treatments-xml/data/03/CD/BE/03CDBE74938DB5ADCDABFA12FB2CFAF5.xml

<document id="08EEE7FF29C9CB3207146E0889DB56AE" ID-DOI="10.5252/zoosystema2023v45a9" ID-ISSN="1638-9387" ID-Zenodo-Dep="8071253" ID-ZooBank="urn:lsid:zoobank.org:pub:69C34731-8C25-4A1E-B336-B222CD3CBAC3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1686747199180" checkinUser="felipe" docAuthor="Guinot, Danièle" docDate="2023" docId="03CDBE74938DB5ADCDABFA12FB2CFAF5" docLanguage="en" docName="Zoosystema.45.9.225-372.pdf" docOrigin="Zoosystema 45 (9)" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.7:Zoosystema.2020-.journal_article" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article" docStyleVersion="7" docTitle="Heikeopsis tuberculata" docType="treatment" docVersion="5" lastPageNumber="360" masterDocId="FFF4C60C9304B524CD15FFC0FF9AFFF8" masterDocTitle="A new subfamily classification of the highly diversified Dorippidae H. Milne Edwards, 1837 (Crustacea, Decapoda, Brachyura, Dorippoidea), using morphological, molecular and palaeotonlogical data, with special emphasis on its unique female reproductive system" masterLastPageNumber="372" masterPageNumber="225" pageNumber="360" updateTime="1699524371485" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="80FDB7FFE0817BF4DF3551CC5311A9E4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="34746D10EB0C3F2AC195EB7C96216D18">
<mods:title id="BCEC40323605EB2F52AE4DFC744A00BE">A new subfamily classification of the highly diversified Dorippidae H. Milne Edwards, 1837 (Crustacea, Decapoda, Brachyura, Dorippoidea), using morphological, molecular and palaeotonlogical data, with special emphasis on its unique female reproductive system</mods:title>
</mods:titleInfo>
<mods:name id="A6BBA99346594808476ACAF0E4DE54A8" type="personal">
<mods:role id="23408709A0BA2039FAAFC26D6A304C1F">
<mods:roleTerm id="21F95DA1054D18C2875E6B7905A09111">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8E5C6915826C2FD0C2F35BE288B50244">Guinot, Danièle</mods:namePart>
<mods:affiliation id="6EBBED4F299184B3172E2B1B6D859FEB">Institut de Systématique, Évolution, Biodiversité (ISYEB), Muséum national d’Histoire naturelle, CNRS, Sorbonne Université, EPHE, Université des Antilles, case postale 53, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) daniele. guinot @ mnhn. fr Dedicated to the memory of my colleague and dearest friend Ngan Kee NG (1966 - 2022)</mods:affiliation>
<mods:nameIdentifier id="26ACE73FC1A9F70AE358891FB4C6391B" type="email">guinot@mnhn.fr</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="2DCACBBA2A7E3E5BC00C8793E45323F4">text</mods:typeOfResource>
<mods:relatedItem id="21B87C1A44E95669BAB1A09C2504F97D" type="host">
<mods:titleInfo id="38BDF9E01C01800D5FF53C30AB14A588">
<mods:title id="C06CAB89BB75A16736F711C608871A1E">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="8227365597C3839920FCC9BDC53A9D49">
<mods:date id="9D7509BCAD0C4779EBD9AED69CA0F733">2023</mods:date>
<mods:detail id="A671BAC481976DE04769158B0D44BE00" type="pubDate">
<mods:number id="9CD1367D095BFB7D38BA5C407AA19BCA">2023-06-05</mods:number>
</mods:detail>
<mods:detail id="9E51A44A0026936317CC0C7CFDE5B2AE" type="volume">
<mods:number id="0D7489D1B026DEA51F0DB2B026A831D4">45</mods:number>
</mods:detail>
<mods:detail id="43EB63A242EB3AC795E7773858B6C5D2" type="issue">
