
|
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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