<mods:number id="CE1B6EF6AB4373D1AA1460A12870CDE9">9</mods:number>
</mods:detail>
<mods:extent id="9B8ED226EE6542471D846C61136426E5" unit="page">
<mods:start id="8641AAA51356605E090958803722EBB8">225</mods:start>
<mods:end id="1122C8A547954D2016270DEB3467C305">372</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="443609B6D84D3F6ABCA2186C412E3036">journal article</mods:classification>
<mods:identifier id="1B2B550877C25DEB42B17E80883CE5DC" type="DOI">10.5252/zoosystema2023v45a9</mods:identifier>
<mods:identifier id="11593512E6A30C8C7902988559294670" type="ISSN">1638-9387</mods:identifier>
<mods:identifier id="F478EBD4817A02F26DD93BFA581F17B0" type="Zenodo-Dep">8071253</mods:identifier>
<mods:identifier id="4D7EA7F86268EAFD9D08ACCED0A8EC28" type="ZooBank">urn:lsid:zoobank.org:pub:69C34731-8C25-4A1E-B336-B222CD3CBAC3</mods:identifier>
</mods:mods>
<treatment id="03CDBE74938DB5ADCDABFA12FB2CFAF5" ID-DOI="http://doi.org/10.5281/zenodo.8038674" ID-Zenodo-Dep="8038674" LSID="urn:lsid:plazi:treatment:03CDBE74938DB5ADCDABFA12FB2CFAF5" httpUri="http://treatment.plazi.org/id/03CDBE74938DB5ADCDABFA12FB2CFAF5" lastPageNumber="360" pageId="137" pageNumber="360">
<subSubSection id="C37E5CE9938DB5ADCDABFA12FD57FA15" box="[190,717,1490,1517]" pageId="137" pageNumber="360" type="nomenclature">
<paragraph id="8BDB0F62938DB5ADCDABFA12FD57FA15" blockId="137.[190,717,1490,1517]" box="[190,717,1490,1517]" pageId="137" pageNumber="360">
<heading id="D093B80E938DB5ADCDABFA12FD57FA15" box="[190,717,1490,1517]" centered="true" fontSize="11" level="2" pageId="137" pageNumber="360" reason="2">
<taxonomicName id="4C6474E1938DB5ADCDABFA12FD57FA15" ID-CoL="f1e74684-3b65-4d04-b30a-b7fd892a7d1e" authority="(Morris &amp; Collins, 1991)" baseAuthorityName="Morris &amp; Collins" baseAuthorityYear="1991" box="[190,717,1490,1517]" class="Malacostraca" family="Dorippidae" genus="Heikeopsis" kingdom="Animalia" order="Decapoda" pageId="137" pageNumber="360" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="B910D370938DB5ADCDABFA12FE28FA15" bold="true" box="[190,434,1490,1517]" italics="true" pageId="137" pageNumber="360">Heikeopsis tuberculata</emphasis>
(Morris &amp; Collins, 1991)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37E5CE9938DB5ADCD91F9CCFD23F8FA" pageId="137" pageNumber="360" type="reference_group">
<paragraph id="8BDB0F62938DB5ADCD91F9CCFF7AF9C6" blockId="137.[132,775,1548,1598]" pageId="137" pageNumber="360">
<taxonomicName id="4C6474E1938DB5ADCD91F9CCFC98F9DC" authority="Morris &amp; Collins, 1991: 5" authorityName="Morris &amp; Collins" authorityPageNumber="5" authorityYear="1991" box="[132,770,1548,1572]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="137" pageNumber="360" phylum="Arthropoda" rank="subSpecies" species="frascone" subGenus="Dorippe" subSpecies="tuberculata">
<emphasis id="B910D370938DB5ADCD91F9CCFF4AF9DB" box="[132,208,1548,1571]" italics="true" pageId="137" pageNumber="360">Dorippe</emphasis>
(
<emphasis id="B910D370938DB5ADCDCAF9CCFEB1F9DB" box="[223,299,1548,1571]" italics="true" pageId="137" pageNumber="360">Dorippe</emphasis>
) 
<emphasis id="B910D370938DB5ADCC2EF9CCFE6EF9DC" box="[315,500,1548,1572]" italics="true" pageId="137" pageNumber="360">frascone tuberculata</emphasis>
Morris &amp; Collins, 1991: 5
</taxonomicName>
, fig. 1a, b.
</paragraph>
<paragraph id="8BDB0F62938DB5ADCD91F997FD08F996" blockId="137.[132,658,1622,1646]" box="[132,658,1622,1646]" pageId="137" pageNumber="360">
<taxonomicName id="4C6474E1938DB5ADCD91F997FE0AF996" authorityName="Morris &amp; Collins" authorityYear="1991" box="[132,400,1622,1646]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="137" pageNumber="360" phylum="Arthropoda" rank="subSpecies" species="frascone" subSpecies="tuberculata">
<emphasis id="B910D370938DB5ADCD91F997FE0AF996" box="[132,400,1622,1646]" italics="true" pageId="137" pageNumber="360">Dorippe frascone tuberculata</emphasis>
</taxonomicName>
– Karasawa 2000: table 1.
</paragraph>
<paragraph id="8BDB0F62938DB5ADCD91F945FD2EF965" blockId="137.[132,692,1669,1694]" box="[132,692,1669,1694]" pageId="137" pageNumber="360">
<taxonomicName id="4C6474E1938DB5ADCD91F945FEACF965" baseAuthorityName="Morris &amp; Collins" baseAuthorityYear="1991" box="[132,310,1669,1693]" class="Malacostraca" family="Dorippidae" genus="Heikea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="137" pageNumber="360" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="B910D370938DB5ADCD91F945FEACF965" box="[132,310,1669,1693]" italics="true" pageId="137" pageNumber="360">Heikea tuberculata</emphasis>
</taxonomicName>
– Collins 
<emphasis id="B910D370938DB5ADCC89F946FE51F965" box="[412,459,1669,1693]" italics="true" pageId="137" pageNumber="360">et al.</emphasis>
2003: 200, pl. 1, fig. 5.
</paragraph>
<paragraph id="8BDB0F62938DB5ADCD91F975FD23F8FA" blockId="137.[132,776,1717,1794]" pageId="137" pageNumber="360">
<emphasis id="B910D370938DB5ADCD91F975FF44F935" box="[132,222,1717,1741]" italics="true" pageId="137" pageNumber="360">Heikeopsi</emphasis>
s 
<taxonomicName id="4C6474E1938DB5ADCDF9F975FECCF935" baseAuthorityName="Morris &amp; Collins" baseAuthorityYear="1991" box="[236,342,1717,1741]" class="Malacostraca" family="Dorippidae" genus="Heikea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="137" pageNumber="360" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="B910D370938DB5ADCDF9F975FECCF935" box="[236,342,1717,1741]" italics="true" pageId="137" pageNumber="360">tuberculata</emphasis>
</taxonomicName>
– Schweitzer 
<emphasis id="B910D370938DB5ADCCCFF976FD92F935" box="[474,520,1717,1741]" italics="true" pageId="137" pageNumber="360">et al.</emphasis>
2010: 79. — Guinot 
<emphasis id="B910D370938DB5ADCFCFF976FC92F935" box="[730,776,1717,1741]" italics="true" pageId="137" pageNumber="360">et al.</emphasis>
2013: 187; 2019: 300. — Sasaki 2019: 7790. — Van Bakel 
<emphasis id="B910D370938DB5ADCFCDF910FC92F91F" box="[728,776,1743,1767]" italics="true" pageId="137" pageNumber="360">et al.</emphasis>
2020: fig. 10.27. — Schweitzer 
<emphasis id="B910D370938DB5ADCCA8F92BFE76F8FA" box="[445,492,1770,1794]" italics="true" pageId="137" pageNumber="360">et al.</emphasis>
2021: 2, fig. 1.5a, b.
</paragraph>
</subSubSection>
<subSubSection id="C37E5CE9938DB5ADCD91F8F2FB2CFAF5" pageId="137" pageNumber="360" type="discussion">
<paragraph id="8BDB0F62938DB5ADCD91F8F2FF72F8B2" blockId="137.[130,776,1842,2028]" box="[132,232,1842,1868]" pageId="137" pageNumber="360">
<smallCapsWord id="8D3D99BE938DB5ADCD91F8F2FF72F8B2" baselines="1861,1861" box="[132,232,1842,1868]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remarks" pageId="137" pageNumber="360">REMARKS</smallCapsWord>
</paragraph>
<paragraph id="8BDB0F62938DB5ADCD91F891FB2CFAF5" blockId="137.[130,776,1842,2028]" lastBlockId="137.[811,1458,215,1294]" pageId="137" pageNumber="360">
<taxonomicName id="4C6474E1938DB5ADCD91F891FEE8F893" baseAuthorityName="Morris &amp; Collins" baseAuthorityYear="1991" box="[132,370,1873,1899]" class="Malacostraca" family="Dorippidae" genus="Heikeopsis" kingdom="Animalia" order="Decapoda" pageId="137" pageNumber="360" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="B910D370938DB5ADCD91F891FEE8F893" box="[132,370,1873,1899]" italics="true" pageId="137" pageNumber="360">Heikeopsis tuberculata</emphasis>
</taxonomicName>
, from the Lower Miocene Miri Formation of 
<collectingCountry id="F3734FF2938DB5ADCDEEF8B1FEC3F873" box="[251,345,1905,1931]" name="Malaysia" pageId="137" pageNumber="360">Sarawak</collectingCountry>
, established without knowledge of the contemporary work of Holthuis &amp; Manning (1990), is one of the few fossil dorippids with a well-preserved ventral part (Morris &amp; Collins 1991: 5, fig. 1a, 1b). Based on examination of the type series (
<typeStatus id="54DFB1C0938DB5ADC909FF17FBE4FF09" box="[1052,1150,215,241]" pageId="137" pageNumber="360" type="holotype">holotype</typeStatus>
and 
<specimenCount id="9D62C4EB938DB5ADC9A7FF18FAD3FF09" box="[1202,1353,215,242]" pageId="137" pageNumber="360" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
) and better preserved specimens, the new description and figures of Collins 
<emphasis id="B910D370938DB5ADCE97FED8FC2EFEC9" box="[898,948,279,305]" italics="true" pageId="137" pageNumber="360">et al.</emphasis>
(2003: 200, pl. 1, fig. 5a-d, as 
<taxonomicName id="4C6474E1938DB5ADC9E8FED7FA3CFEC9" authorityName="von Siebold" authorityYear="1824" box="[1277,1446,279,305]" class="Malacostraca" family="Dorippidae" genus="Heikea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="137" pageNumber="360" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B910D370938DB5ADC9E8FED7FA3CFEC9" box="[1277,1446,279,305]" italics="true" pageId="137" pageNumber="360">Heikea japonica</emphasis>
</taxonomicName>
) confirm all the characteristic features of the family 
<taxonomicName id="4C6474E1938DB5ADC877FEF7FCF1FE89" class="Malacostraca" family="Dorippidae" kingdom="Animalia" order="Decapoda" pageId="137" pageNumber="360" phylum="Arthropoda" rank="family">Dorippidae</taxonomicName>
, e.g. the clearly recognisable Milne Edwards openings and the curvature of the sternal suture 5/6 including the press-button for pleonal locking. This crab is undeniably a dorippid, with the two typical main carapace grooves: a ‘hepatic furrow’ that corresponds to the precervical groove; and a ‘wide, fairly deep, and broadly V-shaped’ cervical groove (see 
<emphasis id="B910D370938DB5ADCEA0FDD7FB8EFDA8" italics="true" pageId="137" pageNumber="360">
Interpretation of grooves on the carapace dorsal surface in 
<taxonomicName id="4C6474E1938DB5ADCE8EFDF6FB8EFDA8" box="[923,1044,566,592]" class="Malacostraca" family="Dorippidae" kingdom="Animalia" order="Decapoda" pageId="137" pageNumber="360" phylum="Arthropoda" rank="family">Dorippidae</taxonomicName>
</emphasis>
). The well-marked posterior rim, described as ‘a broad ridge [that] bounds the deeply sinuous, not concave posterior margin’, is similar to that of the extant 
<taxonomicName id="4C6474E1938DB5ADCE3EFD56FC03FD48" box="[811,921,662,688]" class="Malacostraca" family="Dorippidae" genus="Heikeopsis" kingdom="Animalia" order="Decapoda" pageId="137" pageNumber="360" phylum="Arthropoda" rank="genus">
<emphasis id="B910D370938DB5ADCE3EFD56FC03FD48" box="[811,921,662,688]" italics="true" pageId="137" pageNumber="360">Heikeopsis</emphasis>
</taxonomicName>
. The thoracic sternum, depicted upside down by Morris &amp; Collins (1991) but correctly oriented by Collins 
<emphasis id="B910D370938DB5ADCE38FD16FCF9FD17" box="[813,867,725,751]" italics="true" pageId="137" pageNumber="360">et al.</emphasis>
(2003) (see Guinot 
<emphasis id="B910D370938DB5ADC95FFD16FBE5FD17" box="[1098,1151,725,751]" italics="true" pageId="137" pageNumber="360">et al.</emphasis>
2013: 187), was shown by Schweitzer 
<emphasis id="B910D370938DB5ADCEBCFD36FC47FCF7" box="[937,989,757,783]" italics="true" pageId="137" pageNumber="360">et al.</emphasis>
(2021: 2, fig. 1.5b). The shape of the carapace, the sculpted and ornamented dorsal surface and the ridged thoracic sternum are not reminiscent of 
<taxonomicName id="4C6474E1938DB5ADC854FCF5FC1DFC97" baseAuthorityName="von Siebold" baseAuthorityYear="1824" class="Malacostraca" family="Dorippidae" genus="Heikeopsis" kingdom="Animalia" order="Decapoda" pageId="137" pageNumber="360" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B910D370938DB5ADC854FCF5FC1DFC97" italics="true" pageId="137" pageNumber="360">Heikeopsis japonica</emphasis>
</taxonomicName>
(
<figureCitation id="135F13E7938DB5ADCE81FC95FC4CFC97" box="[916,982,853,879]" captionStart="FIG" captionStartId="6.[132,143,1917,1935]" captionTargetBox="[317,1283,239,1875]" captionTargetId="figure-16@6.[317,1292,212,1875]" captionTargetPageId="6" captionText="FIG. 1. — A typical dorippid crab, Heikeopsis japonica (von Siebold, 1824), the iconic ‘samurai crab’ known in Japan as ’heike-gani’, whose carapace looks like a human face: A, the species represented by De Haan (1839: pl. 31, fig. 1, as Dorippe japonica) in Fauna Japonica: top, a female; below, chelae of a male; B, lectotype selected by Yamaguchi &amp; Baba (1993: 300, fig. 90-A.a-2, as Heikea japonica) from type series material labelled “Types, Japan, 1823, Leg. P. H. von Siebold”, RMNH CRUS D 822 (see Fransen et al. 1997: 83); C, ♂ 25.0 × 27.3 mm, ♀ 23.5 × 26.7 mm, Japan, Chaffanjon, 174-96, E.-L. Bouvier det. 1899 Dorippe japonica, dry condition, MNHN-IU-2000-4091 (= MNHN-B4091)." figureDoi="http://doi.org/10.5281/zenodo.8071255" httpUri="https://zenodo.org/record/8071255/files/figure.png" pageId="137" pageNumber="360">Figs 1</figureCitation>
; 
<figureCitation id="135F13E7938DB5ADCEF1FC95FB9BFC97" box="[996,1025,853,879]" captionStart="FIG" captionStartId="44.[132,143,1709,1727]" captionTargetBox="[132,1455,216,1666]" captionTargetId="figure-124@44.[132,1455,216,1666]" captionTargetPageId="44" captionText="FIG. 19. — Heikeopsinae n. subfam. A-D, Heikeopsis japonica (von Siebold, 1824): A, ♂ 22.5 × 24.3 mm, Japan, Tokushima, SMF 57856: habitus; B, ♀ 23.5 × 26.7 mm, Japan, Chaffanjon, MNHN-IU-2000-4091 (= MNHN-B4091): habitus; C, ♂ 25.0 × 27.3 mm, same data as B: ventral view; D, ♂ 24.2 × 25.4 mm, same data as B and C: thoracic sternum and pleon. E, F,?Heikeopsis aff. japonica, China, seas off Qingdao, ZRC 2002.0491: habitus: E, ♂ 21.9 × 24 mm; F, ♀ 23.4 × 26.4 mm." figureDoi="http://doi.org/10.5281/zenodo.8071302" httpUri="https://zenodo.org/record/8071302/files/figure.png" pageId="137" pageNumber="360">19</figureCitation>
A-D), just as the posterolateral tubercle that becomes a sharp spine in some large adults casts doubt on this generic identification. Furthermore, as the present distribution of 
<taxonomicName id="4C6474E1938DB5ADCEC8FC75FBFAFC36" baseAuthorityName="von Siebold" baseAuthorityYear="1824" box="[989,1120,949,974]" class="Malacostraca" family="Dorippidae" genus="Heikeopsis" kingdom="Animalia" order="Decapoda" pageId="137" pageNumber="360" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B910D370938DB5ADCEC8FC75FBFAFC36" box="[989,1120,949,974]" italics="true" pageId="137" pageNumber="360">H. japonica</emphasis>
</taxonomicName>
most probably includes only 
<collectingCountry id="F3734FF2938DB5ADCE3AFC15FCF7FC17" box="[815,877,981,1007]" name="Japan" pageId="137" pageNumber="360">Japan</collectingCountry>
and the nearby region, it may be preferable to assign it to another genus and species. The 
<typeStatus id="54DFB1C0938DB5ADC9D2FC34FAB3FBF6" box="[1223,1321,1012,1038]" pageId="137" pageNumber="360" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C6474E1938DB5ADC85AFC34FC53FBD6" baseAuthorityName="Morris &amp; Collins" baseAuthorityYear="1991" class="Malacostraca" family="Dorippidae" genus="Heikeopsis" kingdom="Animalia" order="Decapoda" pageId="137" pageNumber="360" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="B910D370938DB5ADC85AFC34FC53FBD6" italics="true" pageId="137" pageNumber="360">Heikeopsis tuberculata</emphasis>
</taxonomicName>
represented by Schweitzer 
<emphasis id="B910D370938DB5ADC811FBD5FAA1FBD6" box="[1284,1339,1044,1070]" italics="true" pageId="137" pageNumber="360">et al.</emphasis>
(2021: 2, fig. 1.5a, b), with the correct registration number (NHMUK In. 61853), has a sculptured carapace and ridged thoracic sternum that do not conform to the material assigned to 
<taxonomicName id="4C6474E1938DB5ADCE3BFB55FC37FB56" baseAuthorityName="von Siebold" baseAuthorityYear="1824" box="[814,941,1173,1198]" class="Malacostraca" family="Dorippidae" genus="Heikeopsis" kingdom="Animalia" order="Decapoda" pageId="137" pageNumber="360" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B910D370938DB5ADCE3BFB55FC37FB56" box="[814,941,1173,1198]" italics="true" pageId="137" pageNumber="360">H. japonica</emphasis>
</taxonomicName>
by Morris &amp; Collins (1991) and Collins 
<emphasis id="B910D370938DB5ADC86FFB55FA2AFB56" box="[1402,1456,1172,1198]" italics="true" pageId="137" pageNumber="360">et al.</emphasis>
(2003), and thus clearly correspond to another genus, most likely in 
<taxonomicName id="4C6474E1938DB5ADCE9EFB14FB91FB16" authority="Guinot, 2023" authorityName="Guinot" authorityYear="2023" box="[907,1035,1236,1262]" class="Malacostraca" family="Dorippidae" kingdom="Animalia" order="Decapoda" pageId="137" pageNumber="360" phylum="Arthropoda" rank="subFamily" status="subfam. nov." subFamily="Dorippinae">Dorippinae</taxonomicName>
<taxonomicNameLabel id="A2236E0B938DB5ADC907FB14FB13FB16" box="[1042,1161,1236,1262]" pageId="137" pageNumber="360" rank="subFamily">n. subfam.</taxonomicNameLabel>
(if the anterior part of the carapace is proven to be narrowed).
</paragraph>
</subSubSection>
</treatment>
</document>